244 lines
28 KiB
XML
244 lines
28 KiB
XML
<document id="8EB0A7094DEC39F90255FBAED9B359F7" ID-DOI="10.5281/zenodo.191476" ID-GBIF-Dataset="8603d4d1-efc8-44bb-84ae-76d3d8dd8161" ID-ISSN="1175-5326" ID-Zenodo-Dep="191476" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461128727612" checkinUser="plazi" docAuthor="Passos, Paulo, Mueses-Cisneros, Jonh Jairo, Lynch, John D. & Fernandes, Ronaldo" docDate="2009" docId="03D087ECAA5C7751FF11B70714542648" docLanguage="en" docName="zt02293p034.pdf" docOrigin="Zootaxa 2293" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atractus boulengerii Peracca 1896" docType="treatment" docVersion="8" lastPageNumber="5" masterDocId="FFE9FF94AA5E7755FF86B32815112046" masterDocTitle="Pacific lowland snakes of the genus Atractus (Serpentes: Dipsadidae), with description of three new species" masterLastPageNumber="34" masterPageNumber="1" pageNumber="3" updateTime="1698597252759" updateUser="plazi">
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<mods:title id="C87BD2D70F5F6D3E6BAA87D869F5C7E5">Pacific lowland snakes of the genus Atractus (Serpentes: Dipsadidae), with description of three new species</mods:title>
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<mods:name id="A2D70467F692FDCAD12B46B34E1A1096" type="personal">
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<mods:namePart id="B43C4ACBB998C9DBDFD181AC0D41CB47">Passos, Paulo</mods:namePart>
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<mods:namePart id="11E13D46AA85CD8C74A3753E46965ABC">Mueses-Cisneros, Jonh Jairo</mods:namePart>
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<mods:name id="510E4EA5F7EA9B872FB18BEFC73B7FEA" type="personal">
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<mods:roleTerm id="FA11D38EFE257C3829161D89DA17E495">Author</mods:roleTerm>
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<mods:namePart id="45F3CDEA411F8B4DC8585A53DC542B86">Lynch, John D.</mods:namePart>
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<mods:name id="1E3A6C0C33832E45DE7684A5265A7F74" type="personal">
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<mods:roleTerm id="DF6CE0D8968D8DBDB82B9F26688DEE0B">Author</mods:roleTerm>
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<mods:namePart id="B14699DEC20DB0BCE2E8617F08EEBFD6">Fernandes, Ronaldo</mods:namePart>
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<mods:title id="5061D3F5DE1B640722187065AF08F1E1">Zootaxa</mods:title>
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<mods:date id="C715BDB238D89B572AB5653EE53597D5">2009</mods:date>
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<mods:detail id="2F151D3F7EF58C7136B9CF1868FFBE62" type="volume">
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<mods:number id="57EA19C9FDB202E4F283E6F7A2906FF2">2293</mods:number>
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<mods:identifier id="C054AA0039B2F78DC6941F0265B4D8FB" type="DOI">10.5281/zenodo.191476</mods:identifier>
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<treatment id="03D087ECAA5C7751FF11B70714542648" ID-DOI="http://doi.org/10.5281/zenodo.6214653" ID-GBIF-Taxon="119626673" ID-Zenodo-Dep="6214653" LSID="urn:lsid:plazi:treatment:03D087ECAA5C7751FF11B70714542648" httpUri="http://treatment.plazi.org/id/03D087ECAA5C7751FF11B70714542648" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<subSubSection id="C3636571AA5C7757FF11B70715CA242A" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BC636FAAA5C7757FF11B7071752240F" blockId="2.[151,579,1071,1132]" box="[151,579,1071,1097]" pageId="2" pageNumber="3">
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<heading id="D08E8196AA5C7757FF11B7071752240F" bold="true" box="[151,579,1071,1097]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<taxonomicName id="4C794D79AA5C7757FF11B7071752240F" authority="Peracca, 1896" authorityName="Peracca" authorityYear="1896" box="[151,579,1071,1097]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="boulengerii">
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<emphasis id="B90DEAE8AA5C7757FF11B7071752240F" bold="true" box="[151,579,1071,1097]" pageId="2" pageNumber="3">
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<emphasis id="B90DEAE8AA5C7757FF11B707149E240F" bold="true" box="[151,399,1071,1097]" italics="true" pageId="2" pageNumber="3">Atractus boulengerii</emphasis>
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<bibRefCitation id="EFE84B0BAA5C7757FE10B7071752240F" author="Peracca" box="[406,579,1071,1097]" pageId="2" pageNumber="33" refString="Peracca, M. G. (1896) Sopra alcuni ofidii nuovi o poco noti dell'America meridionale. Bolletin Museo di Torino, 11, 1 - 12." type="journal article" year="1896">Peracca, 1896</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BC636FAAA5C7757FF11B77A15CA242A" blockId="2.[151,579,1071,1132]" box="[151,219,1106,1132]" pageId="2" pageNumber="3">
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<figureCitation id="13422A7FAA5C7757FF11B77A15CA242A" box="[151,219,1106,1132]" captionStart="FIGURE 1" captionStartId="2.[151,255,1950,1974]" captionTargetBox="[195,1392,1415,1923]" captionTargetId="figure@2.[195,1393,1415,1924]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal (A) and ventral (B) views of Atractus boulengerii (UV-C 6591). SVL 310 mm, CL 52 mm." httpUri="https://zenodo.org/record/191477/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3636571AA5C7757FF11B788113024F1" box="[151,1057,1184,1207]" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="8BC636FAAA5C7757FF11B788113024F1" blockId="2.[151,1057,1184,1207]" box="[151,1057,1184,1207]" pageId="2" pageNumber="3">
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<taxonomicName id="4C794D79AA5C7757FF11B788170324F1" authority="Peracca, 1896" authorityName="Peracca" authorityYear="1896" box="[151,530,1184,1207]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="boulengerii">
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<emphasis id="B90DEAE8AA5C7757FF11B788146224F1" box="[151,371,1184,1207]" italics="true" pageId="2" pageNumber="3">Atractus boulengerii</emphasis>
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<bibRefCitation id="EFE84B0BAA5C7757FEFFB788170324F1" author="Peracca" box="[377,530,1184,1207]" pageId="2" pageNumber="33" refString="Peracca, M. G. (1896) Sopra alcuni ofidii nuovi o poco noti dell'America meridionale. Bolletin Museo di Torino, 11, 1 - 12." type="journal article" year="1896">Peracca, 1896</bibRefCitation>
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</taxonomicName>
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; Boll. Mus. Zool. Anat. Comp. Torino 11(252):1.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3636571AA5C7757FF11B7CE11D82508" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8BC636FAAA5C7757FF11B7CE11D82508" blockId="2.[151,1437,1254,1358]" pageId="2" pageNumber="3">
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<emphasis id="B90DEAE8AA5C7757FF11B7CE14072546" bold="true" box="[151,278,1254,1280]" pageId="2" pageNumber="3">
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<typeStatus id="54C28858AA5C7757FF11B7CE14002546" box="[151,273,1254,1280]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Adult male, from “
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<collectingCountry id="F36E766AAA5C7757FD85B7CE177F2546" box="[515,622,1254,1280]" name="United States of America" pageId="2" pageNumber="3">America</collectingCountry>
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Meridionali” (= South
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<collectingCountry id="F36E766AAA5C7757FC08B7CE16EA2546" box="[910,1019,1254,1280]" name="United States of America" pageId="2" pageNumber="3">America</collectingCountry>
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), originally deposited in Museo di Zoologia dell’ Università degli Studi di Torino, Turin
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<collectionCode id="ED68AE3FAA5C7757FCB7B62516972561" box="[817,902,1293,1319]" httpUri="http://grbio.org/cool/yjj8-is06" name="Museo di Zoologia, Instituto di Zoologia e Anatomia Comparata Universita di Torino" pageId="2" pageNumber="3">MZUT</collectionCode>
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R1832, now housed at Museo Regionali di Scienze Naturali, Turin (
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<bibRefCitation id="EFE84B0BAA5C7757FE31B61C17FE2508" author="Andreone" box="[439,751,1332,1358]" pageId="2" pageNumber="31" refString="Andreone, F. & Gavetti, E. (2007) The Life and Herpetological Contributions of Mario Ciacinto Peracca (1861 - 1923). Society of the Study of Amphibians and Reptiles Press, Villanova, Pennsylvania, U. S. A., 570 pp." type="book chapter" year="2007">Andreone & Gavetti 2007</bibRefCitation>
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), (specimen examined via photographs).
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</paragraph>
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</subSubSection>
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<caption id="DF066672AA5C7757FF11B4B6102827F3" box="[151,1337,1950,1974]" httpUri="https://zenodo.org/record/191477/files/figure.png" pageId="2" pageNumber="3" targetBox="[195,1392,1415,1923]" targetPageId="2">
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<paragraph id="8BC636FAAA5C7757FF11B4B6102827F3" blockId="2.[151,1337,1950,1974]" box="[151,1337,1950,1974]" pageId="2" pageNumber="3">
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<emphasis id="B90DEAE8AA5C7757FF11B4B6140827F0" bold="true" box="[151,281,1950,1974]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
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Dorsal (A) and ventral (B) views of
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<taxonomicName id="4C794D79AA5C7757FD25B4B7166E27F0" box="[675,895,1951,1974]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="boulengerii">
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<emphasis id="B90DEAE8AA5C7757FD25B4B7166E27F0" box="[675,895,1951,1974]" italics="true" pageId="2" pageNumber="3">Atractus boulengerii</emphasis>
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</taxonomicName>
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(UV-C 6591). SVL 310 mm, CL 52 mm.
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</paragraph>
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</caption>
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<subSubSection id="C3636571AA5D7756FF40B3BF1167223E" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="8BC636FAAA5D7756FF40B3BF112321D3" blockId="3.[151,1438,151,2033]" pageId="3" pageNumber="4">
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<emphasis id="B90DEAE8AA5D7756FF40B3BF145920F7" bold="true" box="[198,328,151,177]" pageId="3" pageNumber="4">Diagnosis:</emphasis>
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<taxonomicName id="4C794D79AA5D7756FED7B3B0175220F7" box="[337,579,152,177]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="boulengerii">
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<emphasis id="B90DEAE8AA5D7756FED7B3B0175220F7" box="[337,579,152,177]" italics="true" pageId="3" pageNumber="4">Atractus boulengerii</emphasis>
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</taxonomicName>
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is distinguished from all congeners by the combination of the following characters: (1) 17/17/17 smooth dorsals; (2) two postoculars; (3) loreal long; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) six infralabials, first three contacting chinshields; (7) five maxillary teeth; (8) four gular scale rows; (9) four preventrals; (10) 176–189 ventrals in males; (11) 41–44 subcaudals in males; (12) dorsum beige with black round blotches decreasing in size posteriorly; (13) venter uniformly creamish white, tail brown pigmented on the median suture of subcaudals; (14) moderate body size, with male reaching
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<quantity id="4C819B1FAA5D7756FEF9B25314F121D3" box="[383,480,379,405]" metricMagnitude="-1" metricUnit="m" metricValue="3.16" pageId="3" pageNumber="4" unit="mm" value="316.0">316 mm</quantity>
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SVL; (15) long tail size in the male (16.8% SVL).
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</paragraph>
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<paragraph id="8BC636FAAA5D7756FF40B2881167223E" blockId="3.[151,1438,151,2033]" pageId="3" pageNumber="4">
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<emphasis id="B90DEAE8AA5D7756FF40B288146721FC" bold="true" box="[198,374,416,442]" pageId="3" pageNumber="4">Comparisons:</emphasis>
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Among all congeners,
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<taxonomicName id="4C794D79AA5D7756FD0DB289162521FC" box="[651,820,417,442]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="boulengerii">
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<emphasis id="B90DEAE8AA5D7756FD0DB289162521FC" box="[651,820,417,442]" italics="true" pageId="3" pageNumber="4">A. boulengerii</emphasis>
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</taxonomicName>
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shares only with
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<taxonomicName id="4C794D79AA5D7756FB80B289119321FC" box="[1030,1154,417,442]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="medusa" status="sp. nov.">
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<emphasis id="B90DEAE8AA5D7756FB80B289119321FC" box="[1030,1154,417,442]" italics="true" pageId="3" pageNumber="4">A. medusa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23E5793AA5D7756FB0CB28911F421FD" box="[1162,1253,417,443]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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(see below) 17 dorsal scale rows, five well-spaced maxillary teeth with a single postdiastemal tooth, more than 40 subcaudals in males, and dorsum with black nuchal collar and round body blotches decreasing in size posteriorly.
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<taxonomicName id="4C794D79AA5D7756FF11B13A1485226D" box="[151,404,530,555]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="boulengerii">
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<emphasis id="B90DEAE8AA5D7756FF11B13A1485226D" box="[151,404,530,555]" italics="true" pageId="3" pageNumber="4">Atractus boulengerii</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C794D79AA5D7756FDBFB13A17A8226D" box="[569,697,530,555]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="medusa">
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<emphasis id="B90DEAE8AA5D7756FDBFB13A17A8226D" box="[569,697,530,555]" italics="true" pageId="3" pageNumber="4">A. medusa</emphasis>
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</taxonomicName>
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in having 176–189 ventrals in males, first three infralabials contacting chinshields, four gular scale rows, and four preventrals (vs. 133 ventrals in the single male, first four infralabials contacting chinshields, three gular scale rows, and two preventrals).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3636571AA5D7756FF40B1AB173C27C6" pageId="3" pageNumber="4" type="description">
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<paragraph id="8BC636FAAA5D7756FF40B1AB14BD24F6" blockId="3.[151,1438,151,2033]" pageId="3" pageNumber="4">
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<emphasis id="B90DEAE8AA5D7756FF40B1AB144622DB" bold="true" box="[198,343,643,669]" pageId="3" pageNumber="4">Description</emphasis>
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: Head twice as long as wide, arched in lateral view, rounded in dorsal view; snout truncate in lateral view, rounded in dorsal view; cervical constriction barely defined; rostral sub-triangular in frontal view, broader than high, poorly visible in dorsal view; internasal longer than wide; internasal suture sinistral with respect to prefrontal suture; prefrontal as long as wide; supraocular sub-trapezoidal, twice as long as wide; frontal pentagonal, as long as wide; parietal about 1.5 times as long as wide; nasal divided; nostril located between prenasal and postnasal; prenasal twice as high as long; postnasal as high as long; loreal long, contacting second and third supralabials; pupil round; two postoculars of similar size; upper postocular slightly longer than lower postocular; temporals 1+2; first temporal twice as long as high; upper posterior temporal elongate, four times as long as wide; seven supralabials, third and fourth contacting orbit; first three supralabials of similar size and lower than fourth; fifth supralabial higher than first four and lower than sixth supralabial; sixth supralabial higher and seventh longer than remaining supralabials; symphisial subtriangular, twice as broad as long; six infralabials, first three contacting chinshields; first pair of infralabials in contact behind symphisial, preventing symphisial/chinshields contact; four gular scale rows; four preventrals; 17/17/17 smooth dorsals; dorsals lacking apical pits, supra-anal tubercles, and keels; terminal spine moderate, conical, and acuminate.
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</paragraph>
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<paragraph id="8BC636FAAA5D7756FF40B7941469250E" blockId="3.[151,1438,151,2033]" pageId="3" pageNumber="4">
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<emphasis id="B90DEAE8AA5D7756FF40B794149D2490" bold="true" box="[198,396,1212,1238]" pageId="3" pageNumber="4">Maxillary arch:</emphasis>
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Arched in dorsal view, with four prediastemal and one postdiastemal teeth; prediastemal teeth large, well spaced, curved posteriorly, angular in cross section, robust at base, narrower at apices, decreasing abruptly in size; maxillary diastema short; postdiastemal tooth slightly smaller than last prediastemal tooth.
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</paragraph>
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<paragraph id="8BC636FAAA5D7756FF40B67B16C02748" blockId="3.[151,1438,151,2033]" pageId="3" pageNumber="4">
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<emphasis id="B90DEAE8AA5D7756FF40B67B14FC252B" bold="true" box="[198,493,1363,1389]" pageId="3" pageNumber="4">Colour in preservative:</emphasis>
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(based on
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<typeStatus id="54C28858AA5D7756FDF2B67B17CB252B" box="[628,730,1363,1389]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
original description and specimen UV-C 6591) Dorsum of head with black cephalic cap extending from rostral to anterior third of parietals; wide occipital band creamish yellow, situated between anterior region of parietals and second dorsal scale rows; background of head black to dorsal margin of supralabials ventrally and to the level of postoculars posteriorly; temporal and occipital regions creamish yellow; supralabials, infralabials, chinshields, and gular region cream; venter uniformly cream; tail cream with small dark brown dots concentrated along the median sutures of subcaudals, forming an irregular line; dorsal ground colour of body beige with about 60 round black blotches paired above paravertebral region; body anteriorly with a wide black collar (six scales long) and large transversal blotches (four or five scales wide); transverse blotches paired or alternating on the flanks, decreasing gradually in size; blotches reaching second dorsal scale rows anteriorly and restricted to paravertebral region posteriorly; vertebral line (one scale wide) distinct on the anterior and posterior thirds of body and tail; vertebral line connected to dorsal blotches; tail dorsally with pattern similar to body.
|
||
</paragraph>
|
||
<paragraph id="8BC636FAAA5D7756FF40B431173C27C6" blockId="3.[151,1438,151,2033]" pageId="3" pageNumber="4">
|
||
<emphasis id="B90DEAE8AA5D7756FF40B43114522775" bold="true" box="[198,323,1817,1843]" pageId="3" pageNumber="4">Variation:</emphasis>
|
||
Additional male specimen have
|
||
<quantity id="4C819B1FAA5D7756FD48B43216232772" box="[718,818,1818,1844]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="4" unit="mm" value="310.0">310 mm</quantity>
|
||
SVL,
|
||
<quantity id="4C819B1FAA5D7756FC04B43216C72772" box="[898,982,1818,1844]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="4" unit="mm" value="52.0">52 mm</quantity>
|
||
CL; tail 16.8% of SVL; 176 ventrals; 141 subcaudals; four gular scale rows; four preventrals; eight or nine dorsal scales on the level of second subcaudal; body diameter
|
||
<quantity id="4C819B1FAA5D7756FE4DB44E173827C6" box="[459,553,1894,1920]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="4" unit="mm" value="5.2">5.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3636571AA5D7756FF40B4A316E527B7" pageId="3" pageNumber="4" type="distribution">
|
||
<paragraph id="8BC636FAAA5D7756FF40B4A316E527B7" blockId="3.[151,1438,151,2033]" pageId="3" pageNumber="4">
|
||
<emphasis id="B90DEAE8AA5D7756FF40B4A3147827E3" bold="true" box="[198,361,1931,1957]" pageId="3" pageNumber="4">Distribution:</emphasis>
|
||
Rediscovered at the lower Anchicayá River, on the old road between Cali and Buenaventura (
|
||
<geoCoordinate id="EE4D503DAA5D7756FECCB49914A0278D" box="[330,433,1969,1995]" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="3.7333333">03º44’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4D503DAA5D7756FE3BB4991737278D" box="[445,550,1969,1995]" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="-76.96667">76º58’W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="4C819B1FAA5D7756FDDBB49917BB278D" box="[605,682,1969,1995]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="m" value="100.0">100 m</quantity>
|
||
), department of Valle del Cauca, Pacific lowlands of
|
||
<collectingCountry id="F36E766AAA5D7756FAA7B4991088278D" box="[1313,1433,1969,1995]" name="Colombia" pageId="3" pageNumber="4">Colombia</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C794D79AA5D7756FF11B4F0149927B7" box="[151,392,2008,2033]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="boulengerii">
|
||
<emphasis id="B90DEAE8AA5D7756FF11B4F0149927B7" box="[151,392,2008,2033]" italics="true" pageId="3" pageNumber="4">Atractus boulengerii</emphasis>
|
||
</taxonomicName>
|
||
apparently inhabits rainforest near sea level (
|
||
<figureCitation id="13422A7FAA5D7756FC18B4FF16F727B7" box="[926,998,2007,2033]" captionStart="FIGURE 2" captionStartId="4.[151,254,1062,1086]" captionTargetBox="[200,1389,156,1061]" captionTargetId="figure@4.[198,1404,145,1061]" captionTargetPageId="4" captionText="FIGURE 2. Geographic distribution of the Atractus multicinctus species group." httpUri="https://zenodo.org/record/191478/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF066672AA5A7751FF11B70E16FA247B" box="[151,1003,1062,1086]" httpUri="https://zenodo.org/record/191478/files/figure.png" pageId="4" pageNumber="5" targetBox="[200,1389,156,1061]" targetPageId="4">
|
||
<paragraph id="8BC636FAAA5A7751FF11B70E16FA247B" blockId="4.[151,1003,1062,1086]" box="[151,1003,1062,1086]" pageId="4" pageNumber="5">
|
||
<emphasis id="B90DEAE8AA5A7751FF11B70E14092478" bold="true" box="[151,280,1062,1086]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
|
||
Geographic distribution of the
|
||
<taxonomicName id="4C794D79AA5A7751FDEEB70E165A247B" box="[616,843,1062,1085]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="multicinctus">
|
||
<emphasis id="B90DEAE8AA5A7751FDEEB70E165A247B" box="[616,843,1062,1085]" italics="true" pageId="4" pageNumber="5">Atractus multicinctus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3636571AA5A7751FF40B7B014542648" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="8BC636FAAA5A7751FF40B7B014542648" blockId="4.[151,1437,1176,1550]" pageId="4" pageNumber="5">
|
||
<emphasis id="B90DEAE8AA5A7751FF40B7B0145324F4" bold="true" box="[198,322,1176,1202]" pageId="4" pageNumber="5">Remarks:</emphasis>
|
||
<bibRefCitation id="EFE84B0BAA5A7751FECFB7B014E824F4" author="Peracca" box="[329,505,1176,1202]" pageId="4" pageNumber="33" refString="Peracca, M. G. (1896) Sopra alcuni ofidii nuovi o poco noti dell'America meridionale. Bolletin Museo di Torino, 11, 1 - 12." type="journal article" year="1896">Peracca (1896)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C794D79AA5A7751FDF1B7B1167824F4" box="[631,873,1177,1202]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="boulengerii">
|
||
<emphasis id="B90DEAE8AA5A7751FDF1B7B1167824F4" box="[631,873,1177,1202]" italics="true" pageId="4" pageNumber="5">Atractus boulengerii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C794D79AA5A7751FC27B7B1112924F4" box="[929,1080,1177,1202]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="iridescens">
|
||
<emphasis id="B90DEAE8AA5A7751FC27B7B1112924F4" box="[929,1080,1177,1202]" italics="true" pageId="4" pageNumber="5">A. iridescens</emphasis>
|
||
</taxonomicName>
|
||
on the basis of two specimens from “
|
||
<collectingCountry id="F36E766AAA5A7751FF64B797145A249F" box="[226,331,1215,1241]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
|
||
Meridionali”, without precise collection data.
|
||
<bibRefCitation id="EFE84B0BAA5A7751FCFFB797115A249F" author="Boulenger" box="[889,1099,1215,1241]" pageId="4" pageNumber="31" refString="Boulenger, G. A. (1913) A collection of batrachians and reptiles made by Dr. H. G. F. Spurrell, F. Z. S., in the Choco. Colombia. Proceedings Zollogical Society, London, 1913, 1019 - 1038." type="proceedings paper" year="1913">Boulenger (1913)</bibRefCitation>
|
||
associated three individuals from Peña Lisa and Condoto in the department of Chocó in
|
||
<collectingCountry id="F36E766AAA5A7751FCE0B7CE16CA2546" box="[870,987,1254,1280]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
|
||
with
|
||
<taxonomicName id="4C794D79AA5A7751FBA6B7CE11A824B9" box="[1056,1209,1254,1279]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="iridescens">
|
||
<emphasis id="B90DEAE8AA5A7751FBA6B7CE11A824B9" box="[1056,1209,1254,1279]" italics="true" pageId="4" pageNumber="5">A. iridescens</emphasis>
|
||
</taxonomicName>
|
||
(see remarks of
|
||
<taxonomicName id="4C794D79AA5A7751FA05B7CE141E2560" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="iridescens">
|
||
<emphasis id="B90DEAE8AA5A7751FA05B7CE141E2560" italics="true" pageId="4" pageNumber="5">A. iridescens</emphasis>
|
||
</taxonomicName>
|
||
), but
|
||
<taxonomicName id="4C794D79AA5A7751FED0B62517112560" box="[342,512,1293,1318]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="boulengerii">
|
||
<emphasis id="B90DEAE8AA5A7751FED0B62517112560" box="[342,512,1293,1318]" italics="true" pageId="4" pageNumber="5">A. boulengerii</emphasis>
|
||
</taxonomicName>
|
||
remained known only from the
|
||
<typeStatus id="54C28858AA5A7751FC04B62416FF2560" box="[898,1006,1292,1318]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
. In the course of this revision of the Pacific
|
||
<taxonomicName id="4C794D79AA5A7751FF72B61B144A250A" box="[244,347,1331,1356]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DEAE8AA5A7751FF72B61B144A250A" box="[244,347,1331,1356]" italics="true" pageId="4" pageNumber="5">Atractus</emphasis>
|
||
</taxonomicName>
|
||
, we found a specimen (UV-C 6591, adult male), from the municipality of Buenaventura, department of Valle del Cauca,
|
||
<collectingCountry id="F36E766AAA5A7751FD89B67217972532" box="[527,646,1370,1396]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
|
||
, which matches the original species description, except for having 176 ventrals and 41 subcaudals (vs.189 ventrals and 44 subcaudals in the
|
||
<typeStatus id="54C28858AA5A7751FB85B6A8117B25DC" box="[1027,1130,1408,1434]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C794D79AA5A7751FB15B6A9102C25DC" box="[1171,1341,1409,1434]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="boulengerii">
|
||
<emphasis id="B90DEAE8AA5A7751FB15B6A9102C25DC" box="[1171,1341,1409,1434]" italics="true" pageId="4" pageNumber="5">A. boulengerii</emphasis>
|
||
</taxonomicName>
|
||
). These small count differences are entirely acceptable in accordance with the pattern of variation of ventral and subcaudals reported for the genus (Savage 1960; Passos 2008), and therefore we refer herein this specimen to
|
||
<taxonomicName id="4C794D79AA5A7751FF11B6DD142F2648" box="[151,318,1525,1550]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="boulengerii">
|
||
<emphasis id="B90DEAE8AA5A7751FF11B6DD142F2648" box="[151,318,1525,1550]" italics="true" pageId="4" pageNumber="5">A. boulengerii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |