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<document id="FADF74421B74A59CAFD11FA6C67606C7" ID-CLB-Dataset="24935" ID-DOI="10.11646/zootaxa.4695.3.1" ID-GBIF-Dataset="256591b2-6d7b-465d-a818-9a158017ca09" ID-ISSN="1175-5326" ID-Zenodo-Dep="3532515" ID-ZooBank="32E17629-F29A-47C8-ACCB-69A32C17B147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573197328991" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03ED0167FFA3FF9BFF1DF130FA01FB3D" docLanguage="en" docName="zootaxa.4695.3.1.pdf" docOrigin="Zootaxa 4695 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus villosus Matthews 1974" docType="key" docVersion="8" lastPageNumber="239" masterDocId="FFD4791FFF85FFBDFF8AF379FF9DFFA8" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups" masterLastPageNumber="252" masterPageNumber="201" pageNumber="239" updateTime="1698764404900" updateUser="ExternalLinkService">
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<mods:title id="62AFE01851F8AFFAD5F839E7FC081020">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups</mods:title>
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<mods:namePart id="54FD6B1771AA50F6824CDE08ACF22EC2">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="9922CB0D323F12FC49A088B50DC1F03C">Weir, Thomas A.</mods:namePart>
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<mods:title id="577B1C117A2B43BC3EA64AE9F8DCB3BF">Zootaxa</mods:title>
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<mods:date id="C9A8E01B1A9BE77F4D5C8F90F75B8A97">2019</mods:date>
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<treatment id="03ED0167FFA3FF9BFF1DF130FA01FB3D" ID-GBIF-Taxon="160217996" LSID="urn:lsid:plazi:treatment:03ED0167FFA3FF9BFF1DF130FA01FB3D" httpUri="http://treatment.plazi.org/id/03ED0167FFA3FF9BFF1DF130FA01FB3D" lastPageNumber="239" pageId="38" pageNumber="239">
<subSubSection id="C35EE3FAFFA3FF9BFF1DF130FD55FDCB" box="[151,712,585,612]" pageId="38" pageNumber="239" type="nomenclature">
<paragraph id="8BFBB071FFA3FF9BFF1DF130FD55FDCB" blockId="38.[151,712,585,612]" box="[151,712,585,612]" pageId="38" pageNumber="239">
<heading id="D0B3071DFFA3FF9BFF1DF130FD55FDCB" bold="true" box="[151,712,585,612]" fontSize="11" level="1" pageId="38" pageNumber="239" reason="1">
<emphasis id="B9306C63FFA3FF9BFF1DF130FD55FDCB" bold="true" box="[151,712,585,612]" pageId="38" pageNumber="239">
Key to species
<taxonomicName id="4C44CBF2FFA3FF9BFEC4F130FD84FDCC" ID-CoL="6P89S" authorityName="Matthews" authorityYear="1974" box="[334,537,585,612]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="B9306C63FFA3FF9BFEC4F130FD84FDCC" bold="true" box="[334,537,585,612]" italics="true" pageId="38" pageNumber="239">Lepanus villosus</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFFA3FF9BFF1DF1F7FA01FB3D" pageId="38" pageNumber="239" type="key">
<keyStep id="30B0A8D4FFA3FF9BFF1DF1F7FA01FD68" pageId="38" pageNumber="239">
<paragraph id="8BFBB071FFA3FF9BFF1DF1F7FA01FD0A" blockId="38.[151,1437,653,1174]" box="[151,1436,653,675]" pageId="38" pageNumber="239">
<keyLead id="30B51344FFA3FF9BFF1DF1F7FA01FD0A" box="[151,1436,653,675]" pageId="38" pageNumber="239">
1. Elytra with stria 8 ending in a deep pit basally and lateral carina indented at this point (
<figureCitation id="137FACF4FFA3FF9BFBAEF1F4FBE1FD0B" box="[1060,1148,653,675]" captionStart="FIGURE 16" captionStartId="39.[151,250,1781,1806]" captionTargetBox="[177,1410,166,1762]" captionTargetId="figure@39.[177,1410,166,1762]" captionTargetPageId="39" captionText="FIGURE 16. Some diagnostic characters of the Lepanus villosus species group. A, lateral view of elytron of Lepanus villosus, white arrow points to deep pit at the base of stria 8; B, lateral view of elytron of Lepanus vestitus, note the absence of any deep pit; C, elytron of Lepanus villosus dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5; D, male metatibiae of Lepanus vestitus, notch on inner edge; E, male protibia of Lepanus pseudovillosus, showing typical configuration for the species group including the small comb of setae (grey arrow); F, close up of elytral interval surface of Lepanus reticulatus, showing reticulation of short longitudinal striations; G, close up of elytral interval surface of Lepanus vestitus showing reticulations of fine meshes." figureDoi="http://doi.org/10.5281/zenodo.3532547" httpUri="https://zenodo.org/record/3532547/files/figure.png" pageId="38" pageNumber="239">Fig. 16A</figureCitation>
)......................
<emphasis id="B9306C63FFA3FF9BFA1AF1F4FA01FD0A" bold="true" box="[1424,1436,653,674]" pageId="38" pageNumber="239">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFBB071FFA3FF9BFF1DF1D2FA01FD68" blockId="38.[151,1437,653,1174]" box="[151,1436,683,705]" pageId="38" pageNumber="239">
<keyLead id="30B51344FFA3FF9BFF1DF1D2FA01FD68" box="[151,1436,683,705]" pageId="38" pageNumber="239">
- Elytra with stria 8 not ending in a deep pit basally and lateral carina not indented at this point (
<figureCitation id="137FACF4FFA3FF9BFBE4F1D2FB5EFD69" box="[1134,1219,683,705]" captionStart="FIGURE 16" captionStartId="39.[151,250,1781,1806]" captionTargetBox="[177,1410,166,1762]" captionTargetId="figure@39.[177,1410,166,1762]" captionTargetPageId="39" captionText="FIGURE 16. Some diagnostic characters of the Lepanus villosus species group. A, lateral view of elytron of Lepanus villosus, white arrow points to deep pit at the base of stria 8; B, lateral view of elytron of Lepanus vestitus, note the absence of any deep pit; C, elytron of Lepanus villosus dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5; D, male metatibiae of Lepanus vestitus, notch on inner edge; E, male protibia of Lepanus pseudovillosus, showing typical configuration for the species group including the small comb of setae (grey arrow); F, close up of elytral interval surface of Lepanus reticulatus, showing reticulation of short longitudinal striations; G, close up of elytral interval surface of Lepanus vestitus showing reticulations of fine meshes." figureDoi="http://doi.org/10.5281/zenodo.3532547" httpUri="https://zenodo.org/record/3532547/files/figure.png" pageId="38" pageNumber="239">Fig. 16B</figureCitation>
)................
<emphasis id="B9306C63FFA3FF9BFA1AF1D2FA01FD68" bold="true" box="[1424,1436,683,704]" pageId="38" pageNumber="239">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B0A8D4FFA3FF9BFF1DF1B1FA00FC26" pageId="38" pageNumber="239">
<paragraph id="8BFBB071FFA3FF9BFF1DF1B1FA01FCB0" blockId="38.[151,1437,653,1174]" pageId="38" pageNumber="239">
<keyLead id="30B51344FFA3FF9BFF1DF1B1FA01FCB0" pageId="38" pageNumber="239">
2. Elytra with a well-developed humeral callus basally between striae 6 and 7 and a distinct depression basally between striae 4 and 5 (
<figureCitation id="137FACF4FFA3FF9BFEACF19CFE1DFD53" box="[294,384,741,763]" captionStart="FIGURE 16" captionStartId="39.[151,250,1781,1806]" captionTargetBox="[177,1410,166,1762]" captionTargetId="figure@39.[177,1410,166,1762]" captionTargetPageId="39" captionText="FIGURE 16. Some diagnostic characters of the Lepanus villosus species group. A, lateral view of elytron of Lepanus villosus, white arrow points to deep pit at the base of stria 8; B, lateral view of elytron of Lepanus vestitus, note the absence of any deep pit; C, elytron of Lepanus villosus dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5; D, male metatibiae of Lepanus vestitus, notch on inner edge; E, male protibia of Lepanus pseudovillosus, showing typical configuration for the species group including the small comb of setae (grey arrow); F, close up of elytral interval surface of Lepanus reticulatus, showing reticulation of short longitudinal striations; G, close up of elytral interval surface of Lepanus vestitus showing reticulations of fine meshes." figureDoi="http://doi.org/10.5281/zenodo.3532547" httpUri="https://zenodo.org/record/3532547/files/figure.png" pageId="38" pageNumber="239">Fig. 16C</figureCitation>
); pygidium flat throughout (
<figureCitation id="137FACF4FFA3FF9BFD1FF19CFD6DFD54" box="[661,752,741,764]" captionStart="FIGURE 19" captionStartId="42.[151,250,859,884]" captionTargetBox="[159,1427,184,840]" captionTargetId="figure@42.[151,1436,174,840]" captionTargetPageId="42" captionText="FIGURE 19. Pygidial configuration of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532553" httpUri="https://zenodo.org/record/3532553/files/figure.png" pageId="38" pageNumber="239">Fig. 19D</figureCitation>
); mesepimeron punctate; hind wings fully developed; male metatibiae with a notch on inner edge; aedeagus as in Fig 22D; northern
<collectingRegion id="49807E93FFA3FF9BFCB8F07AFC3BFCB1" box="[818,934,771,793]" country="Australia" name="Queensland" pageId="38" pageNumber="239">Queensland</collectingRegion>
, Wet Tropics...
<taxonomicName id="4C44CBF2FFA3FF9BFBC5F07AFA01FCB0" ID-CoL="6P89S" authority="Matthews, 1974" authorityName="Matthews" authorityYear="1974" box="[1103,1436,771,793]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="B9306C63FFA3FF9BFBC5F07AFA01FCB0" bold="true" box="[1103,1436,771,793]" pageId="38" pageNumber="239">
<emphasis id="B9306C63FFA3FF9BFBC5F07AFB68FCB1" bold="true" box="[1103,1269,771,793]" italics="true" pageId="38" pageNumber="239">Lepanus villosus</emphasis>
<bibRefCitation id="EFD5CD80FFA3FF9BFB71F07AFA01FCB0" author="Matthews, E. G." box="[1275,1436,771,792]" pageId="38" pageNumber="239" pagination="1 - 211" refId="ref56271" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211. https: // doi. org / 10.1071 / AJZS 024" type="journal article" year="1974">Matthews, 1974</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFBB071FFA3FF9BFF1DF058FA00FC26" blockId="38.[151,1437,653,1174]" pageId="38" pageNumber="239">
<keyLead id="30B51344FFA3FF9BFF1DF058FA00FC26" pageId="38" pageNumber="239">
- Elytra without a well-developed humeral callus basally between striae 6 and 7 or a distinct depression basally between striae 4 and 5; pygidium with a raised shiny knob apically – more so in males than females (
<figureCitation id="137FACF4FFA3FF9BFB82F044FBFFFCFC" box="[1032,1122,829,852]" captionStart="FIGURE 19" captionStartId="42.[151,250,859,884]" captionTargetBox="[159,1427,184,840]" captionTargetId="figure@42.[151,1436,174,840]" captionTargetPageId="42" captionText="FIGURE 19. Pygidial configuration of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532553" httpUri="https://zenodo.org/record/3532553/files/figure.png" pageId="38" pageNumber="239">Fig. 19A</figureCitation>
); mesepimeron impunctate; hind wings vestigial; male metatibiae gently curved on inner edge; aedeagus as in
<figureCitation id="137FACF4FFA3FF9BFC4DF022FB82FCD8" box="[967,1055,859,881]" captionStart="FIGURE 18" captionStartId="41.[151,250,1897,1922]" captionTargetBox="[153,1430,180,1864]" captionTargetId="figure@41.[151,1436,166,1876]" captionTargetPageId="41" captionText="FIGURE 18. Left, dorsal, and right view of aedeagi of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532551" httpUri="https://zenodo.org/record/3532551/files/figure.png" pageId="38" pageNumber="239">Fig. 18A</figureCitation>
; northern
<collectingRegion id="49807E93FFA3FF9BFB0AF022FB69FCD9" box="[1152,1268,859,881]" country="Australia" name="Queensland" pageId="38" pageNumber="239">Queensland</collectingRegion>
, Wet Tropics...........................................................
<emphasis id="B9306C63FFA3FF9BFC05F000FA00FC26" bold="true" box="[911,1437,888,910]" pageId="38" pageNumber="239">
<taxonomicName id="4C44CBF2FFA3FF9BFC05F000FA81FC26" ID-CoL="3T8VH" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[911,1308,888,910]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="pseudovillosus" status="sp. nov.">
<emphasis id="B9306C63FFA3FF9BFC05F000FBE7FC26" bold="true" box="[911,1146,888,910]" italics="true" pageId="38" pageNumber="239">Lepanus pseudovillosus</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FFA3FF9BFAACF001FA00FC26" box="[1318,1437,888,910]" pageId="38" pageNumber="239" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B0A8D4FFA3FF9BFF1DF0EFFA01FB3D" pageId="38" pageNumber="239">
<paragraph id="8BFBB071FFA3FF9BFF1DF0EFFA00FB89" blockId="38.[151,1437,653,1174]" pageId="38" pageNumber="239">
<keyLead id="30B51344FFA3FF9BFF1DF0EFFA00FB89" pageId="38" pageNumber="239">
3. Larger species, about
<quantity id="4CBC1D94FFA3FF9BFE3FF0ECFE63FC03" box="[437,510,917,939]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="38" pageNumber="239" unit="mm" value="3.5">3.5 mm</quantity>
in length; clypeal teeth large and markedly upturned; reticulation strongly developed on all elytral intervals, consisting of dense, short, longitudinally aligned striae (
<figureCitation id="137FACF4FFA3FF9BFC06F0CAFC7EFC61" box="[908,995,947,969]" captionStart="FIGURE 16" captionStartId="39.[151,250,1781,1806]" captionTargetBox="[177,1410,166,1762]" captionTargetId="figure@39.[177,1410,166,1762]" captionTargetPageId="39" captionText="FIGURE 16. Some diagnostic characters of the Lepanus villosus species group. A, lateral view of elytron of Lepanus villosus, white arrow points to deep pit at the base of stria 8; B, lateral view of elytron of Lepanus vestitus, note the absence of any deep pit; C, elytron of Lepanus villosus dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5; D, male metatibiae of Lepanus vestitus, notch on inner edge; E, male protibia of Lepanus pseudovillosus, showing typical configuration for the species group including the small comb of setae (grey arrow); F, close up of elytral interval surface of Lepanus reticulatus, showing reticulation of short longitudinal striations; G, close up of elytral interval surface of Lepanus vestitus showing reticulations of fine meshes." figureDoi="http://doi.org/10.5281/zenodo.3532547" httpUri="https://zenodo.org/record/3532547/files/figure.png" pageId="38" pageNumber="239">Fig. 16F</figureCitation>
); mesoventrite punctate; pygidium somewhat convex, with an indication of a raised longitudinal midline (
<figureCitation id="137FACF4FFA3FF9BFCA3F0A9FC1CFC4E" box="[809,897,976,998]" captionStart="FIGURE 19" captionStartId="42.[151,250,859,884]" captionTargetBox="[159,1427,184,840]" captionTargetId="figure@42.[151,1436,174,840]" captionTargetPageId="42" captionText="FIGURE 19. Pygidial configuration of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532553" httpUri="https://zenodo.org/record/3532553/files/figure.png" pageId="38" pageNumber="239">Fig. 19B</figureCitation>
); hind wings vestigial; male metatibiae with inner edge virtually straight; aedeagus as in
<figureCitation id="137FACF4FFA3FF9BFD94F094FDE8FBAB" box="[542,629,1005,1027]" captionStart="FIGURE 18" captionStartId="41.[151,250,1897,1922]" captionTargetBox="[153,1430,180,1864]" captionTargetId="figure@41.[151,1436,166,1876]" captionTargetPageId="41" captionText="FIGURE 18. Left, dorsal, and right view of aedeagi of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532551" httpUri="https://zenodo.org/record/3532551/files/figure.png" pageId="38" pageNumber="239">Fig. 18B</figureCitation>
; northern
<collectingRegion id="49807E93FFA3FF9BFD5CF094FCD7FBAB" box="[726,842,1005,1027]" country="Australia" name="Queensland" pageId="38" pageNumber="239">Queensland</collectingRegion>
, Mount Lewis area..............................................................................................
<emphasis id="B9306C63FFA3FF9BFC3FF772FA00FB89" bold="true" box="[949,1437,1035,1057]" pageId="38" pageNumber="239">
<taxonomicName id="4C44CBF2FFA3FF9BFC3FF772FA81FB89" ID-CoL="3T8VM" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[949,1308,1035,1057]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<emphasis id="B9306C63FFA3FF9BFC3FF772FBE7FB89" bold="true" box="[949,1146,1035,1057]" italics="true" pageId="38" pageNumber="239">Lepanus reticulatus</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FFA3FF9BFAACF772FA00FB89" box="[1318,1437,1035,1057]" pageId="38" pageNumber="239" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BFBB071FFA3FF9BFF1DF750FA01FB3D" blockId="38.[151,1437,653,1174]" pageId="38" pageNumber="239">
<keyLead id="30B51344FFA3FF9BFF1DF750FA01FB3D" pageId="38" pageNumber="239">
- Smaller species, less than 3.0 mm in length; clypeal teeth small and not markedly upturned; reticulation on elytral intervals less strongly developed and often confined to apical and lateral areas of elytra, consisting of meshes (
<figureCitation id="137FACF4FFA3FF9BFB3FF73CFA92FBF3" box="[1205,1295,1093,1115]" captionStart="FIGURE 16" captionStartId="39.[151,250,1781,1806]" captionTargetBox="[177,1410,166,1762]" captionTargetId="figure@39.[177,1410,166,1762]" captionTargetPageId="39" captionText="FIGURE 16. Some diagnostic characters of the Lepanus villosus species group. A, lateral view of elytron of Lepanus villosus, white arrow points to deep pit at the base of stria 8; B, lateral view of elytron of Lepanus vestitus, note the absence of any deep pit; C, elytron of Lepanus villosus dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5; D, male metatibiae of Lepanus vestitus, notch on inner edge; E, male protibia of Lepanus pseudovillosus, showing typical configuration for the species group including the small comb of setae (grey arrow); F, close up of elytral interval surface of Lepanus reticulatus, showing reticulation of short longitudinal striations; G, close up of elytral interval surface of Lepanus vestitus showing reticulations of fine meshes." figureDoi="http://doi.org/10.5281/zenodo.3532547" httpUri="https://zenodo.org/record/3532547/files/figure.png" pageId="38" pageNumber="239">Fig. 16G</figureCitation>
); mesoventrite impunctate; pygidium virtually flat throughout (
<figureCitation id="137FACF4FFA3FF9BFD3BF71AFC94FBD1" box="[689,777,1123,1145]" captionStart="FIGURE 19" captionStartId="42.[151,250,859,884]" captionTargetBox="[159,1427,184,840]" captionTargetId="figure@42.[151,1436,174,840]" captionTargetPageId="42" captionText="FIGURE 19. Pygidial configuration of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532553" httpUri="https://zenodo.org/record/3532553/files/figure.png" pageId="38" pageNumber="239">Fig. 19C</figureCitation>
); hind wings fully developed; male metatibiae with a notch on inner edge (
<figureCitation id="137FACF4FFA3FF9BFE96F7F9FEE8FB3E" box="[284,373,1152,1174]" captionStart="FIGURE 16" captionStartId="39.[151,250,1781,1806]" captionTargetBox="[177,1410,166,1762]" captionTargetId="figure@39.[177,1410,166,1762]" captionTargetPageId="39" captionText="FIGURE 16. Some diagnostic characters of the Lepanus villosus species group. A, lateral view of elytron of Lepanus villosus, white arrow points to deep pit at the base of stria 8; B, lateral view of elytron of Lepanus vestitus, note the absence of any deep pit; C, elytron of Lepanus villosus dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5; D, male metatibiae of Lepanus vestitus, notch on inner edge; E, male protibia of Lepanus pseudovillosus, showing typical configuration for the species group including the small comb of setae (grey arrow); F, close up of elytral interval surface of Lepanus reticulatus, showing reticulation of short longitudinal striations; G, close up of elytral interval surface of Lepanus vestitus showing reticulations of fine meshes." figureDoi="http://doi.org/10.5281/zenodo.3532547" httpUri="https://zenodo.org/record/3532547/files/figure.png" pageId="38" pageNumber="239">Fig. 16D</figureCitation>
); aedeagus as in
<figureCitation id="137FACF4FFA3FF9BFD92F7F9FDF3FB3D" box="[536,622,1152,1174]" captionStart="FIGURE 18" captionStartId="41.[151,250,1897,1922]" captionTargetBox="[153,1430,180,1864]" captionTargetId="figure@41.[151,1436,166,1876]" captionTargetPageId="41" captionText="FIGURE 18. Left, dorsal, and right view of aedeagi of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532551" httpUri="https://zenodo.org/record/3532551/files/figure.png" pageId="38" pageNumber="239">Fig. 18C</figureCitation>
; central
<collectingRegion id="49807E93FFA3FF9BFD4AF7F9FCA9FB3E" box="[704,820,1152,1174]" country="Australia" name="Queensland" pageId="38" pageNumber="239">Queensland</collectingRegion>
, Eungella area...........
<taxonomicName id="4C44CBF2FFA3FF9BFBDAF7F8FA01FB3D" ID-CoL="3T8VT" authority="Matthews, 1974" authorityName="Matthews" authorityYear="1974" box="[1104,1436,1152,1174]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B9306C63FFA3FF9BFBDAF7F8FAFDFB3D" bold="true" box="[1104,1376,1152,1174]" pageId="38" pageNumber="239">
<emphasis id="B9306C63FFA3FF9BFBDAF7F8FB68FB3D" bold="true" box="[1104,1269,1152,1174]" italics="true" pageId="38" pageNumber="239">Lepanus vestitus</emphasis>
<bibRefCitation id="EFD5CD80FFA3FF9BFB71F7F9FAFDFB3D" author="Matthews, E. G." box="[1275,1376,1152,1173]" pageId="38" pageNumber="239" pagination="1 - 211" refId="ref56271" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211. https: // doi. org / 10.1071 / AJZS 024" type="journal article" year="1974">Matthews</bibRefCitation>
</emphasis>
, 1974
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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