214 lines
21 KiB
XML
214 lines
21 KiB
XML
<document id="D4518BBCCA6933F96DC115F2AC88C71B" ID-DOI="10.3897/zookeys.1215.128795" ID-ZooBank="11120B98-7E4D-4E1B-980A-2975059B1C9C" ID-publisher-id="128795" URI-arpha="F77C60F0-518A-5C9B-9098-123E707CCFB9" URI-zoobank="http://zoobank.org/11120B98-7E4D-4E1B-980A-2975059B1C9C" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1729177022173" checkinUser="pensoft" docAuthor="Hurtado-Gómez, Juan Pablo, Daza, Juan M., Vargas-Ramírez, Mario, Deepak, V. & Fritz, Uwe" docDate="2024" docId="9389EAF2ACA4510F8CF8D7A44C8602D4" docLanguage="en" docName="ZooKeys 1215: 335-358" docOrigin="ZooKeys 1215" docSource="https://zookeys.pensoft.net/article/128795/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Hydrops (Tachynectes) Fitzinger 1843" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="358" masterDocId="F77C60F0518A5C9B9098123E707CCFB9" masterDocTitle="Phylogeography of the Colombian water snake Helicops danieli Amaral, 1938 (Reptilia, Squamata, Dipsadidae) with comments on the systematics and evolution of the genus Helicops Wagler, 1828" masterLastPageNumber="358" masterPageNumber="335" pageNumber="335" updateTime="1729186367996" updateUser="admin">
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<mods:titleInfo id="4F218EB0FB3470393DCFF4E4AF9D7BCA">
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<mods:title id="0626EAF55A47197113D8092723286DE3">Phylogeography of the Colombian water snake Helicops danieli Amaral, 1938 (Reptilia, Squamata, Dipsadidae) with comments on the systematics and evolution of the genus Helicops Wagler, 1828</mods:title>
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<mods:namePart id="B87237A7410DD030153DA1056EE40875">Hurtado-Gómez, Juan Pablo</mods:namePart>
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<mods:nameIdentifier id="7812F60723023986B304158A203762C7" type="ORCID">0000-0003-4351-2834</mods:nameIdentifier>
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<mods:affiliation id="9D302662FEB7D8D3F0E3476788A20ACB">Museum of Zoology, Senckenberg Natural History Collections Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="C226A2D86C03506E21902A8CEF1DAD35">Daza, Juan M.</mods:namePart>
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<mods:affiliation id="C780150E653883B16CBB9FF01A94DE37">Grupo Herpetológico de Antioquia (GHA), Instituto de Biología, Universidad de Antioquia, Medellín, Colombia</mods:affiliation>
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<mods:namePart id="4550AEDE9E717BA5AABBCEA4687FC84F">Vargas-Ramírez, Mario</mods:namePart>
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<mods:nameIdentifier id="321898E5F529F73DF672935B3081C322" type="ORCID">0000-0001-8974-3430</mods:nameIdentifier>
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<mods:affiliation id="75B4FAAB377C11728CC604EADFA0D3DC">Grupo Biodiversidad y Conservación Genética, Instituto de Genética, Universidad Nacional de Colombia, Bogotá, Colombia & Estación de Biología Tropical Roberto Franco (EBTRF), Universidad Nacional de Colombia, Villavicencio, Colombia</mods:affiliation>
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<mods:namePart id="B9DC271297C1FB6EE37BE50FD7646075">Deepak, V.</mods:namePart>
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<mods:nameIdentifier id="509DC78F809955A0A45C95005A919546" type="ORCID">0000-0002-8826-9367</mods:nameIdentifier>
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<mods:affiliation id="3FD36CC454F221D4CA9211BCF5384F50">Museum of Zoology, Senckenberg Natural History Collections Dresden, A. B. Meyer Building, 01109 Dresden, Germany & School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
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<mods:namePart id="49E3B0D9752286E0FB218A6219251DCC">Fritz, Uwe</mods:namePart>
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<mods:affiliation id="1C5AFACBDD869CFCFF9B471D561B69A2">Museum of Zoology, Senckenberg Natural History Collections Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:title id="3D7860AE3C0024A71D6DE4C403209F44">ZooKeys</mods:title>
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<mods:part id="976896C5EC91EE8C93EA29F0AB7FA3C5">
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<mods:date id="B50E7F65E46B7F781A9AF27605AA2FD3">2024</mods:date>
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<mods:number id="C508D2BF0BFF139D56BF0C65CB2438A6">2024-10-17</mods:number>
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<mods:number id="01C403BA49E8F499A58BCCB1634A8AD1">1215</mods:number>
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<mods:start id="E5C73B18D85D45638273254AECF92FAD">335</mods:start>
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<mods:classification id="E93BC55F58997273B1F5886A97BB2315">journal article</mods:classification>
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<mods:identifier id="48D7E13FE8B9CF5D82C10146479ED5D7" type="DOI">10.3897/zookeys.1215.128795</mods:identifier>
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<mods:identifier id="DDB1A8A31850AA2D06151098B5EDBD1A" type="ZooBank">11120B98-7E4D-4E1B-980A-2975059B1C9C</mods:identifier>
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<treatment id="9389EAF2ACA4510F8CF8D7A44C8602D4" ID-DOI="http://doi.org/10.5281/zenodo.13946718" ID-Zenodo-Dep="13946718" ID-arpha="9389EAF2-ACA4-510F-8CF8-D7A44C8602D4" LSID="urn:lsid:plazi:treatment:9389EAF2ACA4510F8CF8D7A44C8602D4" httpUri="http://treatment.plazi.org/id/9389EAF2ACA4510F8CF8D7A44C8602D4">
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<subSubSection id="D3CCD35E9505B67FBD83CC0F39DB4FEF" type="nomenclature">
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<paragraph id="7EC444F0BEBB63E71403E552C6F916B9">
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<heading id="A7529FD29F34CFE1B7DEADC12604A5FB" reason="label">Subgenus:</heading>
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<taxonomicName id="B9FD6DA97ADBDC2293A01B4B5E219313" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" class="Squamata" family="Dipsadidae" genus="Hydrops" kingdom="Animalia" phylum="Chordata" rank="subGenus" subGenus="Tachynectes">
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<emphasis id="55CB8A11B80883C6511C3C5984BB98DE" italics="true">Tachynectes</emphasis>
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<bibRefCitation id="12B1C8BB46771A62CE4AB3C62A114E25" DOI="10.5962/bhl.title.4694" author="Fitzinger LJ" journalOrPublisher="Braumüller et Seidel, Vindobona [Vienna]" refId="B17" refString="Fitzinger LJ (1843) Systema reptilium. Fasciculus primus. Amblyglossae. Braumuller et Seidel, Vindobona [Vienna], 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" title="Systema reptilium. Fasciculus primus. Amblyglossae." year="1843">Fitzinger, 1843</bibRefCitation>
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</taxonomicName>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EWTDI" type="type_taxon">
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<paragraph id="AEF12B6F6FB1E798A04439ACEA8D8EE0">
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<heading id="D113FDB0BEDE172211264F2CE98E3847" reason="title">Type species.</heading>
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||
</paragraph>
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<paragraph id="380CC6BA52E49CDC92ADA54298A4DAF1">
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<taxonomicName id="EACF59C530FABE0B664292ACF18EA0B8" authority="Schlegel, 1837" authorityName="Schlegel" authorityYear="1837" class="Squamata" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Cyprinodontiformes" phylum="Chordata" rank="species" species="leopardina">
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<emphasis id="BD7DB641AACDB4798AB6402EE76E417D" italics="true">Homalopsis leopardina</emphasis>
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Schlegel, 1837
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="SECID0EHUDI" type="diagnosis">
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<paragraph id="F7412FB60070F5A0BEFB487C4D360B30">
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<heading id="27ADBC5F10FF4167BABC7177538C4E74" reason="title">Diagnosis.</heading>
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</paragraph>
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<paragraph id="3488C8EF844E19C5D9BD779823677DAE">
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Members of the
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<emphasis id="40C0293EC6BD5574721D93A95258B938" italics="true">
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subgenus
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<taxonomicName id="C39F393C642D878736FF24BA66F808E2" rank="subGenus" subGenus="Tachynectes">Tachynectes</taxonomicName>
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</emphasis>
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have short hemipenial lobes decorated with spinules. The hemipenial body is homogeneously covered with spinules (Fig.
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<figureCitation id="99F9CE916AC8FCD3BF161ED750159FC1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemipenes of A H. leopardinus (based on UFMTR 1504 from Moraes-da-Silva et al. 2021) and B H. danieli (based on MHUAR 15565) in sulcate (left) and asulcate (right) views. Scale bars: 10 mm. The hemipenis of H. leopardinus represents the morphology with short lobes characteristic for the subgenus Tachynectes; the hemipenis of H. danieli represents the morphology with long lobes characteristic for the subgenus Helicops. Note that the hemipenis is homogeneously covered with spinules in (A), whereas large spines are confined to the hemipenial body (i. e., excluding the lobes) on the sulcate surface and the lateral regions in (B). Drawings: V. Deepak." figureDoi="10.3897/zookeys.1215.128795.figure4" httpUri="https://binary.pensoft.net/fig/1155736">4 A</figureCitation>
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), occasionally a few enlarged spines may occur, e. g., in
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<taxonomicName id="AA12AE78210E47497B07CDA4FACB11BA" authorityName="Moraes-Da-Silva, Cecilia Amaro, Sales-Nunes, Rodrigues & Curcio" authorityYear="2021" class="Squamata" family="Colubridae" genus="Helicops" kingdom="Animalia" phylum="Chordata" rank="species" species="phantasma" subGenus="Tachynectes">
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<emphasis id="042371D54C89D6E6AC31A7D19A78623A" italics="true">H.</emphasis>
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(
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||
<emphasis id="6C8EEB59B29E10162BDBC95B8BAF9C48" italics="true">T.</emphasis>
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||
)
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||
<emphasis id="D2F5C62E67ED0DD8060BDDAA4D7A7421" italics="true">phantasma</emphasis>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EIVDI" type="Content">
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||
<paragraph id="E4048849F95F3FEAD7216332AE19CEA5">
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<heading id="E32A69854FAEB19AA2152C375EB83D52" reason="title">Content.</heading>
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||
</paragraph>
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<paragraph id="999CF129726F330A285886BC9B4817A8">
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Five species,
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<taxonomicName id="5E9D4AC92EE7C09731818ABF3E4A150B" authority="Jan, 1865" authorityName="Jan" authorityYear="1865" class="Squamata" family="Colubridae" genus="Helicops" kingdom="Animalia" phylum="Chordata" rank="species" species="infrataeniatus" subGenus="Tachynectes">
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<emphasis id="BB3E62CEABAE896F6D0266E3B5717F6C" italics="true">Helicops</emphasis>
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(
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<emphasis id="B75DA28648E62D28A8BE1F422B438943" italics="true">Tachynectes</emphasis>
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)
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<emphasis id="570591AF1CE83F5C041C7C51FD274059" italics="true">infrataeniatus</emphasis>
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Jan, 1865
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</taxonomicName>
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,
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<taxonomicName id="1F22D13E54482CD3E8B1B629295BB929" authority="(Schlegel, 1837)" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Squamata" family="Colubridae" genus="Helicops" kingdom="Animalia" phylum="Chordata" rank="species" species="leopardinus" subGenus="Tachynectes">
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<emphasis id="03EFADA4FCD30AF6630003D9FEA2950E" italics="true">H.</emphasis>
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(
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<emphasis id="0A0A12EBD8D55B3BD7AD51E0603C7A2A" italics="true">T.</emphasis>
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)
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<emphasis id="FEF3108184FDBD3D2943A30EEE5EB9BF" italics="true">leopardinus</emphasis>
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(Schlegel, 1837)
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</taxonomicName>
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,
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<taxonomicName id="CA9673EDAE60B0B33AC35141D58D1410" authority="Gunther, 1861" authorityName="Gunther" authorityYear="1861" class="Squamata" family="Colubridae" genus="Helicops" kingdom="Animalia" phylum="Chordata" rank="species" species="modestus" subGenus="Tachynectes">
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<emphasis id="3D2287A9D62E9DC73E10876FDA8F9F54" italics="true">H.</emphasis>
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(
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<emphasis id="88E85359666009D63B62875E37CF8103" italics="true">T.</emphasis>
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)
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<emphasis id="D427A2032B1AF7A0B376B023695B1119" italics="true">modestus</emphasis>
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Günther, 1861
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</taxonomicName>
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,
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<taxonomicName id="97A33FDF8BB86744F1B5466A77891E5A" authorityName="Moraes-Da-Silva, Cecilia Amaro, Sales-Nunes, Rodrigues & Curcio" authorityYear="2021" class="Squamata" family="Colubridae" genus="Helicops" kingdom="Animalia" phylum="Chordata" rank="species" species="phantasma" subGenus="Tachynectes">
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<emphasis id="676C895520E7CA11CF1A7060F62A02C2" italics="true">H.</emphasis>
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(
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<emphasis id="EBE638E4F52053E821438A4E2CA59139" italics="true">T.</emphasis>
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)
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<emphasis id="0D2C14067425EB904BBDAB8BF4A5A534" italics="true">phantasma</emphasis>
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</taxonomicName>
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<bibRefCitation id="16AFC908BB6A5D8EE7567D50A018B2B9" DOI="10.11646/zootaxa.4903.2.3" author="Moraes-da-Silva" etAl="et al." firstAuthor="Moraes-da-Silva" journalOrPublisher="Zootaxa" pagination="151-193" refId="B41" refString="Moraes-da-Silva A, Amaro RC, Nunes PMS, Rodrigues MT, Curcio FF (2021) Long known, brand new, and possibly threatened: A new species of watersnake of the genus Helicops Wagler, 1828 (Serpentes; Xenodontinae) from the Tocantins-Araguaia River Basin, Brazil. Zootaxa 4903: 151–193. https://doi.org/10.11646/zootaxa.4903.2.3" title="Long known, brand new, and possibly threatened: A new species of watersnake of the genus Helicops Wagler, 1828 (Serpentes; Xenodontinae) from the Tocantins-Araguaia River Basin, Brazil." volume="4903" year="2021">Moraes-da-Silva et al., 2021</bibRefCitation>
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||
,
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<taxonomicName id="E4C9FF39A00028B8CD524418962C5F70" authority="da Frota, 2005" authorityName="da Frota" authorityYear="2005" class="Squamata" family="Colubridae" genus="Helicops" kingdom="Animalia" phylum="Chordata" rank="species" species="tapajonicus" subGenus="Tachynectes">
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||
<emphasis id="C4FC1C2F32621F44EF3F7DEC723796C8" italics="true">H.</emphasis>
|
||
(
|
||
<emphasis id="7E7CECB4565512001B41F82D70AA76D8" italics="true">T.</emphasis>
|
||
)
|
||
<emphasis id="92D8EE326E85F90B5B10376AD1B09EF3" italics="true">tapajonicus</emphasis>
|
||
<bibRefCitation id="FF099A71D699412BE0020F64A0C6E6A8" DOI="10.11606/issn.2316-9079.v4i1p61-67" author="da Frota JG" journalOrPublisher="Phyllomedusa" pagination="61-68" refId="B12" refString="da Frota JG (2005) Nova especie de Helicops Wagler, 1830 (Serpentes, Colubridae) do rio Tapajos, Amazonia, Brasil. Phyllomedusa 4: 61 - 68. https: // doi. org / 10.11606 / issn. 2316 - 9079. v 4 i 1 p 61 - 67" title="Nova espécie de Helicops Wagler, 1830 (Serpentes, Colubridae) do rio Tapajós, Amazônia, Brasil." volume="4" year="2005">da Frota, 2005</bibRefCitation>
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||
</taxonomicName>
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.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EYXDI" type="comments">
|
||
<paragraph id="4578B6040471A66538622DB6BD82B0C2">
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||
<heading id="43E3A18B1CDB067C35B46223A58B1998" reason="title">Comments.</heading>
|
||
</paragraph>
|
||
<paragraph id="607A19AE8F7E934EC4217D202FFC9E54">
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||
<taxonomicName id="4CF505A96BA73272294C3A025B0207E7" authority="von der Mark, 1863" authorityName="von der Mark" authorityYear="1863" class="Squamata" family="Colubridae" genus="Tachynectes" kingdom="Animalia" phylum="Chordata" rank="genus">
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||
<emphasis id="A157A8619A1590608612ADBFAEB5C8B7" italics="true">Tachynectes</emphasis>
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von der Mark, 1863
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</taxonomicName>
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, erected for a genus of fossil fishes, is a primary junior homonym of
|
||
<taxonomicName id="5B155AB33788FBB4F7939DD47A43D90F" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" class="Squamata" family="Colubridae" genus="Tachynectes" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2AA1DFC4E68539F38F9D4833336BFA6" italics="true">Tachynectes</emphasis>
|
||
<bibRefCitation id="B16618D400C8F7DEF802BD8DEAE0FD82" DOI="10.5962/bhl.title.4694" author="Fitzinger LJ" journalOrPublisher="Braumüller et Seidel, Vindobona [Vienna]" refId="B17" refString="Fitzinger LJ (1843) Systema reptilium. Fasciculus primus. Amblyglossae. Braumuller et Seidel, Vindobona [Vienna], 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" title="Systema reptilium. Fasciculus primus. Amblyglossae." year="1843">Fitzinger, 1843</bibRefCitation>
|
||
</taxonomicName>
|
||
. As
|
||
<taxonomicName id="FD44B9D7635B3F68F2FAA38555C4E4CE" authority="von der Mark, 1863" authorityName="von der Mark" authorityYear="1863" class="Squamata" family="Colubridae" genus="Tachynectes" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||
<emphasis id="96DE3160D6964A947F62102A060D26FD" italics="true">Tachynectes</emphasis>
|
||
von der Mark, 1863
|
||
</taxonomicName>
|
||
has only been used four times in the past 50 years according to our searches (Google Scholar, Zoological Record:
|
||
<bibRefCitation id="C7101A752C9A00B6183893C76FAF5D26" author="Sepkoski" firstAuthor="Sepkoski" journalOrPublisher="Bulletin of American Paleontology" pagination="1-500" refId="B64" refString="Sepkoski JJ (2002) A compendium of fossil marine animal genera. Bulletin of American Paleontology 363: 1–500." title="A compendium of fossil marine animal genera." volume="363" year="2002">Sepkoski 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="276B07842BB1ED7E030DD3E166F2D84A" DOI="10.1111/j.1463-6395.2008.00364.x" author="Albert" etAl="et al." firstAuthor="Albert" journalOrPublisher="Acta Zoologica" pagination="357-384" refId="B1" refString="Albert JS, Johnson DM, Knouft JH (2009) Fossils provide better estimates of ancestral body size than do extant taxa in fishes. Acta Zoologica 90: 357–384. https://doi.org/10.1111/j.1463-6395.2008.00364.x" title="Fossils provide better estimates of ancestral body size than do extant taxa in fishes." volume="90" year="2009">Albert et al. 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="246021879426670126ED3B549C152743" DOI="10.1016/j.cretres.2008.11.001" author="Dietze" firstAuthor="Dietze" journalOrPublisher="Cretaceous Research" pagination="559-574" refId="B14" refString="Dietze K (2009) Morphology and phylogenetic relationships of certain neoteleostean fishes from the Upper Cretaceous of Sendenhorst, Germany. Cretaceous Research 30: 559–574. https://doi.org/10.1016/j.cretres.2008.11.001" title="Morphology and phylogenetic relationships of certain neoteleostean fishes from the Upper Cretaceous of Sendenhorst, Germany." volume="30" year="2009">Dietze 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="955D118D10DEDD19705E25F5E2D95C77" DOI="10.1017/pab.2021.2" author="Schwarzhans and Carnevale" firstAuthor="Schwarzhans" journalOrPublisher="Paleobiology" pagination="446-463" refId="B63" refString="Schwarzhans W, Carnevale G (2021) The rise to dominance of lanternfishes (Teleostei: Myctophidae) in the oceanic ecosystems: A paleontological perspective. Paleobiology 47: 446–463. https://doi.org/10.1017/pab.2021.2" title="The rise to dominance of lanternfishes (Teleostei: Myctophidae) in the oceanic ecosystems: A paleontological perspective." volume="47" year="2021">Schwarzhans and Carnevale 2021</bibRefCitation>
|
||
), it fails to meet the criterion in Article 23.9. 2 of the International Code of Zoological Nomenclature (
|
||
<collectionCode id="77322478D97F74259F7AF4A92D4D621B">ICZN</collectionCode>
|
||
1999) for prevailing usage and is therefore unavailable. As a replacement name for
|
||
<taxonomicName id="D17B370B3251CC47AE32DDEB6D5D7021" authority="von der Mark, 1863" authorityName="von der Mark" authorityYear="1863" class="Squamata" family="Colubridae" genus="Tachynectes" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||
<emphasis id="C055D3F3B350F9AAEF1A602457C17CE4" italics="true">Tachynectes</emphasis>
|
||
von der Mark, 1863
|
||
</taxonomicName>
|
||
, we propose
|
||
<taxonomicName id="318BD605A055B5C29E88910DB45F876E" authorityName="Hurtado-Gómez & Daza & Vargas-Ramírez & Deepak & Fritz" authorityYear="2024" family="Myctophidae" genus="Ichthyotachynectes" kingdom="Animalia" order="Myctophiformes" phylum="Chordata" rank="genus" status="nom. nov.">
|
||
<emphasis id="B45E5C630E212CFD26A47C0913C04AEB" italics="true">Ichthyotachynectes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="21C333ADA0EE5EFE972D642DE4F98463">nom. nov.</taxonomicNameLabel>
|
||
to accommodate the fossil fish species previously assigned to
|
||
<taxonomicName id="DD7D74A69F0B15AAD2FF955936D1CBF3" authority="von der Mark, 1863" authorityName="von der Mark" authorityYear="1863" class="Squamata" family="Colubridae" genus="Tachynectes" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||
<emphasis id="2989C69E0B3D9E92A6B011B97BA7FDDE" italics="true">Tachynectes</emphasis>
|
||
von der Mark, 1863
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |