681 lines
70 KiB
XML
681 lines
70 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.973.51435" ID-GBIF-Dataset="5812c79e-7c58-402d-9c8f-bc44bb258e57" ID-PMC="PMC7550391" ID-Pensoft-Pub="1313-2970-973-1" ID-Pensoft-UUID="78B73376DDD85D65A79F30D1A830200E" ID-PubMed="33110370" ID-ZooBank="6E1E443A737449129462E267B49EEA2E" ModsDocID="1313-2970-973-1" checkinTime="1601957171786" checkinUser="pensoft" docAuthor="Rohacek, Jindrich" docDate="2020" docId="E5AF0B3A36C3549AAAC79C1FE665BEE0" docLanguage="en" docName="ZooKeys 973: 1-15" docOrigin="ZooKeys 973" docSource="http://dx.doi.org/10.3897/zookeys.973.51435" docTitle="Protanthomyza grimaldii Roháček 2020, sp. nov." docType="treatment" docUuid="D95A90D3-4F55-46E2-814A-4D8117555C47" docUuidSource="ZooBank" docVersion="4" id="78B73376DDD85D65A79F30D1A830200E" lastPageNumber="1" masterDocId="78B73376DDD85D65A79F30D1A830200E" masterDocTitle="Protanthomyza grimaldii sp. nov., a further member of the extinct subfamily Protanthomyzinae (Diptera, Anthomyzidae) from Baltic amber" masterLastPageNumber="15" masterPageNumber="1" pageNumber="1" updateTime="1668169559776" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Protanthomyza grimaldii sp. nov., a further member of the extinct subfamily Protanthomyzinae (Diptera, Anthomyzidae) from Baltic amber</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rohacek, Jindrich</mods:namePart>
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<mods:affiliation>Silesian Museum, Nadrazni okruh 31, CZ- 746 01 Opava, Czech Republic</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3311-2087</mods:nameIdentifier>
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<mods:nameIdentifier type="email">rohacek@szm.cz</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>973</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>15</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.973.51435</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.973.51435</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-973-1</mods:identifier>
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<mods:identifier type="ZooBank">6E1E443A737449129462E267B49EEA2E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">78B73376DDD85D65A79F30D1A830200E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168237486" LSID="urn:lsid:zoobank.org:act:D95A90D3-4F55-46E2-814A-4D8117555C47" httpUri="http://treatment.plazi.org/id/E5AF0B3A36C3549AAAC79C1FE665BEE0" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="E5AF0B3A-36C3-549A-AAC7-9C1FE665BEE0" authority="Roháček, 2020" authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">Protanthomyza grimaldii</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">Figures 1-2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F2" captionText="Figure 3. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Photograph by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figure3" httpUri="https://binary.pensoft.net/fig/459226" pageId="0" pageNumber="1">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">, 4</figureCitation>
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<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">, 5-9</figureCitation>
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<figureCitation captionStart="Figures 10, 11" captionStartId="F5" captionText="Figures 10, 11. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 10 entire specimen, subdorsal view 11 ditto, lateroventral view. Body length ca 2.9 mm. Photographs by J. Rohacek (adapted from Rohacek 2013 a, fig. 7 E, F)." figureDoi="10.3897/zookeys.973.51435.figures10-11" httpUri="https://binary.pensoft.net/fig/459229" pageId="0" pageNumber="1">, 10, 11</figureCitation>
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<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">, 12-14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Hennig" authorityYear="1965" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Protanthomyza</taxonomicName>
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sp. nov.:
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<bibRefCitation DOI="https://doi.org/10.1080/14772019.2012.691907" author="Rohacek, J" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="431 - 473" refId="B10" refString="Rohacek, J, 2013a. New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity. Journal of Systematic Palaeontology 11: 431 - 473, DOI: https://doi.org/10.1080/14772019.2012.691907" title="New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity." url="https://doi.org/10.1080/14772019.2012.691907" volume="11" year="2013 a">
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<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
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2013a
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</bibRefCitation>
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: 451-452, fig. 7E, F (diagnosis, female only).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species is dedicated to Prof. Dr. David Grimaldi (New York, U. S. A.), the distinguished American dipterist and palaeontologist, in recognition of his valuable contribution to the knowledge of amber fossil flies, including acalyptrates (largely from Dominican amber).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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♂ labelled "Faszination Bernstein, Christel Hoffeins, Hans Werner Hoffeins" (framed on obverse), "1040-5a
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<taxonomicName authorityName="Hennig" authorityYear="1965" lsidName="" order="Diptera" pageId="0" pageNumber="1" rank="order">Diptera</taxonomicName>
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:
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<taxonomicName authorityName="Macquart" authorityYear="1835" class="Globothalamea" family="Spiroplectamminidae" genus="Acalyptratae" higherTaxonomySource="GBIF" kingdom="Chromista" lsidName="" order="Lituolida" pageId="0" pageNumber="1" phylum="Foraminifera" rank="genus">Acalyptratae</taxonomicName>
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,
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<taxonomicName family="Anthomyzidae" lsidName="" pageId="0" pageNumber="1" rank="family">Anthomyzidae</taxonomicName>
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♂" (handwritten by C. Hoffeins, on reverse), "Baltic amber, Russia: Kaliningrad region, Yantarny", "obtained in May 2010 from Dr. Andrey Krylov, Kaliningrad, Russia", and "Holotypus ♂,
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<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
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<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
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</taxonomicName>
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sp.n., J.
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<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
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det. 2020" (red label) [amber piece embedded in polyester resin, size 8.6
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<normalizedToken originalValue="×">x</normalizedToken>
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7.9
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<normalizedToken originalValue="×">x</normalizedToken>
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4.8 mm], deposited in SDEI (inventory number Dip-00821). The original amber stone (in form of an icicle = Zapfenschlaube in German), size about 45
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<normalizedToken originalValue="×">x</normalizedToken>
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20
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<normalizedToken originalValue="×">x</normalizedToken>
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11 mm, with multiple layers and aggregation of inclusions, was cut in two pieces; that with the inclusion of
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<taxonomicName lsidName="P. grimaldii" pageId="0" pageNumber="1" rank="species" species="grimaldii">
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<emphasis italics="true" pageId="0" pageNumber="1">P. grimaldii</emphasis>
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</taxonomicName>
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(No. 1040-5a) was separated, manually prepared and embedded in polyester resin by H. W. Hoffeins in August 2015. Syninclusions in 1040-5a: 1 stellate hair, pollen grains. Syninclusions in the remaining part (1040-5b, deposited in CCHH):
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<taxonomicName lsidName="" order="Diptera" pageId="0" pageNumber="1" rank="order">Diptera</taxonomicName>
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:
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<taxonomicName family="Empididae" lsidName="" pageId="0" pageNumber="1" rank="family">Empididae</taxonomicName>
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: 1
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<taxonomicName class="Insecta" family="Empididae" genus="Rhamphomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhamphomyia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhamphomyia</emphasis>
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</taxonomicName>
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sp. female; 2
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<taxonomicName family="Mycetophilidae" lsidName="" pageId="0" pageNumber="1" rank="family">Mycetophilidae</taxonomicName>
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males + fragment; 1
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<taxonomicName family="Simuliidae" lsidName="" pageId="0" pageNumber="1" rank="family">Simuliidae</taxonomicName>
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; 2
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<taxonomicName family="Chironomidae" lsidName="" pageId="0" pageNumber="1" rank="family">Chironomidae</taxonomicName>
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female and male;
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<taxonomicName lsidName="" order="Trichoptera" pageId="0" pageNumber="1" rank="order">Trichoptera</taxonomicName>
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, not identifiable;
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<taxonomicName lsidName="" order="Coleoptera" pageId="0" pageNumber="1" rank="order">Coleoptera</taxonomicName>
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fragment;
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<taxonomicName lsidName="" order="Araneae" pageId="0" pageNumber="1" rank="order">Araneae</taxonomicName>
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fragment; stellate hairs. Paratype ♀ labelled "47b-1, (Baltic A.), 15.1
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<normalizedToken originalValue="×">x</normalizedToken>
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10.7
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<normalizedToken originalValue="×">x</normalizedToken>
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3.3 [mm]" (handwritten), "
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<taxonomicName authorityName="Hennig" authorityYear="1965" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza</emphasis>
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</taxonomicName>
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sp.n. ♀, J.
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<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
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det. 2011" (yellow label) and "Paratypus ♀,
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<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
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<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
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</taxonomicName>
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sp.n., J.
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<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
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det. 2020" (yellow label) [shape of stone irregularly pentagonal, thin], temporarily held in CMTB; it will be deposited in AMNH. Syninclusions: only stellate hairs.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality and age.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Russia: Kaliningrad region, Yantarny mine. Middle to Late Eocene, 48-34 Ma (cf.
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<bibRefCitation DOI="https://doi.org/10.1111/brv.12414" author="Seyfullah, LJ" journalOrPublisher="Biological Reviews" pageId="0" pageNumber="1" pagination="1684 - 1714" refId="B14" refString="Seyfullah, LJ, Beimforde, C, Dal Corso, J, Perrichot, V, Rikkinen, J, Schmidt, AR, 2018. Production and preservation of resins - past and present. Biological Reviews 93: 1684 - 1714, DOI: https://doi.org/10.1111/brv.12414" title="Production and preservation of resins - past and present." url="https://doi.org/10.1111/brv.12414" volume="93" year="2018">Seyfullah et al. 2018</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Ocellar triangle delimited by a groove; frontal triangle not delimited; 1st antennal flagellomere normal, not enlarged; arista bare; 3 or 4
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<emphasis italics="true" pageId="0" pageNumber="1">dc</emphasis>
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macrosetae;
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<emphasis italics="true" pageId="0" pageNumber="1">f1</emphasis>
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without a ctenidial spine; male epandrium elongate and posteriorly tapered, with a robust, hook-like, curved anteroventral process; female T7 short; female S6 and S7 broadly transverse.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
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(Figs
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<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">1</figureCitation>
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,
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<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">2</figureCitation>
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). Total body length ca 2.6 mm; general colour brown to blackish brown; only head and some extremities partly ochreous to yellow.
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</paragraph>
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<caption doi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1" start="Figures 1, 2" startId="F1">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figures 1, 2.</emphasis>
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<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
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<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
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</taxonomicName>
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sp. nov., holotype male (Baltic amber)
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<emphasis bold="true" pageId="0" pageNumber="1">1</emphasis>
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entire specimen, left laterodorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">2</emphasis>
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ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J.
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<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
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.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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(Figs
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<figureCitation captionStart="Figure 3" captionStartId="F2" captionText="Figure 3. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Photograph by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figure3" httpUri="https://binary.pensoft.net/fig/459226" pageId="0" pageNumber="1">3</figureCitation>
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-
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<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">5</figureCitation>
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) higher than long, anteriorly somewhat angularly protruding in front of eye margin (Fig.
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<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">5</figureCitation>
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). Occiput very slightly concave, blackish brown. Frons relatively narrow, blackish brown only posteriorly around ocellar triangle, pale brown in anterior half to ochreous yellow at anterior marginal area surrounding frontal lunule. Frontal triangle not developed; ocellar triangle blackish brown, distinctly protruding and delimited by marginal groove. Ocelli large (Figs
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<figureCitation captionStart="Figure 3" captionStartId="F2" captionText="Figure 3. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Photograph by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figure3" httpUri="https://binary.pensoft.net/fig/459226" pageId="0" pageNumber="1">3</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">4</figureCitation>
|
||
). Orbital plate lighter (ochreous yellow) anteriorly, becoming pale brown posteriorly where contrasting with blackish-brown vicinity of ocellar triangle. Frontal lunule long, ochreous yellow. Face relatively narrow, ochreous, medially lighter and somewhat depressed; parafacialia and anterior half of gena dirty yellow and narrowly brown bordered (gena ventrally); posterior half of gena and postgena brown; mouthparts yellow to ochreous, clypeus pale brown, palpus dirty yellow. Cephalic chaetotaxy (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">5</figureCitation>
|
||
):
|
||
<emphasis italics="true" pageId="0" pageNumber="1">pvt</emphasis>
|
||
(only left one visible) relatively short, convergent but not crossed;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">vti</emphasis>
|
||
longest of cephalic setae, slightly inclinate;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">vte</emphasis>
|
||
strongly exclinate and only slightly shorter than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">vti</emphasis>
|
||
; 3 distinct
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ors</emphasis>
|
||
, all slightly reclinate, posterior
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ors</emphasis>
|
||
longest (about as long as
|
||
<emphasis italics="true" pageId="0" pageNumber="1">vte</emphasis>
|
||
), others becoming slightly shorter anteriorly;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">oc</emphasis>
|
||
relatively thin (not longer than middle
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ors</emphasis>
|
||
), proclinate (and unnaturally crossed in holotype), arising inside ocellar triangle; anterior half of frons with about 5 or 6 pairs of microsetae, mostly medially in front of ocellar triangle but a few (1 or 2) also between anterior and middle
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ors</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">vi</emphasis>
|
||
distinct (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">5</figureCitation>
|
||
), about 3 times as long as foremost peristomal setula; no subvibrissa; 4 or 5 weak proclinate peristomals; postocular setulae in two rows as usual but with only 3 setulae in inner row (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">4</figureCitation>
|
||
); outer row of postoculars long, reaching ventral eye margin; postgena with 2 setae, anterior short, posterior longer. Palpus slender, elongate, with a few (3 visible) minute setulae distally (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">5</figureCitation>
|
||
). Mouthparts relatively short. Eye bare, relatively large and strongly convex, suboval, anteriorly regularly rounded, with only posterior margin somewhat straighter; its longest diameter almost vertical and 1.16 times as long as shortest diameter. Gena low, its shortest height about 0.08 times as long as shortest eye diameter. Antenna medium-sized, generally porrect (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">5</figureCitation>
|
||
) but directed anteroventrally to ventrally, with dark-brown basal segments and pale-brown to ochreous 1st flagellomere. Pedicel with 1 longer seta and several microsetae; 1st flagellomere oval, laterally compressed, with very short, dense and dark pilosity; arista about 1.8 times as long as antenna, entirely bare (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">5</figureCitation>
|
||
), 2 basal segments slightly widened.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.973.51435.figure3" httpUri="https://binary.pensoft.net/fig/459226" pageId="0" pageNumber="1" start="Figure 3" startId="F2">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Photograph by J.
|
||
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1" start="Figure 4" startId="F3">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
|
||
hardly narrower than head, brown to blackish brown, with shining mesonotum and duller pleural part. Mesonotum relatively convex, separated from scutellum by deep suture. Scutellum rounded subtriangular, wider than long, convex dorsally; postscutellum not visible. Thoracic chaetotaxy (see Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F2" captionText="Figure 3. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Photograph by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figure3" httpUri="https://binary.pensoft.net/fig/459226" pageId="0" pageNumber="1">3</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">7</figureCitation>
|
||
) rich as usual in the genus: 1 short
|
||
<emphasis italics="true" pageId="0" pageNumber="1">hu</emphasis>
|
||
(markedly shorter than anterior
|
||
<emphasis italics="true" pageId="0" pageNumber="1">npl</emphasis>
|
||
) and only 2 or 3 setulae on humeral callus (= postpronotal lobe); 2 long
|
||
<emphasis italics="true" pageId="0" pageNumber="1">npl</emphasis>
|
||
, anterior slightly longer; 1 distinct
|
||
<emphasis italics="true" pageId="0" pageNumber="1">prs</emphasis>
|
||
(about as long as posterior
|
||
<emphasis italics="true" pageId="0" pageNumber="1">npl</emphasis>
|
||
); 1 long
|
||
<emphasis italics="true" pageId="0" pageNumber="1">sa</emphasis>
|
||
; 2
|
||
<emphasis italics="true" pageId="0" pageNumber="1">pa</emphasis>
|
||
, external very long (longer than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">sa</emphasis>
|
||
), internal shorter (not visible on left side in Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">4</figureCitation>
|
||
); 3 or 4 postsutural
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dc</emphasis>
|
||
(number different on left and right sides) becoming shorter anteriorly, the hindmost longest (together with apical
|
||
<emphasis italics="true" pageId="0" pageNumber="1">sc</emphasis>
|
||
longest thoracic setae);
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ac</emphasis>
|
||
microsetae dense, in 9 or 10 rows on suture, those in medial rows reaching up to posterior
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dc</emphasis>
|
||
; prescutellar
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ac</emphasis>
|
||
macrosetae long and strong, as long as
|
||
<emphasis italics="true" pageId="0" pageNumber="1">sa</emphasis>
|
||
; 2
|
||
<emphasis italics="true" pageId="0" pageNumber="1">sc</emphasis>
|
||
, laterobasal shorter (about as long as
|
||
<emphasis italics="true" pageId="0" pageNumber="1">prs</emphasis>
|
||
), apical very long; no additional setulae on scutellum; 2 upcurved
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ppl</emphasis>
|
||
, anterior distinct (as long as
|
||
<emphasis italics="true" pageId="0" pageNumber="1">hu</emphasis>
|
||
), posterior small. Mesopleuron (anepisternum) and sternopleuron (katepisternum) setose (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">7</figureCitation>
|
||
) as in most other species of
|
||
<taxonomicName authorityName="Hennig" authorityYear="1965" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza</emphasis>
|
||
</taxonomicName>
|
||
: 3 long
|
||
<emphasis italics="true" pageId="0" pageNumber="1">mspl</emphasis>
|
||
(1 dorsal upcurved and shortest, 1 posterodorsal and 1 posterior longest, cf. Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">7</figureCitation>
|
||
) and numerous setulae in posterodorsal half of mesopleuron; 1 long posterior
|
||
<emphasis italics="true" pageId="0" pageNumber="1">stpl</emphasis>
|
||
and about 9 or 10 setulae in posterior half of sternopleuron; other sclerites of pleural part of thorax bare.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1" start="Figures 5–9" startId="F4">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figures 5-9.</emphasis>
|
||
<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (Baltic amber)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
|
||
head, left sublateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">6</emphasis>
|
||
base of left wing
|
||
<emphasis bold="true" pageId="0" pageNumber="1">7</emphasis>
|
||
right mesopleuron, sternopleuron and pteropleuron, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">8</emphasis>
|
||
abdomen and haltere, right lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">9</emphasis>
|
||
terminalia, right lateral view. Scale bars: 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
|
||
); 0.3 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">6-8</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">9</emphasis>
|
||
). For abbreviations see p. 4.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
|
||
brown to ochreous, femora darkest, fore coxa pale ochreous.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">f1</emphasis>
|
||
lacking ctenidial spine; 5 (2 longer) distinct widely-spaced setae in posterodorsal row; setae in posteroventral row more numerous but short and weak.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">f2</emphasis>
|
||
with 2 or 3 anterior setae near middle (cf. Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">14</figureCitation>
|
||
), otherwise shortly setulose as is
|
||
<emphasis italics="true" pageId="0" pageNumber="1">f3</emphasis>
|
||
.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">t2</emphasis>
|
||
with distinct ventroapical seta (about as long as maximum width of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">t2</emphasis>
|
||
) and 2 or 3 small setae adjacent to the latter;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">t1</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">t3</emphasis>
|
||
and all tarsi simply setulose but basitarsi of all legs with ventrobasal setulae somewhat longer than others.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">6</figureCitation>
|
||
) moderately long and narrow, widest at distal third; veins brown to pale brown, membrane unicolourous, pale-brown tinged;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C</emphasis>
|
||
with more or less distinct subcostal break and somewhat attenuated at humeral cross-vein.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C</emphasis>
|
||
extended to apex of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M</emphasis>
|
||
, densely uniformly setulose on
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cs2</emphasis>
|
||
(from subcostal break to apex of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R2+3</emphasis>
|
||
), finely short-pilose more distally (on
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cs3</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cs4</emphasis>
|
||
);
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sc</emphasis>
|
||
distinct, separate almost along its entire length, only apically fused with
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R1</emphasis>
|
||
to form preapical kink (see Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">6</figureCitation>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">pk</emphasis>
|
||
);
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R1</emphasis>
|
||
short, dilated distally due to fusion with
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sc</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R2+3</emphasis>
|
||
long, very slightly sinuate to almost straight and only its extreme apex slightly upcurved to
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R4+5</emphasis>
|
||
slightly recurved in distal half, divergent from
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R2+3</emphasis>
|
||
and apically slightly convergent with
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M</emphasis>
|
||
almost straight. Cell
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm</emphasis>
|
||
of moderate length, narrow proximally and much widened distally, with angle of anterior outer corner obtuse while that of posterior outer corner distinctly acute (cf. Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">12</figureCitation>
|
||
);
|
||
<emphasis italics="true" pageId="0" pageNumber="1">r-m</emphasis>
|
||
situated in basal third of cell
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm-cu</emphasis>
|
||
straight; apical portion of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">CuA1</emphasis>
|
||
much shorter than distance between
|
||
<emphasis italics="true" pageId="0" pageNumber="1">r-m</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm-cu</emphasis>
|
||
, distinctly longer than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm-cu</emphasis>
|
||
and almost reaching wing margin;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A1</emphasis>
|
||
relatively long but ending far from wing margin;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A2</emphasis>
|
||
well developed, slightly shorter than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A1</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">6</figureCitation>
|
||
); alula distinct but narrow (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">6</figureCitation>
|
||
). Wing measurements: length ca 2.4 mm, width ca 0.8 mm,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cs3</emphasis>
|
||
:
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cs4</emphasis>
|
||
= 1.38,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">r-m\dm-cu</emphasis>
|
||
:
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm-cu</emphasis>
|
||
= 3.50,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">r-m\dm-cu</emphasis>
|
||
:
|
||
<emphasis italics="true" pageId="0" pageNumber="1">CuA1</emphasis>
|
||
= 2.50. Haltere (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber). Head and thorax, left laterodorsal view. Scale bar: 0.3 mm. For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figure4" httpUri="https://binary.pensoft.net/fig/459227" pageId="0" pageNumber="1">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
) pale ochreous, knob relatively large, darker dorsally.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
) relatively short, robust but not very broad. Preabdominal terga blackish brown;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T1</emphasis>
|
||
fused with
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T2</emphasis>
|
||
but delimited by a groove (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
);
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T1</emphasis>
|
||
finely short-setose,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T2-T5</emphasis>
|
||
with relatively long and dense setae (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
);
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T2</emphasis>
|
||
somewhat shorter than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T3</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T3-T5</emphasis>
|
||
subequal in length, all relatively short and transverse (Figs
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
). Preabdominal sterna mostly invisible (Figs
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
) but probably small (narrow) and pale-pigmented; only
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S5</emphasis>
|
||
discernible (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
), paler brown and shorter than adjacent
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T5</emphasis>
|
||
, with sparse setae. Also 2 setae of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S4</emphasis>
|
||
visible on right side of abdomen (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
) but margins of sclerite are not recognizable. Postabdomen:
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S6</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S7</emphasis>
|
||
not visible because situated on obscured left side of abdomen (cf. Fig.
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">1</figureCitation>
|
||
), probably asymmetrical and (partly) fused together (dorsally also with
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S8</emphasis>
|
||
) as in other
|
||
<taxonomicName authorityName="Hennig" authorityYear="1965" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza</emphasis>
|
||
</taxonomicName>
|
||
species (cf.
|
||
<bibRefCitation DOI="https://doi.org/10.1080/14772019.2012.691907" author="Rohacek, J" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="431 - 473" refId="B10" refString="Rohacek, J, 2013a. New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity. Journal of Systematic Palaeontology 11: 431 - 473, DOI: https://doi.org/10.1080/14772019.2012.691907" title="New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity." url="https://doi.org/10.1080/14772019.2012.691907" volume="11" year="2013 a">
|
||
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
|
||
2013a
|
||
</bibRefCitation>
|
||
, fig. 4H).
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S8</emphasis>
|
||
relatively long, blackish brown, situated dorsally and readily visible on right side (Figs
|
||
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 1 entire specimen, left laterodorsal view 2 ditto, right lateroventral view. Body length ca 2.6 mm. Photographs by J. Rohacek." figureDoi="10.3897/zookeys.973.51435.figures1-2" httpUri="https://binary.pensoft.net/fig/459225" pageId="0" pageNumber="1">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
), almost bare, with only a few setulae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Epandrium (Figs
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">9</figureCitation>
|
||
) relatively long but tapered posteriorly, with anteroventral corner modified to distinct, flat and somewhat hook-like projection (see Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">9</figureCitation>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">avp</emphasis>
|
||
) distinctly different from those in other
|
||
<taxonomicName authorityName="Hennig" authorityYear="1965" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza</emphasis>
|
||
</taxonomicName>
|
||
species where known (cf.
|
||
<bibRefCitation DOI="https://doi.org/10.1080/14772019.2012.691907" author="Rohacek, J" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="431 - 473" refId="B10" refString="Rohacek, J, 2013a. New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity. Journal of Systematic Palaeontology 11: 431 - 473, DOI: https://doi.org/10.1080/14772019.2012.691907" title="New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity." url="https://doi.org/10.1080/14772019.2012.691907" volume="11" year="2013 a">
|
||
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
|
||
2013a
|
||
</bibRefCitation>
|
||
, figs 1F, 2E, 4H, 6B, 12C). Epandrium anterodorsally with 3 pairs of long erect setae (the most lateral markedly shorter), otherwise with scattered short setae. Anal fissure relatively small and cercus reduced (small and narrow) with fine short pubescence (Fig.
|
||
<figureCitation captionStart="Figures 5–9" captionStartId="F4" captionText="Figures 5 - 9. Protanthomyza grimaldii sp. nov., holotype male (Baltic amber) 5 head, left sublateral view 6 base of left wing 7 right mesopleuron, sternopleuron and pteropleuron, lateral view 8 abdomen and haltere, right lateral view 9 terminalia, right lateral view. Scale bars: 0.5 mm (5); 0.3 mm (6 - 8); 0.1 mm (9). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures5-9" httpUri="https://binary.pensoft.net/fig/459228" pageId="0" pageNumber="1">9</figureCitation>
|
||
). Gonostylus not discernible (on any side) but probably small and pale. No structures of internal genitalia visible.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 10, 11" captionStartId="F5" captionText="Figures 10, 11. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 10 entire specimen, subdorsal view 11 ditto, lateroventral view. Body length ca 2.9 mm. Photographs by J. Rohacek (adapted from Rohacek 2013 a, fig. 7 E, F)." figureDoi="10.3897/zookeys.973.51435.figures10-11" httpUri="https://binary.pensoft.net/fig/459229" pageId="0" pageNumber="1">10</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 10, 11" captionStartId="F5" captionText="Figures 10, 11. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 10 entire specimen, subdorsal view 11 ditto, lateroventral view. Body length ca 2.9 mm. Photographs by J. Rohacek (adapted from Rohacek 2013 a, fig. 7 E, F)." figureDoi="10.3897/zookeys.973.51435.figures10-11" httpUri="https://binary.pensoft.net/fig/459229" pageId="0" pageNumber="1">11</figureCitation>
|
||
). Similar to male unless mentioned otherwise. Total body length ca 2.9 mm.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">pvt</emphasis>
|
||
short, strongly convergent, with apices meeting medially;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">oc</emphasis>
|
||
proclinate and divergent; setae in posteroventral corner of postgena subequal. Palpus with more setulae (5 or 6) visible ventrally, subapically and apically. Mesonotum with 3 strong
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dc</emphasis>
|
||
in left row and with 4
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dc</emphasis>
|
||
(2 posterior strong, 2 anterior short and weaker) in right row; prescutellar
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ac</emphasis>
|
||
macrosetae longer than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">sa</emphasis>
|
||
; about 8 rows of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">ac</emphasis>
|
||
microsetae on suture. Legs more slender but with same chaetotaxy as in male, except t2 (Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">14</figureCitation>
|
||
) with ventroapical seta longer. Wing (Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">12</figureCitation>
|
||
) venation resembling that of male but cell
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm</emphasis>
|
||
shorter and cross-vein r-m situated more distally. Wing measurements: length 2.58 mm, width 0.95 mm,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cs3</emphasis>
|
||
:
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cs4</emphasis>
|
||
= 1.44,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">rm\dm-cu</emphasis>
|
||
:
|
||
<emphasis italics="true" pageId="0" pageNumber="1">dm-cu</emphasis>
|
||
= 2.37,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">rm\dm-cu</emphasis>
|
||
:
|
||
<emphasis italics="true" pageId="0" pageNumber="1">CuA1</emphasis>
|
||
= 1.97.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.973.51435.figures10-11" httpUri="https://binary.pensoft.net/fig/459229" pageId="0" pageNumber="1" start="Figures 10, 11" startId="F5">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figures 10, 11.</emphasis>
|
||
<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., female paratype (Baltic amber)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">10</emphasis>
|
||
entire specimen, subdorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">11</emphasis>
|
||
ditto, lateroventral view. Body length ca 2.9 mm. Photographs by J.
|
||
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
|
||
(adapted from
|
||
<bibRefCitation DOI="https://doi.org/10.1080/14772019.2012.691907" author="Rohacek, J" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="431 - 473" refId="B10" refString="Rohacek, J, 2013a. New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity. Journal of Systematic Palaeontology 11: 431 - 473, DOI: https://doi.org/10.1080/14772019.2012.691907" title="New amber fossil Anthomyzidae (Diptera): an unexpected Eocene diversity." url="https://doi.org/10.1080/14772019.2012.691907" volume="11" year="2013 a">
|
||
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
|
||
2013a
|
||
</bibRefCitation>
|
||
, fig. 7E, F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 10, 11" captionStartId="F5" captionText="Figures 10, 11. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 10 entire specimen, subdorsal view 11 ditto, lateroventral view. Body length ca 2.9 mm. Photographs by J. Rohacek (adapted from Rohacek 2013 a, fig. 7 E, F)." figureDoi="10.3897/zookeys.973.51435.figures10-11" httpUri="https://binary.pensoft.net/fig/459229" pageId="0" pageNumber="1">11</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">13</figureCitation>
|
||
) only partly visible. Preabdomen with terga (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T1+2, T3-T5</emphasis>
|
||
) obviously darker brown than sterna, relatively narrow, hence pleural part of preabdominal segments large; setae on
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T3-T5</emphasis>
|
||
(and also on
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T6</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T7</emphasis>
|
||
) longer than those on adjacent sterna. Preabdominal sterna pale brown to ochreous.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S1-S5</emphasis>
|
||
becoming distinctly wider posteriorly,
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S5</emphasis>
|
||
widest and largest.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postabdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T6</emphasis>
|
||
distinctly shorter than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T5</emphasis>
|
||
, transverse;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T7</emphasis>
|
||
yet shorter and also narrower than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T6</emphasis>
|
||
.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S6</emphasis>
|
||
strikingly broad and transverse, wider but shorter than
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S5</emphasis>
|
||
and apparently larger than adjacent
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T6</emphasis>
|
||
and laterally almost reaching the latter;
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S7</emphasis>
|
||
only half length of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S6</emphasis>
|
||
, strongly transverse and laterally meeting with sides of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T7</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">13</figureCitation>
|
||
); both
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S6</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S7</emphasis>
|
||
finely setulose. Apex of postabdomen obscured, only
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T8</emphasis>
|
||
discernible as small bare(?) sclerite (Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F6" captionText="Figures 12 - 14. Protanthomyza grimaldii sp. nov., female paratype (Baltic amber) 12 left wing (invisible parts omitted) 13 abdomen, right lateroventral view 14 right f 2, t 2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (12), 0.3 mm (13, 14). For abbreviations see p. 4." figureDoi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1">13</figureCitation>
|
||
), less than half length of
|
||
<emphasis italics="true" pageId="0" pageNumber="1">T7</emphasis>
|
||
. Cercus not visible.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.973.51435.figures12-14" httpUri="https://binary.pensoft.net/fig/459230" pageId="0" pageNumber="1" start="Figures 12–14" startId="F6">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figures 12-14.</emphasis>
|
||
<taxonomicName authorityName="Roháček" authorityYear="2020" class="Insecta" family="Anthomyzidae" genus="Protanthomyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanthomyza grimaldii" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Protanthomyza grimaldii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., female paratype (Baltic amber)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">12</emphasis>
|
||
left wing (invisible parts omitted)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">13</emphasis>
|
||
abdomen, right lateroventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">14</emphasis>
|
||
right f2, t2 and mid basitarsus, anterior view. Scale bars: 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">12</emphasis>
|
||
), 0.3 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">13</emphasis>
|
||
,
|
||
<emphasis bold="true" pageId="0" pageNumber="1">14</emphasis>
|
||
). For abbreviations see p. 4.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |