125 lines
13 KiB
XML
125 lines
13 KiB
XML
<document id="0BBD0B658D521B9A2A51286FE1D62EC5" ID-CLB-Dataset="25961" ID-DOI="10.11646/zootaxa.4658.1.1" ID-GBIF-Dataset="31d6e081-95c3-4086-a45a-c11555dda3aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373040" ID-ZooBank="ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566364910789" checkinUser="plazi" docAuthor="Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed & Barbero, Enrico" docDate="2019" docId="8861532B88561827FF760BE2FCFFC37C" docLanguage="en" docName="zootaxa.4658.1.1.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euonthophagus Balthasar 1959" docType="treatment" docVersion="12" lastPageNumber="6" masterDocId="74582B5388531822FFE10957FFA3C74B" masterDocTitle="An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="6" updateTime="1698751869822" updateUser="ExternalLinkService">
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<mods:title id="DAD29E9B0703A1621B554F2B3445F162">An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
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<mods:namePart id="D051810FD36C66949EEBB3ED91144B1A">Ziani, Stefano</mods:namePart>
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<mods:namePart id="66BF57620C81A5EA6B5A1B265814479F">Abdel-Dayem, Mahmoud Saleh</mods:namePart>
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<mods:namePart id="9760B78FF973A5104D39696525EAD9A8">Aldhafer, Hathal Mohammed</mods:namePart>
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<mods:namePart id="BB83B3621FF54B714028A85DA01A53B4">Barbero, Enrico</mods:namePart>
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<mods:title id="CA98A593C0E3F29234835322D2559C8D">Zootaxa</mods:title>
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<mods:part id="693518F09922A74D6B7DCBCA86CF854F">
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<mods:date id="11C4DFDB5ABF000524A5BD4F884847BF">2019</mods:date>
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<mods:number id="AD4FEFCFF1F10F7A3A23A352F8BECCCD">2019-08-21</mods:number>
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<mods:number id="993433159B8991E125997A9812FDA843">4658</mods:number>
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<treatment id="8861532B88561827FF760BE2FCFFC37C" ID-DOI="http://doi.org/10.5281/zenodo.5667055" ID-GBIF-Taxon="159217211" ID-Zenodo-Dep="5667055" LSID="urn:lsid:plazi:treatment:8861532B88561827FF760BE2FCFFC37C" httpUri="http://treatment.plazi.org/id/8861532B88561827FF760BE2FCFFC37C" lastPageNumber="6" pageId="5" pageNumber="6">
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<subSubSection id="48D2B1B688561827FF760BE2FDDAC584" box="[151,633,693,720]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="0077E23D88561827FF760BE2FDDAC584" blockId="5.[151,633,693,720]" box="[151,633,693,720]" pageId="5" pageNumber="6">
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<heading id="5B3F555188561827FF760BE2FDDAC584" bold="true" box="[151,633,693,720]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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<emphasis id="32BC3E2F88561827FF760BE2FDDAC584" bold="true" box="[151,633,693,720]" pageId="5" pageNumber="6">
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Genus
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<taxonomicName id="C7C899BE88561827FF0F0BE2FDDAC584" ID-CoL="4G5D" authority="Balthasar, 1959" authorityName="Balthasar" authorityYear="1959" box="[238,633,693,720]" class="Insecta" family="Scarabaeidae" genus="Euonthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F88561827FF0F0BE2FE13C59B" bold="true" box="[238,432,693,720]" italics="true" pageId="5" pageNumber="6">Euonthophagus</emphasis>
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Balthasar, 1959
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="48D2B1B688561827FF760BAAFCFFC37C" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="0077E23D88561827FF760BAAFCFFC37C" blockId="5.[151,1437,764,1079]" pageId="5" pageNumber="6">
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<emphasis id="32BC3E2F88561827FF760BAAFEA7C45C" bold="true" box="[151,260,765,791]" pageId="5" pageNumber="6">Remarks</emphasis>
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.
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<taxonomicName id="C7C899BE88561827FEF00BAAFD1DC45C" authority="Klug, 1855" authorityName="Klug" authorityYear="1855" box="[273,702,764,791]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="carbonarius">
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<emphasis id="32BC3E2F88561827FEF00BAAFD9BC45C" box="[273,568,765,791]" italics="true" pageId="5" pageNumber="6">Onthophagus carbonarius</emphasis>
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Klug, 1855
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</taxonomicName>
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,
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<taxonomicName id="C7C899BE88561827FD280BAAFBBFC45C" authority="d'Orbigny, 1898" authorityName="d'Orbigny" authorityYear="1898" box="[713,1052,765,791]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maindroni">
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<emphasis id="32BC3E2F88561827FD280BAAFCFDC45C" box="[713,862,765,791]" italics="true" pageId="5" pageNumber="6">O. maindroni</emphasis>
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d’Orbigny, 1898
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</taxonomicName>
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and
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<taxonomicName id="C7C899BE88561827FBB30BAAFA3FC45C" authority="Balthasar, 1946" authorityName="Balthasar" authorityYear="1946" box="[1106,1436,765,791]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yemenicus">
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<emphasis id="32BC3E2F88561827FBB30BAAFB4BC45C" box="[1106,1256,765,791]" italics="true" pageId="5" pageNumber="6">O. yemenicus</emphasis>
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Balthasar, 1946
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</taxonomicName>
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have been placed in
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<taxonomicName id="C7C899BE88561827FE9F0A76FD8DC470" authorityName="Balthasar" authorityYear="1959" box="[382,558,801,827]" class="Insecta" family="Scarabaeidae" genus="Euonthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F88561827FE9F0A76FD8DC470" box="[382,558,801,827]" italics="true" pageId="5" pageNumber="6">Euonthophagus</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="64599FCC88561827FDB80A76FCB4C477" author="Balthasar, V." box="[601,791,801,828]" pageId="5" pageNumber="6" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar (1963)</bibRefCitation>
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, probably for the presence on the head of only one transverse carina in male or a middle tubercle in female, not homologous either with the clypeofrontal carina or the occipital one, and located either at the middle of the head or slightly forward of the middle. On the basis of the morphology of the male genital armature, in the first two species at least, this placement does not seem to have a phylogenetic basis. Most probably
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<taxonomicName id="C7C899BE88561827FDC20AE6FD6CC480" authorityName="Klug" authorityYear="1855" box="[547,719,945,971]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="carbonarius">
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<emphasis id="32BC3E2F88561827FDC20AE6FD6CC480" box="[547,719,945,971]" italics="true" pageId="5" pageNumber="6">O. carbonarius</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="C7C899BE88561827FCE60AE6FC3DC480" authorityName="d'Orbigny" authorityYear="1898" box="[775,926,945,971]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maindroni">
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<emphasis id="32BC3E2F88561827FCE60AE6FC3DC480" box="[775,926,945,971]" italics="true" pageId="5" pageNumber="6">O. maindroni</emphasis>
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</taxonomicName>
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belong to two different phylogenetic groups.
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<bibRefCitation id="64599FCC88561827FF760A82FEF2C4BB" author="Kabakov, O. N." box="[151,337,981,1008]" pageId="5" pageNumber="6" refId="ref38133" refString="Kabakov, O. N. (2006) Plastinchatousye zhuki podsemeystva Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) fauny Rossii i sopredel'nykh stran. [The lamellicorn beetle subfamily Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) in the fauna of Russia and adjacent countries]. Tovarishchestvo Nauchnykh Izdanii KMK, Moskva, 374 pp." type="book" year="2006">Kabakov (2006)</bibRefCitation>
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inserted
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<taxonomicName id="C7C899BE88561827FE5F0A82FDCEC4A4" authorityName="Klug" authorityYear="1855" box="[446,621,981,1007]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="carbonarius">
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<emphasis id="32BC3E2F88561827FE5F0A82FDCEC4A4" box="[446,621,981,1007]" italics="true" pageId="5" pageNumber="6">O. carbonarius</emphasis>
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</taxonomicName>
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in the subgenus
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<taxonomicName id="C7C899BE88561827FCD70A82FB69C4A4" authority="Balthasar, 1963" authorityName="Balthasar" authorityYear="1963" box="[822,1226,981,1007]" class="Insecta" family="Scarabaeidae" genus="Colobonthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F88561827FCD70A82FBA8C4A4" box="[822,1035,981,1007]" italics="true" pageId="5" pageNumber="6">Colobonthophagus</emphasis>
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<bibRefCitation id="64599FCC88561827FBF40A82FB69C4A4" author="Balthasar, V." box="[1045,1226,981,1007]" pageId="5" pageNumber="6" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar, 1963</bibRefCitation>
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</taxonomicName>
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, but not even this nomenclatorial act seems to be satisfactory. The placement of the three species is in great need of further study, and until such time we provisionally leave them in
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<taxonomicName id="C7C899BE88561827FD440D4AFCF6C37C" authorityName="Balthasar" authorityYear="1959" box="[677,853,1053,1079]" class="Insecta" family="Scarabaeidae" genus="Euonthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F88561827FD440D4AFCF6C37C" box="[677,853,1053,1079]" italics="true" pageId="5" pageNumber="6">Euonthophagus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |