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<document id="50DFE72A7A4AFD138B1880F84B9CA119" ID-DOI="10.5252/geodiversitas2019v41a1" ID-GBIF-Dataset="ed35d9a8-8d9f-4c2a-a43e-cd1f5ca7e064" ID-ISSN="1638-9395" ID-Zenodo-Dep="3678724" ID-ZooBank="urn:lsid:zoobank.org:pub:86084BFE-6ECC-4F8B-9B38-D2C203483CA7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582313579726" checkinUser="valdenar" docAuthor="Esu, Daniela &amp; Girotti, Odoardo" docDate="2019" docId="9954879CE760560FFEF140C5FD22FBF6" docLanguage="en" docName="GEODIVERSITAS.41.1.1-9.pdf.imf" docOrigin="Geodiversitas 41 (1)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Hydrobia ionica Esu &amp; Girotti, 2019, n. sp." docType="treatment" docUuid="8D0FEDB0-E195-4827-A3EF-C2D25A445D1B" docUuidSource="ZooBank" docVersion="11" lastPageNumber="7" masterDocId="656DFFE4E7675607FFC14676FFDCFFA7" masterDocTitle="Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="6" updateTime="1698721032456" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="C78CB930EFAC86297F25BC4C29E905E8">Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy)</mods:title>
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<mods:namePart id="989AA7BA6315FEBEF552A3E36CEAC9C8">Esu, Daniela</mods:namePart>
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<mods:namePart id="BDC5A86B0A39FB2498E52D5160059A08">Girotti, Odoardo</mods:namePart>
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<mods:title id="3311F2BA4FD21B1A5EB6AB509DEB0AC4">Geodiversitas</mods:title>
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<mods:date id="C67688EE768CAD24ED8AA8E0AFA7029A">2019</mods:date>
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<mods:number id="DA0D6E3585E4706B665B3CB634BADF36">2019-01-10</mods:number>
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<mods:number id="DAF8B81E0B1838414C30172F1B0BB1CB">41</mods:number>
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<treatment id="9954879CE760560FFEF140C5FD22FBF6" ID-DOI="http://doi.org/10.5281/zenodo.3682305" ID-GBIF-Taxon="162247367" ID-Zenodo-Dep="3682305" LSID="urn:lsid:zoobank.org:act:8D0FEDB0-E195-4827-A3EF-C2D25A445D1B" httpUri="http://treatment.plazi.org/id/9954879CE760560FFEF140C5FD22FBF6" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<subSubSection id="59E76501E7605600FEF140C5FD87F968" box="[304,603,1715,1743]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="1142368AE7605600FEF140C5FD87F968" blockId="7.[304,603,1715,1774]" box="[304,603,1715,1743]" pageId="7" pageNumber="6">
<heading id="4A0A81E6E7605600FEF140C5FD87F968" box="[304,603,1715,1743]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="6" reason="2">
<taxonomicName id="D6FD4D09E7605600FEF140C5FDC4F969" ID-CoL="3N5H3" authorityName="Hartmann" authorityYear="1821" box="[304,536,1715,1743]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="ionica" status="sp. nov.">
<emphasis id="2389EA98E7605600FEF140C5FE44F969" bold="true" box="[304,408,1715,1742]" italics="true" pageId="7" pageNumber="6">Hydrobia</emphasis>
(s.l.)
<emphasis id="2389EA98E7605600FE1740C2FDC4F969" bold="true" box="[470,536,1716,1742]" italics="true" pageId="7" pageNumber="6">ionica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38BA57E3E7605600FDE140C3FD87F968" box="[544,603,1717,1743]" pageId="7" pageNumber="6">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59E76501E7605600FE9D40A2FDF3F949" box="[348,559,1748,1774]" pageId="7" pageNumber="6" type="description">
<paragraph id="1142368AE7605600FE9D40A2FDF3F949" blockId="7.[304,603,1715,1774]" box="[348,559,1748,1774]" pageId="7" pageNumber="6">
<heading id="4A0A81E6E7605600FE9D40A2FDF3F949" box="[348,559,1748,1774]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="6" reason="5">
(
<figureCitation id="89C62A0FE7605600FEA240A2FE03F949" box="[355,479,1748,1774]" captionStart="FIG" captionStartId="5.[132,143,1471,1489]" captionTargetBox="[146,1442,215,1424]" captionTargetId="figure@5.[146,1442,214,1424]" captionTargetPageId="5" captionText="FIG. 2. — A, B, Hydrobia (s.l.) galatoniana n. sp., Galatone (Lecce, Apulia), Late Oligocene, paratype MPUR7-4198: A, apical view of the shell; B, detail of the protoconch; C, D, Hydrobia (s.l.) ionica n. sp., Galatone (Lecce, Apulia), Late Oligocene, paratype MPUR7-4199: C, apical view of the shell; D, detail of the protoconch. SEM image. Scale bars: A, C, 500 µm; B, D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3678726" httpUri="https://zenodo.org/record/3678726/files/figure.png" pageId="7" pageNumber="6">Figs 2C, D</figureCitation>
;
<figureCitation id="89C62A0FE7605600FE2A40A2FE25F949" box="[491,505,1748,1774]" captionStart="FIG" captionStartId="7.[132,143,1015,1033]" captionTargetBox="[132,1455,215,970]" captionTargetId="figure@7.[132,1455,214,970]" captionTargetPageId="7" captionText="FIG. 4. — A-C, Hydrobia (s.l.) ionica n. sp.,Galatone (Lecce,Apulia),Late Oligocene:Aa-Ac, holotype MPUR7-3899;Ba-Bc, paratype MPUR7-4197/2a; Ca-Cc, paratype MPUR7-4197/2b. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3678730" httpUri="https://zenodo.org/record/3678730/files/figure.png" pageId="7" pageNumber="6">4</figureCitation>
A-C)
</heading>
</paragraph>
</subSubSection>
<paragraph id="1142368AE7605600FF45408BFCDBF8B6" blockId="7.[132,775,1789,1809]" box="[132,775,1789,1809]" pageId="7" pageNumber="6">
<uri id="656C3A88E7605600FF45408BFCDBF8B6" box="[132,775,1789,1809]" pageId="7" pageNumber="6">
urn:lsid:zoobank.org:act:
<uuid id="655B0C5FE7605600FE9E408BFCDBF8B6" box="[351,775,1789,1809]" pageId="7" pageNumber="6">
8D0FEDB0-
<collectionCode id="77ECAE4FE7605600FE0A408BFE0BF8B6" box="[459,471,1789,1809]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="7" pageNumber="6" type="Herbarium">E</collectionCode>
195-4827-
<collectionCode id="77ECAE4FE7605600FDF9408BFD9BF8B6" box="[568,583,1789,1809]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="7" pageNumber="6" type="Herbarium">A</collectionCode>
3EF-C2D25
<collectionCode id="77ECAE4FE7605600FD75408BFD1FF8B6" box="[692,707,1789,1809]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="7" pageNumber="6" type="Herbarium">A</collectionCode>
445D1
<collectionCode id="77ECAE4FE7605600FD3B408BFCDBF8B6" box="[762,775,1789,1809]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="7" pageNumber="6" type="Herbarium">B</collectionCode>
</uuid>
</uri>
</paragraph>
<subSubSection id="59E76501E7605600FF45414DFD01F8F3" box="[132,733,1851,1876]" pageId="7" pageNumber="6" type="reference_group">
<paragraph id="1142368AE7605600FF45414DFD01F8F3" blockId="7.[132,733,1851,1876]" box="[132,733,1851,1876]" pageId="7" pageNumber="6">
<treatmentCitationGroup id="31ED11A4E7605600FF45414DFD01F8F3" box="[132,733,1851,1876]" pageId="7" pageNumber="6">
<taxonomicName id="D6FD4D09E7605600FF45414DFE89F8F3" authority="" authorityName="Brusina" authorityYear="1897" box="[132,341,1851,1876]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="6" phylum="Mollusca" rank="subGenus" subGenus="Staja">
<emphasis id="2389EA98E7605600FF45414DFED3F8F4" box="[132,271,1851,1875]" italics="true" pageId="7" pageNumber="6">Pseudamnicola</emphasis>
(
<emphasis id="2389EA98E7605600FEDF414AFE92F8F4" box="[286,334,1852,1875]" italics="true" pageId="7" pageNumber="6">Staja</emphasis>
)
</taxonomicName>
sp. –
<treatmentCitation id="905C109BE7605600FE51414AFD8AF8F3" author="ESU D. &amp; GIROTTI O. &amp; IANNONE A. &amp; PIGNATTI J. S. &amp; RICCHETTI G." box="[400,598,1851,1876]" page="189" pageId="7" pageNumber="6" year="1994">
<bibRefCitation id="756C4B7BE7605600FE51414AFD8AF8F3" author="Esu" box="[400,598,1851,1876]" etAl="et al." firstAuthor="Esu" pageId="7" pageNumber="6" pagination="183 - 195" refId="ref6499" refString="ESU D., GIROTTI O., IANNONE A., PIGNATTI J. S. &amp; RICCHETTI G. 1994. - Lagoonal-continental Oligocene of southern Apulia (Italy). Bollettino della Societa Paleontologica Italiana 33: 183 - 195." type="journal article" year="1994">
Esu
<emphasis id="2389EA98E7605600FE7B414AFE35F8F4" box="[442,489,1851,1875]" italics="true" pageId="7" pageNumber="6">et al.</emphasis>
1994: 189
</bibRefCitation>
</treatmentCitation>
, pl. 2, fig. 4a.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="59E76501E7605600FF45411BFB62FB5A" pageId="7" pageNumber="6" type="materials_examined">
<paragraph id="1142368AE7605600FF45411BFD26F807" blockId="7.[132,775,1901,1953]" pageId="7" pageNumber="6">
<collectionCode id="77ECAE4FE7605600FF45411BFECEF824" box="[132,274,1901,1925]" pageId="7" pageNumber="6">TYPE MATERIAL</collectionCode>
. —
<materialsCitation id="A1953CD7E7605600FEFB4118FED7F807" ID-GBIF-Occurrence="2571181192" collectionCode="A" pageId="7" pageNumber="6" specimenCode="MPUR7-3899" specimenCount="1" typeStatus="holotype">
<typeStatus id="CE468828E7605600FEFB4118FE4BF821" box="[314,407,1902,1926]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
:
<specimenCode id="415B9EF1E7605600FE5F4118FDF7F821" box="[414,555,1902,1926]" pageId="7" pageNumber="6">MPUR7-3899</specimenCode>
(
<figureCitation id="89C62A0FE7605600FDF64118FDA2F821" box="[567,638,1902,1926]" captionStart="FIG" captionStartId="7.[132,143,1015,1033]" captionTargetBox="[132,1455,215,970]" captionTargetId="figure@7.[132,1455,214,970]" captionTargetPageId="7" captionText="FIG. 4. — A-C, Hydrobia (s.l.) ionica n. sp.,Galatone (Lecce,Apulia),Late Oligocene:Aa-Ac, holotype MPUR7-3899;Ba-Bc, paratype MPUR7-4197/2a; Ca-Cc, paratype MPUR7-4197/2b. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3678730" httpUri="https://zenodo.org/record/3678730/files/figure.png" pageId="7" pageNumber="6">
Fig. 4
<collectionCode id="77ECAE4FE7605600FDAA4118FDA2F821" box="[619,638,1902,1926]" pageId="7" pageNumber="6">A</collectionCode>
</figureCitation>
), H =
<quantity id="D6059B6FE7605600FD77411BFCDFF822" box="[694,771,1901,1925]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="6" unit="mm" value="4.5">4.5 mm</quantity>
, W =
<quantity id="D6059B6FE7605600FF7741FEFEDBF807" box="[182,263,1928,1952]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="6" unit="mm" value="2.5">2.5 mm</quantity>
.
</materialsCitation>
—
<materialsCitation id="A1953CD7E7605600FEF141FFFD26F807" ID-GBIF-Occurrence="2571181191" box="[304,762,1928,1953]" collectionCode="MPUR" pageId="7" pageNumber="6" specimenCode="MPUR7" specimenCount="1" typeStatus="paratype">
<typeStatus id="CE468828E7605600FEF141FFFE53F806" box="[304,399,1929,1953]" pageId="7" pageNumber="6" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="415B9EF1E7605600FE5941FEFE2DF807" box="[408,497,1928,1952]" pageId="7" pageNumber="6">MPUR7</specimenCode>
-4197/2, 4199/1, 3900/40.
</materialsCitation>
</paragraph>
<paragraph id="1142368AE7605600FF4541CCFE49F84B" blockId="7.[132,775,1978,2029]" pageId="7" pageNumber="6">
<collectionCode id="77ECAE4FE7605600FF4541CCFE89F877" box="[132,341,1978,2002]" pageId="7" pageNumber="6">DERIVATION OF NAME</collectionCode>
. — Named after the Ionian Sea, whose shoreline is near the
<typeStatus id="CE468828E7605600FED841A3FE9FF84A" box="[281,323,2005,2029]" pageId="7" pageNumber="6">type</typeStatus>
locality.
</paragraph>
<paragraph id="1142368AE7605600FCEC4226FB63FB24" blockId="7.[813,1456,1104,1155]" pageId="7" pageNumber="6">
<collectionCode id="77ECAE4FE7605600FCEC4226FC62FBC1" box="[813,958,1104,1128]" pageId="7" pageNumber="6">
<typeStatus id="CE468828E7605600FCEC4226FC81FBC2" box="[813,861,1104,1128]" pageId="7" pageNumber="6">TYPE</typeStatus>
HORIZON
</collectionCode>
. — Limy mudstone layers of Galatone Fm at Galatone (
<bibRefCitation id="756C4B7BE7605600FCA7421AFC24FB24" author="Esu" box="[870,1016,1131,1155]" etAl="et al." firstAuthor="Esu" pageId="7" pageNumber="6" pagination="183 - 195" refId="ref6499" refString="ESU D., GIROTTI O., IANNONE A., PIGNATTI J. S. &amp; RICCHETTI G. 1994. - Lagoonal-continental Oligocene of southern Apulia (Italy). Bollettino della Societa Paleontologica Italiana 33: 183 - 195." type="journal article" year="1994">
Esu
<emphasis id="2389EA98E7605600FC50421AFC1CFB24" box="[913,960,1131,1155]" italics="true" pageId="7" pageNumber="6">et al.</emphasis>
1994
</bibRefCitation>
;
<bibRefCitation id="756C4B7BE7605600FBC4421DFB6DFB24" author="Bossio" box="[1029,1201,1131,1155]" etAl="et al." firstAuthor="Bossio" pageId="7" pageNumber="6" pagination="151 - 156" refId="ref5979" refString="BOSSIO A., ESU D., FORESI L. M., GIROTTI O., IANNONE A., LUPERTO E., MARGIOTTA S., MAZZEI R., MONTEFORTI B., RICCHETTI G. &amp; SALVATORINI G. 1998. - Formazione di Galatone, nuovo nome per un'unita litostratigrafica del Salento (Puglia, Italia meridionale). Atti della Societa Toscana di Scenze Naturali, Memorie S. A 105: 151 - 156." type="journal article" year="1998">
Bossio
<emphasis id="2389EA98E7605600FB8B421AFBA5FB24" box="[1098,1145,1131,1155]" italics="true" pageId="7" pageNumber="6">et al.</emphasis>
1998
</bibRefCitation>
).
</paragraph>
<paragraph id="1142368AE7605600FCEC42ECFC1DFB6A" blockId="7.[813,1455,1178,1229]" pageId="7" pageNumber="6">
<collectionCode id="77ECAE4FE7605600FCEC42ECFC62FB17" box="[813,958,1178,1202]" pageId="7" pageNumber="6">
<typeStatus id="CE468828E7605600FCEC42ECFC81FB17" box="[813,861,1178,1202]" pageId="7" pageNumber="6">TYPE</typeStatus>
LOCALITY
</collectionCode>
. —
<materialsCitation id="A1953CD7E7605600FC2842EDFC1DFB6A" ID-GBIF-Occurrence="2571181195" country="Italy" latitude="40.140842" location="Apulia" longLatPrecision="1" longitude="18.050102" pageId="7" pageNumber="6" specimenCount="1" stateProvince="Apulia">
Galatone (Lecce,
<collectingRegion id="D339F868E7605600FB5042EDFB08FB14" box="[1169,1236,1179,1203]" country="Italy" name="Puglia" pageId="7" pageNumber="6">Apulia</collectingRegion>
,
<collectingCountry id="69EA761AE7605600FB1C42EDFAD6FB14" box="[1245,1290,1179,1203]" name="Italy" pageId="7" pageNumber="6">Italy</collectingCountry>
) (
<geoCoordinate id="74C9504DE7605600FADC42EDFA77FB14" box="[1309,1451,1179,1203]" degrees="40" direction="north" minutes="8" orientation="latitude" pageId="7" pageNumber="6" precision="1" seconds="27.03" value="40.140842">40°8’27.03”N</geoCoordinate>
,
<geoCoordinate id="74C9504DE7605600FCEC42C3FC6AFB6A" box="[813,950,1205,1229]" degrees="18" direction="east" minutes="03" orientation="longitude" pageId="7" pageNumber="6" precision="1" seconds="0.37" value="18.050102">18°03’0.37”E</geoCoordinate>
).
</materialsCitation>
</paragraph>
<paragraph id="1142368AE7605600FCEC4292FB62FB5A" blockId="7.[813,1214,1252,1277]" box="[813,1214,1252,1277]" pageId="7" pageNumber="6">
<collectionCode id="77ECAE4FE7605600FCEC4292FC55FB5D" box="[813,905,1252,1276]" pageId="7" pageNumber="6">
<typeStatus id="CE468828E7605600FCEC4292FC81FB5D" box="[813,861,1252,1276]" pageId="7" pageNumber="6">TYPE</typeStatus>
AGE
</collectionCode>
. — Chattian (Late Oligocene).
</paragraph>
</subSubSection>
<subSubSection id="59E76501E7605600FCEC4362FA94FADB" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="1142368AE7605600FCEC4362FA94FADB" blockId="7.[813,1456,1300,1404]" pageId="7" pageNumber="6">DIAGNOSIS. — Small-sized shell, ovately conical, with up to 5 slightly convex whorls, the last large and elongate. Aperture pear-shaped, relatively high and elongate, adapically barely angulate, somewhat flared and slightly thickened inside. Umbilicus slit-like.</paragraph>
</subSubSection>
<subSubSection id="59E76501E760560FFCEC43C2FD22FBF6" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6" type="description">
<paragraph id="1142368AE7605600FCEC43C2FC61FA6A" blockId="7.[813,1456,1460,2030]" box="[813,957,1460,1486]" pageId="7" pageNumber="6">DESCRIPTION</paragraph>
<paragraph id="1142368AE7605600FCEC43A3FB2DF84A" blockId="7.[813,1456,1460,2030]" pageId="7" pageNumber="6">
Shell small, solid, thick-walled, ovately conical, with up to 5 slightly convex whorls, the last large, elongate, exceeding two-thirds of shell height. Early whorls slowly increasing, the last faster and somewhat descending (
<figureCitation id="89C62A0FE7605600FB264042FAE2F9E8" box="[1255,1342,1588,1615]" captionStart="FIG" captionStartId="5.[132,143,1471,1489]" captionTargetBox="[146,1442,215,1424]" captionTargetId="figure@5.[146,1442,214,1424]" captionTargetPageId="5" captionText="FIG. 2. — A, B, Hydrobia (s.l.) galatoniana n. sp., Galatone (Lecce, Apulia), Late Oligocene, paratype MPUR7-4198: A, apical view of the shell; B, detail of the protoconch; C, D, Hydrobia (s.l.) ionica n. sp., Galatone (Lecce, Apulia), Late Oligocene, paratype MPUR7-4199: C, apical view of the shell; D, detail of the protoconch. SEM image. Scale bars: A, C, 500 µm; B, D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3678726" httpUri="https://zenodo.org/record/3678726/files/figure.png" pageId="7" pageNumber="6">Figs 2C</figureCitation>
;
<figureCitation id="89C62A0FE7605600FA884043FA8BF9E8" box="[1353,1367,1589,1615]" captionStart="FIG" captionStartId="7.[132,143,1015,1033]" captionTargetBox="[132,1455,215,970]" captionTargetId="figure@7.[132,1455,214,970]" captionTargetPageId="7" captionText="FIG. 4. — A-C, Hydrobia (s.l.) ionica n. sp.,Galatone (Lecce,Apulia),Late Oligocene:Aa-Ac, holotype MPUR7-3899;Ba-Bc, paratype MPUR7-4197/2a; Ca-Cc, paratype MPUR7-4197/2b. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3678730" httpUri="https://zenodo.org/record/3678730/files/figure.png" pageId="7" pageNumber="6">4</figureCitation>
Ab, Ba). Suture shallow, oblique between the penultimate and ultimate whorl. Protoconch of
<emphasis id="2389EA98E7605600FB9A4003FBB0F929" box="[1115,1132,1653,1678]" italics="true" pageId="7" pageNumber="6">c.</emphasis>
<quantity id="D6059B6FE7605600FBB24002FB16F928" box="[1139,1226,1652,1679]" metricMagnitude="-1" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="6" unit="mm" value="410.0">410 µm</quantity>
maximum diameter, smooth, consisting of one slightly convex whorl. Boundary to teleoconch distinct by the formation of closely spaced growth lines (
<figureCitation id="89C62A0FE7605600FC7E40A2FBC9F949" box="[959,1045,1748,1774]" captionStart="FIG" captionStartId="5.[132,143,1471,1489]" captionTargetBox="[146,1442,215,1424]" captionTargetId="figure@5.[146,1442,214,1424]" captionTargetPageId="5" captionText="FIG. 2. — A, B, Hydrobia (s.l.) galatoniana n. sp., Galatone (Lecce, Apulia), Late Oligocene, paratype MPUR7-4198: A, apical view of the shell; B, detail of the protoconch; C, D, Hydrobia (s.l.) ionica n. sp., Galatone (Lecce, Apulia), Late Oligocene, paratype MPUR7-4199: C, apical view of the shell; D, detail of the protoconch. SEM image. Scale bars: A, C, 500 µm; B, D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3678726" httpUri="https://zenodo.org/record/3678726/files/figure.png" pageId="7" pageNumber="6">Fig. 2D</figureCitation>
). Teleoconch glossy, with very delicate and thin prosocline growth lines. An axial swelling occurs sometimes in the last whorl towards the aperture. Aperture pear-shaped, high, elongate, adapically barely angulate and abapically rounded, somewhat flared and slightly thickened inside; peristome continuous, columellar lip almost completely adherent to the last whorl, outer lip straight in lateral profile. Umbilicus slit-like. Range dimensions: H =
<quantity id="D6059B6FE7605600FAB541C5FC81F849" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="7" pageNumber="6" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4-4.5 mm</quantity>
, W =
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, H/W = 1.48-1.8.
</paragraph>
<paragraph id="1142368AE76F560FFF45468FFD22FBF6" blockId="8.[132,775,217,1105]" pageId="8" pageNumber="7">
Provisionally,
<bibRefCitation id="756C4B7BE76F560FFEDF468CFE0BFEB3" author="Esu" box="[286,471,249,276]" etAl="et al." firstAuthor="Esu" pageId="8" pageNumber="7" pagination="183 - 195" refId="ref6499" refString="ESU D., GIROTTI O., IANNONE A., PIGNATTI J. S. &amp; RICCHETTI G. 1994. - Lagoonal-continental Oligocene of southern Apulia (Italy). Bollettino della Societa Paleontologica Italiana 33: 183 - 195." type="journal article" year="1994">
Esu
<emphasis id="2389EA98E76F560FFE8E468CFE58FEB4" box="[335,388,249,275]" italics="true" pageId="8" pageNumber="7">et al.</emphasis>
(1994)
</bibRefCitation>
assigned this stout form to the subgenus
<taxonomicName id="D6FD4D09E76F560FFED4476FFD47FE94" authority="Brusina, 1897" authorityName="Brusina" authorityYear="1897" box="[277,667,281,308]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="7" phylum="Mollusca" rank="subGenus" subGenus="Staja">
<emphasis id="2389EA98E76F560FFED4476FFE6CFE94" box="[277,432,281,307]" italics="true" pageId="8" pageNumber="7">Pseudamnicola</emphasis>
(
<emphasis id="2389EA98E76F560FFE7F476FFE2FFE94" box="[446,499,281,307]" italics="true" pageId="8" pageNumber="7">Staja</emphasis>
)
<bibRefCitation id="756C4B7BE76F560FFDC0476FFD47FE94" author="Brusina" box="[513,667,281,308]" firstAuthor="Brusina" pageId="8" pageNumber="7" pagination="1 - 43" refId="ref6173" refString="BRUSINA S. 1897. - Materiaux pour la faune malacologique neogene de la Dalmatie, de la Croatie et de la Slavonie avec des especes de la Bosnie, de l'Herzegovine et de la Serbie. Djela Jugoslavenske Akademije Znanosti i Umjetnosti 18: i-xxi, 1 - 43." type="book chapter" year="1897">Brusina, 1897</bibRefCitation>
</taxonomicName>
, currently considered a separate genus of the family
<taxonomicName id="D6FD4D09E76F560FFDFA474FFD18FEF4" authorityName="Stimpson" authorityYear="1865" box="[571,708,313,339]" class="Gastropoda" family="Hydrobiidae" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="7" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
(
<bibRefCitation id="756C4B7BE76F560FFD13474FFE8EFED3" author="Harzhauser" etAl="et al." firstAuthor="Harzhauser" pageId="8" pageNumber="7" pagination="1 - 53" refId="ref6847" refString="HARZHAUSER M., NEUBAUER T. A., GEORGOPOULOU E., ESU D., D'AMICO C., PAVIA G., GIUNTELLI P. &amp; CARNEVALE G. 2015. - Late Messinian continental and Lago-Mare gastropods from the Tertiary Piedmont Basin, NW Italy. Bollettino della Societa Paleontologica Italiana 54: 1 - 53. https: // doi. org / 10.4435 / BSPI. 2015.1" type="journal article" year="2015">
Harzhauser
<emphasis id="2389EA98E76F560FFF1C472FFECEFED4" box="[221,274,345,371]" italics="true" pageId="8" pageNumber="7">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="756C4B7BE76F560FFEA1472FFD98FED4" author="MolluscaBase" box="[352,580,345,371]" firstAuthor="MolluscaBase" pageId="8" pageNumber="7" refId="ref7497" refString="MOLLUSCABASE 2018 b. - Staja Brusina, 1897 †. Accessed through: World Register of Marine Species at: http: // www. marinespecies. org / aphia. php rp = taxdetails &amp; id = 820431 on 2018 - 05 - 03" type="url" year="2018" yearSuffix="b">MolluscaBase 2018b</bibRefCitation>
). Examination of further material allowed distinguishing this species from
<emphasis id="2389EA98E76F560FFD0C470FFCDCFE34" box="[717,768,377,403]" italics="true" pageId="8" pageNumber="7">Staja</emphasis>
, which is a Miocene Eastern European taxon rich in several species (cf.
<bibRefCitation id="756C4B7BE76F560FFF3647CEFE13FE75" author="MolluscaBase" box="[247,463,440,466]" firstAuthor="MolluscaBase" pageId="8" pageNumber="7" refId="ref7497" refString="MOLLUSCABASE 2018 b. - Staja Brusina, 1897 †. Accessed through: World Register of Marine Species at: http: // www. marinespecies. org / aphia. php rp = taxdetails &amp; id = 820431 on 2018 - 05 - 03" type="url" year="2018" yearSuffix="b">MolluscaBase 2018b</bibRefCitation>
), characterised by a very small size (dimensional range: H =
<quantity id="D6059B6FE76F560FFE0747AEFD91FE54" box="[454,589,472,499]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="8" pageNumber="7" unit="mm" value="2.05" valueMax="2.5" valueMin="1.6">1.6-2.5 mm</quantity>
, W =
<quantity id="D6059B6FE76F560FFD4E47AEFD23FE54" box="[655,767,472,499]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="8" pageNumber="7" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1-1.2 mm</quantity>
) and a mainly conical shell with regularly growing whorls (cf.
<bibRefCitation id="756C4B7BE76F560FFF45446EFEC6FD95" author="Brusina" box="[132,282,536,562]" firstAuthor="Brusina" pageId="8" pageNumber="7" pagination="1 - 43" refId="ref6173" refString="BRUSINA S. 1897. - Materiaux pour la faune malacologique neogene de la Dalmatie, de la Croatie et de la Slavonie avec des especes de la Bosnie, de l'Herzegovine et de la Serbie. Djela Jugoslavenske Akademije Znanosti i Umjetnosti 18: i-xxi, 1 - 43." type="book chapter" year="1897">Brusina 1897</bibRefCitation>
;
<bibRefCitation id="756C4B7BE76F560FFEE8446EFE62FD95" author="Jekelius" box="[297,446,536,562]" firstAuthor="Jekelius" pageId="8" pageNumber="7" pagination="1 - 167" refId="ref7089" refString="JEKELIUS E. 1944. - Sarmat und Pont von Soceni. Memoriile Institutului Geologic al Romaniei 5: 1 - 167." type="journal article" year="1944">Jekelius 1944</bibRefCitation>
). The assignment of
<emphasis id="2389EA98E76F560FFD68446FFD35FD95" box="[681,745,537,562]" italics="true" pageId="8" pageNumber="7">ionica</emphasis>
n. sp. to the same genus as
<emphasis id="2389EA98E76F560FFE45444EFDDCFDF5" box="[388,512,568,594]" italics="true" pageId="8" pageNumber="7">galatoniana</emphasis>
n. sp. is likely supported by similar conchological characters between the two species, such as hydrobioid shell shape with slightly convex whorls, last whorl higher, ovate-elongate and barely flared aperture, and smooth protoconch.
<taxonomicName id="D6FD4D09E76F560FFE5B44C1FE22FD76" authorityName="Hartmann" authorityYear="1821" box="[410,510,695,721]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="2389EA98E76F560FFE5B44C1FE22FD76" box="[410,510,695,721]" italics="true" pageId="8" pageNumber="7">Hydrobia</emphasis>
</taxonomicName>
(s.l.)
<emphasis id="2389EA98E76F560FFDFA44CEFDA7FD76" box="[571,635,696,721]" italics="true" pageId="8" pageNumber="7">ionica</emphasis>
n. sp. differs from
<emphasis id="2389EA98E76F560FFF0244AEFF3CFD56" box="[195,224,728,753]" italics="true" pageId="8" pageNumber="7">H.</emphasis>
(s.l.)
<emphasis id="2389EA98E76F560FFEE344A1FE43FD56" box="[290,415,727,753]" italics="true" pageId="8" pageNumber="7">galatoniana</emphasis>
n. sp. in having a more globose shell, a wider apical angle, a smaller protoconch, the last whorl proportionally wider and higher, the aperture more elongate, and a smaller H/W (max) ratio (1.8 vs 2.08). It differs from the coeval
<emphasis id="2389EA98E76F560FFE45452EFD43FCD6" box="[388,671,855,881]" italics="true" pageId="8" pageNumber="7">H. dubuissonii hydruntina</emphasis>
from the Upper Oligocene marls of Otranto bauxite quarry (
<bibRefCitation id="756C4B7BE76F560FFD7E4501FED0FC16" author="Esu &amp; Girotti" firstAuthor="Esu" pageId="8" pageNumber="7" pagination="137 - 174" refId="ref6441" refString="ESU D. &amp; GIROTTI O. 2010. - The Late Oligocene molluscan fauna from Otranto (Apulia, southern Italy): an example of alternating freshwater, lagoonal and emerged environments. Palaeontology 53: 137 - 174. https: // doi. org / 10.1111 / j. 1475 - 4983.2009.00923. x" type="journal article" year="2010">Esu &amp; Girotti 2010</bibRefCitation>
) in being conical-ovate in shape, in having a less acute spire, a proportionally higher last whorl and wider and more elongate aperture.
<emphasis id="2389EA98E76F560FFE5545A0FDA4FC57" box="[404,632,982,1008]" italics="true" pageId="8" pageNumber="7">
<taxonomicName id="D6FD4D09E76F560FFE5545A0FE24FC57" authorityName="Hartmann" authorityYear="1821" box="[404,504,982,1008]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="7" phylum="Mollusca" rank="genus">Hydrobia</taxonomicName>
dubuissonii
</emphasis>
from Oligocene deposits of
<collectingCountry id="69EA761AE76F560FFEF34581FEA6FBB6" box="[306,378,1015,1041]" name="France" pageId="8" pageNumber="7">France</collectingCountry>
and
<collectingCountry id="69EA761AE76F560FFE6F4580FDD6FBB7" box="[430,522,1014,1040]" name="Belgium" pageId="8" pageNumber="7">Belgium</collectingCountry>
(see above) differs from
<emphasis id="2389EA98E76F560FFF454261FF7EFB97" box="[132,162,1047,1072]" italics="true" pageId="8" pageNumber="7">H.</emphasis>
(s.l.)
<emphasis id="2389EA98E76F560FFF224261FEFFFB97" box="[227,291,1047,1072]" italics="true" pageId="8" pageNumber="7">ionica</emphasis>
n. sp. in having a conical-pyramidal shape, a more acute spire and a smaller and more ovate aperture.
</paragraph>
</subSubSection>
</treatment>
</document>
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