263 lines
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263 lines
40 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.987.55317" ID-GBIF-Dataset="17f5a541-c279-4823-8c0e-2af5a2ebc943" ID-PMC="PMC7666079" ID-Pensoft-Pub="1313-2970-987-115" ID-Pensoft-UUID="A1D8404DABD3516296CD9F36436A73E3" ID-PubMed="33223887" ID-ZooBank="BF3D30382C5A4EC4BAB47700D0DFB374" ModsDocID="1313-2970-987-115" checkinTime="1604753987840" checkinUser="pensoft" docAuthor="Bashir, Nawaz Haider, Ma, Li & Li, Qiang" docDate="2020" docId="F1C3EC6A76675EEFB318B24A80339D46" docLanguage="en" docName="ZooKeys 987: 115-134" docOrigin="ZooKeys 987" docSource="http://dx.doi.org/10.3897/zookeys.987.55317" docTitle="Carinostigmus vesulcatus Bashir & Ma 2020, sp. nov." docType="treatment" docUuid="0FDD9981-3372-4E78-8C16-BB9D7ECC7353" docUuidSource="ZooBank" docVersion="4" id="A1D8404DABD3516296CD9F36436A73E3" lastPageNumber="115" masterDocId="A1D8404DABD3516296CD9F36436A73E3" masterDocTitle="Chinese species of Carinostigmus Tsuneki (Hymenoptera, Crabronidae), including three new species and a new record to China" masterLastPageNumber="134" masterPageNumber="115" pageNumber="115" updateTime="1668169775777" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Chinese species of Carinostigmus Tsuneki (Hymenoptera, Crabronidae), including three new species and a new record to China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bashir, Nawaz Haider</mods:namePart>
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<mods:affiliation>Department of Entomology, College of Plant Protection, Yunnan Agricultural University, Kunming, Yunnan, 650201, China</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-9627-9920</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ma, Li</mods:namePart>
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<mods:affiliation>Department of Entomology, College of Plant Protection, Yunnan Agricultural University, Kunming, Yunnan, 650201, China</mods:affiliation>
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<mods:nameIdentifier type="email">maliwasps@aliyun.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Qiang</mods:namePart>
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<mods:affiliation>Department of Entomology, College of Plant Protection, Yunnan Agricultural University, Kunming, Yunnan, 650201, China</mods:affiliation>
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<mods:nameIdentifier type="email">liqiangkm@126.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>987</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>115</mods:start>
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<mods:end>134</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.987.55317</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.987.55317</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-987-115</mods:identifier>
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<mods:identifier type="ZooBank">BF3D30382C5A4EC4BAB47700D0DFB374</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A1D8404DABD3516296CD9F36436A73E3</mods:identifier>
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<treatment ID-GBIF-Taxon="168821861" LSID="urn:lsid:zoobank.org:act:0FDD9981-3372-4E78-8C16-BB9D7ECC7353" httpUri="http://treatment.plazi.org/id/F1C3EC6A76675EEFB318B24A80339D46" lastPageNumber="115" pageId="0" pageNumber="115">
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<subSubSection pageId="0" pageNumber="115" type="nomenclature">
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<paragraph pageId="0" pageNumber="115">
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<taxonomicName LSID="F1C3EC6A-7667-5EEF-B318-B24A80339D46" authority="Bashir & Ma" authorityName="Bashir & Ma" authorityYear="2020" class="Insecta" family="Crabronidae" genus="Carinostigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinostigmus vesulcatus" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="vesulcatus" status="sp. nov.">Carinostigmus vesulcatus Bashir & Ma</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="115">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">Figs 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a Carinostigmus frontirugatus Bashir & Ma, sp. nov., (female) b, c C. latidentatus Bashir & Ma, sp. nov., (b female c male) d, e C. vesulcatus Bashir & Ma, sp. nov., (d female e male) f C. palawanensis (Tsuneki, 1976) (female) a-f lateral view." figureDoi="10.3897/zookeys.987.55317.figure5" httpUri="https://binary.pensoft.net/fig/472420" pageId="0" pageNumber="115">, 5d, e</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="115" type="type material">
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<paragraph pageId="0" pageNumber="115">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Holotype</emphasis>
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: ♀, China: Yunnan: Jinghong: Menghai: Bulang Mountain,
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<geoCoordinate degrees="21" direction="north" minutes="57" orientation="latitude" precision="925" value="21.95">21°57'N</geoCoordinate>
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,
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<geoCoordinate degrees="100" direction="east" minutes="27" orientation="longitude" precision="925" value="100.45">100°27'E</geoCoordinate>
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, 17-VI.21.V.2018, No. 2019000099, Malaise trap (YNAU).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Paratypes</emphasis>
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: 2♂♂, China: Yunnan: Dehong: Nabang,
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<geoCoordinate degrees="24" direction="north" minutes="26" orientation="latitude" precision="925" value="24.433332">24°26'N</geoCoordinate>
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,
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<geoCoordinate degrees="98" direction="east" minutes="35" orientation="longitude" precision="925" value="98.583336">98°35'E</geoCoordinate>
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, 15.V.2009, No. 201005191, coll. Jie Zeng, No. 201005205, coll. Manman Wang (YNAU); 1♂, China: Yunnan: Dehong: Yinjiang: Tongbiguan,
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<geoCoordinate degrees="24" direction="north" minutes="42" orientation="latitude" precision="925" value="24.7">24°42'N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="55" orientation="longitude" precision="925" value="97.916664">97°55'E</geoCoordinate>
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, 18.V.2009, No. 201005224, coll. Manman Wang (YNAU); 1♀, China: Yunnan: Nujiang: Fugong: Yueliang Village,
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<geoCoordinate degrees="25" direction="north" minutes="49" orientation="latitude" precision="925" value="25.816668">25°49'N</geoCoordinate>
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,
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<geoCoordinate degrees="98" direction="east" minutes="51" orientation="longitude" precision="925" value="98.85">98°51'E</geoCoordinate>
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, 27.V.2007, coll. Feng Yuan (CAS); 1♀, China: Yunnan: Kaiyuan: Nandong,
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<geoCoordinate degrees="23" direction="north" minutes="40" orientation="latitude" precision="925" value="23.666666">23°40'N</geoCoordinate>
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,
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<geoCoordinate degrees="103" direction="east" minutes="15" orientation="longitude" precision="925" value="103.25">103°15'E</geoCoordinate>
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, 16.VII.2003, coll. Qiang Li (YNAU).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="115" type="diagnosis">
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<paragraph pageId="0" pageNumber="115">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="115">
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This species resembles
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<taxonomicName lsidName="C. congruus" pageId="0" pageNumber="115" rank="species" species="congruus">
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<emphasis italics="true" pageId="0" pageNumber="115">C. congruus</emphasis>
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</taxonomicName>
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(Walker, 1860) in sharing: median and lower frons microstriate, with a sturdy frontal median longitudinal carina (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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); upper frons with fine and coarse punctures, frontal carina distinct anteriorly, not reaching anterior ocellus (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4b</figureCitation>
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); ocellar triangle flat, gena with several sturdy oblique transverse rugae medially, lower gena with weak striations; outer-orbital sulcus broad; notaulus deeply grooved (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4j</figureCitation>
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); propodeal enclosure triangular, with sturdy longitudinal rugae anteriorly, slender dense, longitudinal rugae laterally (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4k</figureCitation>
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); pygidial area oval shaped (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4h</figureCitation>
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). It can be differentiated from
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<taxonomicName lsidName="C. congruus" pageId="0" pageNumber="115" rank="species" species="congruus">
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<emphasis italics="true" pageId="0" pageNumber="115">C. congruus</emphasis>
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</taxonomicName>
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by labrum, in female, pentagonal, longer than wide (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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), in male, wider as long, deeply emarginated apically forming two rounded lobes (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4d</figureCitation>
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); clypeus slightly convex; free margin of clypeal lobe nearly truncate medially, teeth inconspicuous (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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); inter-antennal tubercle without T-shaped at apex (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4b</figureCitation>
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); inner-orbital sulcus narrowed, inconspicuously foveolate (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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); occipital carina foveolate (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4b</figureCitation>
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); anterior pronotal ridge strongly marked (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4f</figureCitation>
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); in female, fovea absent on scutum posterior margin (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4f</figureCitation>
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); admedian line conspicuous in female, inconspicuous in male; parapsidal line conspicuous, and metanotum with inconspicuous rugae medially (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4f</figureCitation>
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).
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<taxonomicName baseAuthorityName="Walker" baseAuthorityYear="1860" class="Insecta" family="Crabronidae" genus="Carinostigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carinostigmus congruus" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="congruus">
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<emphasis italics="true" pageId="0" pageNumber="115">Carinostigmus congruus</emphasis>
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</taxonomicName>
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(Walker) has the following characters: labrum triangular, broadly rounded at apex; clypeus strongly convex at middle; free margin of clypeal lobe slightly emarginate medially, with two distinct lateral small teeth (
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<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.396" author="Krombein, KV" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="115" pagination="1 - 37" refId="B4" refString="Krombein, KV, 1984. Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae). Smithsonian Contributions to Zoology 396: 1 - 37, DOI: https://doi.org/10.5479/si.00810282.396" title="Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae)." url="https://doi.org/10.5479/si.00810282.396" volume="396" year="1984">Krombein 1984</bibRefCitation>
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: fig. 9); inter-antennal tubercle with T-shaped at apex (
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<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.396" author="Krombein, KV" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="115" pagination="1 - 37" refId="B4" refString="Krombein, KV, 1984. Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae). Smithsonian Contributions to Zoology 396: 1 - 37, DOI: https://doi.org/10.5479/si.00810282.396" title="Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae)." url="https://doi.org/10.5479/si.00810282.396" volume="396" year="1984">Krombein 1984</bibRefCitation>
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: fig. 15); inner-orbital sulcus broad, distinctly foveolate (
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<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.396" author="Krombein, KV" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="115" pagination="1 - 37" refId="B4" refString="Krombein, KV, 1984. Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae). Smithsonian Contributions to Zoology 396: 1 - 37, DOI: https://doi.org/10.5479/si.00810282.396" title="Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae)." url="https://doi.org/10.5479/si.00810282.396" volume="396" year="1984">Krombein 1984</bibRefCitation>
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: fig. 3); occipital carina not foveolate; pronotal collar ridged weakly marked; in female, fovea present on scutum posterior margin; admedian and parapsidal lines inconspicuous, and metanotum smooth medially (
|
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<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.396" author="Krombein, KV" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="115" pagination="1 - 37" refId="B4" refString="Krombein, KV, 1984. Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae). Smithsonian Contributions to Zoology 396: 1 - 37, DOI: https://doi.org/10.5479/si.00810282.396" title="Biosystematic studies of Ceylonese wasps, XIV: a revision of Carinostigmus Tsuneki (Hymenoptera: Sphecoidea: Pemphredonidae)." url="https://doi.org/10.5479/si.00810282.396" volume="396" year="1984">Krombein 1984</bibRefCitation>
|
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: fig. 39).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="115" type="description">
|
|
<paragraph pageId="0" pageNumber="115">Description.</paragraph>
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<paragraph pageId="0" pageNumber="115">
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|
Female (Figs
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|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a-c, f-k</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a Carinostigmus frontirugatus Bashir & Ma, sp. nov., (female) b, c C. latidentatus Bashir & Ma, sp. nov., (b female c male) d, e C. vesulcatus Bashir & Ma, sp. nov., (d female e male) f C. palawanensis (Tsuneki, 1976) (female) a-f lateral view." figureDoi="10.3897/zookeys.987.55317.figure5" httpUri="https://binary.pensoft.net/fig/472420" pageId="0" pageNumber="115">5d</figureCitation>
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): Based on holotype, if any variation in paratypes described in square brackets.
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</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Measurements.</emphasis>
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</paragraph>
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<paragraph pageId="0" pageNumber="115">♀, BL 4.6 [4.5-4.9] mm;</paragraph>
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<paragraph pageId="0" pageNumber="115">HW:HLD:HLF = 60:38:40;</paragraph>
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<paragraph pageId="0" pageNumber="115">HWmax:HWmin = 60:31;</paragraph>
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|
<paragraph pageId="0" pageNumber="115">HW:EWd:IOW:EW:OOW:TW:OCW:EL = 60:14:1:16:2:22:1:41;</paragraph>
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<paragraph pageId="0" pageNumber="115">AOD:WAS:IAD = 5:5:10;</paragraph>
|
|
<paragraph pageId="0" pageNumber="115">POD:OOD:OCD:IODc:IODv:IODmin = 4:12:14:28:36:28;</paragraph>
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<paragraph pageId="0" pageNumber="115">LS:LP:LFI:WFI:LFII:WFII = 18:7:8:3:8:3;</paragraph>
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<paragraph pageId="0" pageNumber="115">LC:WC = 25:5;</paragraph>
|
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<paragraph pageId="0" pageNumber="115">PL:PW:LTI:WTI = 45:6:30:23.</paragraph>
|
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<paragraph pageId="0" pageNumber="115">♂, BL 4-4.6 mm;</paragraph>
|
|
<paragraph pageId="0" pageNumber="115">HW:HLD:HLF = 65:35:43;</paragraph>
|
|
<paragraph pageId="0" pageNumber="115">HWmax:HWmin = 65:31;</paragraph>
|
|
<paragraph pageId="0" pageNumber="115">HW:EWd:IOW:EW:OOW:TW:OCW:EL = 65:18:1:18:2:20:1:40;</paragraph>
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<paragraph pageId="0" pageNumber="115">AOD:WAS:IAD = 3:3:8;</paragraph>
|
|
<paragraph pageId="0" pageNumber="115">POD:OOD:OCD:IODc:IODv:IODmin = 4:10:12:20:36:20;</paragraph>
|
|
<paragraph pageId="0" pageNumber="115">LS:LP:LFI:WFI:LFII:WFII = 20:8:8:2:8:2;</paragraph>
|
|
<paragraph pageId="0" pageNumber="115">LC:WC = 23:5;</paragraph>
|
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<paragraph pageId="0" pageNumber="115">PL:PW:LTI:WTI = 46:6:27:25.</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Color pattern.</emphasis>
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Body black with shiny aspect, except the following: mandible yellowish, apically dark; labrum yellowish with ivory marked medially; palpi pale; scape and pedicel extensively yellowish; flagellomeres I-III yellowish, remaining darker; pronotal lobe ivory with yellowish spot; tegula fulvous; forewing veins dark brown; legs fulvous except mid and hind femur reddish brown; pale setae on clypeal margin and on mandible.
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</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Head.</emphasis>
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|
Mandible tridentate apically; setae on mandible sparsed, long, longer than labrum length. Labrum pentagonal, rounded toward apex (Fig.
|
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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). Clypeus slightly convex, with coarse punctures; setae on margin of clypeus sparsed, long as labrum length; free margin of clypeal lobe nearly truncate, sinuous, not forming reflexed teeth (Fig.
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|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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). Median and lower frons microstriate, with a sturdy frontal median longitudinal carina (Fig.
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|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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), inter-antennal tubercle short, less than midocellus diameter (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4b</figureCitation>
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); upper frons smooth, with fine and coarse punctures, frontal carina distinct anteriorly, not reaching anterior ocellus (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4b</figureCitation>
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). Ocellar triangle nearly flat, finely punctate [impunctate]; vertex behind ocelli, finely punctate [impunctate]; gena with several sturdy transverse rugae medially; lower gena with weak striations. Occipital carina broad and foveolate as Fig.
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|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4b</figureCitation>
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; inner-orbital sulcus narrow as flagellomere I width, with inner marginal carina distinct, inconspicuously foveolate (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4a</figureCitation>
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); outer-orbital sulcus broad as pedicel width, hind marginal carina distinct, inconspicuously foveolate.
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</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Mesosoma.</emphasis>
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Pronotal collar smooth, strongly elevated mesally, ridged inconspicuous anteriorly, weakly marked, lateral angles blunt (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4f</figureCitation>
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|
). Scutum with fine punctures, sparsely distributed, several slender transverse striations anteriorly (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4j</figureCitation>
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|
); admedian line conspicuous, extending to one third of scutum length; notaulus deeply grooved and foveolate, extending to only anterior of scutum length (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4j</figureCitation>
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|
); parapsidal line inconspicuous (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4f</figureCitation>
|
|
). Scutellum dull aspect densely micropunctated; metanotum with dense, slender, longitudinal rugae laterally, weak medially (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4f</figureCitation>
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). Mesopleuron with fine punctures, sparsely distributed; omaulus and hypersternaulus narrowed as pedicel width, inconspicuously foveate [distinctly foveolate]; scrobal sulcus absent (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4g</figureCitation>
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). Propodeal enclosure shallowly impressed, triangular, with sturdy longitudinal rugae anteriorly, slender dense, longitudinal rugae laterally (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4k</figureCitation>
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); propodeum posteriorly with oval median groove, slender dense transverse rugae anteriorly, smooth medially (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4k</figureCitation>
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); propodeal side presenting obliques and regular striae, sparsely distributed (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4g</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Legs.</emphasis>
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Outer surface of hindtibia without spines.
|
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</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="115">Metasoma.</emphasis>
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Petiole dorsal surface sub quadrate (cross section), basal width narrower than apically (Fig.
|
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4c</figureCitation>
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), side smooth. Gaster segments III-V [IV-VI] with fine sparsed punctures, rest impunctate (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4i</figureCitation>
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). Pygidial area punctate anteriorly, oval shaped (Fig.
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|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4h</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="115">
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<emphasis bold="true" pageId="0" pageNumber="115">Male</emphasis>
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(Figs
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|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinostigmus vesulcatus Bashir & Ma, sp. nov. (a-c, f-k female d, e male): a, d head (frontal view) b, e head (dorsal view) c petiole (dorsal view) f scutum, scutellum and metanotum (dorsal view) g thorax (lateral view) h pygidial plate i metasoma (dorsal view) j scutum anterior k propodeum (dorso-posterior view)." figureDoi="10.3897/zookeys.987.55317.figure4" httpUri="https://binary.pensoft.net/fig/472419" pageId="0" pageNumber="115">4d, e</figureCitation>
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,
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|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. a Carinostigmus frontirugatus Bashir & Ma, sp. nov., (female) b, c C. latidentatus Bashir & Ma, sp. nov., (b female c male) d, e C. vesulcatus Bashir & Ma, sp. nov., (d female e male) f C. palawanensis (Tsuneki, 1976) (female) a-f lateral view." figureDoi="10.3897/zookeys.987.55317.figure5" httpUri="https://binary.pensoft.net/fig/472420" pageId="0" pageNumber="115">5e</figureCitation>
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). Same as female except mandible ivory, reddish brown apically; labrum, scape beneath and pedicel ivory, remaining scape and flagellomeres fulvous; flagellomeres without tyloids; forewing veins brown to dark brown; legs ivory to fulvous; setae on margin of clypeus and on mandible fulvous and short, less than pedicel length; mandible bidentate apically; labrum notched, deeply emarginated at apex; clypeus slightly microstriate, convex, impunctate; gena with several sturdy oblique transverse rugae near eye; fovea present on scutum posterior margin; parapsidal line inconspicuous.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="115" type="distribution">
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<paragraph pageId="0" pageNumber="115">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="115">China (Yunnan).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="115" type="etymology">
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<paragraph pageId="0" pageNumber="115">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="115">
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The name
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<taxonomicName authorityName="Bashir & Ma" authorityYear="2020" class="Insecta" family="Crabronidae" genus="Carinostigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="vesulcatus" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="vesulcatus">
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<emphasis italics="true" pageId="0" pageNumber="115">vesulcatus</emphasis>
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</taxonomicName>
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, is derived from the Latin words
|
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<emphasis italics="true" pageId="0" pageNumber="115">ve</emphasis>
|
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- (= without) and
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<emphasis italics="true" pageId="0" pageNumber="115">sulcatus</emphasis>
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(= sulcate), referring to the scrobal sulcus absent.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |