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<document id="9C7661C0621A589A44BA49C5754DAD2A" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFFFEFFC0FCBEF8AB5B6D3827" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Hydraenopsis) pantitillata Perkins 2023, new species" docType="treatment" docVersion="2" lastPageNumber="50" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="47" updateTime="1700168056660" updateUser="ExternalLinkService">
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<mods:title id="698CD3B9C10CB1881CF0FFD96B03A295">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
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<mods:role id="F33730607617D049163BB42D21740237">
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<mods:namePart id="89BA151E09932CB24AD6C400E32DDB3E">Perkins, Philip D.</mods:namePart>
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<mods:title id="F118C1FAEF01DAC101F206795BD86974">Zootaxa</mods:title>
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<mods:part id="7E9DC6C1C91284F08EB23212BAD50EF5">
<mods:date id="45FCD1F66F06DBE9BF700FBF7128723F">2023</mods:date>
<mods:detail id="6C39080FC7A0626A81CB7EB93BFD9548" type="pubDate">
<mods:number id="D84B6ECCF11187A566E1013F07E11998">2023-11-07</mods:number>
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<mods:number id="DD91A0D6074144EE256BC094694A9C88">5367</mods:number>
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<mods:number id="42814C5D65B65F3D4FD31F6E1A18CB37">1</mods:number>
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<mods:url id="020A6F12A09C7B2AEFC95876A626CADB">https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225</mods:url>
</mods:location>
<mods:classification id="BF9B0CF689F9A2936A62F94927E0CC24">journal article</mods:classification>
<mods:identifier id="DA306A686C8D6E81FB7DF367E5ABCBA5" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
<mods:identifier id="B1CE3DDD6F2600B38B9E5993A824A7C6" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03FD878BFFFEFFC0FCBEF8AB5B6D3827" LSID="urn:lsid:plazi:treatment:03FD878BFFFEFFC0FCBEF8AB5B6D3827" httpUri="http://treatment.plazi.org/id/03FD878BFFFEFFC0FCBEF8AB5B6D3827" lastPageId="49" lastPageNumber="50" pageId="46" pageNumber="47">
<subSubSection id="C34E6516FFFEFFDFFCBEF8AB5B943B1F" box="[151,763,1806,1833]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BEB369DFFFEFFDFFCBEF8AB5B943B1F" blockId="46.[151,778,1806,1869]" box="[151,763,1806,1833]" pageId="46" pageNumber="47">
<heading id="D0A381F1FFFEFFDFFCBEF8AB5B943B1F" bold="true" box="[151,763,1806,1833]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
<emphasis id="B920EA8FFFFEFFDFFCBEF8AB5B943B1F" bold="true" box="[151,763,1806,1833]" pageId="46" pageNumber="47">
<taxonomicName id="4C544D1EFFFEFFDFFCBEF8AB5B333B1E" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,604,1806,1833]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pantitillata" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B920EA8FFFFEFFDFFCBEF8AB5B333B1E" bold="true" box="[151,604,1806,1833]" italics="true" pageId="46" pageNumber="47">Hydraena (Hydraenopsis) pantitillata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21357F4FFFEFFDFFE42F8AA5B943B1F" box="[619,763,1806,1833]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFFEFFDFFCBEF8965C243B8E" pageId="46" pageNumber="47" type="description">
<paragraph id="8BEB369DFFFEFFDFFCBEF8965A653B7A" blockId="46.[151,778,1806,1869]" box="[151,778,1843,1869]" pageId="46" pageNumber="47">
<figureCitation id="136F2A18FFFEFFDFFCBEF896598C3B7A" box="[151,227,1843,1869]" captionStart="FIGURE 2" captionStartId="6.[151,250,1867,1891]" captionTargetBox="[228,1360,181,1844]" captionTargetId="figure-35@6.[228,1360,181,1844]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.10145110" httpUri="https://zenodo.org/record/10145110/files/figure.png" pageId="46" pageNumber="47">Figs. 2</figureCitation>
(habitus comparison), 28 (habitus and aedeagus)
</paragraph>
<paragraph id="8BEB369DFFFEFFDFFCBEF8DE5C243B8E" blockId="46.[151,1436,1914,1977]" pageId="46" pageNumber="47">
Type Material.
<materialsCitation id="3B3C3CC0FFFEFFDFFD6CF8DE5B823B8E" collectionCode="FMNH" collectorName="A. Newton" country="Panama" location="El Hato del Volcan" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<typeStatus id="54EF883FFFFEFFDFFD6CF8DE58C23BA2" box="[325,429,1915,1941]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="F343760DFFFEFFDFFE31F8DE5BF93BA2" box="[536,662,1915,1941]" name="Panama" pageId="46" pageNumber="47">PANAMA</collectingCountry>
:
<collectingRegion id="4990F87FFFFEFFDFFE8BF8DE5B903BA2" box="[674,767,1915,1941]" country="Panama" name="Chiriqui" pageId="46" pageNumber="47">Chiriqui</collectingRegion>
<quantity id="4CAC9B78FFFEFFDFFF2EF8DE5A273BA2" box="[775,840,1915,1941]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="46" pageNumber="47" unit="km" value="24.0">24km</quantity>
W
<location id="8E8B6046FFFEFFDFFF58F8DE5D273BA2" LSID="urn:lsid:plazi:treatment:03FD878BFFFEFFC0FCBEF8AB5B6D3827:8E8B6046FFFEFFDFFF58F8DE5D273BA2" box="[881,1096,1915,1941]" country="Panama" name="El Hato del Volcan" pageId="46" pageNumber="47" stateProvince="Chiriqui">El Hato del Volcan</location>
3800’ VI.26–27.1976 // flood debris forest stream //
<collectorName id="26A1534BFFFEFFDFFDB9F83A5B643B8E" box="[400,523,1951,1977]" pageId="46" pageNumber="47">A. Newton</collectorName>
collector” (
<collectionCode id="ED45AE58FFFEFFDFFEA6F83A5B8D3B8E" box="[655,738,1951,1977]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="46" pageNumber="47" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3C3CC0FFFEFFDFFED0F83A5C293B8E" box="[761,1350,1951,1977]" collectionCode="FMNH" collectorName="A. Newton" country="Panama" location="El Hato del Volcan" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui" typeStatus="paratype">
<emphasis id="B920EA8FFFFEFFDFFED0F83A5A003B8E" bold="true" box="[761,879,1951,1977]" pageId="46" pageNumber="47">
<typeStatus id="54EF883FFFFEFFDFFED0F83A5A053B8E" box="[761,874,1951,1977]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
:
</emphasis>
Same data as holotype (
<specimenCount id="9D52FD14FFFEFFDFF8AAF83A5D8C3B8E" box="[1155,1251,1951,1977]" count="1" pageId="46" pageNumber="47" type="female">1 female</specimenCount>
<collectionCode id="ED45AE58FFFEFFDFF8C3F83A5C543B8E" box="[1258,1339,1951,1977]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="46" pageNumber="47" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2B6615FFFFFFDEFCBEF8A55DFF3B2F" ID-DOI="http://doi.org/10.5281/zenodo.10145158" ID-Zenodo-Dep="10145158" box="[151,1168,1792,1816]" httpUri="https://zenodo.org/record/10145158/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1792,1816]" targetBox="[291,1295,181,1768]" targetPageId="47" targetType="figure">
<paragraph id="8BEB369DFFFFFFDEFCBEF8A55DFF3B2F" blockId="47.[151,1168,1792,1816]" box="[151,1168,1792,1816]" pageId="47" pageNumber="48">
<emphasis id="B920EA8FFFFFFFDEFCBEF8A5584F3B2F" bold="true" box="[151,288,1792,1816]" pageId="47" pageNumber="48">FIGURE 27.</emphasis>
<taxonomicName id="4C544D1EFFFFFFDEFD0FF8A55BAC3B2F" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[294,707,1792,1816]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="panelephanta" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B920EA8FFFFFFFDEFD0FF8A558E43B2F" box="[294,395,1792,1816]" italics="true" pageId="47" pageNumber="48">Hydraena</emphasis>
(
<emphasis id="B920EA8FFFFFFFDEFDB3F8A55B483B2F" box="[410,551,1792,1816]" italics="true" pageId="47" pageNumber="48">Hydraenopsis</emphasis>
)
<emphasis id="B920EA8FFFFFFFDEFE1FF8A55BAC3B2F" box="[566,707,1792,1816]" italics="true" pageId="47" pageNumber="48">panelephanta</emphasis>
</taxonomicName>
<emphasis id="B920EA8FFFFFFFDEFEEBF8A55BA93B2F" box="[706,710,1792,1816]" italics="true" pageId="47" pageNumber="48">,</emphasis>
<taxonomicNameLabel id="A21357F4FFFFFFDEFEE5F8A45A293B2F" box="[716,838,1792,1816]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
, holotype habitus and aedeagus.
</paragraph>
</caption>
<caption id="DF2B6615FFE0FFC1FCBEF8A55D183B2F" ID-DOI="http://doi.org/10.5281/zenodo.10145160" ID-Zenodo-Dep="10145160" box="[151,1143,1792,1816]" httpUri="https://zenodo.org/record/10145160/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1792,1816]" targetBox="[291,1295,181,1768]" targetPageId="48" targetType="figure">
<paragraph id="8BEB369DFFE0FFC1FCBEF8A55D183B2F" blockId="48.[151,1143,1792,1816]" box="[151,1143,1792,1816]" pageId="48" pageNumber="49">
<emphasis id="B920EA8FFFE0FFC1FCBEF8A5584F3B2F" bold="true" box="[151,288,1792,1816]" pageId="48" pageNumber="49">FIGURE 28.</emphasis>
<taxonomicName id="4C544D1EFFE0FFC1FD0FF8A55BC63B2F" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[294,681,1792,1816]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pantitillata" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B920EA8FFFE0FFC1FD0FF8A558E43B2F" box="[294,395,1792,1816]" italics="true" pageId="48" pageNumber="49">Hydraena</emphasis>
(
<emphasis id="B920EA8FFFE0FFC1FDB3F8A55B483B2F" box="[410,551,1792,1816]" italics="true" pageId="48" pageNumber="49">Hydraenopsis</emphasis>
)
<emphasis id="B920EA8FFFE0FFC1FE1FF8A55BC63B2F" box="[566,681,1792,1816]" italics="true" pageId="48" pageNumber="49">pantitillata</emphasis>
</taxonomicName>
<emphasis id="B920EA8FFFE0FFC1FE80F8A55BC23B2F" box="[681,685,1792,1816]" italics="true" pageId="48" pageNumber="49">,</emphasis>
<taxonomicNameLabel id="A21357F4FFE0FFC1FE9AF8A45A433B2F" box="[691,812,1792,1816]" pageId="48" pageNumber="49" rank="species">new species</taxonomicNameLabel>
, holotype habitus and aedeagus.
</paragraph>
</caption>
<subSubSection id="C34E6516FFE0FFC1FCEEF8CF5DD03BC6" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BEB369DFFE0FFC1FCEEF8CF5DD03BC6" blockId="48.[151,1437,1898,2033]" pageId="48" pageNumber="49">
<emphasis id="B920EA8FFFE0FFC1FCEEF8CF58BB3BB3" bold="true" box="[199,468,1898,1924]" pageId="48" pageNumber="49">Differential Diagnosis.</emphasis>
This new species, which is a member of the
<taxonomicName id="4C544D1EFFE0FFC1FFF5F8CE5C653BB3" authority="Subgroup" authorityName="Subgroup" box="[988,1290,1898,1924]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B920EA8FFFE0FFC1FFF5F8CE5DFB3BB3" box="[988,1172,1898,1924]" italics="true" pageId="48" pageNumber="49">H. marginicollis</emphasis>
Subgroup
</taxonomicName>
(see
<bibRefCitation id="EFC54B6CFFE0FFC1F961F8CF59933B9F" author="Perkins, P. D." pageId="48" pageNumber="49" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 69</bibRefCitation>
), can be differentiated from others by the combination of the small body size (ca.
<quantity id="4CAC9B78FFE0FFC1F8A2F82A5D9D3B9E" box="[1163,1266,1935,1961]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="48" pageNumber="49" unit="mm" value="1.3">1.30 mm</quantity>
), the very wide metaventral intercoxal process, the carinate metaventral plaques (P2/w/l/s ratios ca. 6/1/6/6), and the aedeagus (
<figureCitation id="136F2A18FFE0FFC1F946F81759D83BC6" captionStart="FIGURE 28" captionStartId="48.[151,250,1792,1816]" captionTargetBox="[291,1295,181,1768]" captionTargetId="figure-35@48.[291,1295,181,1768]" captionTargetPageId="48" captionText="FIGURE 28. Hydraena (Hydraenopsis) pantitillata, new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145160" httpUri="https://zenodo.org/record/10145160/files/figure.png" pageId="48" pageNumber="49">Fig. 28</figureCitation>
). The aedeagus cannot be favorably compared with any other known New World
<taxonomicName id="4C544D1EFFE0FFC1F861F8735DD73BC7" box="[1096,1208,2006,2032]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B920EA8FFFE0FFC1F861F8735DD73BC7" box="[1096,1208,2006,2032]" italics="true" pageId="48" pageNumber="49">Hydraena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFE1FFC0FCEEFF335BAE3F6A" pageId="49" pageNumber="50" type="description">
<paragraph id="8BEB369DFFE1FFC0FCEEFF335A353DBE" blockId="49.[151,1437,150,933]" pageId="49" pageNumber="50">
<emphasis id="B920EA8FFFE1FFC0FCEEFF33583E3C87" bold="true" box="[199,337,150,176]" pageId="49" pageNumber="50">Description</emphasis>
. Size:
<typeStatus id="54EF883FFFE1FFC0FD8DFF325B693C86" box="[420,518,151,177]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.30/0.59; head width 0.33; pronotum 0.33/0.45, PA 0.38, PB 0.38; elytra 0.78/0.59. Habitus as illustrated (
<figureCitation id="136F2A18FFE1FFC0F810FF1E5DFE3CE2" box="[1081,1169,187,213]" captionStart="FIGURE 28" captionStartId="48.[151,250,1792,1816]" captionTargetBox="[291,1295,181,1768]" captionTargetId="figure-35@48.[291,1295,181,1768]" captionTargetPageId="48" captionText="FIGURE 28. Hydraena (Hydraenopsis) pantitillata, new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145160" httpUri="https://zenodo.org/record/10145160/files/figure.png" pageId="49" pageNumber="50">Fig. 28</figureCitation>
). Dorsum brown, head and pronotal disc darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum finely sparsely punctulate, interstices shining; punctures of frons disc and pronotal disc ca. 1–2xef, interstices ca. 2–3xpd or slightly larger, each puncture with short, recumbent seta; punctures of clypeus very fine. Labrum apicomedially deeply excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal fovea shallow, other usual foveae absent.
</paragraph>
<paragraph id="8BEB369DFFE1FFC0FCEEFE365DB63E2E" blockId="49.[151,1437,150,933]" pageId="49" pageNumber="50">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 2–3xpd or slightly greater, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="8BEB369DFFE1FFC0FCEEFD865BAE3F6A" blockId="49.[151,1437,150,933]" pageId="49" pageNumber="50">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely punctulate, shining, with concavity on each side. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 6/1/6/6. P1 moderately wide; median carina sinuate in profile. P2 very wide, l/w ca. 1/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques carinate, straight, slightly converging toward one another anteriorly, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin slightly less than width of P2. Protibia (male) with very small, acute process on inner margin slightly distal to midlength. Mesotibia straight. Metatibia inner margin slightly widened. Abdominal apex symmetrical; last tergite with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFE1FFC0FCEEFCC35A323FB6" box="[199,861,870,897]" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8BEB369DFFE1FFC0FCEEFCC35A323FB6" blockId="49.[151,1437,150,933]" box="[199,861,870,897]" pageId="49" pageNumber="50">
<emphasis id="B920EA8FFFE1FFC0FCEEFCC3580E3FB7" bold="true" box="[199,353,870,896]" pageId="49" pageNumber="50">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54EF883FFFE1FFC0FEE5FCC25B933FB6" box="[716,764,871,897]" pageId="49" pageNumber="50">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFE1FFC0FCEEFC2E5B6D3827" pageId="49" pageNumber="50" type="etymology">
<paragraph id="8BEB369DFFE1FFC0FCEEFC2E5A5F3F92" blockId="49.[151,1437,150,933]" box="[199,816,907,933]" pageId="49" pageNumber="50">
<emphasis id="B920EA8FFFE1FFC0FCEEFC2E58213F92" bold="true" box="[199,334,907,933]" pageId="49" pageNumber="50">Etymology.</emphasis>
Named in reference to the spiny aedeagus.
</paragraph>
<paragraph id="8BEB369DFFE1FFC0FCBEFC525B6D3827" blockId="49.[151,514,1014,1041]" box="[151,514,1014,1041]" pageId="49" pageNumber="50">
<emphasis id="B920EA8FFFE1FFC0FCBEFC525B6D3827" bold="true" box="[151,514,1014,1041]" pageId="49" pageNumber="50">
New species from
<collectingCountry id="F343760DFFE1FFC0FD51FC535B6D3827" box="[376,514,1014,1040]" name="Saint Vincent and the Grenadines" pageId="49" pageNumber="50">St. Vincent</collectingCountry>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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