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<document id="175E5F228E0516F86BE14E7BB1603787" ID-DOI="10.11646/zootaxa.5319.3.9" ID-ISSN="1175-5326" ID-ZooBank="692A03A9-75CB-402F-9060-6A779E88782B" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1690882982572" checkinUser="plazi" docAuthor="Borkenhagen, Kai &amp; Freyhof, Jörg" docDate="2023" docId="E511CE49FFEEFFF17FC7972A709CF904" docLanguage="en" docName="zootaxa.5319.3.9.pdf" docOrigin="Zootaxa 5319 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.3.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atlantor Borkenhagen &amp; Freyhof 2023, new genus" docType="treatment" docVersion="2" lastPageNumber="432" masterDocId="1928B631FFECFFF27F5095417179FFAE" masterDocTitle="Atlantor, a new generic name for Barbus reinii Günther, 1874 from Morocco (Teleostei: Cyprinidae)" masterLastPageNumber="434" masterPageNumber="429" pageNumber="431" updateTime="1691003432666" updateUser="tatiana" zenodo-license-document="CLOSED">
<mods:mods id="C61D5CDB9C7C457326380E2D4B63F228" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9507F755366BFFFEF34BB5B27BD1C7D1">
<mods:title id="DA64C3836CAB6CB6C858C17EB472F59A">Atlantor, a new generic name for Barbus reinii Günther, 1874 from Morocco (Teleostei: Cyprinidae)</mods:title>
</mods:titleInfo>
<mods:name id="3253C3E6A832342A73BE6068CE15CA15" type="personal">
<mods:role id="D4BBECBA404EC3EEFD247EB7A0829D60">
<mods:roleTerm id="1DC941264AED8B235E25343FFE4CF0C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E45E6072ED45E57DBDD9BCF558FE934C">Borkenhagen, Kai</mods:namePart>
<mods:affiliation id="16287EC147E72AA41A30ECA5065E345B">Federation of German Avifaunists, 48157 M ̧ nster, Germany. &amp; Senckenberg Research Institute and Museum of Nature, Senckenberganlage 25, 60325 Frankfurt am Main, Germany.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="FDD8EE9F2292F3CE280978F17A8354B6">Freyhof, Jörg</mods:namePart>
<mods:nameIdentifier id="D1499B7A20E1CE27BC28A3D3037E7E60" type="ORCID">/0000-0002-7042-312</mods:nameIdentifier>
<mods:affiliation id="B23E10DABCDEA5308A85C6EF20BD8088">Museum f ̧ r Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, 10115 Berlin, Germany. joerg. freyhof @ mfn. berlin; https: // orcid. org / 0000 - 0002 - 7042 - 3127</mods:affiliation>
<mods:nameIdentifier id="21B46AC6E0E09AD71504DB2DC89605CD" type="email">joerg.freyhof@mfn.berli</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="2BAB545A8053EF29E290962AFEBF6E13">text</mods:typeOfResource>
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<mods:title id="53B20AB9FA67ED4FAE145041E3119811">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E595BA0FB24DB9875697DDB786C8E52B">
<mods:date id="F0E5ED3F76457C6FE1275161179469FF">2023</mods:date>
<mods:detail id="5B7AFD6BB3EC4FA20EFE1D58D5201BE1" type="pubDate">
<mods:number id="E9DC7FBBCD2D8A033DAF69A87F1C8171">2023-07-26</mods:number>
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<mods:detail id="485AF551948B5FFB5E2F39BD3857D677" type="volume">
<mods:number id="C6C69B46D6741D83C5B7B381E55B62AA">5319</mods:number>
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<mods:number id="47215B8744C541D64780E486A70AAA74">3</mods:number>
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<mods:start id="F80094765832F972225BAD8BCD116FA9">429</mods:start>
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<mods:url id="04B28EE514741A8DC76F6409A7E2D4B2">http://dx.doi.org/10.11646/zootaxa.5319.3.9</mods:url>
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<mods:classification id="7EA45000221E52A3F6D3E1A98F8691D9">journal article</mods:classification>
<mods:identifier id="7C382144466284EB0A266E1DF9386D63" type="DOI">10.11646/zootaxa.5319.3.9</mods:identifier>
<mods:identifier id="BFBA53E17670DC04A567B89AD1964FD3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="569858FD02D2532BE3DF389151CB44BF" type="ZooBank">692A03A9-75CB-402F-9060-6A779E88782B</mods:identifier>
</mods:mods>
<treatment id="E511CE49FFEEFFF17FC7972A709CF904" LSID="urn:lsid:plazi:treatment:E511CE49FFEEFFF17FC7972A709CF904" httpUri="http://treatment.plazi.org/id/E511CE49FFEEFFF17FC7972A709CF904" lastPageId="3" lastPageNumber="432" pageId="2" pageNumber="431">
<subSubSection id="25A22CD4FFEEFFF07FC7972A70F4FD2B" box="[151,397,619,646]" pageId="2" pageNumber="431" type="nomenclature">
<paragraph id="6D077F5FFFEEFFF07FC7972A70F4FD2B" blockId="2.[151,397,619,681]" box="[151,397,619,646]" pageId="2" pageNumber="431">
<heading id="364FC833FFEEFFF07FC7972A70F4FD2B" bold="true" box="[151,397,619,646]" fontSize="11" level="1" pageId="2" pageNumber="431" reason="1">
<emphasis id="5FCCA34DFFEEFFF07FC7972A70F4FD2B" bold="true" box="[151,397,619,646]" pageId="2" pageNumber="431">
<taxonomicName id="AAB804DCFFEEFFF07FC7972A7185FD28" authority="Borkenhagen &amp; Freyhof, 2023" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[151,252,619,646]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus" status="new genus">
<emphasis id="5FCCA34DFFEEFFF07FC7972A7185FD28" bold="true" box="[151,252,619,646]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="44FF1E36FFEEFFF07E5B972A70F4FD2B" box="[267,397,619,645]" pageId="2" pageNumber="431" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25A22CD4FFEEFFF07FC797CE7181FD07" box="[151,248,655,681]" pageId="2" pageNumber="431" type="description">
<paragraph id="6D077F5FFFEEFFF07FC797CE7181FD07" blockId="2.[151,397,619,681]" box="[151,248,655,681]" pageId="2" pageNumber="431">
<figureCitation id="F58363DAFFEEFFF07FC797CE7181FD07" box="[151,248,655,681]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,741,765]" captionStartId-1="3.[151,250,1574,1598]" captionTargetBox-0="[152,1436,179,715]" captionTargetBox-1="[152,1436,790,1550]" captionTargetId-0="figure-26@3.[152,1436,179,715]" captionTargetId-1="figure-52@3.[152,1436,790,1550]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionText-0="FIGURE 1. Atlantor reinii, BMNH 1901.7.26.2-3, 1, 146 mm SL; Oued Oum er Rbia, Morocco. Photo: S. Tränkner." captionText-1="FIGURE 2. Atlantor reinii, BMNH 1901.7.26.2-3, 1, 147 mm SL; Oued Oum er Rbia, Morocco. Photo: S. Tränkner." pageId="2" pageNumber="431">Figs 1–2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25A22CD4FFEEFFF07FC797967219FC67" pageId="2" pageNumber="431" type="materials_examined">
<paragraph id="6D077F5FFFEEFFF07FC797967219FC67" blockId="2.[151,1437,727,2014]" pageId="2" pageNumber="431">
<emphasis id="5FCCA34DFFEEFFF07FC7979670FBFD5F" bold="true" box="[151,386,727,753]" pageId="2" pageNumber="431">Material examined.</emphasis>
<materialsCitation id="DDD07502FFEEFFF07EDA979973FAFCBB" collectionCode="BMNH" collectorName="K. v. Fritsch &amp; J. Rein" country="Morocco" latitude="32.032776" location="Oued Tensift" longLatPrecision="20" longitude="-9.344167" pageId="2" pageNumber="431" specimenCount="1" typeStatus="syntype">
<collectionCode id="0BA9E79AFFEEFFF07EDA979970A7FD5C" box="[394,478,728,754]" pageId="2" pageNumber="431">BMNH</collectionCode>
1874.1.30.22-24, 3,
<typeStatus id="B203C1FDFFEEFFF07D9C97997248FD5C" box="[716,817,728,754]" pageId="2" pageNumber="431">syntypes</typeStatus>
,
<quantity id="AA40D2BAFFEEFFF07C6D979672BCFD5C" box="[829,965,727,754]" metricMagnitude="-1" metricUnit="m" metricValue="1.255" metricValueMax="1.55" metricValueMin="0.96" pageId="2" pageNumber="431" unit="mm" value="125.5" valueMax="155.0" valueMin="96.0">96–155 mm</quantity>
SL;
<collectingCountry id="15AF3FCFFFEEFFF07CAE97997511FD5C" box="[1022,1128,728,754]" name="Morocco" pageId="2" pageNumber="431">Morocco</collectingCountry>
:
<location id="68672984FFEEFFF07B2397967471FD5F" LSID="urn:lsid:plazi:treatment:E511CE49FFEEFFF17FC7972A709CF904:68672984FFEEFFF07B2397967471FD5F" box="[1139,1288,727,753]" country="Morocco" latitude="32.032776" longLatPrecision="20" longitude="-9.344167" name="Oued Tensift" pageId="2" pageNumber="431">Oued Tensift</location>
, [
<geoCoordinate id="088C1998FFEEFFF07A4C979874EEFD5A" box="[1308,1431,729,756]" degrees="32" direction="north" minutes="1" orientation="latitude" pageId="2" pageNumber="431" precision="15" seconds="58" value="32.032776">32°1′58″N</geoCoordinate>
,
<geoCoordinate id="088C1998FFEEFFF07FC797BC7062FCB6" box="[151,283,765,792]" degrees="9" direction="west" minutes="20" orientation="longitude" pageId="2" pageNumber="431" precision="15" seconds="39" value="-9.344167">9°20′39″W</geoCoordinate>
];
<collectorName id="C04D1A89FFEEFFF07E6097BD70B8FCBB" box="[304,449,763,790]" pageId="2" pageNumber="431">K. v. Fritsch</collectorName>
&amp;
<collectorName id="C04D1A89FFEEFFF07EBA97BD7347FCBB" box="[490,574,763,790]" pageId="2" pageNumber="431">J. Rein</collectorName>
, 1872
</materialsCitation>
.—
<materialsCitation id="DDD07502FFEEFFF07DF597BA73E0FC94" collectionCode="BMNH" collectorName="E. Hartert" country="Morocco" latitude="33.327778" location="Oued Oum" longLatPrecision="20" longitude="-8.333889" municipality="Rbia" pageId="2" pageNumber="431" specimenCount="1" typeStatus="syntype">
<collectionCode id="0BA9E79AFFEEFFF07DF597BA7385FCBB" box="[677,764,763,789]" pageId="2" pageNumber="431">BMNH</collectionCode>
1901.7.26.2-3, 2,
<quantity id="AA40D2BAFFEEFFF07C8297BA7513FCB8" box="[978,1130,763,790]" metricMagnitude="-1" metricUnit="m" metricValue="1.355" metricValueMax="1.46" metricValueMin="1.25" pageId="2" pageNumber="431" unit="mm" value="135.5" valueMax="146.0" valueMin="125.0">125–146 mm</quantity>
SL;
<collectingCountry id="15AF3FCFFFEEFFF07BF697BD7469FCB8" box="[1190,1296,764,790]" name="Morocco" pageId="2" pageNumber="431">Morocco</collectingCountry>
:
<location id="68672984FFEEFFF07A4D97BA74E5FCBB" LSID="urn:lsid:plazi:treatment:E511CE49FFEEFFF17FC7972A709CF904:68672984FFEEFFF07A4D97BA74E5FCBB" box="[1309,1436,763,789]" country="Morocco" latitude="33.327778" longLatPrecision="20" longitude="-8.333889" municipality="Rbia" name="Oued Oum" pageId="2" pageNumber="431">Oued Oum</location>
er
<collectingMunicipality id="8D63E525FFEEFFF07FE8965E7194FC97" box="[184,237,799,825]" pageId="2" pageNumber="431">Rbia</collectingMunicipality>
[
<geoCoordinate id="088C1998FFEEFFF07FAF966070F0FC92" box="[255,393,801,828]" degrees="33" direction="north" minutes="19" orientation="latitude" pageId="2" pageNumber="431" precision="15" seconds="40" value="33.327778">33°19′40″N</geoCoordinate>
,
<geoCoordinate id="088C1998FFEEFFF07EC996607377FC92" box="[409,526,801,828]" degrees="8" direction="west" minutes="20" orientation="longitude" pageId="2" pageNumber="431" precision="15" seconds="2" value="-8.333889">8°20′2″W</geoCoordinate>
];
<collectorName id="C04D1A89FFEEFFF07D75966173E0FC94" box="[549,665,800,826]" pageId="2" pageNumber="431">E. Hartert</collectorName>
</materialsCitation>
.—
<materialsCitation id="DDD07502FFEEFFF07DE996617542FC97" box="[697,1083,799,826]" collectionCode="MNHN" country="Morocco" pageId="2" pageNumber="431" specimenCode="MNHN 1912-94" specimenCount="1" typeStatus="syntype">
<specimenCode id="3D1ED724FFEEFFF07DE996617205FC97" box="[697,892,799,826]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="431">MNHN 1912-94</specimenCode>
,
<quantity id="AA40D2BAFFEEFFF07CDB965E7575FC94" box="[907,1036,799,826]" metricMagnitude="0" metricUnit="m" metricValue="1.1199999999999999" pageId="2" pageNumber="431" unit="mm" value="1120.0">1, 120 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="DDD07502FFEEFFF07B1B966171C2FCF3" collectionCode="MNHN" country="Morocco" pageId="2" pageNumber="431" specimenCode="MNHN 1912-95" specimenCount="1" typeStatus="syntype">
<specimenCode id="3D1ED724FFEEFFF07B1B96617475FC94" box="[1099,1292,799,826]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="431">MNHN 1912-95</specimenCode>
,
<quantity id="AA40D2BAFFEEFFF07A4B965E74E4FC94" box="[1307,1437,799,826]" metricMagnitude="0" metricUnit="m" metricValue="1.124" pageId="2" pageNumber="431" unit="mm" value="1124.0">1, 124 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="DDD07502FFEEFFF07F9A96057022FC2F" collectionCode="MNHN" collectorName="C. du Gast" country="Morocco" latitude="33.289444" location="Oued Oum" longLatPrecision="20" longitude="-8.3425" municipality="Rbia" pageId="2" pageNumber="431" specimenCode="MNHN 1912-96, 1" specimenCount="1" typeStatus="syntype">
<specimenCode id="3D1ED724FFEEFFF07F9A960570F2FCF3" box="[202,395,835,862]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="431">MNHN 1912-96</specimenCode>
,
<specimenCode id="3D1ED724FFEEFFF07ECA960270D1FCF3" box="[410,424,835,861]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="431">1</specimenCode>
,
<quantity id="AA40D2BAFFEEFFF07EEA96027375FCF0" box="[442,524,835,862]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="431" unit="mm" value="92.0">92 mm</quantity>
SL;
<collectingCountry id="15AF3FCFFFEEFFF07D1A960573CDFCF0" box="[586,692,836,862]" name="Morocco" pageId="2" pageNumber="431">Morocco</collectingCountry>
:
<location id="68672984FFEEFFF07D9296027238FCF3" LSID="urn:lsid:plazi:treatment:E511CE49FFEEFFF17FC7972A709CF904:68672984FFEEFFF07D9296027238FCF3" box="[706,833,835,861]" country="Morocco" latitude="33.289444" longLatPrecision="20" longitude="-8.3425" municipality="Rbia" name="Oued Oum" pageId="2" pageNumber="431">Oued Oum</location>
er
<collectingMunicipality id="8D63E525FFEEFFF07C3C960272D8FCF3" box="[876,929,835,861]" pageId="2" pageNumber="431">Rbia</collectingMunicipality>
near Azemmour [
<geoCoordinate id="088C1998FFEEFFF07B2596047479FCCE" box="[1141,1280,837,864]" degrees="33" direction="north" minutes="17" orientation="latitude" pageId="2" pageNumber="431" precision="15" seconds="22" value="33.289444">33°17′22″N</geoCoordinate>
,
<geoCoordinate id="088C1998FFEEFFF07A5F960474E8FCCE" box="[1295,1425,837,864]" degrees="8" direction="west" minutes="20" orientation="longitude" pageId="2" pageNumber="431" precision="15" seconds="33" value="-8.3425">8°20′33″W</geoCoordinate>
],
<collectorName id="C04D1A89FFEEFFF07FC79626706FFC2F" box="[151,278,871,897]" pageId="2" pageNumber="431">C. du Gast</collectorName>
, 1912
</materialsCitation>
.—
<materialsCitation id="DDD07502FFEEFFF07E2C962670F4FC0B" collectionCode="MNHN" collectorName="A. Gruvel" country="Morocco" county="Khenifra" latitude="32.939167" location="Oued Oum" longLatPrecision="20" longitude="-5.668611" municipality="Rbia" pageId="2" pageNumber="431" specimenCode="MNHN 1927-98" specimenCount="1" typeStatus="syntype">
<specimenCode id="3D1ED724FFEEFFF07E2C96267347FC2F" box="[380,574,871,897]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="431">MNHN 1927-98</specimenCode>
,
<quantity id="AA40D2BAFFEEFFF07D1C962673B1FC2C" box="[588,712,871,898]" metricMagnitude="0" metricUnit="m" metricValue="1.111" pageId="2" pageNumber="431" unit="mm" value="1111.0">1, 111 mm</quantity>
SL;
<collectingCountry id="15AF3FCFFFEEFFF07C5396297214FC2C" box="[771,877,872,898]" name="Morocco" pageId="2" pageNumber="431">Morocco</collectingCountry>
:
<location id="68672984FFEEFFF07C2A96267281FC2F" LSID="urn:lsid:plazi:treatment:E511CE49FFEEFFF17FC7972A709CF904:68672984FFEEFFF07C2A96267281FC2F" box="[890,1016,871,897]" country="Morocco" county="Khenifra" latitude="32.939167" longLatPrecision="20" longitude="-5.668611" municipality="Rbia" name="Oued Oum" pageId="2" pageNumber="431">Oued Oum</location>
er
<collectingMunicipality id="8D63E525FFEEFFF07B719626752FFC2F" box="[1057,1110,871,897]" pageId="2" pageNumber="431">Rbia</collectingMunicipality>
near
<collectingCounty id="846607D3FFEEFFF07BC996267584FC2F" box="[1177,1277,871,897]" pageId="2" pageNumber="431">Khenifra</collectingCounty>
[
<geoCoordinate id="088C1998FFEEFFF07A5D962874EEFC2A" box="[1293,1431,873,900]" degrees="32" direction="north" minutes="56" orientation="latitude" pageId="2" pageNumber="431" precision="15" seconds="21" value="32.939167">32°56′21″N</geoCoordinate>
,
<geoCoordinate id="088C1998FFEEFFF07FC796CC7075FC06" box="[151,268,909,936]" degrees="5" direction="west" minutes="40" orientation="longitude" pageId="2" pageNumber="431" precision="15" seconds="7" value="-5.668611">5°40′7″W</geoCoordinate>
],
<collectorName id="C04D1A89FFEEFFF07E4D96CA70F4FC0B" box="[285,397,907,933]" pageId="2" pageNumber="431">A. Gruvel</collectorName>
</materialsCitation>
&amp;
<materialsCitation id="DDD07502FFEEFFF07EE196CD7312FC0B" box="[433,619,907,934]" collectionCode="R" country="Morocco" location="Dollfus" pageId="2" pageNumber="431" specimenCount="1" typeStatus="syntype">
<collectionCode id="0BA9E79AFFEEFFF07EE196CD70BEFC08" box="[433,455,908,934]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="431">R</collectionCode>
.
<location id="68672984FFEEFFF07E8296CA7350FC0B" LSID="urn:lsid:plazi:treatment:E511CE49FFEEFFF17FC7972A709CF904:68672984FFEEFFF07E8296CA7350FC0B" box="[466,553,907,933]" country="Morocco" name="Dollfus" pageId="2" pageNumber="431">Dollfus</location>
, 1927
</materialsCitation>
.—
<materialsCitation id="DDD07502FFEEFFF07DDD96CA72E4FC0B" box="[653,925,907,934]" collectionCode="SMF" country="Morocco" pageId="2" pageNumber="431" specimenCode="SMF 636, 1" specimenCount="1" typeStatus="syntype">
<specimenCode id="3D1ED724FFEEFFF07DDD96CA7382FC0B" box="[653,763,907,933]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="431">SMF 636</specimenCode>
,
<specimenCode id="3D1ED724FFEEFFF07C5696CA726DFC0B" box="[774,788,907,933]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="431">1</specimenCode>
,
<quantity id="AA40D2BAFFEEFFF07C7396CA720BFC08" box="[803,882,907,934]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="431" unit="mm" value="82.0">82 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="DDD07502FFEEFFF07CF896CA7222FC67" collectionCode="SMF" collectorName="K. v. Fritsch &amp; J. Rein" country="Morocco" latitude="31.483055" location="Oued Ksob" longLatPrecision="20" longitude="-9.7675" pageId="2" pageNumber="431" specimenCode="SMF 952, 2" specimenCount="1" typeStatus="syntype">
<specimenCode id="3D1ED724FFEEFFF07CF896CA756DFC08" box="[936,1044,907,934]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="431">SMF 952</specimenCode>
,
<specimenCode id="3D1ED724FFEEFFF07B7096CA7549FC0B" box="[1056,1072,907,933]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="431">2</specimenCode>
,
<quantity id="AA40D2BAFFEEFFF07B6C96CA75CCFC08" box="[1084,1205,907,934]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" metricValueMax="8.3" metricValueMin="4.9" pageId="2" pageNumber="431" unit="mm" value="66.0" valueMax="83.0" valueMin="49.0">49–83 mm</quantity>
SL;
<collectingCountry id="15AF3FCFFFEEFFF07BBC96CD742FFC08" box="[1260,1366,908,934]" name="Morocco" pageId="2" pageNumber="431">Morocco</collectingCountry>
:
<location id="68672984FFEEFFF07A3096CA71ABFC67" LSID="urn:lsid:plazi:treatment:E511CE49FFEEFFF17FC7972A709CF904:68672984FFEEFFF07A3096CA71ABFC67" country="Morocco" latitude="31.483055" longLatPrecision="20" longitude="-9.7675" name="Oued Ksob" pageId="2" pageNumber="431">Oued Ksob</location>
[
<geoCoordinate id="088C1998FFEEFFF07FB096F07012FC62" box="[224,363,945,972]" degrees="31" direction="north" minutes="28" orientation="latitude" pageId="2" pageNumber="431" precision="15" seconds="59" value="31.483055">31°28′59″N</geoCoordinate>
, W
<geoCoordinate id="088C1998FFEEFFF07EC396F0737DFC62" box="[403,516,945,972]" degrees="9" direction="west" minutes="46" orientation="longitude" pageId="2" pageNumber="431" precision="15" seconds="3" value="-9.7675">9°46′3″W</geoCoordinate>
],
<collectorName id="C04D1A89FFEEFFF07D4796F173D8FC67" box="[535,673,943,970]" pageId="2" pageNumber="431">K. v. Fritsch</collectorName>
&amp;
<collectorName id="C04D1A89FFEEFFF07D9496F1726CFC67" box="[708,789,943,970]" pageId="2" pageNumber="431">J. Rein</collectorName>
, 1872
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="25A22CD4FFEEFFF07F97969272A0FA30" pageId="2" pageNumber="431" type="diagnosis">
<paragraph id="6D077F5FFFEEFFF07F97969272A0FA30" blockId="2.[151,1437,727,2014]" pageId="2" pageNumber="431">
<emphasis id="5FCCA34DFFEEFFF07F979692703BFC43" bold="true" box="[199,322,979,1005]" pageId="2" pageNumber="431">Diagnosis.</emphasis>
<taxonomicName id="AAB804DCFFEEFFF07E1B969270D1FC43" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[331,424,979,1005]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07E1B969270D1FC43" box="[331,424,979,1005]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="AAB804DCFFEEFFF07DFC96927235FC43" authorityName="Karaman" authorityYear="1971" box="[684,844,979,1005]" family="Cyprinidae" genus="Carasobarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07DFC96927235FC43" box="[684,844,979,1005]" italics="true" pageId="2" pageNumber="431">Carasobarbus</emphasis>
</taxonomicName>
by having 5½ branched anal-fin rays (vs. 6½), and modally 8½ branched dorsal-fin ray (vs. 9½–10½), and from
<taxonomicName id="AAB804DCFFEEFFF07C6196B6757AFBBF" box="[817,1027,1015,1041]" family="Cyprinidae" genus="Mesopotamichthys" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07C6196B6757AFBBF" box="[817,1027,1015,1041]" italics="true" pageId="2" pageNumber="431">Mesopotamichthys</emphasis>
</taxonomicName>
and
<taxonomicName id="AAB804DCFFEEFFF07B6596B975B8FBBC" authorityName="Banister &amp; Bunni" authorityYear="1980" box="[1077,1217,1016,1042]" family="Cyprinidae" genus="Caecocypris" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07B6596B975B8FBBC" box="[1077,1217,1016,1042]" italics="true" pageId="2" pageNumber="431">Caecocypris</emphasis>
</taxonomicName>
by having two pairs of long barbels (vs. no barbels). There is no consistent morphological character that clearly distinguishes
<taxonomicName id="AAB804DCFFEEFFF07A6F915A74E5FB9B" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[1343,1436,1051,1077]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07A6F915A74E5FB9B" box="[1343,1436,1051,1077]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
from all species of
<taxonomicName id="AAB804DCFFEEFFF07E3E917E7085FBF7" authorityName="Borkenhagen" authorityYear="2014" box="[366,508,1087,1113]" family="Cyprinidae" genus="Arabibarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07E3E917E7085FBF7" box="[366,508,1087,1113]" italics="true" pageId="2" pageNumber="431">Arabibarbus</emphasis>
</taxonomicName>
and
<taxonomicName id="AAB804DCFFEEFFF07D63917E73BEFBF7" baseAuthorityName="Borkenhagen" baseAuthorityYear="2014" box="[563,711,1087,1113]" family="Cyprinidae" genus="Labeobarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07D63917E73BEFBF7" box="[563,711,1087,1113]" italics="true" pageId="2" pageNumber="431">Labeobarbus</emphasis>
</taxonomicName>
, and the new genus can only be distinguished from
<taxonomicName id="AAB804DCFFEEFFF07A5F917E74E4FBF7" authorityName="Borkenhagen" authorityYear="2014" box="[1295,1437,1087,1113]" family="Cyprinidae" genus="Arabibarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07A5F917E74E4FBF7" box="[1295,1437,1087,1113]" italics="true" pageId="2" pageNumber="431">Arabibarbus</emphasis>
</taxonomicName>
and
<taxonomicName id="AAB804DCFFEEFFF07F9991227024FBD3" baseAuthorityName="Borkenhagen" baseAuthorityYear="2014" box="[201,349,1123,1149]" family="Cyprinidae" genus="Labeobarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07F9991227024FBD3" box="[201,349,1123,1149]" italics="true" pageId="2" pageNumber="431">Labeobarbus</emphasis>
</taxonomicName>
by its distinct phylogenetic position and genetic distance (p-distance in cytochrome
<emphasis id="5FCCA34DFFEEFFF07A7A91227441FBD3" box="[1322,1336,1123,1149]" italics="true" pageId="2" pageNumber="431">b</emphasis>
of 5.3% and 6.7% respectively; Kai Borkenhagen, unpublished).
<taxonomicName id="AAB804DCFFEEFFF07C4191C67217FB0F" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[785,878,1159,1185]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07C4191C67217FB0F" box="[785,878,1159,1185]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
differs from
<taxonomicName id="AAB804DCFFEEFFF07B5391C975E0FB0F" authorityName="Günther" authorityYear="1902" box="[1027,1177,1160,1185]" family="Cyprinidae" genus="Pterocapoeta" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07B5391C975E0FB0F" box="[1027,1177,1160,1185]" italics="true" pageId="2" pageNumber="431">Pterocapoeta</emphasis>
</taxonomicName>
by having two pairs of long barbels (vs. one pair of minute maxillary barbels hidden in a skin fold), dorsal fin with 8½–9½ branched rays (vs. 10½–11½), 25–32 total lateral line scales (vs. 43–46), well developed, fleshy lips and a small median lobe on the lower lip (vs. lower lip with cornified sheath), mouth arched (vs. straight). Current knowledge indicates that all Western Asian, and African Torinae are evolutionarily hexaploid, and as
<taxonomicName id="AAB804DCFFEEFFF07CA79056752DFA9F" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[1015,1108,1303,1329]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07CA79056752DFA9F" box="[1015,1108,1303,1329]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
belongs to this phylogenetic group of species, it is assumed, that they are also hexaploid. As a consequence,
<taxonomicName id="AAB804DCFFEEFFF07B4F907A7505FAFB" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[1055,1148,1339,1365]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07B4F907A7505FAFB" box="[1055,1148,1339,1365]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
is distinguished from the Indomalayan genera
<taxonomicName id="AAB804DCFFEEFFF07ED2901E7355FAD7" box="[386,556,1375,1401]" family="Cyprinidae" genus="Hypselobarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07ED2901E7355FAD7" box="[386,556,1375,1401]" italics="true" pageId="2" pageNumber="431">Hypselobarbus</emphasis>
</taxonomicName>
,
<taxonomicName id="AAB804DCFFEEFFF07D6A901E7393FAD7" authorityName="Raj" authorityYear="1941" box="[570,746,1375,1401]" family="Cyprinidae" genus="Lepidopygopsis" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07D6A901E7393FAD7" box="[570,746,1375,1401]" italics="true" pageId="2" pageNumber="431">Lepidopygopsis</emphasis>
</taxonomicName>
,
<taxonomicName id="AAB804DCFFEEFFF07DA790217227FAD7" authorityName="Mirza &amp; Javed" authorityYear="1985" box="[759,862,1376,1401]" family="Cyprinidae" genus="Naziritor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07DA790217227FAD7" box="[759,862,1376,1401]" italics="true" pageId="2" pageNumber="431">Naziritor</emphasis>
</taxonomicName>
,
<taxonomicName id="AAB804DCFFEEFFF07C3C901E7576FAD7" box="[876,1039,1375,1401]" family="Cyprinidae" genus="Neolissochilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07C3C901E7576FAD7" box="[876,1039,1375,1401]" italics="true" pageId="2" pageNumber="431">Neolissochilus</emphasis>
</taxonomicName>
,
<taxonomicName id="AAB804DCFFEEFFF07B4E901E75A0FAD7" authorityName="Hora" authorityYear="1942" box="[1054,1241,1375,1401]" family="Cyprinidae" genus="Osteochilichthys" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07B4E901E75A0FAD7" box="[1054,1241,1375,1401]" italics="true" pageId="2" pageNumber="431">Osteochilichthys</emphasis>
</taxonomicName>
, and from
<taxonomicName id="AAB804DCFFEEFFF07A0390217400FAD7" box="[1363,1401,1376,1401]" family="Cyprinidae" genus="Tor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07A0390217400FAD7" box="[1363,1401,1376,1401]" italics="true" pageId="2" pageNumber="431">Tor</emphasis>
</taxonomicName>
by being evolutionarily hexaploid (vs. tetraploid (2n≈100) (
<bibRefCitation id="092902AEFFEEFFF07C5F90C272B7FA30" author="Yang, L. &amp; Sado, T. &amp; Vincent Hirt, M. &amp; Pasco-Viel, E. &amp; Arunachalam, M. &amp; Li, J. &amp; Wang, X. &amp; Freyhof, J. &amp; Saitoh, K. &amp; Simons, A. M. &amp; Miya, M. &amp; He, S. &amp; Mayden, R. L." box="[783,974,1411,1438]" pageId="2" pageNumber="431" pagination="97 - 116" refId="ref4923" refString="Yang, L., Sado, T., Vincent Hirt, M., Pasco-Viel, E., Arunachalam, M., Li, J., Wang, X., Freyhof, J., Saitoh, K., Simons, A. M., Miya, M., He, S. &amp; Mayden, R. L. (2015) Phylogeny and polyploidy: resolving the classification of cyprinine fishes (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution, 85, 97 - 116. https: // doi. org / 10.1016 / j. ympev. 2015.01.014" type="journal article" year="2015">
Yang
<emphasis id="5FCCA34DFFEEFFF07C0290C572FAFA33" box="[850,899,1411,1437]" italics="true" pageId="2" pageNumber="431">et al</emphasis>
. 2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="25A22CD4FFEEFFF07F9790E6727CFA4B" pageId="2" pageNumber="431" type="type_taxon">
<paragraph id="6D077F5FFFEEFFF07F9790E673A4FA6F" blockId="2.[151,1437,727,2014]" box="[199,733,1447,1473]" pageId="2" pageNumber="431">
<emphasis id="5FCCA34DFFEEFFF07F9790E6701BFA6F" bold="true" box="[199,354,1447,1473]" pageId="2" pageNumber="431">
<typeStatus id="B203C1FDFFEEFFF07F9790E6707BFA6F" box="[199,258,1447,1473]" pageId="2" pageNumber="431">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="AAB804DCFFEEFFF07E3990E673A4FA6F" authority="Gunther, 1874: 231" authorityName="Gunther" authorityYear="1874" box="[361,733,1447,1473]" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="species" species="reinii">
<emphasis id="5FCCA34DFFEEFFF07E3990E67084FA6F" box="[361,509,1447,1473]" italics="true" pageId="2" pageNumber="431">Barbus reinii</emphasis>
<bibRefCitation id="092902AEFFEEFFF07D5490E673A4FA6F" author="Gunther, A." box="[516,733,1447,1473]" pageId="2" pageNumber="431" pagination="230 - 232" refId="ref4079" refString="Gunther, A. (1874) Notice of some new species of fishes from Morocco. Annals and Magazine of Natural History, Series 4, 13 (75), 230 - 232. https: // doi. org / 10.1080 / 00222937408680848" type="journal article" year="1874">Günther, 1874: 231</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="6D077F5FFFEEFFF07F97908A727CFA4B" blockId="2.[151,1437,727,2014]" box="[199,773,1483,1509]" pageId="2" pageNumber="431">
<emphasis id="5FCCA34DFFEEFFF07F97908A70EBFA4B" bold="true" box="[199,402,1483,1509]" pageId="2" pageNumber="431">Included species.</emphasis>
<taxonomicName id="AAB804DCFFEEFFF07EC8908A727CFA4B" authority="Gunther, 1874: 231" authorityName="Gunther" authorityPageNumber="231" authorityYear="1874" box="[408,773,1483,1509]" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="species" species="reinii">
<emphasis id="5FCCA34DFFEEFFF07EC8908A7355FA4B" box="[408,556,1483,1509]" italics="true" pageId="2" pageNumber="431">Barbus reinii</emphasis>
<bibRefCitation id="092902AEFFEEFFF07D63908A727CFA4B" author="Gunther, A." box="[563,773,1483,1509]" pageId="2" pageNumber="431" pagination="230 - 232" refId="ref4079" refString="Gunther, A. (1874) Notice of some new species of fishes from Morocco. Annals and Magazine of Natural History, Series 4, 13 (75), 230 - 232. https: // doi. org / 10.1080 / 00222937408680848" type="journal article" year="1874">Günther, 1874:231</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="25A22CD4FFEEFFF07F9790AE7080F983" pageId="2" pageNumber="431" type="etymology">
<paragraph id="6D077F5FFFEEFFF07F9790AE7080F983" blockId="2.[151,1437,727,2014]" pageId="2" pageNumber="431">
<emphasis id="5FCCA34DFFEEFFF07F9790AE7037F9A7" bold="true" box="[199,334,1519,1545]" pageId="2" pageNumber="431">Etymology.</emphasis>
The name is derived from the Atlas Mountains in
<collectingCountry id="15AF3FCFFFEEFFF07CD890B17294F9A4" box="[904,1005,1520,1546]" name="Morocco" pageId="2" pageNumber="431">Morocco</collectingCountry>
and
<taxonomicName id="AAB804DCFFEEFFF07B7490B17533F9A7" box="[1060,1098,1520,1545]" family="Cyprinidae" genus="Tor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07B7490B17533F9A7" box="[1060,1098,1520,1545]" italics="true" pageId="2" pageNumber="431">Tor</emphasis>
</taxonomicName>
, the generic name of the tribe that includes
<taxonomicName id="AAB804DCFFEEFFF07E7C935270F0F983" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[300,393,1555,1581]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07E7C935270F0F983" box="[300,393,1555,1581]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
(neutral).
</paragraph>
</subSubSection>
<subSubSection id="25A22CD4FFEEFFF07F9793767070F8AB" pageId="2" pageNumber="431" type="distribution">
<paragraph id="6D077F5FFFEEFFF07F9793767070F8AB" blockId="2.[151,1437,727,2014]" pageId="2" pageNumber="431">
<emphasis id="5FCCA34DFFEEFFF07F9793767306F9FF" bold="true" box="[199,639,1591,1617]" pageId="2" pageNumber="431">Distribution and conservation status.</emphasis>
The
<typeStatus id="B203C1FDFFEEFFF07DE893797391F9FC" box="[696,744,1592,1618]" pageId="2" pageNumber="431">type</typeStatus>
locality of
<taxonomicName id="AAB804DCFFEEFFF07C37937672B8F9FF" authorityName="Gunther" authorityYear="1874" box="[871,961,1591,1617]" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="species" species="reinii">
<emphasis id="5FCCA34DFFEEFFF07C37937672B8F9FF" box="[871,961,1591,1617]" italics="true" pageId="2" pageNumber="431">A. reinii</emphasis>
</taxonomicName>
is the Tennsift River in
<collectingCountry id="15AF3FCFFFEEFFF07B9C93797448F9FC" box="[1228,1329,1592,1618]" name="Morocco" pageId="2" pageNumber="431">Morocco</collectingCountry>
(
<bibRefCitation id="092902AEFFEEFFF07A10937671AAF9DB" author="Gunther, A." pageId="2" pageNumber="431" pagination="230 - 232" refId="ref4079" refString="Gunther, A. (1874) Notice of some new species of fishes from Morocco. Annals and Magazine of Natural History, Series 4, 13 (75), 230 - 232. https: // doi. org / 10.1080 / 00222937408680848" type="journal article" year="1874">Günther 1874</bibRefCitation>
). Records from the Oum er Rbia (
<bibRefCitation id="092902AEFFEEFFF07D18931A739DF9DB" author="Gunther, A." box="[584,740,1627,1653]" pageId="2" pageNumber="431" pagination="446 - 448" refId="ref4125" refString="Gunther, A. (1902) Third notice of new species of fishes from Morocco. Novitates Zoologicae. A Journal of Zoology in Connection with the Tring Museum, 9, 446 - 448." type="journal article" year="1902">Günther 1902</bibRefCitation>
,
<bibRefCitation id="092902AEFFEEFFF07DBE931A72E8F9DB" author="Pellegrin, J." box="[750,913,1627,1653]" pageId="2" pageNumber="431" pagination="1" refId="ref4384" refString="Pellegrin, J. (1921) Les poissons des eaux douces de l'Afrique du Nord francaise Maroc (Algerie, Tunisie, Sahara). Memoires de la Societe des Sciences Naturelles du Maroc, 1921, 1." type="journal article" year="1921">Pellegrin 1921</bibRefCitation>
) are confirmed by vouchers (MNHN 1912-94– 96, MNHN 1927-98). In recent years, the species was only found in the Ksob River, a tributary to Oued Igrounzar, by
<bibRefCitation id="092902AEFFEEFFF07FEE93E2735BF913" author="Machordom, A. &amp; Doadrio, I." box="[190,546,1699,1725]" pageId="2" pageNumber="431" pagination="1297 - 1306" refId="ref4336" refString="Machordom, A. &amp; Doadrio, I. (2001) Evolutionary history and speciation modes in the cyprinid genus Barbus. Proceedings of The Royal Society B, 268, 1297 - 1306. https: // doi. org / 10.1098 / rspb. 2001.1654" type="journal article" year="2001">Machordom &amp; Doadrio (2001)</bibRefCitation>
(I. Doadrio, pers. communication, 2019).
<taxonomicName id="AAB804DCFFEEFFF07B4093E275CDF913" box="[1040,1204,1699,1725]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="species" species="reinii">
<emphasis id="5FCCA34DFFEEFFF07B4093E275CDF913" box="[1040,1204,1699,1725]" italics="true" pageId="2" pageNumber="431">Atlantor reinii</emphasis>
</taxonomicName>
has not been found since 2001, despite a number of attempts to confirm its continued existence.
<bibRefCitation id="092902AEFFEEFFF07CAA93867479F94F" author="Freyhof, J. &amp; Ford, M." box="[1018,1280,1735,1761]" pageId="2" pageNumber="431" pagination="137272327" refId="ref3718" refString="Freyhof, J. &amp; Ford, M. (2022) Labeobarbus reinii. The IUCN Red List of Threatened Species 2022: e. T 60765 A 137272327. https: // doi. org / 10.2305 / IUCN. UK. 2022 - 1. RLTS. T 60765 A 137272327. en" type="journal article" year="2022">Freyhof &amp; Ford (2022)</bibRefCitation>
treat
<taxonomicName id="AAB804DCFFEEFFF07A11938674E4F94F" box="[1345,1437,1735,1761]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="species" species="reinii">
<emphasis id="5FCCA34DFFEEFFF07A11938674E4F94F" box="[1345,1437,1735,1761]" italics="true" pageId="2" pageNumber="431">A. reinii</emphasis>
</taxonomicName>
as extinct.
</paragraph>
</subSubSection>
<subSubSection id="25A22CD4FFEEFFF17F97924E709CF904" lastPageId="3" lastPageNumber="432" pageId="2" pageNumber="431" type="discussion">
<paragraph id="6D077F5FFFEEFFF07F97924E7283F870" blockId="2.[151,1437,727,2014]" pageId="2" pageNumber="431">
<emphasis id="5FCCA34DFFEEFFF07F97924E7042F887" bold="true" box="[199,315,1807,1833]" pageId="2" pageNumber="431">Remarks.</emphasis>
There are only two cytochrome
<emphasis id="5FCCA34DFFEEFFF07D9B924E73A0F887" box="[715,729,1807,1833]" italics="true" pageId="2" pageNumber="431">b</emphasis>
sequences available, which were published by
<bibRefCitation id="092902AEFFEEFFF07A43924E701EF8E3" author="Machordom, A. &amp; Doadrio, I." pageId="2" pageNumber="431" pagination="1297 - 1306" refId="ref4336" refString="Machordom, A. &amp; Doadrio, I. (2001) Evolutionary history and speciation modes in the cyprinid genus Barbus. Proceedings of The Royal Society B, 268, 1297 - 1306. https: // doi. org / 10.1098 / rspb. 2001.1654" type="journal article" year="2001">Machordom &amp; Doadrio (2001)</bibRefCitation>
and
<bibRefCitation id="092902AEFFEEFFF07EF4927273ECF8E3" author="Durand, J. - D. &amp; Tsigenopoulos, C. S. &amp; Unlu, E. &amp; Berrebi, P." box="[420,661,1843,1869]" pageId="2" pageNumber="431" pagination="1" refId="ref3556" refString="Durand, J. - D., Tsigenopoulos, C. S., Unlu, E. &amp; Berrebi, P. (2002) Phylogeny and Biogeography of the Family Cyprinidae in the Middle East Inferred from Cytochrome b DNA - Evolutionary Significance of this region. Molecular Phylogenetics and Evolution, 22 (1), 91 - I 00. https: // doi. org / 10.1006 / mpev. 2001.1040" type="journal article" year="2002">
Durand
<emphasis id="5FCCA34DFFEEFFF07D5292757347F8E3" box="[514,574,1843,1869]" italics="true" pageId="2" pageNumber="431">et al.</emphasis>
(2002)
</bibRefCitation>
, and later included in phylogenetic studies by
<bibRefCitation id="092902AEFFEEFFF07BE49272719DF8DF" author="Tsigenopoulos, C. S. &amp; Berrebi, P." pageId="2" pageNumber="431" pagination="851 - 856" refId="ref4654" refString="Tsigenopoulos, C. S., Kasapidis. P. &amp; Berrebi, P. (2010) Phylogenetic relationships of hexaploid large-sized barbs (genus Labeobarbus, Cyprinidae) based on mtDNA data. Molecular Phylogenetics and Evolution, 56 (2), 851 - 856. https: // doi. org / 10.1016 / j. ympev. 2010.02.006" type="journal article" year="2010">
Tsigenopoulos
<emphasis id="5FCCA34DFFEEFFF07A31927574ECF8E3" box="[1377,1429,1843,1869]" italics="true" pageId="2" pageNumber="431">et al</emphasis>
. (2010)
</bibRefCitation>
,
<bibRefCitation id="092902AEFFEEFFF07FA0921670A1F8DF" author="Berrebi, P. &amp; Chenuil, A. &amp; Kotlik, P. &amp; Machordom, A. &amp; Tsigenopoulos, C. S." box="[240,472,1879,1906]" pageId="2" pageNumber="431" pagination="29 - 55" refId="ref2946" refString="Berrebi, P., Chenuil, A., Kotlik, P., Machordom, A. &amp; Tsigenopoulos, C. S. (2014) Disentangling the evolutionary history of the genus Barbus sensu lato, a twenty years adventure. In: Alves, M. J., Cartaxana, A., Correia, A. M. &amp; Lopes, L. F. (Eds.), Professor Carlos Almaca (1934 - 2010). Estado da Arte em Areas Cienticas que Desenvolveu. Museu Nacional de Historia Natural e da Ciencia, Lisboa, pp. 29 - 55." type="book chapter" year="2014">
Berrebi
<emphasis id="5FCCA34DFFEEFFF07E1B92197004F8DF" box="[331,381,1879,1905]" italics="true" pageId="2" pageNumber="431">et al</emphasis>
. (2014)
</bibRefCitation>
,
<bibRefCitation id="092902AEFFEEFFF07EB4921673B4F8DF" author="Borkenhagen, K." box="[484,717,1879,1905]" pageId="2" pageNumber="431" pagination="1179 - 1195" refId="ref3225" refString="Borkenhagen, K. (2014) A new genus and species of cyprinid fish (Actinopterygii, Cyprinidae) from the Arabian Peninsula, and its phylogenetic and zoogeographic affinities. Environmental Biology of Fishes, 97, 1179 - 1195. https: // doi. org / 10.1007 / s 10641 - 011 - 9787 - 1" type="journal article" year="2014">Borkenhagen (2014)</bibRefCitation>
,
<bibRefCitation id="092902AEFFEEFFF07D88921972DFF8DC" author="Yang, L. &amp; Sado, T. &amp; Vincent Hirt, M. &amp; Pasco-Viel, E. &amp; Arunachalam, M. &amp; Li, J. &amp; Wang, X. &amp; Freyhof, J. &amp; Saitoh, K. &amp; Simons, A. M. &amp; Miya, M. &amp; He, S. &amp; Mayden, R. L." box="[728,934,1879,1906]" pageId="2" pageNumber="431" pagination="97 - 116" refId="ref4923" refString="Yang, L., Sado, T., Vincent Hirt, M., Pasco-Viel, E., Arunachalam, M., Li, J., Wang, X., Freyhof, J., Saitoh, K., Simons, A. M., Miya, M., He, S. &amp; Mayden, R. L. (2015) Phylogeny and polyploidy: resolving the classification of cyprinine fishes (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution, 85, 97 - 116. https: // doi. org / 10.1016 / j. ympev. 2015.01.014" type="journal article" year="2015">
Yang
<emphasis id="5FCCA34DFFEEFFF07C4992197233F8DF" box="[793,842,1879,1905]" italics="true" pageId="2" pageNumber="431">et al</emphasis>
. (2015)
</bibRefCitation>
. All these studies show
<taxonomicName id="AAB804DCFFEEFFF07BE392167469F8DF" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[1203,1296,1879,1905]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07BE392167469F8DF" box="[1203,1296,1879,1905]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
as a distinct phylogenetic lineage from related genera such as
<taxonomicName id="AAB804DCFFEEFFF07D81923A7226F83B" authorityName="Borkenhagen" authorityYear="2014" box="[721,863,1915,1941]" family="Cyprinidae" genus="Arabibarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07D81923A7226F83B" box="[721,863,1915,1941]" italics="true" pageId="2" pageNumber="431">Arabibarbus</emphasis>
</taxonomicName>
,
<taxonomicName id="AAB804DCFFEEFFF07C3F923A7576F83B" authorityName="Karaman" authorityYear="1971" box="[879,1039,1915,1941]" family="Cyprinidae" genus="Carasobarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07C3F923A7576F83B" box="[879,1039,1915,1941]" italics="true" pageId="2" pageNumber="431">Carasobarbus</emphasis>
</taxonomicName>
,
<taxonomicName id="AAB804DCFFEEFFF07B4F923A75CAF83B" baseAuthorityName="Borkenhagen" baseAuthorityYear="2014" box="[1055,1203,1915,1941]" family="Cyprinidae" genus="Labeobarbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07B4F923A75CAF83B" box="[1055,1203,1915,1941]" italics="true" pageId="2" pageNumber="431">Labeobarbus</emphasis>
</taxonomicName>
,
<taxonomicName id="AAB804DCFFEEFFF07B93923A74ECF83B" box="[1219,1429,1915,1941]" family="Cyprinidae" genus="Mesopotamichthys" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07B93923A74ECF83B" box="[1219,1429,1915,1941]" italics="true" pageId="2" pageNumber="431">Mesopotamichthys</emphasis>
</taxonomicName>
, and
<taxonomicName id="AAB804DCFFEEFFF07F9792E17036F817" box="[199,335,1952,1977]" genus="Ptercapoeta" pageId="2" pageNumber="430" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07F9792E17036F817" box="[199,335,1952,1977]" italics="true" pageId="2" pageNumber="431">Ptercapoeta</emphasis>
</taxonomicName>
. We have not reproduced the phylogenetic trees here, as they are available open-access from their original sources, which indicate the unique phylogenetic position of
<taxonomicName id="AAB804DCFFEEFFF07CC69282728AF873" authorityName="Borkenhagen &amp; Freyhof" authorityYear="2023" box="[918,1011,1987,2013]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="5FCCA34DFFEEFFF07CC69282728AF873" box="[918,1011,1987,2013]" italics="true" pageId="2" pageNumber="431">Atlantor</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="39C72FD7FFEFFFF17FC797A47442FD50" box="[151,1339,741,766]" pageId="3" pageNumber="432" startId="3.[151,250,741,765]" targetBox="[152,1436,179,715]" targetPageId="3" targetType="figure">
<paragraph id="6D077F5FFFEFFFF17FC797A47442FD50" blockId="3.[151,1339,741,766]" box="[151,1339,741,766]" pageId="3" pageNumber="432">
<emphasis id="5FCCA34DFFEFFFF17FC797A4706AFD53" bold="true" box="[151,275,741,765]" pageId="3" pageNumber="432">FIGURE 1.</emphasis>
<taxonomicName id="AAB804DCFFEFFFF17E4997A7737AFD53" authority=", BMNH" authorityName="BMNH" box="[281,515,742,766]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="432" phylum="Chordata" rank="species" species="reinii">
<emphasis id="5FCCA34DFFEFFFF17E4997A770D3FD53" box="[281,426,742,766]" italics="true" pageId="3" pageNumber="432">Atlantor reinii</emphasis>
, BMNH
</taxonomicName>
1901.7.26.2-3, 1, 146 mm SL; Oued Oum er Rbia, Morocco. Photo: S. Tränkner.
</paragraph>
</caption>
<caption id="39C72FD7FFEFFFF17FC793677442F990" box="[151,1339,1574,1598]" pageId="3" pageNumber="432" startId="3.[151,250,1574,1598]" targetBox="[152,1436,790,1550]" targetPageId="3" targetType="figure">
<paragraph id="6D077F5FFFEFFFF17FC793677442F990" blockId="3.[151,1339,1574,1598]" box="[151,1339,1574,1598]" pageId="3" pageNumber="432">
<emphasis id="5FCCA34DFFEFFFF17FC79367706AF990" bold="true" box="[151,275,1574,1598]" pageId="3" pageNumber="432">FIGURE 2.</emphasis>
<taxonomicName id="AAB804DCFFEFFFF17E499367737AF993" authority=", BMNH" authorityName="BMNH" box="[281,515,1574,1598]" family="Cyprinidae" genus="Atlantor" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="432" phylum="Chordata" rank="species" species="reinii">
<emphasis id="5FCCA34DFFEFFFF17E49936770D3F990" box="[281,426,1574,1598]" italics="true" pageId="3" pageNumber="432">Atlantor reinii</emphasis>
, BMNH
</taxonomicName>
1901.7.26.2-3, 1, 147 mm SL; Oued Oum er Rbia, Morocco. Photo: S. Tränkner.
</paragraph>
</caption>
<paragraph id="6D077F5FFFEFFFF17F97932D709CF904" blockId="3.[151,1436,1644,1707]" pageId="3" pageNumber="432">
<emphasis id="5FCCA34DFFEFFFF17F97932D70AEF928" bold="true" box="[199,471,1644,1670]" pageId="3" pageNumber="432">Comparative material.</emphasis>
See
<bibRefCitation id="092902AEFFEFFFF17D5B932D7222F928" author="Borkenhagen, K. &amp; Krupp, F." box="[523,859,1644,1671]" pageId="3" pageNumber="432" pagination="1 - 53" refId="ref3454" refString="Borkenhagen, K. &amp; Krupp, F. (2013) Taxonomic revision of the genus Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae). ZooKeys, 339, 1 - 53. https: // doi. org / 10.3897 / zookeys. 339.4903" type="journal article" year="2013">Borkenhagen &amp; Krupp (2013)</bibRefCitation>
and
<bibRefCitation id="092902AEFFEFFFF17CDF932D750CF928" author="Borkenhagen, K." box="[911,1141,1644,1670]" pageId="3" pageNumber="432" pagination="1179 - 1195" refId="ref3225" refString="Borkenhagen, K. (2014) A new genus and species of cyprinid fish (Actinopterygii, Cyprinidae) from the Arabian Peninsula, and its phylogenetic and zoogeographic affinities. Environmental Biology of Fishes, 97, 1179 - 1195. https: // doi. org / 10.1007 / s 10641 - 011 - 9787 - 1" type="journal article" year="2014">Borkenhagen (2014)</bibRefCitation>
for materials of additional species and genera examined.
</paragraph>
</subSubSection>
</treatment>
</document>
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