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<document id="79FB55EE25416D44A384D810B3681740" ID-DOI="10.11646/zootaxa.3981.1.7" ID-GBIF-Dataset="5e653810-c17b-46b7-b552-db36e102582a" ID-ISSN="1175-5326" ID-Zenodo-Dep="241815" ID-ZooBank="1DBD87D6-7AA4-456B-878D-C50BF38C173D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461245448865" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Rahayu, Dwi Listyo" docDate="2015" docId="038DFD0AFF98E26FFF0DFF52FC96C8CD" docLanguage="en" docName="zt03981p137.pdf" docOrigin="Zootaxa 3981 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexapus sexpes Fabricius 1798" docType="treatment" docVersion="9" lastPageNumber="126" masterDocId="FFB48572FF99E26EFF9AFFC5FFBBCB55" masterDocTitle="Notes on Hexapodidae (Crustacea, Brachyura) from Indonesia and Malaysia, with description of a new species of Mariaplax Rahayu &amp; Ng, 2014, from Lombok, Indonesia" masterLastPageNumber="137" masterPageNumber="125" pageNumber="126" updateTime="1698606294926" updateUser="plazi">
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<mods:title id="52DEC90D846E21972C4E564917C15B2A">Notes on Hexapodidae (Crustacea, Brachyura) from Indonesia and Malaysia, with description of a new species of Mariaplax Rahayu &amp; Ng, 2014, from Lombok, Indonesia</mods:title>
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<mods:namePart id="575357CA2784BBBC03E007EBD304DB74">Ng, Peter K. L.</mods:namePart>
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<treatment id="038DFD0AFF98E26FFF0DFF52FC96C8CD" ID-DOI="http://doi.org/10.5281/zenodo.5457334" ID-GBIF-Taxon="119634724" ID-Zenodo-Dep="5457334" LSID="urn:lsid:plazi:treatment:038DFD0AFF98E26FFF0DFF52FC96C8CD" httpUri="http://treatment.plazi.org/id/038DFD0AFF98E26FFF0DFF52FC96C8CD" lastPageNumber="126" pageId="1" pageNumber="126">
<subSubSection id="C33E1F97FF98E26FFF0DFF52FE9BCB86" pageId="1" pageNumber="126" type="nomenclature">
<paragraph id="8B9B4C1CFF98E26FFF0DFF52FD89CBE7" blockId="1.[151,562,151,211]" box="[151,562,151,178]" pageId="1" pageNumber="126">
<heading id="D0D3FB70FF98E26FFF0DFF52FD89CBE7" bold="true" box="[151,562,151,178]" fontSize="11" level="1" pageId="1" pageNumber="126" reason="1">
<taxonomicName id="4C24379FFF98E26FFF0DFF52FD89CBE7" ID-CoL="3LGRL" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[151,562,151,178]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFF0DFF52FD89CBE7" bold="true" box="[151,562,151,178]" pageId="1" pageNumber="126">
<emphasis id="B950900EFF98E26FFF0DFF52FEE3CBE4" bold="true" box="[151,344,151,177]" italics="true" pageId="1" pageNumber="126">Hexapus sexpes</emphasis>
(
<bibRefCitation id="EFB531EDFF98E26FFEFDFF5DFD91CBE7" author="Fabricius" box="[359,554,152,178]" pageId="1" pageNumber="126" refString="Fabricius, J. C. (1798) Supplementum Entomologiae systematicae. Proft et Storch, Hafniae, 572 pp." type="book" year="1798">Fabricius, 1798</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9B4C1CFF98E26FFF0DFF7FFE9BCB86" blockId="1.[151,562,151,211]" box="[151,288,186,211]" pageId="1" pageNumber="126">
(
<figureCitation id="131F5099FF98E26FFF05FF7FFF65CB86" box="[159,222,186,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1927,1949]" captionTargetBox="[151,1436,265,1904]" captionTargetId="figure@2.[151,1436,265,1905]" captionTargetPageId="2" captionText="FIGURE 1. Overall habitus. A, Hexapus sexpes (Fabricius, 1798), male (7.3 × 4.5 mm, 6.0 × 3.6 mm) (ZRC), Jeram, Peninsular Malaysia; B, Hexapus sexpes (Fabricius, 1798), female (7.6 × 4.7 mm) (ZRC), Jeram, Peninsular Malaysia; C, Hexapinus latus Rahayu &amp; Ng, 2014, female (7.3 × 4.2 mm) (NNM-ZMA Crust De 241762 b), Paternoster Islands." httpUri="https://zenodo.org/record/241816/files/figure.png" pageId="1" pageNumber="126">Fig. 1</figureCitation>
A, B)
</paragraph>
</subSubSection>
<subSubSection id="C33E1F97FF98E26FFF0DFEC6FD56CA62" pageId="1" pageNumber="126" type="reference_group">
<paragraph id="8B9B4C1CFF98E26FFF0DFEC6FDB3CA4D" blockId="1.[151,749,258,311]" box="[151,520,258,280]" pageId="1" pageNumber="126">
<treatmentCitationGroup id="AB346B32FF98E26FFF0DFEC6FDB3CA4D" box="[151,520,258,280]" pageId="1" pageNumber="126">
<taxonomicName id="4C24379FFF98E26FFF0DFEC6FDBFCA4D" authority="Fabricius, 1798: 334" authorityName="Fabricius" authorityPageNumber="334" authorityYear="1798" box="[151,516,258,280]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFF0DFEC6FE91CA4D" box="[151,298,259,280]" italics="true" pageId="1" pageNumber="126">Cancer sexpes</emphasis>
<treatmentCitation id="0A856A0DFF98E26FFEAAFEC7FDBFCA4D" author="Fabricius" box="[304,516,258,280]" page="334" pageId="1" pageNumber="126" year="1798">
<bibRefCitation id="EFB531EDFF98E26FFEAAFEC7FE6ACA4D" author="Fabricius" box="[304,465,258,280]" pageId="1" pageNumber="126" refString="Fabricius, J. C. (1798) Supplementum Entomologiae systematicae. Proft et Storch, Hafniae, 572 pp." type="book" year="1798">Fabricius, 1798</bibRefCitation>
: 334
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B4C1CFF98E26FFF0DFEE4FD56CA62" blockId="1.[151,749,258,311]" box="[151,749,289,311]" pageId="1" pageNumber="126">
See
<bibRefCitation id="EFB531EDFF98E26FFF58FEE4FE77CA62" author="Rahayu" box="[194,460,289,311]" pageId="1" pageNumber="126" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014: 400)</bibRefCitation>
for the complete synonymy.
</paragraph>
</subSubSection>
<subSubSection id="C33E1F97FF98E26FFF0DFEA1FB3CCA91" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="8B9B4C1CFF98E26FFF0DFEA1FB3CCA91" blockId="1.[151,1437,356,920]" pageId="1" pageNumber="126">
<emphasis id="B950900EFF98E26FFF0DFEA1FEC6CA28" bold="true" box="[151,381,356,381]" pageId="1" pageNumber="126">Material examined</emphasis>
.
<emphasis id="B950900EFF98E26FFE14FEA1FD33CA28" bold="true" box="[398,648,356,381]" pageId="1" pageNumber="126">
Peninsular
<collectingCountry id="F3330C8CFF98E26FFD81FEA1FD33CA28" box="[539,648,356,381]" name="Malaysia" pageId="1" pageNumber="126">Malaysia</collectingCountry>
</emphasis>
:
<specimenCount id="9D228795FF98E26FFD00FEA1FD4FCA28" box="[666,756,356,381]" pageId="1" pageNumber="126" type="male">2 males</specimenCount>
(7.3 ×
<quantity id="4CDCE1F9FF98E26FFCCAFEA1FC17CA29" box="[848,940,356,381]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="126" unit="mm" value="4.5">4.5 mm</quantity>
, 6.0 ×
<quantity id="4CDCE1F9FF98E26FFB9EFEA1FBDACA29" box="[1028,1121,356,381]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="126" unit="mm" value="3.6">3.6 mm</quantity>
),
<specimenCount id="9D228795FF98E26FFBE2FEA1FB66CA28" box="[1144,1245,356,381]" pageId="1" pageNumber="126" type="female">1 female</specimenCount>
(7.6 ×
<quantity id="4CDCE1F9FF98E26FFAA3FEA1FA2FCA29" box="[1337,1428,356,380]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="126" unit="mm" value="4.7">4.7 mm</quantity>
) (
<collectionCode id="ED35D4D9FF98E26FFF05FE4DFF60CAF5" box="[159,219,392,416]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="126">ZRC</collectionCode>
), station D, Jeram Polychaete Reef, intertidal muddy shore and intermittent sabellariid reefs, lower, lowest shore, Jeram, Kuala Selangor,
<geoCoordinate id="EE102ADBFF98E26FFE77FE69FDD9CA91" box="[493,610,428,452]" direction="north" orientation="latitude" pageId="1" pageNumber="126" precision="15" value="3.2241666">3°13'27&quot;N</geoCoordinate>
<geoCoordinate id="EE102ADBFF98E26FFDF3FE69FD41CA91" box="[617,762,428,452]" direction="east" orientation="longitude" pageId="1" pageNumber="126" precision="15" value="101.30361">101°18'13&quot;E</geoCoordinate>
, coll. J. J. Eeo,
<date id="FF9A6ADCFF98E26FFC30FE69FB3FCA91" box="[938,1156,428,453]" pageId="1" pageNumber="126" value="2013-07" valueMax="2013-10-31" valueMin="2013-07-00">July–October 2013</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E1F97FF98E26FFF5DFE0AFC96C8CD" pageId="1" pageNumber="126" type="discussion">
<paragraph id="8B9B4C1CFF98E26FFF5DFE0AFAE2C805" blockId="1.[151,1437,356,920]" pageId="1" pageNumber="126">
<emphasis id="B950900EFF98E26FFF5DFE0AFE8FCABD" bold="true" box="[199,308,463,488]" pageId="1" pageNumber="126">Remarks</emphasis>
.
<bibRefCitation id="EFB531EDFF98E26FFEDBFE0AFD96CABD" author="Rahayu" box="[321,557,463,488]" pageId="1" pageNumber="126" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014)</bibRefCitation>
revised the taxonomy of the species and clarified its identity based on material from Phuket in western
<collectingCountry id="F3330C8CFF98E26FFE28FE31FDA2C958" box="[434,537,500,525]" name="Thailand" pageId="1" pageNumber="126">Thailand</collectingCountry>
. Velip &amp; Rivonker (2014) examined the
<typeStatus id="549FF2BEFF98E26FFC62FE30FB93C958" box="[1016,1064,501,525]" pageId="1" pageNumber="126">type</typeStatus>
material of
<taxonomicName id="4C24379FFF98E26FFB21FE30FEC4C965" authority="Sankarankutty, 1975" authorityName="Sankarankutty" authorityYear="1975" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="estuarinus">
<emphasis id="B950900EFF98E26FFB21FE30FA20C959" box="[1211,1435,501,524]" italics="true" pageId="1" pageNumber="126">Hexapus estuarinus</emphasis>
Sankarankutty, 1975
</taxonomicName>
, from southern
<collectingCountry id="F3330C8CFF98E26FFDABFDD2FDD6C965" box="[561,621,535,560]" name="India" pageId="1" pageNumber="126">India</collectingCountry>
, and commented that it was a valid species and not synonymous with
<taxonomicName id="4C24379FFF98E26FFA1BFDDCFF5BC901" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFA1BFDDCFF5BC901" italics="true" pageId="1" pageNumber="126">H. sexpes</emphasis>
</taxonomicName>
. They argued that the structure of the chela (with a basal tooth on the dactylus of the major chela) was different, that there was a large gap between the closed fingers of the minor chela, setae at the base of the propodal finger was absent, and tubercles were present on the thoracic sternum. The differences observed for the chela are subject to size (the
<typeStatus id="549FF2BEFF98E26FFEEAFD6DFE1BC995" box="[368,416,680,704]" pageId="1" pageNumber="126">type</typeStatus>
of
<taxonomicName id="4C24379FFF98E26FFE5CFD6CFD8AC995" box="[454,561,681,704]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFE5CFD6CFD8AC995" box="[454,561,681,704]" italics="true" pageId="1" pageNumber="126">H. sexpes</emphasis>
</taxonomicName>
is relatively smaller than that of
<taxonomicName id="4C24379FFF98E26FFC3EFD6CFB80C9EA" box="[932,1083,680,704]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="estuarinus">
<emphasis id="B950900EFF98E26FFC3EFD6CFB80C9EA" box="[932,1083,680,704]" italics="true" pageId="1" pageNumber="126">H. estuarinus</emphasis>
</taxonomicName>
), however, and this structure is also known to vary to some degree. We do not regard these differences as substantial; they certainly agree very well with the material identified as
<taxonomicName id="4C24379FFF98E26FFE69FD34FDE4C85D" box="[499,607,753,776]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFE69FD34FDE4C85D" box="[499,607,753,776]" italics="true" pageId="1" pageNumber="126">H. sexpes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB531EDFF98E26FFD16FD2AFCC4C85D" author="Rahayu" box="[652,895,751,776]" pageId="1" pageNumber="126" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014)</bibRefCitation>
, the redescription of the
<typeStatus id="549FF2BEFF98E26FFB01FD35FB70C85D" box="[1179,1227,752,776]" pageId="1" pageNumber="126">type</typeStatus>
male by
<bibRefCitation id="EFB531EDFF98E26FFAADFD2AFF58C879" author="Manning" pageId="1" pageNumber="126" refString="Manning, R. B. (1982) A redescription of Cancer sexpes Fabricius, 1798 (Crustacea, Decapoda, Hexapodidae). Steenstrupia, 8, 157 - 161." type="journal article" year="1982">Manning (1982)</bibRefCitation>
, and the present material here from
<collectingCountry id="F3330C8CFF98E26FFDEEFCD1FD65C878" box="[628,734,788,813]" name="Malaysia" pageId="1" pageNumber="126">Malaysia</collectingCountry>
. As such, we prefer to regard them as synonymous for the time being. The presence of the species in the Straits of
<collectingCountry id="F3330C8CFF98E26FFD4AFCF2FC89C805" box="[720,818,823,848]" name="Malaysia" pageId="1" pageNumber="126">Malacca</collectingCountry>
, Peninsular
<collectingCountry id="F3330C8CFF98E26FFC20FCF2FB9AC805" box="[954,1057,823,848]" name="Malaysia" pageId="1" pageNumber="126">Malaysia</collectingCountry>
is nevertheless noteworthy.
</paragraph>
<paragraph id="8B9B4C1CFF98E26FFF5DFC99FC96C8CD" blockId="1.[151,1437,356,920]" pageId="1" pageNumber="126">
Three species of
<taxonomicName id="4C24379FFF98E26FFE09FC98FE4EC821" box="[403,501,861,884]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B950900EFF98E26FFE09FC98FE4EC821" box="[403,501,861,884]" italics="true" pageId="1" pageNumber="126">Hexapus</emphasis>
</taxonomicName>
are now known:
<taxonomicName id="4C24379FFF98E26FFD50FC98FB6DC821" authority="Velip &amp; Rivonker, 2014" authorityName="Velip &amp; Rivonker" authorityYear="2014" box="[714,1238,860,885]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="B950900EFF98E26FFD50FC98FC16C821" box="[714,941,860,884]" italics="true" pageId="1" pageNumber="126">Hexapus bidentatus</emphasis>
Velip &amp; Rivonker, 2014
</taxonomicName>
,
<taxonomicName id="4C24379FFF98E26FFB7FFC98FEE3C8CD" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFB7FFC98FA27C821" box="[1253,1436,861,884]" italics="true" pageId="1" pageNumber="126">Hexapus sexpes</emphasis>
(
<bibRefCitation id="EFB531EDFF98E26FFF04FCBAFEF4C8CD" author="Fabricius" box="[158,335,895,920]" pageId="1" pageNumber="126" refString="Fabricius, J. C. (1798) Supplementum Entomologiae systematicae. Proft et Storch, Hafniae, 572 pp." type="book" year="1798">Fabricius, 1798</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C24379FFF98E26FFE14FC44FC92C8CD" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[398,809,895,920]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="timika">
<emphasis id="B950900EFF98E26FFE14FC44FD86C8CD" box="[398,573,896,920]" italics="true" pageId="1" pageNumber="126">Hexapus timika</emphasis>
<bibRefCitation id="EFB531EDFF98E26FFDDEFCBAFC92C8CD" author="Rahayu" box="[580,809,895,920]" pageId="1" pageNumber="126" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng, 2014</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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