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<mods:title id="8362ED6FC4A10A9949C8AA1EE70330E5">A new integrated morpho- and molecular systematic classification of Cenozoic radiolarians (Class Polycystinea) - suprageneric taxonomy and logical nomenclatorial acts</mods:title>
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<mods:namePart id="74D922F15A10CD8CA31C4F7E132D7F1A">Caulet, Jean-Pierre</mods:namePart>
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<treatment id="038DDA73FFC9FE6B066EF88FFCFC4CDD" ID-GBIF-Taxon="183733842" LSID="urn:lsid:plazi:treatment:038DDA73FFC9FE6B066EF88FFCFC4CDD" httpUri="http://treatment.plazi.org/id/038DDA73FFC9FE6B066EF88FFCFC4CDD" lastPageId="94" lastPageNumber="497" pageId="93" pageNumber="496">
<subSection id="E2AB728EFFC9FE68066EF88FFA784814" box="[859,1409,1886,1912]" pageId="93" pageNumber="496" type="multiple">
<subSubSection id="C33E38EEFFC9FE68066EF88FFA784814" box="[859,1409,1886,1912]" pageId="93" pageNumber="496" type="nomenclature">
<paragraph id="8B9B6B65FFC9FE68066EF88FFA784814" blockId="93.[859,1409,1886,1912]" box="[859,1409,1886,1912]" pageId="93" pageNumber="496">
Superfamily
<taxonomicName id="4C2410E6FFC9FE6806D1F88FFAC94814" authority="Haeckel, 1882" authorityName="Haeckel" authorityYear="1882" box="[996,1328,1886,1912]" class="Polycystina" kingdom="Chromista" order="Nassellaria" pageId="93" pageNumber="496" phylum="Radiozoa" rank="superFamily" superFamily="Theopilioidea">
THEOPILIOIDEA
<bibRefCitation id="EFB51694FFC9FE6801A6F88FFAC94814" author="HAECKEL E." box="[1171,1328,1886,1912]" pageId="93" pageNumber="496" pagination="418 - 472" refId="ref179241" refString="HAECKEL E. 1882. - Entwurf eines Radiolarien-Systems auf Grund von Studien der Challenger-Radiolarien. Jenaische Zeitschrift fur Naturwissenschaft 15: 418 - 472. https: // www. biodiversitylibrary. org / page / 8700599" type="journal article" year="1882">Haeckel, 1882</bibRefCitation>
</taxonomicName>
n. stat.
</paragraph>
</subSubSection>
</subSection>
<subSubSection id="C33E38EEFFC9FE6B06EEF8AFFD0D4E55" lastPageId="94" lastPageNumber="497" pageId="93" pageNumber="496" type="reference_group">
<paragraph id="8B9B6B65FFC9FE6806EEF8AFFAF848F4" blockId="93.[987,1281,1918,1944]" box="[987,1281,1918,1944]" pageId="93" pageNumber="496">
<heading id="D0D3DC09FFC9FE6806EEF8AFFAF848F4" box="[987,1281,1918,1944]" centered="true" fontSize="11" level="2" pageId="93" pageNumber="496" reason="2">
<taxonomicName id="4C2410E6FFC9FE6806EEF8AFFBA648F4" authority="Suzuki" authorityName="Suzuki" box="[987,1119,1918,1944]" class="Polycystina" family="Theoperidae" genus="Calocyclas" kingdom="Chromista" order="Nassellaria" pageId="93" pageNumber="496" phylum="Radiozoa" rank="species" species="sensu">
<emphasis id="B950B777FFC9FE6806EEF8AFFBF648FB" box="[987,1039,1918,1943]" italics="true" pageId="93" pageNumber="496">sensu</emphasis>
Suzuki
</taxonomicName>
emend. herein
</heading>
</paragraph>
<paragraph id="8B9B6B65FFC9FE680618F869FC544887" blockId="93.[813,1455,1976,2027]" pageId="93" pageNumber="496">
Theopilida
<bibRefCitation id="EFB51694FFC9FE6806A9F869FBA648BC" author="HAECKEL E." box="[924,1119,1976,2000]" pageId="93" pageNumber="496" pagination="418 - 472" refId="ref179241" refString="HAECKEL E. 1882. - Entwurf eines Radiolarien-Systems auf Grund von Studien der Challenger-Radiolarien. Jenaische Zeitschrift fur Naturwissenschaft 15: 418 - 472. https: // www. biodiversitylibrary. org / page / 8700599" type="journal article" year="1882">Haeckel, 1882: 435</bibRefCitation>
[as a tribe]; 1887: 1313, 1315 [as a subfamily].
</paragraph>
<paragraph id="8B9B6B65FFCAFE6B05B1FF06FDF94E65" blockId="94.[132,775,215,265]" pageId="94" pageNumber="497">
Theopiliilae
<bibRefCitation id="EFB51694FFCAFE6B043EFF06FE5A4F83" author="CAMPBELL A. S." box="[267,419,215,239]" pageId="94" pageNumber="497" pagination="11 - 195" refId="ref170808" refString="CAMPBELL A. S. 1954. - Radiolaria, in MOORE R. C. (ed.), Treatise on Invertebrate Paleontology. Vol. Part. D, Protista 3. Geological Society of America and University of Kansas Press, Lawrence / Kansas: 11 - 195." type="book chapter" year="1954">Campbell 1954</bibRefCitation>
: D129 [as a subsuperfamily]. —
<bibRefCitation id="EFB51694FFCAFE6B07D5FF06FEDD4E65" author="NAKASEKO K." pageId="94" pageNumber="497" pagination="20 - 42" refId="ref188110" refString="NAKASEKO K. 1957. - On Radiolaria. Yukochu (Foraminifera) 8: 20 - 42. [in Japanese]" type="journal article" year="1957">Nakaseko 1957: 27</bibRefCitation>
[as a subsuperfamily].
</paragraph>
<paragraph id="8B9B6B65FFCAFE6B05B1FEF0FD0D4E55" blockId="94.[132,756,289,313]" box="[132,756,289,313]" pageId="94" pageNumber="497">
Neosciadiocapsaceae
<bibRefCitation id="EFB51694FFCAFE6B0466FEF0FDC34E55" author="O'DOGHERTY L." box="[339,570,289,313]" pageId="94" pageNumber="497" pagination="1 - 415" refId="ref189547" refString="O'DOGHERTY L. 1994. - Biochronology and Paleontology of Mid-Cretaceous Radiolarians from Northern Apennines (Italy) and Betic Cordillera (Spain). Memoires de Geologie (Lausanne) 21: 1 - 415." type="journal article" year="1994">ODogherty, 1994: 227</bibRefCitation>
[as a superfamily].
</paragraph>
</subSubSection>
<subSubSection id="C33E38EEFFCAFE6B05B1FE83FD984EBE" pageId="94" pageNumber="497" type="diagnosis">
<paragraph id="8B9B6B65FFCAFE6B05B1FE83FD984EBE" blockId="94.[131,777,337,467]" pageId="94" pageNumber="497">DIAGNOSIS. — Skeleton having a very shallow hat-shaped to hatshaped shell with two segments. The thoracic pore frames of the thorax are systematically distributed in both longitudinal and lateral directions. The distal end of the thorax is associated with a velum, a latticed frame, many feet and other ornaments.</paragraph>
</subSubSection>
<subSubSection id="C33E38EEFFCAFE6B05B1FE29FCFC4CDD" pageId="94" pageNumber="497" type="discussion">
<paragraph id="8B9B6B65FFCAFE6B05B1FE29FF114D7C" blockId="94.[130,777,504,945]" box="[132,232,504,530]" pageId="94" pageNumber="497">REMARKS</paragraph>
<paragraph id="8B9B6B65FFCAFE6B05B1FDC6FCFC4CDD" blockId="94.[130,777,504,945]" pageId="94" pageNumber="497">
This superfamily consists of the
<taxonomicName id="4C2410E6FFCAFE6B04FEFDC6FD8F4D5D" authorityName="Haeckel" authorityYear="1882" box="[459,630,535,561]" class="Polycystina" family="Anthocyrtididae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="570" phylum="Radiozoa" rank="family">Anthocyrtididae</taxonomicName>
and
<taxonomicName id="4C2410E6FFCAFE6B0793FDC6FF4A4D3D" class="Polycystinea" family="Theopiliidae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="497" phylum="Retaria" rank="family">Theopiliidae</taxonomicName>
. The taxonomic position of the
<taxonomicName id="4C2410E6FFCAFE6B04CAFDE6FD764D3D" authorityName="Haeckel" authorityYear="1882" box="[511,655,567,593]" class="Polycystina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="570" phylum="Radiozoa" rank="superFamily" superFamily="Theopilioidea">Theopilioidea</taxonomicName>
is based on the molecular phylogenetic position of
<taxonomicName id="4C2410E6FFCAFE6B0705FD86FE5C4DFD" authority="(Lineage III, Sandin et al. 2019)" baseAuthorityName="Lineage III, Sandin" baseAuthorityYear="2019" class="Polycystina" family="Theoperidae" genus="Eucecryphalus" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="497" phylum="Radiozoa" rank="genus">
<emphasis id="B950B777FFCAFE6B0705FD86FD474D1D" box="[560,702,599,625]" italics="true" pageId="94" pageNumber="497">Eucecryphalus</emphasis>
(Lineage III,
<bibRefCitation id="EFB51694FFCAFE6B05EDFDA6FE634DFD" author="SANDIN M. M. &amp; PILLET L. &amp; BIARD T. &amp; POIRIER C. &amp; BIGEARD E. &amp; ROMAC S. &amp; SUZUKI N. &amp; NOT F." box="[216,410,631,657]" pageId="94" pageNumber="497" pagination="187 - 208" refId="ref194897" refString="SANDIN M. M., PILLET L., BIARD T., POIRIER C., BIGEARD E., ROMAC S., SUZUKI N. &amp; NOT F. 2019. - Time Calibrated Morpho-molecular Classification of Nassellaria (Radiolaria). Protist 170 (2): 187 - 208. https: // doi. org / 10.1016 / j. protis. 2019.02.002" type="journal article" year="2019">
Sandin
<emphasis id="B950B777FFCAFE6B041CFDA9FEA54DFD" box="[297,348,631,657]" italics="true" pageId="94" pageNumber="497">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
. They correspond to nassellarians characterized by a very shallow hat-shaped shells, classified into family
<taxonomicName id="4C2410E6FFCAFE6B05FBFD66FE854DBD" authorityName="Haeckel" authorityYear="1882" box="[206,380,695,721]" class="Polycystina" family="Anthocyrtididae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="497" phylum="Radiozoa" rank="family">Anthocyrtididae</taxonomicName>
(=
<taxonomicName id="4C2410E6FFCAFE6B0496FD66FEA74D9D" authority="in De Wever et al. 2001: 233 - 235" authorityName="in De Wever" authorityPageNumber="233 - 235" authorityYear="2001" class="Polycystinea" family="Neosciadiocapsidae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="497" phylum="Retaria" rank="family">
Neosciadiocapsidae in
<bibRefCitation id="EFB51694FFCAFE6B07A9FD66FEA74D9D" author="DE WEVER P. &amp; DUMITRICA P. &amp; CAULET J. P. &amp; NIGRINI C. &amp; CARIDROIT M." pageId="94" pageNumber="497" refId="ref173394" refString="DE WEVER P., DUMITRICA P., CAULET J. P., NIGRINI C. &amp; CARIDROIT M. 2001. - Radiolarians in the sedimentary record, Amsterdam, 533 p. https: // doi. org / 10.1201 / 9781482283181" type="book" year="2001">
De Wever
<emphasis id="B950B777FFCAFE6B05B0FD06FF414D9C" box="[133,184,726,752]" italics="true" pageId="94" pageNumber="497">et al.</emphasis>
2001: 233-235
</bibRefCitation>
</taxonomicName>
) or
<taxonomicName id="4C2410E6FFCAFE6B04BEFD07FF134C7D" authority="(De Wever et al. 2001: 238 - 239)" baseAuthorityName="De Wever" baseAuthorityPageNumber="238 - 239" baseAuthorityYear="2001" class="Polycystinea" family="Theopiliidae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="497" phylum="Retaria" rank="family">
Theopiliidae (
<bibRefCitation id="EFB51694FFCAFE6B072BFD06FF1B4C7D" author="DE WEVER P. &amp; DUMITRICA P. &amp; CAULET J. P. &amp; NIGRINI C. &amp; CARIDROIT M." pageId="94" pageNumber="497" refId="ref173394" refString="DE WEVER P., DUMITRICA P., CAULET J. P., NIGRINI C. &amp; CARIDROIT M. 2001. - Radiolarians in the sedimentary record, Amsterdam, 533 p. https: // doi. org / 10.1201 / 9781482283181" type="book" year="2001">
De Wever
<emphasis id="B950B777FFCAFE6B07A5FD06FD3B4D9C" box="[656,706,726,752]" italics="true" pageId="94" pageNumber="497">et al.</emphasis>
2001: 238-239
</bibRefCitation>
)
</taxonomicName>
. However, molecular phylogenetic data placed a close related morphological group, the
<taxonomicName id="4C2410E6FFCAFE6B04D1FCC7FD6E4C5C" authorityName="Suzuki" authorityYear="2019" box="[484,663,790,816]" class="Polycystina" family="Cycladophoridae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="571" phylum="Radiozoa" rank="family">Cycladophoridae</taxonomicName>
(based on
<taxonomicName id="4C2410E6FFCAFE6B05B1FCE7FEF44C3C" authorityName="Ehrenberg" authorityYear="1846" box="[132,269,822,848]" class="Polycystina" family="Theoperidae" genus="Cycladophora" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="497" phylum="Radiozoa" rank="genus">
<emphasis id="B950B777FFCAFE6B05B1FCE7FEF44C3C" box="[132,269,822,848]" italics="true" pageId="94" pageNumber="497">Cycladophora</emphasis>
</taxonomicName>
) into Lineage IV (
<bibRefCitation id="EFB51694FFCAFE6B04EEFCE7FD5B4C3C" author="SANDIN M. M. &amp; PILLET L. &amp; BIARD T. &amp; POIRIER C. &amp; BIGEARD E. &amp; ROMAC S. &amp; SUZUKI N. &amp; NOT F." box="[475,674,822,848]" pageId="94" pageNumber="497" pagination="187 - 208" refId="ref194897" refString="SANDIN M. M., PILLET L., BIARD T., POIRIER C., BIGEARD E., ROMAC S., SUZUKI N. &amp; NOT F. 2019. - Time Calibrated Morpho-molecular Classification of Nassellaria (Radiolaria). Protist 170 (2): 187 - 208. https: // doi. org / 10.1016 / j. protis. 2019.02.002" type="journal article" year="2019">
Sandin
<emphasis id="B950B777FFCAFE6B071BFCE6FD9A4C3C" box="[558,611,822,848]" italics="true" pageId="94" pageNumber="497">et al.</emphasis>
2019
</bibRefCitation>
). At that time, it was impossible to conceptualize a higher classification position for the
<taxonomicName id="4C2410E6FFCAFE6B0416FCA7FE374CFC" authorityName="Haeckel" authorityYear="1882" box="[291,462,886,912]" class="Polycystina" family="Anthocyrtididae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="94" pageNumber="570" phylum="Radiozoa" rank="family">Anthocyrtididae</taxonomicName>
. The
<taxonomicName id="4C2410E6FFCAFE6B0734FCA7FD534CFC" authorityName="Haeckel" authorityYear="1882" box="[513,682,886,912]" family="Lampromitridae" pageId="94" pageNumber="570" rank="family">Lampromitridae</taxonomicName>
may also belong to this superfamily (see remarks for
<taxonomicName id="4C2410E6FFCAFE6B0772FC47FD004CDD" authorityName="Haeckel" authorityYear="1882" box="[583,761,918,945]" family="Lampromitridae" pageId="94" pageNumber="570" rank="family">Lampromitridae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>