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<mods:title id="E2D02436886B3887282E59ED85D8142E">Morphometric analysis of Eocene nummulitids in western and central Cuba: taxonomy, biostratigraphy and evolutionary trends</mods:title>
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<mods:namePart id="DE6ED64C656F45FCD9FB3F8D2378A9EF">Eder, Wolfgang</mods:namePart>
<mods:affiliation id="3EB6145E7BBDC6B0E9008F5B93469580">Department of Palaeontology, University of Vienna, Althanstrasse 14, 1090, Vienna, Austria;</mods:affiliation>
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<mods:namePart id="EE33B7A86BB048853A3BD102A7AF49E6">Hohenegger, Johann</mods:namePart>
<mods:affiliation id="DDCE2E399997645A3F239DF9AF79D295">Department of Palaeontology, University of Vienna, Althanstrasse 14, 1090, Vienna, Austria;</mods:affiliation>
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<mods:affiliation id="ABD108DF4A1AAFAE47D138F2981B4B7F">Dipartimento di Scienze della, Terra dellAmbiente e della Vita, Universita ́ degli Studi di Genova, Corso Europa, 26, I- 16132 Genova, Italy</mods:affiliation>
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<treatment id="9E6A87F1FFC12828FEFBFB3890958BA7" LSID="urn:lsid:plazi:treatment:9E6A87F1FFC12828FEFBFB3890958BA7" httpUri="http://treatment.plazi.org/id/9E6A87F1FFC12828FEFBFB3890958BA7" lastPageId="27" lastPageNumber="583" pageId="26" pageNumber="582">
<subSubSection id="5ED9656CFFC12829FEFBFB3893F18DF7" box="[264,677,1228,1252]" pageId="26" pageNumber="582" type="nomenclature">
<paragraph id="167C36E7FFC12829FEFBFB3893F18DF7" blockId="26.[264,677,1228,1252]" box="[264,677,1228,1252]" pageId="26" pageNumber="582">
<heading id="4D34818BFFC12829FEFBFB3893F18DF7" box="[264,677,1228,1252]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="582" reason="6">
Genus
<taxonomicName id="D1C34D64FFC12829FEA1FB3893F18DF7" ID-CoL="BFKTC" authority="Hanzawa, 1935" authorityName="Hanzawa" authorityYear="1935" box="[338,677,1228,1252]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FEA1FB3890AD8DF0" bold="true" box="[338,505,1228,1252]" italics="true" pageId="26" pageNumber="582">Operculinoides</emphasis>
<bibRefCitation id="72524B16FFC12829FE0CFB3893F18DF7" author="Hanzawa, S." box="[511,677,1228,1251]" pageId="26" pageNumber="582" pagination="1 - 29" refId="ref21083" refString="Hanzawa, S. 1935. Some fossil Operculina and Miogypsina from Japan and their stratigraphical significance. Science Reports of the Tohoku Imperial University. 2 nd Series, Geology 18, 1 - 29." type="journal article" year="1935">Hanzawa, 1935</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5ED9656CFFC12829FF6CFAFC92A98C2B" pageId="26" pageNumber="582" type="diagnosis">
<paragraph id="167C36E7FFC12829FF6CFAFC925A8C2B" blockId="26.[159,782,1288,1343]" pageId="26" pageNumber="582">
<emphasis id="24B7EAF5FFC12829FF6CFAFC90408C0B" bold="true" box="[159,276,1288,1311]" pageId="26" pageNumber="582">Diagnosis.</emphasis>
Planispiral, involute or partially involute in the nepionic stage, becoming evolute in the adult stage.
</paragraph>
<paragraph id="167C36E7FFC12829FCBEFCBA95218D97" blockId="26.[845,1467,846,1251]" pageId="26" pageNumber="582">Tests with the strongest marginal radius increase and strongest backward bend angles of the investigated individuals, producing rapidly widening coils and highly projecting later chambers. Chambers are up to 4 times as high as wide and are separated by primary operculine septa with septal undulations, which are more pronounced in loosely coiled forms. These forms with the highest values in chamber height in the adult stage have chambers up to 10 times higher than wide. The marginal cord is moderately well developed.</paragraph>
<paragraph id="167C36E7FFC12829FC94FB7894418DF7" blockId="26.[845,1467,846,1251]" pageId="26" pageNumber="582">
Characters and attributes (means and standard deviations) for
<taxonomicName id="D1C34D64FFC12829FC48FB58950B8DD7" authorityName="Hanzawa" authorityYear="1935" box="[955,1119,1196,1219]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FC48FB58950B8DD7" box="[955,1119,1196,1219]" italics="true" pageId="26" pageNumber="582">Operculinoides</emphasis>
</taxonomicName>
and comparison to
<taxonomicName id="D1C34D64FFC12829FAC8FB5894EE8DD7" box="[1339,1466,1196,1219]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FAC8FB5894EE8DD7" box="[1339,1466,1196,1219]" italics="true" pageId="26" pageNumber="582">Nummulites</emphasis>
</taxonomicName>
and
<emphasis id="24B7EAF5FFC12829FC89FB3895678DF7" box="[890,1075,1228,1251]" italics="true" pageId="26" pageNumber="582">Palaeoummulites</emphasis>
are given in
<tableCitation id="5B41035CFFC12829FB4EFB38945A8DF7" box="[1213,1294,1228,1251]" captionStart="Table 6" captionStartId="26.[159,217,1418,1439]" captionText="Table 6. Characters and attributes (means and standard deviations, SD, in mm) for Operculinoides and comparisons with Nummulites and Palaeoummulites. Symbol key: ++, strong positive differences with &lt;1% error probability; 0, no significant differences; ――, strong negative differences with &lt;1% error probability." pageId="26" pageNumber="582">Table 6</tableCitation>
.
</paragraph>
<paragraph id="167C36E7FFC12829FCBEFAFC92A98C2B" blockId="26.[845,1466,1288,1343]" pageId="26" pageNumber="582">
<emphasis id="24B7EAF5FFC12829FCBEFAFC92B78C0B" bold="true" box="[845,995,1288,1311]" pageId="26" pageNumber="582">Occurrences.</emphasis>
<taxonomicName id="D1C34D64FFC12829FC01FAFC95C28C0B" authorityName="Hanzawa" authorityYear="1935" box="[1010,1174,1288,1311]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FC01FAFC95C28C0B" box="[1010,1174,1288,1311]" italics="true" pageId="26" pageNumber="582">Operculinoides</emphasis>
</taxonomicName>
is common in the middle and late Eocene.
</paragraph>
</subSubSection>
<caption id="42BC666FFFC12829FF6CFA7E93E88CC0" pageId="26" pageNumber="582" startId="26.[159,217,1418,1439]" targetType="table">
<paragraph id="167C36E7FFC12829FF6CFA7E93E88CC0" blockId="26.[159,1466,1418,1492]" pageId="26" pageNumber="582">
<emphasis id="24B7EAF5FFC12829FF6CFA7E91A78C8B" bold="true" box="[159,243,1418,1439]" pageId="26" pageNumber="582">Table 6.</emphasis>
Characters and attributes (means and standard deviations, SD, in mm) for
<taxonomicName id="D1C34D64FFC12829FC21FA7E95328C8B" authorityName="Hanzawa" authorityYear="1935" box="[978,1126,1418,1439]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FC21FA7E95328C8B" box="[978,1126,1418,1439]" italics="true" pageId="26" pageNumber="582">Operculinoides</emphasis>
</taxonomicName>
and comparisons with
<taxonomicName id="D1C34D64FFC12829FABBFA7E94EE8C8B" box="[1352,1466,1418,1439]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FABBFA7E94EE8C8B" box="[1352,1466,1418,1439]" italics="true" pageId="26" pageNumber="582">Nummulites</emphasis>
</taxonomicName>
and
<emphasis id="24B7EAF5FFC12829FF38FA50902C8CAD" box="[203,376,1444,1465]" italics="true" pageId="26" pageNumber="582">Palaeoummulites.</emphasis>
Symbol key: ++, strong positive differences with &lt;1% error probability; 0, no significant differences; ――, strong negative differences with &lt;1% error probability.
</paragraph>
</caption>
<paragraph id="167C36E7FFC12829FF6CFA0594C08E63" pageId="26" pageNumber="582">
<table id="64C3C447FFC1D7CCFF6CFA0594EE8E63" box="[159,1466,1521,1911]" gridcols="5" gridrows="12" pageId="26" pageNumber="582" rowsContinueFrom="26.[159,1466,316,732]">
<tr id="A8F334A5FFC1D7CCFF6CFA0594EE8F12" box="[159,1466,1521,1542]" gridrow="0" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFA05909C8F12" box="[159,456,1521,1542]" gridcol="0" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FF6CFA0590678F12" authority="Mean SD" authorityName="Mean SD" box="[159,307,1521,1542]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FF6CFA0590678F12" box="[159,307,1521,1542]" italics="true" pageId="26" pageNumber="582">Operculinoides</emphasis>
</taxonomicName>
</th>
<th id="EB225DD9FFC1D7CCFDA2FA0593CB8F12" box="[593,671,1521,1542]" gridcol="1" gridrow="0" pageId="26" pageNumber="582">Mean</th>
<th id="EB225DD9FFC1D7CCFCDAFA0592D78F12" box="[809,899,1521,1542]" gridcol="2" gridrow="0" pageId="26" pageNumber="582">SD</th>
<th id="EB225DD9FFC1D7CCFBFFFA0595EA8F12" box="[1036,1214,1521,1542]" gridcol="3" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FBFFFA0595EA8F12" box="[1036,1214,1521,1542]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FBFFFA0595EA8F12" box="[1036,1214,1521,1542]" italics="true" pageId="26" pageNumber="582">Palaeonummulites</emphasis>
</taxonomicName>
</th>
<th id="EB225DD9FFC1D7CCFABBFA0594EE8F12" box="[1352,1466,1521,1542]" gridcol="4" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FABBFA0594EE8F12" authority="First" authorityName="First" box="[1352,1466,1521,1542]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FABBFA0594EE8F12" box="[1352,1466,1521,1542]" italics="true" pageId="26" pageNumber="582">Nummulites</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF9D694EE8F2C" box="[159,1466,1570,1592]" gridrow="1" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF9D6909C8F2C" box="[159,456,1570,1592]" gridcol="0" gridrow="1" pageId="26" pageNumber="582">First chamber length</th>
<td id="EB225DD9FFC1D7CCFDA2F9D693CB8F2C" box="[593,671,1570,1592]" gridcol="1" gridrow="1" pageId="26" pageNumber="582">369.0</td>
<td id="EB225DD9FFC1D7CCFCDAF9D692D78F2C" box="[809,899,1570,1592]" gridcol="2" gridrow="1" pageId="26" pageNumber="582">165.69</td>
<td id="EB225DD9FFC1D7CCFBFFF9D695EA8F2C" box="[1036,1214,1570,1592]" gridcol="3" gridrow="1" pageId="26" pageNumber="582">++</td>
<td id="EB225DD9FFC1D7CCFABBF9D694EE8F2C" box="[1352,1466,1570,1592]" gridcol="4" gridrow="1" pageId="26" pageNumber="582">0</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF9B694EE8F4C" box="[159,1466,1602,1624]" gridrow="2" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF9B6909C8F4C" box="[159,456,1602,1624]" gridcol="0" gridrow="2" pageId="26" pageNumber="582">Proloculus nominal diameter</th>
<td id="EB225DD9FFC1D7CCFDA2F9B693CB8F4C" box="[593,671,1602,1624]" gridcol="1" gridrow="2" pageId="26" pageNumber="582">146.3</td>
<td id="EB225DD9FFC1D7CCFCDAF9B692D78F4C" box="[809,899,1602,1624]" gridcol="2" gridrow="2" pageId="26" pageNumber="582">67.08</td>
<td id="EB225DD9FFC1D7CCFBFFF9B695EA8F4C" box="[1036,1214,1602,1624]" gridcol="3" gridrow="2" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFABBF9B694EE8F4C" box="[1352,1466,1602,1624]" gridcol="4" gridrow="2" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF99694EE8F6C" box="[159,1466,1634,1656]" gridrow="3" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF996909C8F6C" box="[159,456,1634,1656]" gridcol="0" gridrow="3" pageId="26" pageNumber="582">Deuteroloculus ratio</th>
<td id="EB225DD9FFC1D7CCFDA2F99693CB8F6C" box="[593,671,1634,1656]" gridcol="1" gridrow="3" pageId="26" pageNumber="582">1.166</td>
<td id="EB225DD9FFC1D7CCFCDAF99692D78F6C" box="[809,899,1634,1656]" gridcol="2" gridrow="3" pageId="26" pageNumber="582">0.1684</td>
<td id="EB225DD9FFC1D7CCFBFFF99695EA8F6C" box="[1036,1214,1634,1656]" gridcol="3" gridrow="3" pageId="26" pageNumber="582">++</td>
<td id="EB225DD9FFC1D7CCFABBF99694EE8F6C" box="[1352,1466,1634,1656]" gridcol="4" gridrow="3" pageId="26" pageNumber="582">++</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF97694EE8F8C" box="[159,1466,1666,1688]" gridrow="4" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF976909C8F8C" box="[159,456,1666,1688]" gridcol="0" gridrow="4" pageId="26" pageNumber="582">Initial marginal radius</th>
<td id="EB225DD9FFC1D7CCFDA2F97693CB8F8C" box="[593,671,1666,1688]" gridcol="1" gridrow="4" pageId="26" pageNumber="582">192.0</td>
<td id="EB225DD9FFC1D7CCFCDAF97692D78F8C" box="[809,899,1666,1688]" gridcol="2" gridrow="4" pageId="26" pageNumber="582">88.59</td>
<td id="EB225DD9FFC1D7CCFBFFF97695EA8F8C" box="[1036,1214,1666,1688]" gridcol="3" gridrow="4" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFABBF97694EE8F8C" box="[1352,1466,1666,1688]" gridcol="4" gridrow="4" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF95594EE8FA3" box="[159,1466,1697,1719]" gridrow="5" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF955909C8FA3" box="[159,456,1697,1719]" gridcol="0" gridrow="5" pageId="26" pageNumber="582">Marginal radius increase</th>
<td id="EB225DD9FFC1D7CCFDA2F95593CB8FA3" box="[593,671,1697,1719]" gridcol="1" gridrow="5" pageId="26" pageNumber="582">0.126</td>
<td id="EB225DD9FFC1D7CCFCDAF95592D78FA3" box="[809,899,1697,1719]" gridcol="2" gridrow="5" pageId="26" pageNumber="582">0.0191</td>
<td id="EB225DD9FFC1D7CCFBFFF95595EA8FA3" box="[1036,1214,1697,1719]" gridcol="3" gridrow="5" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFABBF95594EE8FA3" box="[1352,1466,1697,1719]" gridcol="4" gridrow="5" pageId="26" pageNumber="582">++</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF93594EE8FC3" box="[159,1466,1729,1751]" gridrow="6" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF935909C8FC3" box="[159,456,1729,1751]" gridcol="0" gridrow="6" pageId="26" pageNumber="582">Spiral chamber height increase</th>
<td id="EB225DD9FFC1D7CCFDA2F93593CB8FC3" box="[593,671,1729,1751]" gridcol="1" gridrow="6" pageId="26" pageNumber="582">2.8</td>
<td id="EB225DD9FFC1D7CCFCDAF93592D78FC3" box="[809,899,1729,1751]" gridcol="2" gridrow="6" pageId="26" pageNumber="582">1.15</td>
<td id="EB225DD9FFC1D7CCFBFFF93595EA8FC3" box="[1036,1214,1729,1751]" gridcol="3" gridrow="6" pageId="26" pageNumber="582">++</td>
<td id="EB225DD9FFC1D7CCFABBF93594EE8FC3" box="[1352,1466,1729,1751]" gridcol="4" gridrow="6" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF91594EE8FE3" box="[159,1466,1761,1783]" gridrow="7" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF915909C8FE3" box="[159,456,1761,1783]" gridcol="0" gridrow="7" pageId="26" pageNumber="582">Initial spiral chamber height</th>
<td id="EB225DD9FFC1D7CCFDA2F91593CB8FE3" box="[593,671,1761,1783]" gridcol="1" gridrow="7" pageId="26" pageNumber="582">62.5</td>
<td id="EB225DD9FFC1D7CCFCDAF91592D78FE3" box="[809,899,1761,1783]" gridcol="2" gridrow="7" pageId="26" pageNumber="582">32.84</td>
<td id="EB225DD9FFC1D7CCFBFFF91595EA8FE3" box="[1036,1214,1761,1783]" gridcol="3" gridrow="7" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFABBF91594EE8FE3" box="[1352,1466,1761,1783]" gridcol="4" gridrow="7" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF8F594EE8E03" box="[159,1466,1793,1815]" gridrow="8" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF8F5909C8E03" box="[159,456,1793,1815]" gridcol="0" gridrow="8" pageId="26" pageNumber="582">Backbend angle</th>
<td id="EB225DD9FFC1D7CCFDA2F8F593CB8E03" box="[593,671,1793,1815]" gridcol="1" gridrow="8" pageId="26" pageNumber="582">0.636</td>
<td id="EB225DD9FFC1D7CCFCDAF8F592D78E03" box="[809,899,1793,1815]" gridcol="2" gridrow="8" pageId="26" pageNumber="582">0.1120</td>
<td id="EB225DD9FFC1D7CCFBFFF8F595EA8E03" box="[1036,1214,1793,1815]" gridcol="3" gridrow="8" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFABBF8F594EE8E03" box="[1352,1466,1793,1815]" gridcol="4" gridrow="8" pageId="26" pageNumber="582">++</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF8D594EE8E23" box="[159,1466,1825,1847]" gridrow="9" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF8D5909C8E23" box="[159,456,1825,1847]" gridcol="0" gridrow="9" pageId="26" pageNumber="582">Initial chamber length</th>
<td id="EB225DD9FFC1D7CCFDA2F8D593CB8E23" box="[593,671,1825,1847]" gridcol="1" gridrow="9" pageId="26" pageNumber="582">224.0</td>
<td id="EB225DD9FFC1D7CCFCDAF8D592D78E23" box="[809,899,1825,1847]" gridcol="2" gridrow="9" pageId="26" pageNumber="582">126.60</td>
<td id="EB225DD9FFC1D7CCFBFFF8D595EA8E23" box="[1036,1214,1825,1847]" gridcol="3" gridrow="9" pageId="26" pageNumber="582">++</td>
<td id="EB225DD9FFC1D7CCFABBF8D594EE8E23" box="[1352,1466,1825,1847]" gridcol="4" gridrow="9" pageId="26" pageNumber="582">0</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF8B594EE8E43" box="[159,1466,1857,1879]" gridrow="10" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF8B5909C8E43" box="[159,456,1857,1879]" gridcol="0" gridrow="10" pageId="26" pageNumber="582">Chamber length increase</th>
<td id="EB225DD9FFC1D7CCFDA2F8B593CB8E43" box="[593,671,1857,1879]" gridcol="1" gridrow="10" pageId="26" pageNumber="582">0.048</td>
<td id="EB225DD9FFC1D7CCFCDAF8B592D78E43" box="[809,899,1857,1879]" gridcol="2" gridrow="10" pageId="26" pageNumber="582">0.0220</td>
<td id="EB225DD9FFC1D7CCFBFFF8B595EA8E43" box="[1036,1214,1857,1879]" gridcol="3" gridrow="10" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFABBF8B594EE8E43" box="[1352,1466,1857,1879]" gridcol="4" gridrow="10" pageId="26" pageNumber="582">++</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CF89594EE8E63" box="[159,1466,1889,1911]" gridrow="11" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CF895909C8E63" box="[159,456,1889,1911]" gridcol="0" gridrow="11" pageId="26" pageNumber="582">Perimeter ratio</th>
<td id="EB225DD9FFC1D7CCFDA2F89593CB8E63" box="[593,671,1889,1911]" gridcol="1" gridrow="11" pageId="26" pageNumber="582">1.392</td>
<td id="EB225DD9FFC1D7CCFCDAF89592D78E63" box="[809,899,1889,1911]" gridcol="2" gridrow="11" pageId="26" pageNumber="582">0.1130</td>
<td id="EB225DD9FFC1D7CCFBFFF89595EA8E63" box="[1036,1214,1889,1911]" gridcol="3" gridrow="11" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFABBF89594EE8E63" box="[1352,1466,1889,1911]" gridcol="4" gridrow="11" pageId="26" pageNumber="582">++</td>
</tr>
</table>
</paragraph>
<subSubSection id="5ED9656CFFC02828FF6CFF4990958BA7" pageId="27" pageNumber="583" type="discussion">
<paragraph id="167C36E7FFC02828FF6CFF4990958BA7" blockId="27.[159,782,189,691]" pageId="27" pageNumber="583">
<emphasis id="24B7EAF5FFC02828FF6CFF49905A89C0" bold="true" box="[159,270,189,212]" pageId="27" pageNumber="583">Remarks.</emphasis>
<bibRefCitation id="72524B16FFC02828FEEFFF4990AC89C0" author="Eames, F. E. &amp; Banner, F. T. &amp; Blow, W. H. &amp; Clarke, W. J." box="[284,504,189,212]" pageId="27" pageNumber="583" refId="ref20674" refString="Eames, F. E., Banner, F. T., Blow, W. H. &amp; Clarke, W. J. 1962. Fundamentals of mid-Tertiary stratigraphical correlation. Cambrige University Press, Cambridge, vii + 151 pp." type="book" year="1962">
Eames
<emphasis id="24B7EAF5FFC02828FE9DFF4990F489C0" box="[366,416,189,212]" italics="true" pageId="27" pageNumber="583">et al</emphasis>
. (1962)
</bibRefCitation>
included
<taxonomicName id="D1C34D64FFC02828FD9AFF49901189E0" authority="Hanzawa, 1935" authorityName="Hanzawa" authorityYear="1935" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC02828FD9AFF49925989C0" box="[617,781,189,212]" italics="true" pageId="27" pageNumber="583">Operculinoides</emphasis>
<bibRefCitation id="72524B16FFC02828FF6CFF29901189E0" author="Hanzawa, S." box="[159,325,221,244]" pageId="27" pageNumber="583" pagination="1 - 29" refId="ref21083" refString="Hanzawa, S. 1935. Some fossil Operculina and Miogypsina from Japan and their stratigraphical significance. Science Reports of the Tohoku Imperial University. 2 nd Series, Geology 18, 1 - 29." type="journal article" year="1935">Hanzawa, 1935</bibRefCitation>
</taxonomicName>
as a synonym of
<taxonomicName id="D1C34D64FFC02828FDF0FF29939D89E0" box="[515,713,221,244]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC02828FDF0FF29939D89E0" box="[515,713,221,244]" italics="true" pageId="27" pageNumber="583">Palaeonummulites</emphasis>
</taxonomicName>
based on the
<typeStatus id="C9788845FFC02828FF05FF0990708800" box="[246,292,253,276]" pageId="27" pageNumber="583">type</typeStatus>
species
<taxonomicName id="D1C34D64FFC02828FE7EFF0993E28800" baseAuthorityName="Heilprin" baseAuthorityYear="1882" box="[397,694,253,276]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="species" species="willcoxi">
<emphasis id="24B7EAF5FFC02828FE7EFF0993E28800" box="[397,694,253,276]" italics="true" pageId="27" pageNumber="583">Palaeonummulites willcoxi</emphasis>
</taxonomicName>
with a tight coil producing chambers one and half times higher than long, i.e. almost square. This was followed by
<bibRefCitation id="72524B16FFC02828FF6CFEA990168860" author="Haynes, J. R." box="[159,322,349,372]" pageId="27" pageNumber="583" pagination="215 - 247" refId="ref21168" refString="Haynes, J. R. 1988. Massively cordate nummulitids (Foraminifera) at the Palaeocene / Eocene boundary in Tethys. Indian Journal of Geology 60, 215 - 247." type="journal article" year="1988">Haynes (1988)</bibRefCitation>
,
<bibRefCitation id="72524B16FFC02828FEA0FEA993D58860" author="Robinson, E. &amp; Wright, R. M." box="[339,641,349,372]" pageId="27" pageNumber="583" refId="ref22720" refString="Robinson, E. &amp; Wright, R. M. 1993. Jamaican Paleogene larger foraminifera. Pp. 283 - 345 in R. M. Wright &amp; E. Robinson (eds) Biostratigraphy of Jamaica. Geological Society of American Memoir, Boulder." type="book" year="1993">Robinson &amp; Wright (1993)</bibRefCitation>
and
<bibRefCitation id="72524B16FFC02828FD4EFEA9907C8880" author="Haynes, J. R. &amp; Racey, A. &amp; Whittaker, J. E." pageId="27" pageNumber="583" refId="ref21201" refString="Haynes, J. R., Racey, A. &amp; Whittaker, J. E. 2010. A revision of the Early Palaeogene nummulitids (Foraminifera) from northern Oman, with implications for their classification. Pp. 29 - 89 in J. E. Whittaker &amp; M. B. Hart (eds) Micropalaeontology, sedimentary environments and stratigraphy: a tribute to Dennis Curry (1912 - 2001). Micropalaeontological Society Special Publication, Geological Society, London." type="book" year="2010">
Haynes
<emphasis id="24B7EAF5FFC02828FF6CFE8991848880" box="[159,208,381,404]" italics="true" pageId="27" pageNumber="583">et al</emphasis>
. (2010)
</bibRefCitation>
. However, the other
<taxonomicName id="D1C34D64FFC02828FDFEFE89925D8880" box="[525,777,381,404]" pageId="27" pageNumber="583">
<emphasis id="24B7EAF5FFC02828FDFEFE8993E58880" box="[525,689,381,404]" italics="true" pageId="27" pageNumber="583">Operculinoides</emphasis>
species
</taxonomicName>
, such as the
<taxonomicName id="D1C34D64FFC02828FED7FE6990E288A0" baseAuthorityName="Heilprin" baseAuthorityYear="1885" box="[292,438,413,436]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC02828FED7FE6990E288A0" box="[292,438,413,436]" italics="true" pageId="27" pageNumber="583">O. floridensis</emphasis>
</taxonomicName>
group with a clear operculinid lax coiling and gradational involution, cannot be considered
<taxonomicName id="D1C34D64FFC02828FF25FE2890C888E7" box="[214,412,476,499]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC02828FF25FE2890C888E7" box="[214,412,476,499]" italics="true" pageId="27" pageNumber="583">Palaeonummulites</emphasis>
</taxonomicName>
. It would be necessary to change the
<typeStatus id="C9788845FFC02828FF3EFE0891AF8B07" box="[205,251,508,531]" pageId="27" pageNumber="583">type</typeStatus>
species of
<taxonomicName id="D1C34D64FFC02828FE70FE0893738B07" authorityName="Hanzawa" authorityYear="1935" box="[387,551,508,531]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC02828FE70FE0893738B07" box="[387,551,508,531]" italics="true" pageId="27" pageNumber="583">Operculinoides</emphasis>
</taxonomicName>
to distinguish these forms generically. The species
<taxonomicName id="D1C34D64FFC02828FE07FDE893D18B27" baseAuthorityName="Heilprin" baseAuthorityYear="1885" box="[500,645,540,563]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC02828FE07FDE893D18B27" box="[500,645,540,563]" italics="true" pageId="27" pageNumber="583">O. floridensis</emphasis>
</taxonomicName>
seems to be the best candidate, as has already been suggested by
<bibRefCitation id="72524B16FFC02828FD2DFDC8907D8B67" author="Butterlin, J." pageId="27" pageNumber="583" refId="ref19901" refString="Butterlin, J. 1981. Claves para la determinacion´de macroforamin´iferos de Mexico´y del Caribe, del Cretacico´Superior al Mioceno Medio. Instituto Mexicano del Petroleo, Mexico, 219 pp." type="book" year="1981">Butterlin (1981)</bibRefCitation>
, because the variability of the coiling mode encompasses characteristics of
<taxonomicName id="D1C34D64FFC02828FE04FD8893E98B87" box="[503,701,636,659]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC02828FE04FD8893E98B87" box="[503,701,636,659]" italics="true" pageId="27" pageNumber="583">Palaeonummulites</emphasis>
</taxonomicName>
,
<taxonomicName id="D1C34D64FFC02828FD3DFD8890598BA7" authorityName="Hanzawa" authorityYear="1935" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC02828FD3DFD8890598BA7" italics="true" pageId="27" pageNumber="583">Operculinoides</emphasis>
</taxonomicName>
and
<taxonomicName id="D1C34D64FFC02828FEB2FD6890EE8BA7" authorityName="d'Orbigny" authorityYear="1826" box="[321,442,668,691]" class="Globothalamea" family="Nummulitidae" genus="Operculina" kingdom="Chromista" order="Rotaliida" pageId="27" pageNumber="583" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC02828FEB2FD6890EE8BA7" box="[321,442,668,691]" italics="true" pageId="27" pageNumber="583">Operculina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>