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<document id="9990E5914EFC0A765AC52B855D4365E1" ID-DOI="10.11646/zootaxa.3858.1.1" ID-GBIF-Dataset="d6152690-b0f3-4f6f-9fea-05c3fb53690e" ID-ISSN="1175-5326" ID-Zenodo-Dep="287013" ID-ZooBank="D1CCF02B-4314-4537-A64F-0372715E3F93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460902618942" checkinUser="plazi" docAuthor="Fleming, Alan J., Wood, Monty, Smith, Alex, Hallwachs, Winnie &amp; Janzen, Daniel H." docDate="2014" docId="03D087FFB7018F1CFF1AFA73FA4DFB17" docLanguage="en" docName="zt03858p090.pdf" docOrigin="Zootaxa 3858 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Houghia Reared" docType="key" docVersion="7" lastPageNumber="32" masterDocId="FFE9FF87B71C8F03FF8DFF96FFD1FFFD" masterDocTitle="Revision of the New World species of Houghia Coquillett (Diptera, Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, Costa Rica" masterLastPageNumber="90" masterPageNumber="1" pageNumber="30" updateTime="1698360527715" updateUser="plazi">
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<mods:title id="B6845C3FF86228550CAB8ED481DB6C0E">Revision of the New World species of Houghia Coquillett (Diptera, Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, Costa Rica</mods:title>
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<mods:namePart id="6758EBDE3F190A0F473D9F3D414E8608">Fleming, Alan J.</mods:namePart>
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<mods:namePart id="15B4501177698C3BFFBA9033BA7561BA">Wood, Monty</mods:namePart>
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<mods:namePart id="6FC71F741E5FC8A924D541FB79BE8EB7">Smith, Alex</mods:namePart>
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<mods:namePart id="AD9906EA57B04A40000C6159D1B3FD39">Hallwachs, Winnie</mods:namePart>
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<mods:namePart id="79CBFCD1A5FF007FB0065BE38DDF67CB">Janzen, Daniel H.</mods:namePart>
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<paragraph id="8BC636E9B7018F1EFF1AFA73FB13FA02" blockId="29.[151,1218,1509,1535]" box="[151,1218,1509,1535]" pageId="29" pageNumber="30">
<heading id="D08E8185B7018F1EFF1AFA73FB13FA02" bold="true" box="[151,1218,1509,1535]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B90DEAFBB7018F1EFF1AFA73FB13FA02" bold="true" box="[151,1218,1509,1535]" pageId="29" pageNumber="30">
Key to Males of
<taxonomicName id="4C794D6AB7018F1EFEEFFA73FDE1FA02" ID-CoL="636XL" ID-ENA="695878" authority="Reared" authorityName="Reared" box="[354,560,1509,1535]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEAFBB7018F1EFEEFFA73FE1FFA02" bold="true" box="[354,462,1509,1535]" italics="true" pageId="29" pageNumber="30">Houghia</emphasis>
Reared
</taxonomicName>
from Caterpillars in ACG, Northwestern
<collectingCountry id="F36E7679B7018F1EFBB7FA73FB13FA02" box="[1082,1218,1509,1535]" name="Costa Rica" pageId="29" pageNumber="30">Costa Rica</collectingCountry>
</emphasis>
</heading>
</paragraph>
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<keyStep id="308D2E4CB7018F1EFF1AF9BAFA4AF95D" pageId="29" pageNumber="30">
<paragraph id="8BC636E9B7018F1EFF1AF9BAFA4AF99C" blockId="29.[151,1437,1579,2019]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF9BAFA4AF99C" pageId="29" pageNumber="30">
1 Sex patches (areas of shorter, denser, appressed hairs, often referred to as sex patches or
<taxonomicName id="4C794D6AB7018F1EFBBAF9BDFB53F9BC" ID-CoL="7PLH" box="[1079,1154,1579,1601]" class="Insecta" family="Tachinidae" genus="Sturmia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEAFBB7018F1EFBBAF9BDFB53F9BC" box="[1079,1154,1579,1601]" italics="true" pageId="29" pageNumber="30">Sturmia</emphasis>
</taxonomicName>
spots, present only in males) lacking, undersides of tergites 4 and 5 tomentose as on dorsal surfaces; vein R1 setose along its entire length dorsally....... 2
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7018F1EFF1AF9FDFA4AF95D" blockId="29.[151,1437,1579,2019]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF9FDFA4AF95D" pageId="29" pageNumber="30">
- Sex patches present (
<figureCitation id="13422A6CB7018F1EFE22F9FDFE29F97D" box="[431,504,1643,1664]" captionStart="FIGURE 10" captionStartId="29.[151,250,1315,1337]" captionTargetBox="[340,1246,375,1293]" captionTargetId="figure@29.[340,1247,375,1293]" captionTargetPageId="29" captionText="FIGURE 10. Ventral view of abdomen showing different configurations of sex patches. a. Houghia graciloides sp. nov., showing characteristic sex patch on tergites 3, 4, and 5. b. Houghia longipilosa sp. nov., with sex patch present on tergites 4 and 5 only. c. Houghia omissa sp. nov., with sex patch present on tergite 4 only. d. Houghia confinis sp. nov with sex patch on tergites 4 and 5. e. Houghia macilenta sp. nov., with no sex patch at all." httpUri="https://zenodo.org/record/287023/files/figure.png" pageId="29" pageNumber="30">Fig. 10</figureCitation>
); if in doubt undersurface of tergite 4 differing in appearance from dorsal surface; vein R1 bare dorsally............................................................................................. 5
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<keyStep id="308D2E4CB7018F1EFF1AF93EFA4AF8CE" pageId="29" pageNumber="30">
<paragraph id="8BC636E9B7018F1EFF1AF93EFA4AF905" blockId="29.[151,1437,1579,2019]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF93EFA4AF905" pageId="29" pageNumber="30">
2 Scape and upper third or more of first flagellomere pale orange; first flagellomere slightly to strongly curved or sinuous (
<figureCitation id="13422A6CB7018F1EFAE6F93EFF2AF926" captionStart="FIGURE 35" captionStartId="71.[151,250,996,1018]" captionTargetBox="[340,1246,337,974]" captionTargetId="figure@71.[340,1247,337,975]" captionTargetPageId="71" captionText="FIGURE 35. General morphology of Houghia nigrofemur sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus." httpUri="https://zenodo.org/record/287048/files/figure.png" pageId="29" pageNumber="30">Figs. 35</figureCitation>
c, 36c), its length surpassing lower facial margin by more than length of pedicel; dark stripes on scutum distinct (
<figureCitation id="13422A6CB7018F1EFAB1F950FA5AF926" box="[1340,1419,1734,1755]" captionStart="FIGURE 35" captionStartId="71.[151,250,996,1018]" captionTargetBox="[340,1246,337,974]" captionTargetId="figure@71.[340,1247,337,975]" captionTargetPageId="71" captionText="FIGURE 35. General morphology of Houghia nigrofemur sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus." httpUri="https://zenodo.org/record/287048/files/figure.png" pageId="29" pageNumber="30">Figs. 35</figureCitation>
a, 36a)................................................................................................ 3
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7018F1EFF1AF896FA4AF8CE" blockId="29.[151,1437,1579,2019]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF896FA4AF8CE" pageId="29" pageNumber="30">
- Antenna entirely black except for slightly paler base of first flagellomere; first flagellomere straight, its length barely surpassing lower facial margin; dark scutal stripes indistinct (
<figureCitation id="13422A6CB7018F1EFD6AF888FCE7F8CE" box="[743,822,1822,1843]" captionStart="FIGURE 25" captionStartId="54.[151,250,1774,1796]" captionTargetBox="[340,1246,626,1752]" captionTargetId="figure@54.[340,1247,626,1752]" captionTargetPageId="54" captionText="FIGURE 25. General morphology of Houghia griseifrons sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287038/files/figure.png" pageId="29" pageNumber="30">Figs. 25</figureCitation>
a, 32a)............................................. 4
</keyLead>
</paragraph>
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<keyStep id="308D2E4CB7018F1EFF1AF8ADFA4DF876" pageId="29" pageNumber="30">
<paragraph id="8BC636E9B7018F1EFF1AF8ADFA4DF893" blockId="29.[151,1437,1579,2019]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF8ADFA4DF893" pageId="29" pageNumber="30">
3 Bases of femora orange, contrasting with dark apices of femora, tibiae and tarsi; first flagellomere distinctly S-shaped, the lower curve parallelling strongly curved facial ridge (
<figureCitation id="13422A6CB7018F1EFD6CF8CEFCF6F890" box="[737,807,1880,1901]" captionStart="FIGURE 36" captionStartId="72.[151,250,1917,1939]" captionTargetBox="[340,1246,769,1895]" captionTargetId="figure@72.[340,1247,769,1895]" captionTargetPageId="72" captionText="FIGURE 36. General morphology of Houghia ochrofemur sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287049/files/figure.png" pageId="29" pageNumber="30">Fig. 36</figureCitation>
c)..................................
<emphasis id="B90DEAFBB7018F1EFB59F8CEFA4DF893" bold="true" box="[1236,1436,1880,1902]" pageId="29" pageNumber="30">
<taxonomicName id="4C794D6AB7018F1EFB59F8CEFA98F893" ID-CoL="9HJ9Y" box="[1236,1353,1880,1902]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ochrofemur" status="sp. nov.">
<emphasis id="B90DEAFBB7018F1EFB59F8CEFA98F893" bold="true" box="[1236,1353,1880,1902]" italics="true" pageId="29" pageNumber="30">ochrofemur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7018F1EFAC2F8CEFA4DF893" box="[1359,1436,1880,1902]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7018F1EFF1AF8E0FA4DF876" blockId="29.[151,1437,1579,2019]" box="[151,1436,1909,1931]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF8E0FA4DF876" box="[151,1436,1909,1931]" pageId="29" pageNumber="30">
- Legs entirely black; first flagellomere merely slightly bowed anteriorly (only one male available)......
<emphasis id="B90DEAFBB7018F1EFB55F8E3FA4DF876" bold="true" box="[1240,1436,1909,1931]" pageId="29" pageNumber="30">
<taxonomicName id="4C794D6AB7018F1EFB55F8E3FA98F876" ID-CoL="9J284" box="[1240,1353,1909,1931]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nigrofemur" status="sp. nov.">
<emphasis id="B90DEAFBB7018F1EFB55F8E3FA98F876" bold="true" box="[1240,1353,1909,1931]" italics="true" pageId="29" pageNumber="30">nigrofemur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7018F1EFAC2F8E3FA4DF876" box="[1359,1436,1909,1931]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7018F1EFF1AF805FA4DF81E" pageId="29" pageNumber="30">
<paragraph id="8BC636E9B7018F1EFF1AF805FA4DF83B" blockId="29.[151,1437,1579,2019]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF805FA4DF83B" pageId="29" pageNumber="30">
4 Lateral margins of frontal vitta yellowish tomentose, leaving only a narrow dark median line (
<figureCitation id="13422A6CB7018F1EFBE6F805FB60F855" box="[1131,1201,1939,1960]" captionStart="FIGURE 32" captionStartId="66.[151,250,1340,1362]" captionTargetBox="[340,1246,193,1318]" captionTargetId="figure@66.[340,1247,193,1319]" captionTargetPageId="66" captionText="FIGURE 32. General morphology of Houghia macilenta sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287045/files/figure.png" pageId="29" pageNumber="30">Fig. 32</figureCitation>
b) and appearing to connect ocellar triangle to pale golden fronto-orbital plates..........................................
<emphasis id="B90DEAFBB7018F1EFB65F826FA4DF83B" bold="true" box="[1256,1436,1968,1990]" pageId="29" pageNumber="30">
<taxonomicName id="4C794D6AB7018F1EFB65F826FA98F83B" ID-CoL="9HJ9W" box="[1256,1353,1968,1990]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="macilenta" status="sp. nov.">
<emphasis id="B90DEAFBB7018F1EFB65F826FA98F83B" bold="true" box="[1256,1353,1968,1990]" italics="true" pageId="29" pageNumber="30">macilenta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7018F1EFAC2F826FA4DF83B" box="[1359,1436,1968,1990]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7018F1EFF1AF858FA4DF81E" blockId="29.[151,1437,1579,2019]" box="[151,1436,1997,2019]" pageId="29" pageNumber="30">
<keyLead id="308895DCB7018F1EFF1AF858FA4DF81E" box="[151,1436,1997,2019]" pageId="29" pageNumber="30">
- Frontal vitta wider, entirely brown, separating ocellar triangle from grayish fronto-orbital plates........
<emphasis id="B90DEAFBB7018F1EFB53F85BFA4DF81E" bold="true" box="[1246,1436,1997,2019]" pageId="29" pageNumber="30">
<taxonomicName id="4C794D6AB7018F1EFB53F85BFA98F81E" ID-CoL="9J283" box="[1246,1353,1997,2019]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="griseifrons" status="sp. nov.">
<emphasis id="B90DEAFBB7018F1EFB53F85BFA98F81E" bold="true" box="[1246,1353,1997,2019]" italics="true" pageId="29" pageNumber="30">griseifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7018F1EFAC2F85BFA4DF81E" box="[1359,1436,1997,2019]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
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<keyStep id="308D2E4CB7028F1DFF1AFF01FA4DFEF9" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFF01FA4DFF37" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFF01FA4DFF37" pageId="30" pageNumber="31">
5 Sex patches present on black integument on tergites 3 as well as on tergites 4 and 5; palpus dark gray to black.................................................................................................
<emphasis id="B90DEAFBB7028F1DFB40FF22FA4DFF37" bold="true" box="[1229,1436,180,202]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB40FF22FA98FF37" ID-CoL="9HJB2" box="[1229,1353,180,202]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="sexmaculata" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB40FF22FA98FF37" bold="true" box="[1229,1353,180,202]" italics="true" pageId="30" pageNumber="31">sexmaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FF22FA4DFF37" box="[1359,1436,180,202]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFF44FA4DFEF9" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFF44FA4DFEF9" pageId="30" pageNumber="31">
- Sex patches present on tergite 4, and usually also on tergite 5, but if apparently also on tergite 3 (as in
<taxonomicName id="4C794D6AB7028F1DFB27FF44FAC1FF1A" ID-CoL="9HJ9Q" box="[1194,1296,209,231]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90DEAFBB7028F1DFB27FF44FAC1FF1A" box="[1194,1296,209,231]" italics="true" pageId="30" pageNumber="31">H. gracilis</emphasis>
</taxonomicName>
(
<figureCitation id="13422A6CB7028F1DFA90FF44FAB0FF1A" box="[1309,1377,210,231]" captionStart="FIGURE 10" captionStartId="29.[151,250,1315,1337]" captionTargetBox="[340,1246,375,1293]" captionTargetId="figure@29.[340,1247,375,1293]" captionTargetPageId="29" captionText="FIGURE 10. Ventral view of abdomen showing different configurations of sex patches. a. Houghia graciloides sp. nov., showing characteristic sex patch on tergites 3, 4, and 5. b. Houghia longipilosa sp. nov., with sex patch present on tergites 4 and 5 only. c. Houghia omissa sp. nov., with sex patch present on tergite 4 only. d. Houghia confinis sp. nov with sex patch on tergites 4 and 5. e. Houghia macilenta sp. nov., with no sex patch at all." httpUri="https://zenodo.org/record/287023/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
a) and
<taxonomicName id="4C794D6AB7028F1DFF6CFF66FEBBFEF8" ID-CoL="9J28F" box="[225,362,239,261]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="B90DEAFBB7028F1DFF6CFF66FEBBFEF8" box="[225,362,239,261]" italics="true" pageId="30" pageNumber="31">H. graciloides</emphasis>
</taxonomicName>
) then integument on ventral surface reddish; palpus pale........................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFE9BFA4AFEC2" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFE9BFA4AFEDF" blockId="30.[151,1437,151,2021]" box="[151,1435,269,290]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFE9BFA4AFEDF" box="[151,1435,269,290]" pageId="30" pageNumber="31">6 Pedicel and upper part of first flagellomere pale orange....................................................... 7</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFEBCFA4AFEC2" blockId="30.[151,1437,151,2021]" box="[151,1435,298,319]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFEBCFA4AFEC2" box="[151,1435,298,319]" pageId="30" pageNumber="31">- Pedicel and first flagellomere entirely dark................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFED1FA4DFE48" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFED1FA4DFE87" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFED1FA4DFE87" pageId="30" pageNumber="31">
7 Facial ridge with row of fine white hairs on at least lower half, easily overlooked and best viewed in profile against a dark background; postgena posterior to postoccipital fringe, with a small patch of black setae...............
<emphasis id="B90DEAFBB7028F1DFB60FEF2FA4DFE87" bold="true" box="[1261,1436,356,378]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB60FEF2FA98FE87" ID-CoL="9HJ9P" box="[1261,1353,356,378]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB60FEF2FA98FE87" bold="true" box="[1261,1353,356,378]" italics="true" pageId="30" pageNumber="31">fimbriata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FEF2FA4DFE87" box="[1359,1436,356,378]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFE14FA4DFE48" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFE14FA4DFE48" pageId="30" pageNumber="31">
- Facial ridge bare except for the usual few supravibrissal setae; postgena posterior to postoccipital fringe lacking black setae..........................................................................................
<emphasis id="B90DEAFBB7028F1DFA88FE09FA4DFE48" bold="true" box="[1285,1436,415,437]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFA88FE09FA98FE48" ID-CoL="9J28C" box="[1285,1353,415,437]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pallida" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFA88FE09FA98FE48" bold="true" box="[1285,1353,415,437]" italics="true" pageId="30" pageNumber="31">pallida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FE09FA4DFE48" box="[1359,1436,415,437]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFE2BFA4AFDD7" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFE2BFA4DFE12" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFE2BFA4DFE12" pageId="30" pageNumber="31">8 Eye conspicuously haired, each hair at least as long as width of three or more ommatidia together; setae of anterodorsal quadrant of anepisternum increasing in length anteriorly but lacking row of 3 or 4 distinctly larger setae..................... 9</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFE61FA4AFDD7" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFE61FA4AFDD7" pageId="30" pageNumber="31">- Eye bare or apparently so except at higher magnification, hairs if present, shorter than width of two ommatidia; setae of anterodorsal quadrant of anepisternum short except for row of 3 or 4 distinctly longer and thicker setae..................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFDA4FA4AFD99" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFDA4FA4DFDB5" blockId="30.[151,1437,151,2021]" box="[151,1436,562,584]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFDA4FA4DFDB5" box="[151,1436,562,584]" pageId="30" pageNumber="31">
9 Parafacial and fronto-orbital plate entirely golden tomentose......................................
<emphasis id="B90DEAFBB7028F1DFB75FDA4FA4DFDB5" bold="true" box="[1272,1436,562,584]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB75FDA4FA98FDB5" ID-CoL="9HJ9G" box="[1272,1353,562,584]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="aurifera" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB75FDA4FA98FDB5" bold="true" box="[1272,1353,562,584]" italics="true" pageId="30" pageNumber="31">aurifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FDA4FA4DFDB5" box="[1359,1436,562,584]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFDD9FA4AFD99" blockId="30.[151,1437,151,2021]" box="[151,1435,591,612]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFDD9FA4AFD99" box="[151,1435,591,612]" pageId="30" pageNumber="31">- Parafacial silvery, contrasting with gold of vertex........................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFDFBFA4DFD05" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFDFBFA4DFD40" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFDFBFA4DFD40" pageId="30" pageNumber="31">
10 Posterior dark bands of abdominal tergites 25 grayish-brown tomentose, indistinctly differentiated from paler tomentum of remaining anterior portion of tergites; fronto-orbital plate gold pollinose only to level of anterior proclinate fronto-orbital seta.....................................................................................
<emphasis id="B90DEAFBB7028F1DFB50FD31FA4DFD40" bold="true" box="[1245,1436,679,701]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB50FD31FA98FD40" ID-CoL="9J288" box="[1245,1353,679,701]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="brevipilosa" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB50FD31FA98FD40" bold="true" box="[1245,1353,679,701]" italics="true" pageId="30" pageNumber="31">brevipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FD31FA4DFD40" box="[1359,1436,679,701]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFD53FA4DFD05" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFD53FA4DFD05" pageId="30" pageNumber="31">
- Posterior bands of abdominal tergites black, clearly differentiated from paler anterior portions of tergites; fronto-orbital plate gold tomentose to base of antenna.........................................................
<emphasis id="B90DEAFBB7028F1DFB51FD74FA4DFD05" bold="true" box="[1244,1436,738,760]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB51FD74FA98FD05" ID-CoL="9HJ9T" box="[1244,1353,738,760]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="longipilosa" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB51FD74FA98FD05" bold="true" box="[1244,1353,738,760]" italics="true" pageId="30" pageNumber="31">longipilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FD74FA4DFD05" box="[1359,1436,738,760]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFD69FA4AFCB2" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFD69FA4DFCCF" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFD69FA4DFCCF" pageId="30" pageNumber="31">11 Scutum with 3 pairs of postsutural dorsocentral setae, with a gap between the first seta and the next (suggesting a missing seta)................................................................................................... 12</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFCACFA4AFCB2" blockId="30.[151,1437,151,2021]" box="[151,1435,826,847]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFCACFA4AFCB2" box="[151,1435,826,847]" pageId="30" pageNumber="31">- Scutum with 4 pairs of postsutural dorsocentral setae, more or less equally spaced................................. 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFCC1FA4AFC5A" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFCC1FA4DFC77" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFCC1FA4DFC77" pageId="30" pageNumber="31">
12 Tergite 5 yellowish tomentose ventrally, lacking any trace of sex patch, contrasting with well developed patch on tergite 4 (
<figureCitation id="13422A6CB7028F1DFAF8FCC1FF2BFC77" captionStart="FIGURE 10" captionStartId="29.[151,250,1315,1337]" captionTargetBox="[340,1246,375,1293]" captionTargetId="figure@29.[340,1247,375,1293]" captionTargetPageId="29" captionText="FIGURE 10. Ventral view of abdomen showing different configurations of sex patches. a. Houghia graciloides sp. nov., showing characteristic sex patch on tergites 3, 4, and 5. b. Houghia longipilosa sp. nov., with sex patch present on tergites 4 and 5 only. c. Houghia omissa sp. nov., with sex patch present on tergite 4 only. d. Houghia confinis sp. nov with sex patch on tergites 4 and 5. e. Houghia macilenta sp. nov., with no sex patch at all." httpUri="https://zenodo.org/record/287023/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
c)....................................................................................
<emphasis id="B90DEAFBB7028F1DFA88FCE2FA4DFC77" bold="true" box="[1285,1436,884,906]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFA88FCE2FA98FC77" ID-CoL="9J28D" box="[1285,1353,884,906]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="omissa" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFA88FCE2FA98FC77" bold="true" box="[1285,1353,884,906]" italics="true" pageId="30" pageNumber="31">omissa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FCE2FA4DFC77" box="[1359,1436,884,906]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFC04FA4AFC5A" blockId="30.[151,1437,151,2021]" box="[151,1435,914,935]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFC04FA4AFC5A" box="[151,1435,914,935]" pageId="30" pageNumber="31">- Tergite 5 with sex patch, similar to, although smaller than, patch on tergite 4, on black integument.................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFC39FA4AFBC7" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFC39FA4AFC02" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFC39FA4AFC02" pageId="30" pageNumber="31">13 Scutum with longitudinal dark stripes on either side of dorsocentral row of setae connected to one another by dark brown tomentosity, creating the appearance of a single wide stripe on each side; integument of abdominal tergite 3 more reddish laterally than adjacent tergites.............................................................................. 14</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFB91FA4AFBC7" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFB91FA4AFBC7" pageId="30" pageNumber="31">- Scutum with 4 (2 pairs of) narrow longitudinal dark stripes separated from each other by golden tomentosity concolorous with rest of scutum; abdominal tergite 3 concolorous with adjacent tergites........................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFBD4FA4DFB30" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFBD4FA4DFB88" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFBD4FA4DFB88" pageId="30" pageNumber="31">
14 Fronto-orbital plate gold tomentose from vertex to antennal bases; ocellar setae arising behind anterior ocellus; colour of brown tomentosity surrounding dorsocentral setae nearly as dark as longitudinal stripes.................
<emphasis id="B90DEAFBB7028F1DFB77FBC9FA4DFB88" bold="true" box="[1274,1436,1119,1141]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB77FBC9FA98FB88" ID-CoL="9HJ9J" box="[1274,1353,1119,1141]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="bivittata" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB77FBC9FA98FB88" bold="true" box="[1274,1353,1119,1141]" italics="true" pageId="30" pageNumber="31">bivittata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FBC9FA4DFB88" box="[1359,1436,1119,1141]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFBEBFA4DFB30" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFBEBFA4DFB30" pageId="30" pageNumber="31">
- Fronto-orbital plate silvery tomentose except for a tinge of gold extending from vertex to anterior proclinate fronto-orbital seta; ocellar setae arising slightly in front of anterior ocellus; colour of brown tomentosity surrounding dorsocentral setae paler than longitudinal stripes...................................................................
<emphasis id="B90DEAFBB7028F1DFB79FB21FA4DFB30" bold="true" box="[1268,1436,1207,1229]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB79FB21FA98FB30" ID-CoL="9HJ9H" box="[1268,1353,1207,1229]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="biseriata" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB79FB21FA98FB30" bold="true" box="[1268,1353,1207,1229]" italics="true" pageId="30" pageNumber="31">biseriata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FB21FA4DFB30" box="[1359,1436,1207,1229]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFB43FA4DFABF" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFB43FA4DFAF5" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFB43FA4DFAF5" pageId="30" pageNumber="31">
15 Inner and outer dark longitudinal stripes on each side of scutum converging posteriorly to coalesce at level of second last postsutural dorsocentral seta, thus enclosing, or nearly enclosing, seta with darker tomentosity...............
<emphasis id="B90DEAFBB7028F1DFB74FB64FA4DFAF5" bold="true" box="[1273,1436,1266,1288]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB74FB64FA98FAF5" ID-CoL="9HJ9L" box="[1273,1353,1266,1288]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="confinis" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB74FB64FA98FAF5" bold="true" box="[1273,1353,1266,1288]" italics="true" pageId="30" pageNumber="31">confinis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FB64FA4DFAF5" box="[1359,1436,1266,1288]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFA99FA4DFABF" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFA99FA4DFABF" pageId="30" pageNumber="31">- Inner and outer dark stripes remaining separate posteriorly separated by yellow tomentum concolorous with rest of scutum...................................................................................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFADCFA44FA0F" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFADCFA4DFA67" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFADCFA4DFA67" pageId="30" pageNumber="31">
16 Ocellar triangle, observed from above, sharply pointed anteriorly, the point extending to level of upper fronto-orbital seta; gold tomentosity of fronto-orbital plate extending forward only to same level; first flagellomere extending to lower facial margin......................................................................................
<emphasis id="B90DEAFBB7028F1DFB59FA12FA4DFA67" bold="true" box="[1236,1436,1412,1434]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB59FA12FA98FA67" ID-CoL="9HJB4" box="[1236,1353,1412,1434]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB59FA12FA98FA67" bold="true" box="[1236,1353,1412,1434]" italics="true" pageId="30" pageNumber="31">triangularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2FA12FA4DFA67" box="[1359,1436,1412,1434]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AFA34FA44FA0F" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFA34FA44FA0F" pageId="30" pageNumber="31">- Ocellar triangle rounded, truncate, or notched anteriorly (point concolorous with vitta hence scarcely distinguishable), extending forward only to level of upper reclinate frontal seta; gold tomentosity of fronto-orbital plate extending forward to or nearly to level of upper fronto-orbital seta; first flagellomere shorter................................................. 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AFA6CFA4DF995" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AFA6CFA4AF9D1" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AFA6CFA4AF9D1" pageId="30" pageNumber="31">
17 Height of apical portion (fused part to apex) of cerci three times height of dorsal notch, enclosing anus (
<figureCitation id="13422A6CB7028F1DFB53FA6CFAF4F9F2" box="[1246,1317,1530,1551]" captionStart="FIGURE 19" captionStartId="45.[151,250,1979,2001]" captionTargetBox="[340,1246,831,1956]" captionTargetId="figure@45.[340,1247,831,1957]" captionTargetPageId="45" captionText="FIGURE 19. General morphology of Houghia crypta sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287032/files/figure.png" pageId="30" pageNumber="31">Fig. 19</figureCitation>
d); surstylus dagger-shaped, pointed apically (
<figureCitation id="13422A6CB7028F1DFD92F981FDB5F9D1" box="[543,612,1559,1580]" captionStart="FIGURE 19" captionStartId="45.[151,250,1979,2001]" captionTargetBox="[340,1246,831,1956]" captionTargetId="figure@45.[340,1247,831,1957]" captionTargetPageId="45" captionText="FIGURE 19. General morphology of Houghia crypta sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287032/files/figure.png" pageId="30" pageNumber="31">Fig. 19</figureCitation>
e).....................................................
<taxonomicName id="4C794D6AB7028F1DFB7BF981FA4AF9D1" ID-CoL="R3LG" authority="Townsend" authorityName="Townsend" box="[1270,1435,1559,1581]" class="Insecta" family="Tachinidae" genus="Carceliocephala" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="B90DEAFBB7028F1DFB7BF981FAE2F9D0" bold="true" box="[1270,1331,1559,1581]" italics="true" pageId="30" pageNumber="31">crypta</emphasis>
Townsend
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AF9A3FA4DF995" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF9A3FA4DF995" pageId="30" pageNumber="31">
- Height of fused portion of cerci less than twice height of dorsal notch (Fig, 21d); surstylus bluntly rounded apically...............................................................................................
<emphasis id="B90DEAFBB7028F1DFB78F9C4FA4DF995" bold="true" box="[1269,1436,1618,1640]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB78F9C4FA98F995" ID-CoL="9J28J" box="[1269,1353,1618,1640]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="destituta" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB78F9C4FA98F995" bold="true" box="[1269,1353,1618,1640]" italics="true" pageId="30" pageNumber="31">destituta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2F9C4FA4DF995" box="[1359,1436,1618,1640]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AF9F9FA4AF95F" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AF9F9FA4AF979" blockId="30.[151,1437,151,2021]" box="[151,1435,1647,1668]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF9F9FA4AF979" box="[151,1435,1647,1668]" pageId="30" pageNumber="31">18 Ground colour of ventral surface of abdominal tergites 1 to 4 yellow or orange, somewhat translucent.................. 19</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AF91BFA4AF95F" blockId="30.[151,1437,151,2021]" box="[151,1435,1677,1698]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF91BFA4AF95F" box="[151,1435,1677,1698]" pageId="30" pageNumber="31">- Ground colour of ventral surface of abdomen entirely black, usually obscured by paler tomentosity.................... 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AF93CFA4AF921" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AF93CFA4DF93D" blockId="30.[151,1437,151,2021]" box="[151,1436,1706,1728]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF93CFA4DF93D" box="[151,1436,1706,1728]" pageId="30" pageNumber="31">
19 Katepisternal setae 2....................................................................
<emphasis id="B90DEAFBB7028F1DFB56F93CFA4DF93D" bold="true" box="[1243,1436,1706,1728]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB56F93CFA98F93D" ID-CoL="9HJ9V" box="[1243,1353,1706,1728]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="luteiventris" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB56F93CFA98F93D" bold="true" box="[1243,1353,1706,1728]" italics="true" pageId="30" pageNumber="31">luteiventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2F93CFA4DF93D" box="[1359,1436,1706,1728]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AF951FA4AF921" blockId="30.[151,1437,151,2021]" box="[151,1435,1735,1756]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF951FA4AF921" box="[151,1435,1735,1756]" pageId="30" pageNumber="31">- Katepisternal setae 3.................................................................................. 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AF973FA4AF8C9" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AF973FA4AF8EA" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF973FA4AF8EA" pageId="30" pageNumber="31">20 Sex patches on tergites 3, 4, and 5 consisting of shorter more appressed hairs than elsewhere on abdomen, but these hairs sparse and indistinct because of reddish background; ocellar setae arising behind anterior ocellus..................... 21</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AF889FA4AF8C9" blockId="30.[151,1437,151,2021]" box="[151,1435,1823,1844]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF889FA4AF8C9" box="[151,1435,1823,1844]" pageId="30" pageNumber="31">- Sex patches confined to tergites 4 and 5; ocellar setae arising beside anterior ocellus............................... 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AF8ABFA4DF88D" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AF8ABFA4DF8AF" blockId="30.[151,1437,151,2021]" box="[151,1436,1852,1874]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF8ABFA4DF8AF" box="[151,1436,1852,1874]" pageId="30" pageNumber="31">
21 Eye apparently bare........................................................................
<emphasis id="B90DEAFBB7028F1DFA8DF8AAFA4DF8AF" bold="true" box="[1280,1436,1852,1874]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFA8DF8AAFA98F8AF" ID-CoL="9HJ9Q" box="[1280,1353,1852,1874]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFA8DF8AAFA98F8AF" bold="true" box="[1280,1353,1852,1874]" italics="true" pageId="30" pageNumber="31">gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2F8AAFA4DF8AF" box="[1359,1436,1852,1874]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AF8CCFA4DF88D" blockId="30.[151,1437,151,2021]" box="[151,1436,1882,1904]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF8CCFA4DF88D" box="[151,1436,1882,1904]" pageId="30" pageNumber="31">
- Eye with minute obscure hairs............................................................
<emphasis id="B90DEAFBB7028F1DFB50F8CCFA4DF88D" bold="true" box="[1245,1436,1882,1904]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB50F8CCFA98F88D" ID-CoL="9J28F" box="[1245,1353,1882,1904]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="graciloides" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB50F8CCFA98F88D" bold="true" box="[1245,1353,1882,1904]" italics="true" pageId="30" pageNumber="31">graciloides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2F8CCFA4DF88D" box="[1359,1436,1882,1904]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7028F1DFF1AF8E1FA4AF819" pageId="30" pageNumber="31">
<paragraph id="8BC636E9B7028F1DFF1AF8E1FA4DF857" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF8E1FA4DF857" pageId="30" pageNumber="31">
22 Fronto-orbital plate mostly silvery, with pale golden tomentosity, and area of small hairs, extending from vertex only to level of anterior fronto-orbital seta.............................................................
<emphasis id="B90DEAFBB7028F1DFB52F802FA4DF857" bold="true" box="[1247,1436,1940,1962]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB52F802FA9BF857" ID-CoL="9HJ9N" box="[1247,1354,1940,1962]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="delospilota" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB52F802FA9BF857" bold="true" box="[1247,1354,1940,1962]" italics="true" pageId="30" pageNumber="31">delospilota</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFAC2F802FA4DF857" box="[1359,1436,1940,1962]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7028F1DFF1AF824FA4AF819" blockId="30.[151,1437,151,2021]" pageId="30" pageNumber="31">
<keyLead id="308895DCB7028F1DFF1AF824FA4AF819" pageId="30" pageNumber="31">
- Fronto-orbital plate entirely golden tomentose and more extensively haired, extending from vertex to lower margin of pedicel..............................................................................
<emphasis id="B90DEAFBB7028F1DFB01F859FA99F818" bold="true" box="[1164,1352,1999,2021]" pageId="30" pageNumber="31">
<taxonomicName id="4C794D6AB7028F1DFB01F859FB24F818" ID-CoL="9J289" box="[1164,1269,1999,2021]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pilosifrons" status="sp. nov.">
<emphasis id="B90DEAFBB7028F1DFB01F859FB24F818" bold="true" box="[1164,1269,1999,2021]" italics="true" pageId="30" pageNumber="31">pilosifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7028F1DFB76F859FA99F818" box="[1275,1352,1999,2021]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in part)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFF01FA4AFF37" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFF01FA4AFF51" blockId="31.[151,1436,151,1258]" box="[151,1435,151,172]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFF01FA4AFF51" box="[151,1435,151,172]" pageId="31" pageNumber="32">23 Fronto-orbital plate entirely gold tomentose................................................................ 24</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFF23FA4AFF37" blockId="31.[151,1436,151,1258]" box="[151,1435,181,202]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFF23FA4AFF37" box="[151,1435,181,202]" pageId="31" pageNumber="32">- Gold tomentosity of fronto-orbital plate restricted to vertex................................................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFF44FA4DFEDF" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFF44FA4AFF1A" blockId="31.[151,1436,151,1258]" box="[151,1435,210,232]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFF44FA4AFF1A" box="[151,1435,210,232]" pageId="31" pageNumber="32">
24 Parafacial silver tomentose, changing abruptly at level of antenna to gold..........................
<taxonomicName id="4C794D6AB7038F1CFB53FF44FA96FF15" ID-CoL="9J289" box="[1246,1351,210,232]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pilosifrons">
<emphasis id="B90DEAFBB7038F1CFB53FF44FA96FF15" bold="true" box="[1246,1351,210,232]" italics="true" pageId="31" pageNumber="32">pilosifrons</emphasis>
</taxonomicName>
(in part)
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFF79FA4DFEDF" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFF79FA4DFEDF" pageId="31" pageNumber="32">
- Parafacial pale yellow tomentose gradually changing at level of antenna to darker brassy yellow on fronto-orbital plate...............................................................................................
<emphasis id="B90DEAFBB7038F1CFA86FE9AFA4DFEDF" bold="true" box="[1291,1436,268,290]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFA86FE9AFA98FEDF" ID-CoL="9HJ9D" box="[1291,1353,268,290]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="aerata" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFA86FE9AFA98FEDF" bold="true" box="[1291,1353,268,290]" italics="true" pageId="31" pageNumber="32">aerata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FE9AFA4DFEDF" box="[1359,1436,268,290]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFEBCFA4AFEA1" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFEBCFA4AFEC2" blockId="31.[151,1436,151,1258]" box="[151,1435,298,319]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFEBCFA4AFEC2" box="[151,1435,298,319]" pageId="31" pageNumber="32">25 Height of gena one-tenth height of head in lateral view....................................................... 26</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFED1FA4AFEA1" blockId="31.[151,1436,151,1258]" box="[151,1435,327,348]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFED1FA4AFEA1" box="[151,1435,327,348]" pageId="31" pageNumber="32">- Height of gena one-twelfth or less height of head........................................................... 29</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFEF3FA4AFE2F" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFEF3FA4AFE6A" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFEF3FA4AFE6A" pageId="31" pageNumber="32">
26 Depth of V-shaped dorsal notch of cerci in posterior view one quarter or less height of entire cerci (
<figureCitation id="13422A6CB7038F1CFB36FEF3FADDFE87" box="[1211,1292,357,378]" captionStart="FIGURE 30" captionStartId="63.[151,250,1343,1365]" captionTargetBox="[340,1246,195,1320]" captionTargetId="figure@63.[340,1247,195,1321]" captionTargetPageId="63" captionText="FIGURE 30. General morphology of Houghia longipilosa sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287043/files/figure.png" pageId="31" pageNumber="32">Figs. 30</figureCitation>
d, 28d); lateral scutal stripe converging to meet medial stripe at its posterior end (
<figureCitation id="13422A6CB7038F1CFCD7FE14FC7BFE6A" box="[858,938,386,407]" captionStart="FIGURE 30" captionStartId="63.[151,250,1343,1365]" captionTargetBox="[340,1246,195,1320]" captionTargetId="figure@63.[340,1247,195,1321]" captionTargetPageId="63" captionText="FIGURE 30. General morphology of Houghia longipilosa sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287043/files/figure.png" pageId="31" pageNumber="32">Figs. 30</figureCitation>
a, 28a).................................. 27
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFE09FA4AFE2F" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFE09FA4AFE2F" pageId="31" pageNumber="32">
- Depth of V-shaped notch one third or more height of cerci (
<figureCitation id="13422A6CB7038F1CFCA5FE09FCA9FE49" box="[808,888,415,436]" captionStart="FIGURE 27" captionStartId="58.[151,250,1628,1650]" captionTargetBox="[340,1246,481,1606]" captionTargetId="figure@58.[340,1247,481,1607]" captionTargetPageId="58" captionText="FIGURE 27. General morphology of Houghia latigena sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287040/files/figure.png" pageId="31" pageNumber="32">Figs. 27</figureCitation>
d, 45d); both pairs of scutal stripes separated along their entire lengths (
<figureCitation id="13422A6CB7038F1CFEFDFE2BFE11FE2F" box="[368,448,445,466]" captionStart="FIGURE 27" captionStartId="58.[151,250,1628,1650]" captionTargetBox="[340,1246,481,1606]" captionTargetId="figure@58.[340,1247,481,1607]" captionTargetPageId="58" captionText="FIGURE 27. General morphology of Houghia latigena sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287040/files/figure.png" pageId="31" pageNumber="32">Figs. 27</figureCitation>
a, 45a)........................................................................... 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFE4CFA4DFDB5" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFE4CFA4DFDF0" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFE4CFA4DFDF0" pageId="31" pageNumber="32">
27 Dorsal lobe of syncerus in posterior view almost as broad as high, bluntly rounded dorsally, depth of V-shaped dorsal notch one quarter height of cerci (
<figureCitation id="13422A6CB7038F1CFE66FE61FDE3FDF1" box="[491,562,503,524]" captionStart="FIGURE 28" captionStartId="59.[151,250,1340,1362]" captionTargetBox="[340,1246,193,1318]" captionTargetId="figure@59.[340,1247,193,1319]" captionTargetPageId="59" captionText="FIGURE 28. General morphology of Houghia latilobus sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287041/files/figure.png" pageId="31" pageNumber="32">Fig. 28</figureCitation>
d); apices of cerci splayed apart apically...........................
<emphasis id="B90DEAFBB7038F1CFB79FE61FA4DFDF0" bold="true" box="[1268,1436,503,525]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB79FE61FA98FDF0" ID-CoL="9J287" box="[1268,1353,503,525]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="latilobus" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB79FE61FA98FDF0" bold="true" box="[1268,1353,503,525]" italics="true" pageId="31" pageNumber="32">latilobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FE61FA4DFDF0" box="[1359,1436,503,525]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFD83FA4DFDB5" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFD83FA4DFDB5" pageId="31" pageNumber="32">
- Dorsal lobe of cerci more pointed and attenuated dorsally; depth of V-shaped notch less than one quarter (about 1/4.5) height of cerci (
<figureCitation id="13422A6CB7038F1CFEBBFDA4FEACFDBA" box="[310,381,562,583]" captionStart="FIGURE 30" captionStartId="63.[151,250,1343,1365]" captionTargetBox="[340,1246,195,1320]" captionTargetId="figure@63.[340,1247,195,1321]" captionTargetPageId="63" captionText="FIGURE 30. General morphology of Houghia longipilosa sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287043/files/figure.png" pageId="31" pageNumber="32">Fig. 30</figureCitation>
d); apices of cerci straight and parallel........................................
<emphasis id="B90DEAFBB7038F1CFB5BFDA4FA4DFDB5" bold="true" box="[1238,1436,562,584]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB5BFDA4FA98FDB5" ID-CoL="9HJ9S" box="[1238,1353,562,584]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="longicercus" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB5BFDA4FA98FDB5" bold="true" box="[1238,1353,562,584]" italics="true" pageId="31" pageNumber="32">longicercus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FDA4FA4DFDB5" box="[1359,1436,562,584]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFDD9FA4DFD40" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFDD9FA4DFD7F" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFDD9FA4DFD7F" pageId="31" pageNumber="32">
28 Cerci in posterior view slightly constricted at midlength (
<figureCitation id="13422A6CB7038F1CFCADFDD9FCBBFD99" box="[800,874,591,612]" captionStart="FIGURE 27" captionStartId="58.[151,250,1628,1650]" captionTargetBox="[340,1246,481,1606]" captionTargetId="figure@58.[340,1247,481,1607]" captionTargetPageId="58" captionText="FIGURE 27. General morphology of Houghia latigena sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287040/files/figure.png" pageId="31" pageNumber="32">Fig. 27</figureCitation>
d); posteroventral quadrant of anepimeron sparsely short haired..................................................................................
<emphasis id="B90DEAFBB7038F1CFB74FDFAFA4DFD7F" bold="true" box="[1273,1436,620,642]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB74FDFAFA98FD7F" ID-CoL="9J282" box="[1273,1353,620,642]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="latigena" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB74FDFAFA98FD7F" bold="true" box="[1273,1353,620,642]" italics="true" pageId="31" pageNumber="32">latigena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FDFAFA4DFD7F" box="[1359,1436,620,642]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFD1CFA4DFD40" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFD1CFA4DFD40" pageId="31" pageNumber="32">
- Cerci in posterior view tapering evenly to apex (
<figureCitation id="13422A6CB7038F1CFD3DFD1CFD26FD62" box="[688,759,650,671]" captionStart="FIGURE 45" captionStartId="85.[151,250,1985,2007]" captionTargetBox="[340,1246,837,1962]" captionTargetId="figure@85.[340,1247,837,1963]" captionTargetPageId="85" captionText="FIGURE 45. General morphology of Houghia velutina sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287058/files/figure.png" pageId="31" pageNumber="32">Fig. 45</figureCitation>
d); posteroventral quadrant of anepimeron more densely long haired.........................................................................................
<emphasis id="B90DEAFBB7038F1CFB74FD31FA4DFD40" bold="true" box="[1273,1436,679,701]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB74FD31FA98FD40" ID-CoL="9J28B" box="[1273,1353,679,701]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="velutina" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB74FD31FA98FD40" bold="true" box="[1273,1353,679,701]" italics="true" pageId="31" pageNumber="32">velutina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FD31FA4DFD40" box="[1359,1436,679,701]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFD53FA4AFD0A" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFD53FA4DFD27" blockId="31.[151,1436,151,1258]" box="[151,1436,708,730]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFD53FA4DFD27" box="[151,1436,708,730]" pageId="31" pageNumber="32">
29 Width of frontal vitta one-tenth width of broad shiny silvery fronto-orbital plate.......................
<emphasis id="B90DEAFBB7038F1CFB7AFD52FA4DFD27" bold="true" box="[1271,1436,708,730]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB7AFD52FA98FD27" ID-CoL="9J28H" box="[1271,1353,708,730]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="parmata" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB7AFD52FA98FD27" bold="true" box="[1271,1353,708,730]" italics="true" pageId="31" pageNumber="32">parmata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FD52FA4DFD27" box="[1359,1436,708,730]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFD74FA4AFD0A" blockId="31.[151,1436,151,1258]" box="[151,1435,738,759]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFD74FA4AFD0A" box="[151,1435,738,759]" pageId="31" pageNumber="32">- Width of frontal vitta greater than one-tenth of fronto-orbital plate.............................................. 30</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFD69FA4AFCCF" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFD69FA4AFCE9" blockId="31.[151,1436,151,1258]" box="[151,1435,767,788]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFD69FA4AFCE9" box="[151,1435,767,788]" pageId="31" pageNumber="32">30 Gold tomentosity of fronto-orbital plate confined to vertex, not extending forward farther than upper fronto-orbital setae... 31</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFC8BFA4AFCCF" blockId="31.[151,1436,151,1258]" box="[151,1435,797,818]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFC8BFA4AFCCF" box="[151,1435,797,818]" pageId="31" pageNumber="32">- Gold tomentosity extending at least to lower fronto-orbital setae and often to the antennal bases...................... 34</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFCACFA4AFC91" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFCACFA4AFCB2" blockId="31.[151,1436,151,1258]" box="[151,1435,826,847]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFCACFA4AFCB2" box="[151,1435,826,847]" pageId="31" pageNumber="32">31 First flagellomere extending to lower facial margin.......................................................... 32</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFCC1FA4AFC91" blockId="31.[151,1436,151,1258]" box="[151,1435,855,876]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFCC1FA4AFC91" box="[151,1435,855,876]" pageId="31" pageNumber="32">- First flagellomere terminating above lower facial margin by about half length of pedicel............................ 33</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFCE3FA4DFC1F" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFCE3FA4DFC55" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFCE3FA4DFC55" pageId="31" pageNumber="32">
32 Surstylus distinctly longer than cerci (
<figureCitation id="13422A6CB7038F1CFDC0FCE3FD4AFC77" box="[589,667,885,906]" captionStart="FIGURE 26" captionStartId="56.[151,250,1523,1545]" captionTargetBox="[340,1246,373,1502]" captionTargetId="figure@56.[340,1247,373,1502]" captionTargetPageId="56" captionText="FIGURE 26. General morphology of Houghia inflatipalpis sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287039/files/figure.png" pageId="31" pageNumber="32">Figs. 26</figureCitation>
d, e), its apex surpassing that of apex of cerci by more than the width of the sur- stylus; [palpus of female swollen apically]..................................................
<emphasis id="B90DEAFBB7038F1CFB5EFC04FA4DFC55" bold="true" box="[1235,1436,914,936]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB5EFC04FA98FC55" ID-CoL="9HJ9R" box="[1235,1353,914,936]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="inflatipalpis" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB5EFC04FA98FC55" bold="true" box="[1235,1353,914,936]" italics="true" pageId="31" pageNumber="32">inflatipalpis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FC04FA4DFC55" box="[1359,1436,914,936]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFC39FA4DFC1F" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFC39FA4DFC1F" pageId="31" pageNumber="32">
- Surstylus barely longer than cerci (
<figureCitation id="13422A6CB7038F1CFDA2FC39FDAFFC39" box="[559,638,943,964]" captionStart="FIGURE 33" captionStartId="67.[151,250,1341,1363]" captionTargetBox="[340,1246,193,1319]" captionTargetId="figure@67.[340,1247,193,1319]" captionTargetPageId="67" captionText="FIGURE 33. General morphology of Houghia marini sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287046/files/figure.png" pageId="31" pageNumber="32">Figs. 33</figureCitation>
, d, e); [palpus of female as in all other species (except
<taxonomicName id="4C794D6AB7038F1CFB07FC26FAC1FC38" box="[1162,1296,943,965]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="luteipennis">
<emphasis id="B90DEAFBB7038F1CFB07FC26FAC1FC38" box="[1162,1296,943,965]" italics="true" pageId="31" pageNumber="32">H. luteipennis</emphasis>
</taxonomicName>
), only slightly enlarged apically].........................................................................
<emphasis id="B90DEAFBB7038F1CFA8BFC5AFA4DFC1F" bold="true" box="[1286,1436,972,994]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFA8BFC5AFA99FC1F" ID-CoL="9J28G" box="[1286,1352,972,994]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFA8BFC5AFA99FC1F" bold="true" box="[1286,1352,972,994]" italics="true" pageId="31" pageNumber="32">marini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FC5AFA4DFC1F" box="[1359,1436,972,994]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFC7CFA4DFBC7" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFC7CFA4DFBE0" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFC7CFA4DFBE0" pageId="31" pageNumber="32">
33 Gold tomentosity of fronto-orbital plate restricted to upper margin, not extending forward beyond upper reclinate orbital seta and apex of ocellar triangle..................................................................
<emphasis id="B90DEAFBB7038F1CFA8DFB91FA4DFBE0" bold="true" box="[1280,1436,1031,1053]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFA8DFB91FA9BFBE0" ID-CoL="9J286" box="[1280,1354,1031,1053]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="blancoi" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFA8DFB91FA9BFBE0" bold="true" box="[1280,1354,1031,1053]" italics="true" pageId="31" pageNumber="32">blancoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FB91FA4DFBE0" box="[1359,1436,1031,1053]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFBB3FA4DFBC7" blockId="31.[151,1436,151,1258]" box="[151,1436,1060,1082]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFBB3FA4DFBC7" box="[151,1436,1060,1082]" pageId="31" pageNumber="32">
- Gold tomentosity more extensive, extending to upper fronto-orbital seta............................
<emphasis id="B90DEAFBB7038F1CFB66FBB2FA4DFBC7" bold="true" box="[1259,1436,1060,1082]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB66FBB2FA98FBC7" ID-CoL="9J285" box="[1259,1353,1060,1082]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="romeroae" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB66FBB2FA98FBC7" bold="true" box="[1259,1353,1060,1082]" italics="true" pageId="31" pageNumber="32">romeroae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FBB2FA4DFBC7" box="[1359,1436,1060,1082]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFBD4FA4AFB52" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFBD4FA4DFB88" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFBD4FA4DFB88" pageId="31" pageNumber="32">
34 Surstylus long, straight, and parallel-sided in lateral view (
<figureCitation id="13422A6CB7038F1CFCA3FBD4FCA9FBAA" box="[814,888,1090,1111]" captionStart="FIGURE 43" captionStartId="82.[151,250,1976,1998]" captionTargetBox="[348,1240,868,1976]" captionTargetId="figure@82.[348,1240,868,1976]" captionTargetPageId="82" captionText="FIGURE 43. General morphology of Houghia spathulata sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287056/files/figure.png" pageId="31" pageNumber="32">Fig. 43</figureCitation>
e); apical processes of sternite 5 elongate, slender, and directed medially (
<figureCitation id="13422A6CB7038F1CFE18FBC9FE0DFB89" box="[405,476,1119,1140]" captionStart="FIGURE 43" captionStartId="82.[151,250,1976,1998]" captionTargetBox="[348,1240,868,1976]" captionTargetId="figure@82.[348,1240,868,1976]" captionTargetPageId="82" captionText="FIGURE 43. General morphology of Houghia spathulata sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287056/files/figure.png" pageId="31" pageNumber="32">Fig. 43</figureCitation>
f)...............................................................
<emphasis id="B90DEAFBB7038F1CFB6FFBC9FA4DFB88" bold="true" box="[1250,1436,1119,1141]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB6FFBC9FA9BFB88" ID-CoL="9HJB3" box="[1250,1354,1119,1141]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="spathulata" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB6FFBC9FA9BFB88" bold="true" box="[1250,1354,1119,1141]" italics="true" pageId="31" pageNumber="32">spathulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FBC9FA4DFB88" box="[1359,1436,1119,1141]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFBEBFA4AFB52" blockId="31.[151,1436,151,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFBEBFA4AFB52" pageId="31" pageNumber="32">
- Surstylus in lateral view obovate, widened moderately at mid-length; apical processes of sternite 5 shorter, thicker and directed posteriorly (
<figureCitation id="13422A6CB7038F1CFEDBFB0CFE75FB52" box="[342,420,1178,1199]" captionStart="FIGURE 17" captionStartId="42.[151,250,1341,1363]" captionTargetBox="[340,1246,193,1319]" captionTargetId="figure@42.[340,1247,193,1320]" captionTargetPageId="42" captionText="FIGURE 17. General morphology of Houghia chavarriae sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287030/files/figure.png" pageId="31" pageNumber="32">Figs. 17</figureCitation>
f, and 34f).......................................................................... 35
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308D2E4CB7038F1CFF1AFB21FA4DFB17" pageId="31" pageNumber="32">
<paragraph id="8BC636E9B7038F1CFF1AFB21FA4DFB30" blockId="31.[151,1436,151,1258]" box="[151,1436,1207,1229]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFB21FA4DFB30" box="[151,1436,1207,1229]" pageId="31" pageNumber="32">
35 Posterior margin of surstylus rounded in lateral view (
<figureCitation id="13422A6CB7038F1CFD6AFB21FCFAFB31" box="[743,811,1207,1228]" captionStart="FIGURE 17" captionStartId="42.[151,250,1341,1363]" captionTargetBox="[340,1246,193,1319]" captionTargetId="figure@42.[340,1247,193,1320]" captionTargetPageId="42" captionText="FIGURE 17. General morphology of Houghia chavarriae sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287030/files/figure.png" pageId="31" pageNumber="32">Fig. 17</figureCitation>
d)..................................
<emphasis id="B90DEAFBB7038F1CFB53FB21FA4DFB30" bold="true" box="[1246,1436,1207,1229]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB53FB21FA99FB30" ID-CoL="9HJ9K" box="[1246,1352,1207,1229]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="chavarriae" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB53FB21FA99FB30" bold="true" box="[1246,1352,1207,1229]" italics="true" pageId="31" pageNumber="32">chavarriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FB21FA4DFB30" box="[1359,1436,1207,1229]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC636E9B7038F1CFF1AFB43FA4DFB17" blockId="31.[151,1436,151,1258]" box="[151,1436,1236,1258]" pageId="31" pageNumber="32">
<keyLead id="308895DCB7038F1CFF1AFB43FA4DFB17" box="[151,1436,1236,1258]" pageId="31" pageNumber="32">
- Posterior margin of surstylus slightly concave in lateral view (
<figureCitation id="13422A6CB7038F1CFCB9FB43FCADFB17" box="[820,892,1237,1258]" captionStart="FIGURE 34" captionStartId="69.[151,250,1593,1615]" captionTargetBox="[340,1246,445,1572]" captionTargetId="figure@69.[340,1247,445,1572]" captionTargetPageId="69" captionText="FIGURE 34. General morphology of Houghia matarritai sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8." httpUri="https://zenodo.org/record/287047/files/figure.png" pageId="31" pageNumber="32">Fig. 34</figureCitation>
d).............................
<emphasis id="B90DEAFBB7038F1CFB6BFB42FA4DFB17" bold="true" box="[1254,1436,1236,1258]" pageId="31" pageNumber="32">
<taxonomicName id="4C794D6AB7038F1CFB6BFB42FA98FB17" ID-CoL="9HJ9X" box="[1254,1353,1236,1258]" class="Insecta" family="Tachinidae" genus="Houghia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="matarritai" status="sp. nov.">
<emphasis id="B90DEAFBB7038F1CFB6BFB42FA98FB17" bold="true" box="[1254,1353,1236,1258]" italics="true" pageId="31" pageNumber="32">matarritai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5780B7038F1CFAC2FB42FA4DFB17" box="[1359,1436,1236,1258]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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