155 lines
20 KiB
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155 lines
20 KiB
XML
<document id="05C7534DEF6DF3C76880F20669DEEB1D" ID-CLB-Dataset="5966" ID-DOI="10.5852/ejt.2021.755.1405" ID-GBIF-Dataset="b932c9e5-ddad-4c11-8eee-ff17188d77cf" ID-ISSN="2118-9773" ID-Zenodo-Dep="5032970" ID-ZooBank="F9105E33-3E8B-4B3C-88B3-0316207B70F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1624643559605" checkinUser="felipe" docAuthor="Gale, Andy S. & Jagt, John W. M." docDate="2021" docId="F160A367717BFFBFFD86760FFB7723FA" docLanguage="en" docName="ejt-755_149-190.pdf" docOrigin="European Journal of Taxonomy 755" docSource="http://dx.doi.org/10.5852/ejt.2021.755.1405" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Jurapecten hessi Gale 2011" docType="treatment" docVersion="9" lastPageNumber="162" masterDocId="0D59DB1F7176FFB2FFF77706FFD02038" masterDocTitle="The fossil record of the family Benthopectinidae (Echinodermata, Asteroidea), a reappraisal" masterLastPageNumber="190" masterPageNumber="149" pageNumber="162" updateTime="1698976416854" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="03AB05BD3807619881625088F2151F3A">The fossil record of the family Benthopectinidae (Echinodermata, Asteroidea), a reappraisal</mods:title>
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<mods:namePart id="AED7B1B5F86AD39D571C0A8AFA08BE27">Gale, Andy S.</mods:namePart>
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<mods:affiliation id="7B6E4D1D110FAD53CE5110E7D2C42648">Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. School of the Environment, Geography and Geological Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO 13 QL, United Kingdom. Natuurhistorisch Museum Maastricht, De Bosquetplein 6 - 7, 6211 KJ Maastricht, the Netherlands.</mods:affiliation>
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<mods:namePart id="C474079CB26A25992DD7A8721E220FFD">Jagt, John W. M.</mods:namePart>
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<mods:date id="9152474E35C7C4905DD14AC5CF87E329">2021</mods:date>
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<treatment id="F160A367717BFFBFFD86760FFB7723FA" ID-DOI="http://doi.org/10.5281/zenodo.5036866" ID-GBIF-Taxon="183195111" ID-Zenodo-Dep="5036866" LSID="urn:lsid:plazi:treatment:F160A367717BFFBFFD86760FFB7723FA" httpUri="http://treatment.plazi.org/id/F160A367717BFFBFFD86760FFB7723FA" lastPageNumber="162" pageId="13" pageNumber="162">
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<subSubSection id="31D341FA717BFFBFFD86760FFC13211B" box="[625,963,264,291]" pageId="13" pageNumber="162" type="nomenclature">
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<paragraph id="79761271717BFFBFFD86760FFC13211B" blockId="13.[625,963,264,326]" box="[625,963,264,291]" pageId="13" pageNumber="162">
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<heading id="223EA51D717BFFBFFD86760FFC13211B" box="[625,963,264,291]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="162" reason="2">
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<taxonomicName id="BEC969F2717BFFBFFD86760FFC13211B" ID-CoL="44918a68-9d8f-49fa-8027-bdef49d925f7" authority="Gale, 2011" authorityName="Gale" authorityYear="2011" box="[625,963,264,291]" class="Asteroidea" family="Benthopectinidae" genus="Jurapecten" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="13" pageNumber="162" phylum="Echinodermata" rank="species" species="hessi">
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<emphasis id="4BBDCE63717BFFBFFD86760FFCEB211B" bold="true" box="[625,827,264,291]" italics="true" pageId="13" pageNumber="162">Jurapecten hessi</emphasis>
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Gale, 2011
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</paragraph>
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<subSubSection id="31D341FA717BFFBFFD58762AFC55217E" box="[687,901,299,326]" pageId="13" pageNumber="162" type="description">
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<paragraph id="79761271717BFFBFFD58762AFC55217E" blockId="13.[625,963,264,326]" box="[687,901,299,326]" pageId="13" pageNumber="162">
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<figureCitation id="E1F20EF4717BFFBFFD58762AFCE2217E" box="[687,818,299,326]" captionStart="Fig" captionStartId="13.[189,232,1329,1355]" captionTargetId="figure-14@14.[189,1398,290,1878]" captionTargetPageId="14" captionText="Fig. 6 (next page). A–B, F.? Benthopectinidae. A–B. Inferomarginals, in lateral aspect (MHI 2183/7, MHI 2183/8). F. Oral ossicle in radial view (MHI 2183/9). — C, G. Jurapecten hessi Gale, 2011. C. Inferomarginal, in lateral view (NHMUK PI EE 17988). G. Oral ossicle, in radial view (NHMUK EE 13606; the original of Gale 2011a: pl. 20 fig. 10). — D–E, H. Pontaster tenuispinus (von Düben & Koren, 1846).D–E. Lateral view of inferomarginals.H. Oral ossicle (A.S. Gale collection, unregistered). — I–U. Jurapecten infrajurensis sp. nov. I. Oral ossicle, radial view (SNSB-BGSP2020XLV 31). J–M. Ambulacral ossicles, in actinal views (J = holotype, NHMUK PI EE 17989; K–L = paratypes, NHMUK PI EE 17990–17991; M = SNSB-BGSP2020XLV16). N–O. Marginal ossicles (paratypes, NHMUK PI EE 17992–17993). P, R–T. Adambulacral ossicles (paratypes), in actinal (P, R) and abactinal (S–T) views (R–S = NHMUK PI EE 17994–17995; P, T = SNSB-BGSP2020XLV 17, SNSB- BGSP2020XLV 19). I, Q. Oral ossicles, in radial view (paratypes: SNSB-BGSP2020XLV 15, SNSB- BGSP2020XLV 18). U. Circumoral ossicle (paratype: SNSB-BGSP2020XLV 20). Provenance: A–B, F. Maantang Formation (Carnian, Upper Triassic), sample C30, Jiancougou, Sechuan Province, China. C, G. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). D–E, H. Recent, Porcupine Trough, NE Atlantic. I, M, P–Q, T–U. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; Hess 2014). J–L, N–O, R–S. Middle Toarcian, Le Clapier (Département d’Aveyron), 62 km WNW of Montpellier, southern France (Thuy 2012). See Fig. 3 for detailed morphological terminology. Scale bars: A–H = 0.4 mm; I–U = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5036868" httpUri="https://zenodo.org/record/5036868/files/figure.png" pageId="13" pageNumber="162">Figs 6C, G</figureCitation>
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,
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<figureCitation id="E1F20EF4717BFFBFFCC9762AFC55217E" box="[830,901,300,326]" captionStart="Fig" captionStartId="15.[189,232,1328,1354]" captionTargetId="figure-15@16.[189,1398,283,1870]" captionTargetPageId="16" captionText="Fig. 7 (next page). A–J. Jurapecten infrajurensis sp. nov., various ossicle types (paratypes). A, H. Ambulacrals, in lateral and actinal aspects, respectively (SNSB-BGSP2020XLV 21,SNSB- BGSP2020XLV 28). B, E–F. Adambulacrals, in actinal and abactinal views (SNSB-BGSP2020XLV 22, SNSB-BGSP2020XLV 25–26). C, G, J. Superomarginals, in lateral views (SNSB-BGSP2020XLV 23, SNSB-BGSP2020XLV 27, SNSB-BGSP2020XLV 30). D, I. Inferomarginals, in lateral views (SNSB-BGSP2020XLV 24, SNSB-BGSP2020XLV 29). — K–Y. Jurapecten hessi Gale, 2011. K, P, U. Adambulacral ossicles. K. Abactinal view (NHMUK PI EE 17996). P. NHMUK EE 13598 (the original of Gale 2011a: pl. 19 fig. 11). U. NHMUK EE 13599, the original of Gale (2011a: pl. 20 fig. 12). L. Circumoral ossicle (NHMUK EE 13605, the original of Gale 2011a: pl. 20 fig. 13). M–N. Ambulacral ossicles, in actinal and abactinal views, respectively (NHMUK EE 13595–13596, the originals of Gale 2011a: pl. 19 figs 8–9). O. Ambulacral, in proximal/distal view (NHMUK 13597). Q–S. Superomarginal ossicles (NHMUK EE 13601–13602; the originals of Gale 2011a: pl. 20 figs 6–7). T, V–W. Inferomarginals (NHMUK EE 13600, EE 13603–13604, EE 13608, the originals of Gale 2011a: pl. 20 figs 5, 8, 16). X–Y. Oral ossicles (NHMUK EE 13606– 13607, the originals of Gale 2011a: pl. 20 figs 10, 14). Provenance: A–J. Upper Toarcian–lower Aalenian (possibly aalensis ammonite Zone), Quedlinburg (Landkreis Harz, northern Germany; see Hess 2014). K–Y. Upper Oxfordian (Couches d’Effingen, bifurcatus ammonite Zone, stenocycloides ammonite Subzone), Savigna, near Orgelet, Département du Jura, France (Gale 2011b). Abbreviations: d = distal; p = proximal. See Fig. 3 for detailed morphological terminology. Scale bars: A–J = 0.2 mm; K–Y = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5036870" httpUri="https://zenodo.org/record/5036870/files/figure.png" pageId="13" pageNumber="162">7K–Y</figureCitation>
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</paragraph>
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<subSubSection id="31D341FA717BFFBFFF4A766BFD1F21FF" pageId="13" pageNumber="162" type="reference_group">
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<paragraph id="79761271717BFFBFFF4A766BFB4221BF" blockId="13.[189,1170,364,391]" box="[189,1170,364,391]" pageId="13" pageNumber="162">
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<treatmentCitationGroup id="59D9355F717BFFBFFF4A766BFB4221BF" box="[189,1170,364,391]" pageId="13" pageNumber="162">
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<treatmentCitation id="F8683460717BFFBFFF4A766BFD9521BF" author="Gale A. S." box="[189,581,364,391]" page="84" pageId="13" pageNumber="162" year="2011">
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<taxonomicName id="BEC969F2717BFFBFFF4A766BFD9521BF" ID-CoL="44918a68-9d8f-49fa-8027-bdef49d925f7" authority="Gale, 2011 a: 84" authorityName="Gale" authorityPageNumber="84" authorityYear="2011" box="[189,581,364,391]" class="Asteroidea" family="Benthopectinidae" genus="Jurapecten" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="13" pageNumber="162" phylum="Echinodermata" rank="species" species="hessi">
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<emphasis id="4BBDCE63717BFFBFFF4A766BFE5221BE" box="[189,386,364,390]" italics="true" pageId="13" pageNumber="162">Jurapecten hessi</emphasis>
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<bibRefCitation id="1D586F80717BFFBFFE7E766AFD9521BF" author="Gale A. S." box="[393,581,364,391]" pageId="13" pageNumber="162" pagination="1 - 112" refId="ref16782" refString="Gale A. S. 2011 a. The phylogeny of post-Palaeozoic Asteroidea (Neoasteroidea, Echinodermata). Special Papers in Palaeontology 85: 1 - 112." type="journal article" year="2011">Gale, 2011a: 84</bibRefCitation>
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</taxonomicName>
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, pl. 19 figs 1–6, 8–12, pl. 20 figs 6–10, 12–14, 19.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="79761271717BFFBFFF4A76A8FD1F21FF" blockId="13.[189,719,429,456]" box="[189,719,429,456]" pageId="13" pageNumber="162">
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<treatmentCitationGroup id="59D9355F717BFFBFFF4A76A8FD1F21FF" box="[189,719,429,456]" pageId="13" pageNumber="162">
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<taxonomicName id="BEC969F2717BFFBFFF4A76A8FE5221FF" ID-CoL="44918a68-9d8f-49fa-8027-bdef49d925f7" authorityName="Gale" authorityYear="2011" box="[189,386,429,455]" class="Asteroidea" family="Benthopectinidae" genus="Jurapecten" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="13" pageNumber="162" phylum="Echinodermata" rank="species" species="hessi">
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<emphasis id="4BBDCE63717BFFBFFF4A76A8FE5221FF" box="[189,386,429,455]" italics="true" pageId="13" pageNumber="162">Jurapecten hessi</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="F8683460717BFFBFFE6876ABFD8521FF" author="Gale A. S." box="[415,597,429,455]" page="76" pageId="13" pageNumber="162" year="2011">
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<bibRefCitation id="1D586F80717BFFBFFE6876ABFD8521FF" author="Gale A. S." box="[415,597,429,455]" pageId="13" pageNumber="162" refId="ref16811" refString="Gale A. S. 2011 b. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Department [sic] du Jura, France. In: Meyer C. A. & Costeur L. (eds) Special Issue: Echinoderms - from the early" type="book" year="2011">Gale 2011b: 76</bibRefCitation>
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</treatmentCitation>
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, fig. 6a–h.
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</paragraph>
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</subSubSection>
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<subSubSection id="31D341FA717BFFBFFF4A76E8FD2F2261" pageId="13" pageNumber="162" type="diagnosis">
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<paragraph id="79761271717BFFBFFF4A76E8FEE62230" blockId="13.[189,310,494,520]" box="[189,310,494,520]" pageId="13" pageNumber="162">
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<heading id="223EA51D717BFFBFFF4A76E8FEE62230" bold="true" box="[189,310,494,520]" fontSize="11" level="3" pageId="13" pageNumber="162" reason="3">
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<emphasis id="4BBDCE63717BFFBFFF4A76E8FEE62230" bold="true" box="[189,310,494,520]" pageId="13" pageNumber="162">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="79761271717BFFBFFF4A751BFD2F2261" blockId="13.[189,1399,540,602]" pageId="13" pageNumber="162">
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<taxonomicName id="BEC969F2717BFFBFFF4A751BFEEF220E" authorityName="Gale" authorityYear="2011" box="[189,319,541,566]" class="Asteroidea" family="Benthopectinidae" genus="Jurapecten" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="13" pageNumber="162" phylum="Echinodermata" rank="genus">
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<emphasis id="4BBDCE63717BFFBFFF4A751BFEEF220E" box="[189,319,541,566]" italics="true" pageId="13" pageNumber="162">Jurapecten</emphasis>
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</taxonomicName>
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in which the marginals bear a sculpture of discrete, rounded rugosities and 1–3 larger spine bases on inferomarginals. Ambulacral base short.
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</paragraph>
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</subSubSection>
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<subSubSection id="31D341FA717BFFBFFF4A7586FAF72309" pageId="13" pageNumber="162" type="materials_examined">
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<paragraph id="79761271717BFFBFFF4A7586FE7A22A2" blockId="13.[189,426,640,666]" box="[189,426,640,666]" pageId="13" pageNumber="162">
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<heading id="223EA51D717BFFBFFF4A7586FE7A22A2" bold="true" box="[189,426,640,666]" fontSize="11" level="3" pageId="13" pageNumber="162" reason="3">
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<emphasis id="4BBDCE63717BFFBFFF4A7586FE7A22A2" bold="true" box="[189,426,640,666]" pageId="13" pageNumber="162">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="79761271717BFFBFFF4A75A8FAF72309" blockId="13.[189,1400,686,817]" pageId="13" pageNumber="162">
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<materialsCitation id="C9A1182C717BFFBFFF4A75A8FAF72309" ID-GBIF-Occurrence="3311523310" collectionCode="NHMUK" pageId="13" pageNumber="162" specimenCode="NHMUK EE 13594" specimenCount="1">
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The type specimen,
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<specimenCode id="296FBA0A717BFFBFFE5375A8FD5E22F1" box="[420,654,686,713]" pageId="13" pageNumber="162">NHMUK EE 13594</specimenCode>
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, consists of a set of associated ossicles from the upper Oxfordian (Couches d’Effingen,
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<emphasis id="4BBDCE63717BFFBFFE3475D6FDE922D2" box="[451,569,720,746]" italics="true" pageId="13" pageNumber="162">bifurcatus</emphasis>
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ammonite Zone,
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<emphasis id="4BBDCE63717BFFBFFCF975D6FC6922D2" box="[782,953,720,746]" italics="true" pageId="13" pageNumber="162">stenocycloides</emphasis>
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||
ammonite Subzone) at Savigna, near Orgelet (Département du Jura, France; see
|
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Gale 2011
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||
<bibRefCitation id="1D586F80717BFFBFFCCC75F5FC982335" author="Gale A. S." box="[827,840,755,781]" pageId="13" pageNumber="162" refId="ref16811" refString="Gale A. S. 2011 b. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Department [sic] du Jura, France. In: Meyer C. A. & Costeur L. (eds) Special Issue: Echinoderms - from the early" type="book" year="2011">a</bibRefCitation>
|
||
</bibRefCitation>
|
||
, 2011b). Additional material comprises several hundred ossicles and groups of associated ossicles from the type locality (
|
||
<collectionCode id="1FD88AB4717BFFBFFBE87410FB402308" box="[1055,1168,790,816]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="162" type="Museum">NHMUK</collectionCode>
|
||
collections).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31D341FA717BFFBFFF4A7451FB7723FA" pageId="13" pageNumber="162" type="discussion">
|
||
<paragraph id="79761271717BFFBFFF4A7451FEFF2349" blockId="13.[189,303,855,881]" box="[189,303,855,881]" pageId="13" pageNumber="162">
|
||
<heading id="223EA51D717BFFBFFF4A7451FEFF2349" bold="true" box="[189,303,855,881]" fontSize="11" level="3" pageId="13" pageNumber="162" reason="3">
|
||
<emphasis id="4BBDCE63717BFFBFFF4A7451FEFF2349" bold="true" box="[189,303,855,881]" pageId="13" pageNumber="162">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="79761271717BFFBFFF4A7483FB7723FA" blockId="13.[189,1399,901,962]" pageId="13" pageNumber="162">
|
||
There is little new information or material since the original description by
|
||
<bibRefCitation id="1D586F80717BFFBFFB837483FADA23A7" author="Gale A. S." box="[1140,1290,901,927]" pageId="13" pageNumber="162" pagination="1 - 112" refId="ref16782" refString="Gale A. S. 2011 a. The phylogeny of post-Palaeozoic Asteroidea (Neoasteroidea, Echinodermata). Special Papers in Palaeontology 85: 1 - 112." type="journal article" year="2011">
|
||
Gale (2011
|
||
<bibRefCitation id="1D586F80717BFFBFFB0A7483FADA23A7" author="Gale A. S." box="[1277,1290,901,927]" pageId="13" pageNumber="162" refId="ref16811" refString="Gale A. S. 2011 b. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Department [sic] du Jura, France. In: Meyer C. A. & Costeur L. (eds) Special Issue: Echinoderms - from the early" type="book" year="2011">a</bibRefCitation>
|
||
</bibRefCitation>
|
||
, 2011b); differences with
|
||
<taxonomicName id="BEC969F2717BFFBFFE7574AEFDE223FA" authority="Gale & Jagt, 2021" authorityName="Gale & Jagt" authorityYear="2021" box="[386,562,936,962]" class="Asteroidea" family="Benthopectinidae" genus="Jurapecten" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paxillosida" pageId="13" pageNumber="162" phylum="Echinodermata" rank="species" species="infrajurensis" status="sp. nov.">
|
||
<emphasis id="4BBDCE63717BFFBFFE7574AEFDE223FA" box="[386,562,936,962]" italics="true" pageId="13" pageNumber="162">J. infrajurensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="508E7318717BFFBFFDCE74AEFD4423FA" box="[569,660,936,962]" pageId="13" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are discussed under that species (see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |