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<document id="6161E9BD77CF659AD5AEE94E2C5216EE" ID-DOI="10.11646/zootaxa.5315.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8130409" ID-ZooBank="38EB0CD1-2432-43A9-908D-EF60B3B7EAC2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688980409704" checkinUser="plazi" docAuthor="Pontes Moraes, Isadora M., Polotow, Daniele, Labarque, Facundo M. &amp; Da Silva, Marcio B." docDate="2023" docId="CD69878AFF89B90AFF307188FB2E7CA9" docLanguage="en" docName="zootaxa.5315.2.6.pdf" docOrigin="Zootaxa 5315 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isoctenus botocudos Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva 2023, new species" docType="treatment" docVersion="2" lastPageNumber="184" masterDocId="3150FFF2FF8FB90DFFA7750FFFD77B53" masterDocTitle="Description of three new species of Isoctenus Bertkau, 1880 and new records for I. areia Polotow &amp; Brescovit (Araneae, Ctenidae) from northeastern Brazil" masterLastPageNumber="186" masterPageNumber="177" pageNumber="183" updateTime="1688980845101" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BD0C2BB01BE862691A387E3D1155631A">Description of three new species of Isoctenus Bertkau, 1880 and new records for I. areia Polotow &amp; Brescovit (Araneae, Ctenidae) from northeastern Brazil</mods:title>
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<mods:namePart id="33B519E723324265E1506456C9AB1902">Pontes Moraes, Isadora M.</mods:namePart>
<mods:nameIdentifier id="28E42151C0A843F15544610EBB9AD65C" type="ORCID">0009-0000-6831-2121</mods:nameIdentifier>
<mods:nameIdentifier id="5F881E70162EB6831A3FA579CED924F0" type="email">isadorapontes91@gmail.com</mods:nameIdentifier>
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<mods:namePart id="348A33C676CDD3D5B70C79E00823B36B">Polotow, Daniele</mods:namePart>
<mods:nameIdentifier id="35A6A6CE1BE8FA5508C11B1F6049B36F" type="ORCID">0000-0002-1069-1808</mods:nameIdentifier>
<mods:nameIdentifier id="B2DCFC457A342C8268FA701783D5678C" type="email">danielepolotow@gmail.com</mods:nameIdentifier>
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<mods:namePart id="34910526FDB91E7E76445478578CF8A4">Labarque, Facundo M.</mods:namePart>
<mods:nameIdentifier id="7D77157B1B368989FF90810329C0270E" type="ORCID">0000-0002-1069-1808</mods:nameIdentifier>
<mods:nameIdentifier id="B31734C6F9953FE48F80B6B4C854DC10" type="email">danielepolotow@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A2AA961DF0B35A783031C469E0CC15D3">Da Silva, Marcio B.</mods:namePart>
<mods:nameIdentifier id="8553789E68A2065B6E3354F215B9DBB4" type="ORCID">0009-0000-6831-2121</mods:nameIdentifier>
<mods:nameIdentifier id="4D3F39383D333E015B67DD0E70AB3259" type="email">isadorapontes91@gmail.com</mods:nameIdentifier>
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<mods:title id="36345BD0C77114859D14D8EB9551E46D">Zootaxa</mods:title>
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<mods:identifier id="92E90DE918E5EBCC49F46C458917D726" type="Zenodo-Dep">8130409</mods:identifier>
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<treatment id="CD69878AFF89B90AFF307188FB2E7CA9" LSID="urn:lsid:plazi:treatment:CD69878AFF89B90AFF307188FB2E7CA9" httpUri="http://treatment.plazi.org/id/CD69878AFF89B90AFF307188FB2E7CA9" lastPageId="7" lastPageNumber="184" pageId="6" pageNumber="183">
<subSubSection id="0DDA6517FF89B90BFF307188FDF37FF3" box="[151,548,1158,1185]" pageId="6" pageNumber="183" type="nomenclature">
<paragraph id="457F369CFF89B90BFF307188FDF37FF3" blockId="6.[151,548,1158,1221]" box="[151,548,1158,1185]" pageId="6" pageNumber="183">
<heading id="1E3781F0FF89B90BFF307188FDF37FF3" bold="true" box="[151,548,1158,1185]" fontSize="11" level="1" pageId="6" pageNumber="183" reason="1">
<emphasis id="77B4EA8EFF89B90BFF307188FDF37FF3" bold="true" box="[151,548,1158,1185]" pageId="6" pageNumber="183">
<taxonomicName id="82C04D1FFF89B90BFF307188FE5B7FF2" authority="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva, 2023" authorityName="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva" authorityYear="2023" box="[151,396,1158,1185]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="183" phylum="Arthropoda" rank="species" species="botocudos" status="new species">
<emphasis id="77B4EA8EFF89B90BFF307188FE5B7FF2" bold="true" box="[151,396,1158,1185]" italics="true" pageId="6" pageNumber="183">Isoctenus botocudos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6C8757F5FF89B90BFE337188FDF37FF3" box="[404,548,1158,1185]" pageId="6" pageNumber="183" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DDA6517FF89B90BFF3071A4FECA7F96" box="[151,285,1194,1221]" pageId="6" pageNumber="183" type="description">
<paragraph id="457F369CFF89B90BFF3071A4FECA7F96" blockId="6.[151,548,1158,1221]" box="[151,285,1194,1221]" pageId="6" pageNumber="183">
<figureCitation id="DDFB2A19FF89B90BFF3071A4FECA7F96" box="[151,285,1194,1221]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="7.[152,255,941,965]" captionStartId-1="7.[152,255,1837,1861]" captionStartId-2="8.[152,255,746,770]" captionTargetBox-0="[151,1436,417,917]" captionTargetBox-1="[151,1436,989,1813]" captionTargetBox-2="[408,1180,181,722]" captionTargetId-0="figure-135@7.[151,1436,417,918]" captionTargetId-1="figure-157@7.[151,1436,989,1813]" captionTargetId-2="figure-23@8.[408,1180,181,722]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 5. Isoctenus botocudos new species, habitus. A–B, male; A, dorsal view; B, ventral view." captionText-1="FIGURE 6. Isoctenus botocudos new species. A–B, male palp. A, ventral view; B, retrolateral view." captionText-2="FIGURE 7. A–B, Geographic distribution records of Isoctenus jussara new species, I. cabula new species, and I. botocudos new species, and new records of Isoctenus areia. A, South America with demarcation of northeastern Brazil; B, Close up of the Brazilian northeastern regions where the species are distributed." figureDoi-0="http://doi.org/10.5281/zenodo.8130419" figureDoi-1="http://doi.org/10.5281/zenodo.8130421" figureDoi-2="http://doi.org/10.5281/zenodo.8130425" httpUri-0="https://zenodo.org/record/8130419/files/figure.png" httpUri-1="https://zenodo.org/record/8130421/files/figure.png" httpUri-2="https://zenodo.org/record/8130425/files/figure.png" pageId="6" pageNumber="183">Figures 5–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0DDA6517FF89B90BFF3071FCFD257E92" pageId="6" pageNumber="183" type="materials_examined">
<paragraph id="457F369CFF89B90BFF3071FCFD4A7E62" blockId="6.[151,1437,1266,2013]" pageId="6" pageNumber="183">
<emphasis id="77B4EA8EFF89B90BFF3071FCFE917E5F" bold="true" box="[151,326,1266,1293]" pageId="6" pageNumber="183">Type material.</emphasis>
<materialsCitation id="F5A83CC1FF89B90BFEEA71FCFD4E7E62" collectingDate="1970-06-03" collectorName="Ceplac Team" country="Brazil" latitude="-14.583333" location="Fazenda Santa Tereza" longLatPrecision="1287" longitude="-39.283333" municipality="Urucuca" pageId="6" pageNumber="183" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
Male
<typeStatus id="9A7B883EFF89B90BFE2871FCFE267E5E" box="[399,497,1267,1293]" pageId="6" pageNumber="183">holotype</typeStatus>
from
<location id="401F6047FF89B90BFD9071FCFCF97E5E" LSID="urn:lsid:plazi:treatment:CD69878AFF89B90AFF307188FB2E7CA9:401F6047FF89B90BFD9071FCFCF97E5E" box="[567,814,1267,1293]" country="Brazil" latitude="-14.583333" longLatPrecision="1287" longitude="-39.283333" municipality="Urucuca" name="Fazenda Santa Tereza" pageId="6" pageNumber="183" stateProvince="Bahia">Fazenda Santa Tereza</location>
,
<collectingMunicipality id="A51BACE6FF89B90BFC9D71FCFC4B7E5E" box="[826,924,1267,1293]" pageId="6" pageNumber="183">Uruçuca</collectingMunicipality>
,
<geoCoordinate id="20F4505BFF89B90BFC0071FDFBD37E5E" box="[935,1028,1266,1293]" degrees="14" direction="south" minutes="35" orientation="latitude" pageId="6" pageNumber="183" precision="925" value="-14.583333">14º35’S</geoCoordinate>
,
<geoCoordinate id="20F4505BFF89B90BFBB771FCFBA37E5E" box="[1040,1140,1267,1293]" degrees="39" direction="west" minutes="17" orientation="longitude" pageId="6" pageNumber="183" precision="925" value="-39.283333">39º17’W</geoCoordinate>
,
<collectingRegion id="8704F87EFF89B90BFB2771FCFB137E5E" box="[1152,1220,1267,1293]" country="Brazil" name="Bahia" pageId="6" pageNumber="183">Bahia</collectingRegion>
,
<collectingCountry id="3DD7760CFF89B90BFB7771FCFAC07E5E" box="[1232,1303,1267,1293]" name="Brazil" pageId="6" pageNumber="183">Brazil</collectingCountry>
,
<date id="317E105CFF89B90BFA8371FCFA407E5E" box="[1316,1431,1267,1293]" pageId="6" pageNumber="183" value="1970-06-03">
<collectingDate id="213AE9B4FF89B90BFA8371FCFA407E5E" box="[1316,1431,1267,1293]" pageId="6" pageNumber="183" value="1970-06-03">3.VI.1970</collectingDate>
</date>
,
<collectorName id="E835534AFF89B90BFF307018FEFE7E62" box="[151,297,1303,1329]" pageId="6" pageNumber="183">Ceplac Team</collectorName>
coll., deposited in MNRJ 13435
</materialsCitation>
.
</paragraph>
<paragraph id="457F369CFF89B90BFF607035FD257E92" blockId="6.[151,1437,1266,2013]" pageId="6" pageNumber="183">
<emphasis id="77B4EA8EFF89B90BFF607035FDE87E07" bold="true" box="[199,575,1338,1364]" pageId="6" pageNumber="183">Additional material examined.</emphasis>
<materialsCitation id="F5A83CC1FF89B90BFDEC7034FD397E92" collectingDate="1987-10-15" collectingDateMax="2002-03-10" collectingDateMin="1987-10-15" collectorName="J. Becker &amp; M. G. C. Costa" country="Brazil" county="Belmonte" latitude="-16.433332" location="Estacao Experimental Gregorio Bondar" longLatPrecision="1282" longitude="-39.05" municipality="Barrolandia" pageId="6" pageNumber="183" specimenCount="4" specimenCount-male="4" stateProvince="Bahia">
<collectingCountry id="3DD7760CFF89B90BFDEC7034FD637E06" box="[587,692,1339,1365]" name="Brazil" pageId="6" pageNumber="183">BRAZIL</collectingCountry>
.
<collectingRegion id="8704F87EFF89B90BFD647034FCDE7E06" box="[707,777,1339,1365]" country="Brazil" name="Bahia" pageId="6" pageNumber="183">Bahia</collectingRegion>
:
<collectingCounty id="AC1E4E10FF89B90BFCBE7034FC5E7E06" box="[793,905,1339,1365]" pageId="6" pageNumber="183">Belmonte</collectingCounty>
,
<collectingMunicipality id="A51BACE6FF89B90BFC3E7034FBF77E06" box="[921,1056,1339,1365]" pageId="6" pageNumber="183">Barrolândia</collectingMunicipality>
,
<location id="401F6047FF89B90BFB967034FF3C7E2A" LSID="urn:lsid:plazi:treatment:CD69878AFF89B90AFF307188FB2E7CA9:401F6047FF89B90BFB967034FF3C7E2A" country="Brazil" county="Belmonte" latitude="-16.433332" longLatPrecision="1282" longitude="-39.05" municipality="Barrolandia" name="Estacao Experimental Gregorio Bondar" pageId="6" pageNumber="183" stateProvince="Bahia">Estação Experimental Gregório Bóndar</location>
,
<geoCoordinate id="20F4505BFF89B90BFF5E7051FE827E2A" box="[249,341,1374,1401]" degrees="16" direction="south" minutes="05" orientation="latitude" pageId="6" pageNumber="183" precision="925" value="-16.083334">16º05’S</geoCoordinate>
,
<geoCoordinate id="20F4505BFF89B90BFEC47050FE1F7E2A" box="[355,456,1375,1401]" degrees="39" direction="west" minutes="16" orientation="longitude" pageId="6" pageNumber="183" precision="925" value="-39.266666">39º16’W</geoCoordinate>
,
<specimenCount id="53C6FD15FF89B90BFE727050FDF27E2A" box="[469,549,1375,1401]" pageId="6" pageNumber="183" type="male">1 male</specimenCount>
,
<date id="317E105CFF89B90BFD947051FD7B7E2A" box="[563,684,1374,1401]" pageId="6" pageNumber="183" value="1987-10-15">
<collectingDate id="213AE9B4FF89B90BFD947051FD7B7E2A" box="[563,684,1374,1401]" pageId="6" pageNumber="183" value="1987-10-15">15.X.1987</collectingDate>
</date>
,
<collectorName id="E835534AFF89B90BFD1D7050FCF37E2A" box="[698,804,1375,1401]" pageId="6" pageNumber="183">J. Becker</collectorName>
coll. (MNRJ 13428); Porto Seguro,
<location id="401F6047FF89B90BFB6B7050FF3C7ECE" LSID="urn:lsid:plazi:treatment:CD69878AFF89B90AFF307188FB2E7CA9:401F6047FF89B90BFB6B7050FF3C7ECE" country="Brazil" county="Belmonte" latitude="-16.433332" longLatPrecision="1282" longitude="-39.05" municipality="Barrolandia" name="Estacao Ecologica Veracel" pageId="6" pageNumber="183" stateProvince="Bahia">Estação Ecológica Veracel</location>
/Veracruz (EEVC),
<geoCoordinate id="20F4505BFF89B90BFE6B708CFDF07ECE" box="[460,551,1411,1437]" degrees="16" direction="south" minutes="26" orientation="latitude" pageId="6" pageNumber="183" precision="925" value="-16.433332">16º26’S</geoCoordinate>
,
<geoCoordinate id="20F4505BFF89B90BFD91708CFD4D7ECE" box="[566,666,1411,1437]" degrees="39" direction="west" minutes="03" orientation="longitude" pageId="6" pageNumber="183" precision="925" value="-39.05">39º03’W</geoCoordinate>
,
<specimenCount id="53C6FD15FF89B90BFD0F708CFCD37ECE" box="[680,772,1411,1437]" pageId="6" pageNumber="183" type="male">2 males</specimenCount>
,
<date id="317E105CFF89B90BFCB5708CFC5D7ECE" box="[786,906,1411,1437]" pageId="6" pageNumber="183" value="2001-10-23">
<collectingDate id="213AE9B4FF89B90BFCB5708CFC5D7ECE" box="[786,906,1411,1437]" pageId="6" pageNumber="183" value="2001-10-23">23.X.2001</collectingDate>
</date>
,
<collectorName id="E835534AFF89B90BFC3F708CFBE07ECE" box="[920,1079,1411,1437]" pageId="6" pageNumber="183">M.G.C. Costa</collectorName>
coll. (UFBA 02175; 02184);
<specimenCount id="53C6FD15FF89B90BFA29708CFF077E92" pageId="6" pageNumber="183" type="male">1 male</specimenCount>
,
<date id="317E105CFF89B90BFF7B70A8FE8B7E92" box="[220,348,1447,1473]" pageId="6" pageNumber="183" value="2002-03-10">
<collectingDate id="213AE9B4FF89B90BFF7B70A8FE8B7E92" box="[220,348,1447,1473]" pageId="6" pageNumber="183" value="2002-03-10">10.III.2002</collectingDate>
</date>
,
<collectorName id="E835534AFF89B90BFECF70A8FDD47E92" box="[360,515,1447,1473]" pageId="6" pageNumber="183">M.G.C. Costa</collectorName>
coll. (UFBA 02181)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DDA6517FF89B90BFF6070C4FDB47D02" pageId="6" pageNumber="183" type="etymology">
<paragraph id="457F369CFF89B90BFF6070C4FDB47D02" blockId="6.[151,1437,1266,2013]" pageId="6" pageNumber="183">
<emphasis id="77B4EA8EFF89B90BFF6070C4FE997EB6" bold="true" box="[199,334,1483,1509]" pageId="6" pageNumber="183">Etymology.</emphasis>
The species name is a noun in apposition that honors the Botocudos indigenous people, a generic denomination used by the Portuguese colonizers to several indigenous peoples from the states of
<collectingRegion id="8704F87EFF89B90BFB5570E0FAE17D5A" box="[1266,1334,1519,1545]" country="Brazil" name="Bahia" pageId="6" pageNumber="183">Bahia</collectingRegion>
,
<collectingRegion id="8704F87EFF89B90BFAE470E0FF007D7E" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="183">Espírito Santo</collectingRegion>
and
<collectingRegion id="8704F87EFF89B90BFEB4731CFE787D7E" box="[275,431,1555,1581]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="183">Minas Gerais</collectingRegion>
,
<collectingCountry id="3DD7760CFF89B90BFE1A731CFDD27D7E" box="[445,517,1555,1581]" name="Brazil" pageId="6" pageNumber="183">Brazil</collectingCountry>
. The term
<taxonomicName id="82C04D1FFF89B90BFD22731CFD277D7E" authorityName="Kirkaldy" authorityYear="1904" box="[645,752,1555,1581]" class="Insecta" family="Rhyparochromidae" genus="Botocudo" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="77B4EA8EFF89B90BFD22731CFD277D7E" box="[645,752,1555,1581]" italics="true" pageId="6" pageNumber="183">Botocudo</emphasis>
</taxonomicName>
in Portuguese (from
<emphasis id="77B4EA8EFF89B90BFC4E731CFB947D7E" box="[1001,1091,1555,1581]" italics="true" pageId="6" pageNumber="183">botoque</emphasis>
, a plug), is an allusion to the wooden disks worn in their lips and ears.
</paragraph>
</subSubSection>
<subSubSection id="0DDA6517FF89B90BFF607355FA507C56" pageId="6" pageNumber="183" type="diagnosis">
<paragraph id="457F369CFF89B90BFF607355FA507C56" blockId="6.[151,1437,1266,2013]" pageId="6" pageNumber="183">
<emphasis id="77B4EA8EFF89B90BFF607355FE957D27" bold="true" box="[199,322,1626,1652]" pageId="6" pageNumber="183">Diagnosis.</emphasis>
Males of
<taxonomicName id="82C04D1FFF89B90BFE177354FD447D26" authority="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva, 2023" authorityName="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva" authorityYear="2023" box="[432,659,1627,1653]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="183" phylum="Arthropoda" rank="species" species="botocudos" status="new species">
<emphasis id="77B4EA8EFF89B90BFE177354FD447D26" box="[432,659,1627,1653]" italics="true" pageId="6" pageNumber="183">Isoctenus botocudos</emphasis>
</taxonomicName>
<emphasis id="77B4EA8EFF89B90BFD3E7354FCF57D27" bold="true" box="[665,802,1626,1653]" pageId="6" pageNumber="183">
<taxonomicNameLabel id="6C8757F5FF89B90BFD3E7354FCF57D27" box="[665,802,1626,1653]" pageId="6" pageNumber="183" rank="species">new species</taxonomicNameLabel>
</emphasis>
resemble those of
<taxonomicName id="82C04D1FFF89B90BFC547354FAAD7D26" authority="Polotow &amp; Brescovit, 2009" authorityName="Polotow &amp; Brescovit" authorityYear="2009" box="[1011,1402,1627,1653]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="183" phylum="Arthropoda" rank="species" species="areia">
<emphasis id="77B4EA8EFF89B90BFC547354FB947D27" box="[1011,1091,1627,1652]" italics="true" pageId="6" pageNumber="183">I. areia</emphasis>
<bibRefCitation id="21514B6DFF89B90BFBED7354FAAD7D26" author="Polotow, D. &amp; Brescovit, A. D." box="[1098,1402,1627,1653]" pageId="6" pageNumber="183" pagination="583 - 614" refId="ref5579" refString="Polotow, D. &amp; Brescovit, A. D. (2009) Revision and cladistic analysis of Isoctenus and description of a new neotropical genus (Araneae, Ctenidae, Cteninae). Zoological Journal of Linnean Society, 155, 583 - 614. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00452. x" type="journal article" year="2009">Polotow &amp; Brescovit, 2009</bibRefCitation>
</taxonomicName>
by the short dorsal branch of the RTA, but can be distinguished by the truncated tip of the dorsal branch of the RTA, by the smaller ventral branch of the RTA, and by the smaller ventral retrolateral projection of cymbium (
<figureCitation id="DDFB2A19FF89B90BFAB373ACFA587DEE" box="[1300,1423,1699,1725]" captionStart="FIGURE 6" captionStartId="7.[152,255,1837,1861]" captionTargetBox="[151,1436,989,1813]" captionTargetId="figure-157@7.[151,1436,989,1813]" captionTargetPageId="7" captionText="FIGURE 6. Isoctenus botocudos new species. A–B, male palp. A, ventral view; B, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.8130421" httpUri="https://zenodo.org/record/8130421/files/figure.png" pageId="6" pageNumber="183">Fig. 6A–B</figureCitation>
), whereas
<taxonomicName id="82C04D1FFF89B90BFF5D73C8FE9C7DB3" authorityName="Polotow &amp; Brescovit" authorityYear="2009" box="[250,331,1735,1760]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="183" phylum="Arthropoda" rank="species" species="areia">
<emphasis id="77B4EA8EFF89B90BFF5D73C8FE9C7DB3" box="[250,331,1735,1760]" italics="true" pageId="6" pageNumber="183">I. areia</emphasis>
</taxonomicName>
can be distinguished by elongated tip of the dorsal branch of the RTA, by the larger ventral branch of the RTA, and by the larger ventral retrolateral projection of cymbium (
<bibRefCitation id="21514B6DFF89B90BFC6D73E4FB2C7C56" author="Polotow, D. &amp; Brescovit, A. D." box="[970,1275,1771,1797]" pageId="6" pageNumber="183" pagination="583 - 614" refId="ref5579" refString="Polotow, D. &amp; Brescovit, A. D. (2009) Revision and cladistic analysis of Isoctenus and description of a new neotropical genus (Araneae, Ctenidae, Cteninae). Zoological Journal of Linnean Society, 155, 583 - 614. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00452. x" type="journal article" year="2009">Polotow &amp; Brescovit 2009</bibRefCitation>
: fig. 6A–B).
</paragraph>
</subSubSection>
<subSubSection id="0DDA6517FF89B90AFF607201FB0D7C86" lastPageId="7" lastPageNumber="184" pageId="6" pageNumber="183" type="description">
<paragraph id="457F369CFF89B90AFF607201FC0B7ADA" blockId="6.[151,1437,1266,2013]" lastBlockId="7.[151,1437,150,393]" lastPageId="7" lastPageNumber="184" pageId="6" pageNumber="183">
<emphasis id="77B4EA8EFF89B90BFF607201FE8E7C7B" bold="true" box="[199,345,1806,1832]" pageId="6" pageNumber="183">Description.</emphasis>
<emphasis id="77B4EA8EFF89B90BFEC67200FE4C7C7A" box="[353,411,1807,1833]" italics="true" pageId="6" pageNumber="183">Male</emphasis>
(
<typeStatus id="9A7B883EFF89B90BFE0C7200FDC67C7A" box="[427,529,1807,1833]" pageId="6" pageNumber="183">holotype</typeStatus>
, MNRJ 13435). Prosoma reddish brown, with brown thoracic groove and black rings around eyes (
<figureCitation id="DDFB2A19FF89B90BFED6723CFE1A7C1E" box="[369,461,1842,1869]" captionStart="FIGURE 5" captionStartId="7.[152,255,941,965]" captionTargetBox="[151,1436,417,917]" captionTargetId="figure-135@7.[151,1436,417,918]" captionTargetPageId="7" captionText="FIGURE 5. Isoctenus botocudos new species, habitus. A–B, male; A, dorsal view; B, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8130419" httpUri="https://zenodo.org/record/8130419/files/figure.png" pageId="6" pageNumber="183">Fig. 5A</figureCitation>
). Sternum, endites, labium, and legs light reddish brown (
<figureCitation id="DDFB2A19FF89B90BFBC4723CFB6A7C1E" box="[1123,1213,1842,1869]" captionStart="FIGURE 5" captionStartId="7.[152,255,941,965]" captionTargetBox="[151,1436,417,917]" captionTargetId="figure-135@7.[151,1436,417,918]" captionTargetPageId="7" captionText="FIGURE 5. Isoctenus botocudos new species, habitus. A–B, male; A, dorsal view; B, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8130419" httpUri="https://zenodo.org/record/8130419/files/figure.png" pageId="6" pageNumber="183">Fig. 5B</figureCitation>
). Brownish dorsum of opisthosoma, with a longitudinal light stripe with indented borders and arrow-shaped (
<figureCitation id="DDFB2A19FF89B90BFB3B7258FB2F7C22" box="[1180,1272,1878,1905]" captionStart="FIGURE 5" captionStartId="7.[152,255,941,965]" captionTargetBox="[151,1436,417,917]" captionTargetId="figure-135@7.[151,1436,417,918]" captionTargetPageId="7" captionText="FIGURE 5. Isoctenus botocudos new species, habitus. A–B, male; A, dorsal view; B, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8130419" httpUri="https://zenodo.org/record/8130419/files/figure.png" pageId="6" pageNumber="183">Fig. 5A</figureCitation>
). Light brown venter of opisthosoma (
<figureCitation id="DDFB2A19FF89B90BFE037274FE297CC6" box="[420,510,1914,1941]" captionStart="FIGURE 5" captionStartId="7.[152,255,941,965]" captionTargetBox="[151,1436,417,917]" captionTargetId="figure-135@7.[151,1436,417,918]" captionTargetPageId="7" captionText="FIGURE 5. Isoctenus botocudos new species, habitus. A–B, male; A, dorsal view; B, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8130419" httpUri="https://zenodo.org/record/8130419/files/figure.png" pageId="6" pageNumber="183">Fig. 5B</figureCitation>
). Total length 9.11. Carapace 4.99 long and 3.76 wide. Eye diameter: AME 0.22, ALE 0.17, PME 0.23, PLE 0.21. Leg measurements: I: femur 3.80/ patella 1.53/ tibia 3.74/ metatarsus 3.33/ tarsus 1.70/ total 14.10; II: 3.41/ 1.56/ 3.02/ 3.20/ 1.20/ 12.39; III: 3.03/ 1.32/ 2.51/ 3.15/ 1.06/ 11.07; IV: 4.22/ 1.49/ 3.92/ 5.95/ 1.44/ 17.02. Leg formula 4123. Leg spination: tibia I and II v2-2-2-2-2, r1-1-0, p0-1-0, III v2-2-2, r1- 1-0, p1,1,0 and IV v2-2-2, r1-1-0, p1-1-0; metatarsus I, II and III v2-2-2, r1-1-1, p-1-1-1, IV v1-1-1-1-2-2, r1-1-1, p1- 1-1. Palp: short tibia, approximately half of cymbium length; dorsal branch of RTA truncate and with a rounded tip and ventral branch of RTA short (
<figureCitation id="DDFB2A19FF88B90AFDA0740CFD557A4E" box="[519,642,259,285]" captionStart="FIGURE 6" captionStartId="7.[152,255,1837,1861]" captionTargetBox="[151,1436,989,1813]" captionTargetId="figure-157@7.[151,1436,989,1813]" captionTargetPageId="7" captionText="FIGURE 6. Isoctenus botocudos new species. A–B, male palp. A, ventral view; B, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.8130421" httpUri="https://zenodo.org/record/8130421/files/figure.png" pageId="7" pageNumber="184">Fig. 6A–B</figureCitation>
); ventral apophysis short and slightly curved retrolaterally (
<figureCitation id="DDFB2A19FF88B90AFAB1740CFA467A4E" box="[1302,1425,259,285]" captionStart="FIGURE 6" captionStartId="7.[152,255,1837,1861]" captionTargetBox="[151,1436,989,1813]" captionTargetId="figure-157@7.[151,1436,989,1813]" captionTargetPageId="7" captionText="FIGURE 6. Isoctenus botocudos new species. A–B, male palp. A, ventral view; B, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.8130421" httpUri="https://zenodo.org/record/8130421/files/figure.png" pageId="7" pageNumber="184">Fig. 6A–B</figureCitation>
). Cymbium with an elongated retrolateral dorsal projection and a short retrolateral ventral projection (
<figureCitation id="DDFB2A19FF88B90AFAB47428FA467A12" box="[1299,1425,295,321]" captionStart="FIGURE 6" captionStartId="7.[152,255,1837,1861]" captionTargetBox="[151,1436,989,1813]" captionTargetId="figure-157@7.[151,1436,989,1813]" captionTargetPageId="7" captionText="FIGURE 6. Isoctenus botocudos new species. A–B, male palp. A, ventral view; B, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.8130421" httpUri="https://zenodo.org/record/8130421/files/figure.png" pageId="7" pageNumber="184">Fig. 6A–B</figureCitation>
). Curved embolus, with a strong probasal projection (
<figureCitation id="DDFB2A19FF88B90AFD7A7444FC8E7A36" box="[733,857,331,357]" captionStart="FIGURE 6" captionStartId="7.[152,255,1837,1861]" captionTargetBox="[151,1436,989,1813]" captionTargetId="figure-157@7.[151,1436,989,1813]" captionTargetPageId="7" captionText="FIGURE 6. Isoctenus botocudos new species. A–B, male palp. A, ventral view; B, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.8130421" httpUri="https://zenodo.org/record/8130421/files/figure.png" pageId="7" pageNumber="184">Fig. 6A–B</figureCitation>
). Cup-shaped median apophysis curved prolaterally and with a small distal projection and a wide basal projection (
<figureCitation id="DDFB2A19FF88B90AFCF37460FC077ADA" box="[852,976,367,393]" captionStart="FIGURE 6" captionStartId="7.[152,255,1837,1861]" captionTargetBox="[151,1436,989,1813]" captionTargetId="figure-157@7.[151,1436,989,1813]" captionTargetPageId="7" captionText="FIGURE 6. Isoctenus botocudos new species. A–B, male palp. A, ventral view; B, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.8130421" httpUri="https://zenodo.org/record/8130421/files/figure.png" pageId="7" pageNumber="184">Fig. 6A–B</figureCitation>
).
</paragraph>
<caption id="11BF6614FF88B90AFF3F76A2FB597896" ID-DOI="http://doi.org/10.5281/zenodo.8130419" ID-Zenodo-Dep="8130419" box="[152,1166,941,965]" httpUri="https://zenodo.org/record/8130419/files/figure.png" pageId="7" pageNumber="184" startId="7.[152,255,941,965]" targetBox="[151,1436,417,917]" targetPageId="7" targetType="figure">
<paragraph id="457F369CFF88B90AFF3F76A2FB597896" blockId="7.[152,1166,941,965]" box="[152,1166,941,965]" pageId="7" pageNumber="184">
<emphasis id="77B4EA8EFF88B90AFF3F76A2FECF7896" bold="true" box="[152,280,941,965]" pageId="7" pageNumber="184">FIGURE 5.</emphasis>
<taxonomicName id="82C04D1FFF88B90AFEB876A1FE3A7896" authority="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva, 2023" authorityName="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva" authorityYear="2023" box="[287,493,941,965]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="184" phylum="Arthropoda" rank="species" species="botocudos" status="new species">
<emphasis id="77B4EA8EFF88B90AFEB876A1FE3A7896" box="[287,493,941,965]" italics="true" pageId="7" pageNumber="184">Isoctenus botocudos</emphasis>
</taxonomicName>
<emphasis id="77B4EA8EFF88B90AFE5476A2FDA77896" bold="true" box="[499,624,941,965]" pageId="7" pageNumber="184">
<taxonomicNameLabel id="6C8757F5FF88B90AFE5476A2FDA77896" box="[499,624,941,965]" pageId="7" pageNumber="184" rank="species">new species</taxonomicNameLabel>
</emphasis>
, habitus. A–B, male; A, dorsal view; B, ventral view.
</paragraph>
</caption>
<caption id="11BF6614FF88B90AFF3F7222FB4D7C16" ID-DOI="http://doi.org/10.5281/zenodo.8130421" ID-Zenodo-Dep="8130421" box="[152,1178,1837,1861]" httpUri="https://zenodo.org/record/8130421/files/figure.png" pageId="7" pageNumber="184" startId="7.[152,255,1837,1861]" targetBox="[151,1436,989,1813]" targetPageId="7" targetType="figure">
<paragraph id="457F369CFF88B90AFF3F7222FB4D7C16" blockId="7.[152,1178,1837,1861]" box="[152,1178,1837,1861]" pageId="7" pageNumber="184">
<emphasis id="77B4EA8EFF88B90AFF3F7222FECF7C16" bold="true" box="[152,280,1837,1861]" pageId="7" pageNumber="184">FIGURE 6.</emphasis>
<taxonomicName id="82C04D1FFF88B90AFEB87221FE3A7C16" authority="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva, 2023" authorityName="Pontes Moraes &amp; Polotow &amp; Labarque &amp; Da Silva" authorityYear="2023" box="[287,493,1837,1861]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="184" phylum="Arthropoda" rank="species" species="botocudos" status="new species">
<emphasis id="77B4EA8EFF88B90AFEB87221FE3A7C16" box="[287,493,1837,1861]" italics="true" pageId="7" pageNumber="184">Isoctenus botocudos</emphasis>
</taxonomicName>
<emphasis id="77B4EA8EFF88B90AFE547222FDA77C16" bold="true" box="[499,624,1837,1861]" pageId="7" pageNumber="184">
<taxonomicNameLabel id="6C8757F5FF88B90AFE547222FDA77C16" box="[499,624,1837,1861]" pageId="7" pageNumber="184" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A–B, male palp. A, ventral view; B, retrolateral view.
</paragraph>
</caption>
<paragraph id="457F369CFF88B90AFF607298FE487CE2" blockId="7.[199,1273,1943,2042]" box="[199,415,1943,1970]" pageId="7" pageNumber="184">
<emphasis id="77B4EA8EFF88B90AFF607298FECC7CE2" box="[199,283,1943,1969]" italics="true" pageId="7" pageNumber="184">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="457F369CFF88B90AFF6072B4FB0D7C86" blockId="7.[199,1273,1943,2042]" box="[199,1242,1979,2006]" pageId="7" pageNumber="184">
<emphasis id="77B4EA8EFF88B90AFF6072B4FEE87C86" bold="true" box="[199,319,1979,2005]" pageId="7" pageNumber="184">Variation.</emphasis>
Four males: total length 7.54–9.65, carapace length 4.61–5.53, femur I 4.60–6.11.
</paragraph>
</subSubSection>
<subSubSection id="0DDA6517FF88B90AFF6072D0FB2E7CA9" box="[199,1273,2015,2042]" pageId="7" pageNumber="184" type="distribution">
<paragraph id="457F369CFF88B90AFF6072D0FB2E7CA9" blockId="7.[199,1273,1943,2042]" box="[199,1273,2015,2042]" pageId="7" pageNumber="184">
<emphasis id="77B4EA8EFF88B90AFF6072D0FEB67CAA" bold="true" box="[199,353,2015,2041]" pageId="7" pageNumber="184">Distribution.</emphasis>
Coastal Atlantic Rain Forest, southern state of
<collectingRegion id="8704F87EFF88B90AFCD572D0FC617CAA" box="[882,950,2015,2041]" country="Brazil" name="Bahia" pageId="7" pageNumber="184">Bahia</collectingRegion>
, northeastern
<collectingCountry id="3DD7760CFF88B90AFBF472D0FB407CAA" box="[1107,1175,2015,2041]" name="Brazil" pageId="7" pageNumber="184">Brazil</collectingCountry>
(
<figureCitation id="DDFB2A19FF88B90AFB0072D0FB3C7CA9" box="[1191,1259,2015,2042]" captionStart="FIGURE 7" captionStartId="8.[152,255,746,770]" captionTargetBox="[408,1180,181,722]" captionTargetId="figure-23@8.[408,1180,181,722]" captionTargetPageId="8" captionText="FIGURE 7. A–B, Geographic distribution records of Isoctenus jussara new species, I. cabula new species, and I. botocudos new species, and new records of Isoctenus areia. A, South America with demarcation of northeastern Brazil; B, Close up of the Brazilian northeastern regions where the species are distributed." figureDoi="http://doi.org/10.5281/zenodo.8130425" httpUri="https://zenodo.org/record/8130425/files/figure.png" pageId="7" pageNumber="184">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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