treatments-xml/data/52/4B/87/524B879BFF9FFFA72D605345FC682A16.xml

<document id="675A562489C0159E7A550588463987F4" ID-DOI="10.5281/zenodo.12586908" ID-ISSN="2064-2474" ID-Zenodo-Dep="12586908" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719596761487" checkinUser="felipe" docAuthor="Papp, L." docDate="2004" docId="524B879BFF9FFFA72D605345FC682A16" docLanguage="en" docName="ActaZoolAcadSciH.50.1.45-53.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 50 (1)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Immoderatus foldvarii Papp, 2004, sp. n." docType="treatment" docVersion="2" lastPageNumber="52" masterDocId="AE72FFE3FF9CFFA02F085728FFF32F51" masterDocTitle="Immoderatus Gen. N. Of Copromyzinae (Diptera: Sphaeroceridae)" masterLastPageNumber="53" masterPageNumber="45" pageNumber="48" updateTime="1719602575633" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="62F59FE82B43EE3772309DF05278B494" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B310EB00FCD20B6B4931F65180429874">
<mods:title id="1CBDA00B323EDAF5F29CEFE9648D02C3">Immoderatus Gen. N. Of Copromyzinae (Diptera: Sphaeroceridae)</mods:title>
</mods:titleInfo>
<mods:name id="7412EF2D73BA70186FEAF1F3DF404EE1" type="personal">
<mods:role id="81F2E1A24EFC72166ABC144F2FEE1E70">
<mods:roleTerm id="7B7A20FF60002E72367B37C9E5DD6F8E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E9C6C3F62D94890F557499341F0576C">Papp, L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="19E4F7BC5C45678192B248C0213AC4AC">text</mods:typeOfResource>
<mods:relatedItem id="BF73A29C4AEBD1F7C327E5F0CF5AF028" type="host">
<mods:titleInfo id="A5A3016A5F841FB940CEBE742FEC8E12">
<mods:title id="21E8BAD38E59071E085C458D869E6498">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="3CDC98BB68C4D455723A67A2C5C0A870">
<mods:date id="3891FD3391B5477EA0CCD47DB36062FE">2004</mods:date>
<mods:detail id="84881FFDCC576DF5E369A6F3654030F9" type="pubDate">
<mods:number id="F4FC969F985F8A3D5BC8D18F8C3FDBB4">2004-09-15</mods:number>
</mods:detail>
<mods:detail id="65522E348B9F456569E05114C7FFAA87" type="volume">
<mods:number id="47E10B8A1CE633F4208DA6F5C3E2CCE5">50</mods:number>
</mods:detail>
<mods:detail id="07635B30604F2F984B5B87222EA8E92C" type="issue">
<mods:number id="88BF987F4971262D5C63F5B8B647AF37">1</mods:number>
</mods:detail>
<mods:extent id="8E0997CC6436C5E2EF174A7411CD4939" unit="page">
<mods:start id="35D60D459929DD896B58B10ACB58A871">45</mods:start>
<mods:end id="544A3F317E905F57BA47C6ABEC83F6B9">53</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ED88EF8DEBA9B3A2ECA2E663E7C6E90A">journal article</mods:classification>
<mods:identifier id="AB534E3AFFD2706828A48F1F19B84A62" type="DOI">10.5281/zenodo.12586908</mods:identifier>
<mods:identifier id="C21FF242937499A233A80F780522D9BA" type="ISSN">2064-2474</mods:identifier>
<mods:identifier id="5BD36DE1150345B572264CA0FD5480F6" type="Zenodo-Dep">12586908</mods:identifier>
</mods:mods>
<treatment id="524B879BFF9FFFA72D605345FC682A16" LSID="urn:lsid:plazi:treatment:524B879BFF9FFFA72D605345FC682A16" httpUri="http://treatment.plazi.org/id/524B879BFF9FFFA72D605345FC682A16" lastPageId="7" lastPageNumber="52" pageId="3" pageNumber="48">
<subSubSection id="92F86506FF9FFFA32D605345FC392BD6" box="[616,970,1133,1159]" pageId="3" pageNumber="48" type="nomenclature">
<paragraph id="DA5D368DFF9FFFA32D605345FC392BD6" blockId="3.[616,970,1133,1159]" box="[616,970,1133,1159]" pageId="3" pageNumber="48">
<heading id="811581E1FF9FFFA32D605345FC392BD6" box="[616,970,1133,1159]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="48" reason="2">
<taxonomicName id="1DE24D0EFF9FFFA32D605345FC762BD6" ID-CoL="3PJ93" authorityName="Papp" authorityYear="2004" box="[616,901,1133,1159]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF9FFFA32D605345FC762BD6" bold="true" box="[616,901,1133,1159]" pageId="3" pageNumber="48">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF9FFFA32C845345FC392BD6" box="[908,970,1133,1159]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92F86506FF9FFFA72DDD53B8FC682A16" lastPageId="7" lastPageNumber="52" pageId="3" pageNumber="48" type="description">
<paragraph id="DA5D368DFF9FFFA32DDD53B8FCAE2BFB" blockId="3.[725,861,1168,1194]" box="[725,861,1168,1194]" pageId="3" pageNumber="48">
<heading id="811581E1FF9FFFA32DDD53B8FCAE2BFB" box="[725,861,1168,1194]" centered="true" fontSize="11" level="3" pageId="3" pageNumber="48" reason="3">
(
<figureCitation id="42D92A08FF9FFFA32DD553B8FCA72BFB" box="[733,852,1168,1194]" captionStart-0="Figs 1–3" captionStart-1="Figs 4–8" captionStart-2="Figs 9–15" captionStart-3="Figs 16–17" captionStartId-0="2.[317,360,1745,1767]" captionStartId-1="4.[317,360,1747,1769]" captionStartId-2="5.[317,360,1722,1744]" captionStartId-3="6.[317,360,1775,1797]" captionTargetBox-0="[357,1262,618,1706]" captionTargetBox-1="[342,1247,578,1717]" captionTargetBox-2="[346,1235,435,1687]" captionTargetBox-3="[321,1265,1159,1725]" captionTargetId-0="figure-88@2.[317,1269,614,1711]" captionTargetId-1="figure-74@4.[339,1247,576,1719]" captionTargetId-2="figure-11@5.[344,1243,424,1694]" captionTargetId-3="figure-362@6.[317,1269,1154,1726]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figs 1–3. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 1–2, 0.2 mm" captionText-1="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" captionText-2="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," captionText-3="Figs 16–17. Immoderatus foldvarii sp. n., paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.12586910" figureDoi-1="http://doi.org/10.5281/zenodo.12586912" figureDoi-2="http://doi.org/10.5281/zenodo.12586914" figureDoi-3="http://doi.org/10.5281/zenodo.12586916" httpUri-0="https://zenodo.org/record/12586910/files/figure.png" httpUri-1="https://zenodo.org/record/12586912/files/figure.png" httpUri-2="https://zenodo.org/record/12586914/files/figure.png" httpUri-3="https://zenodo.org/record/12586916/files/figure.png" pageId="3" pageNumber="48">Figs 1–17</figureCitation>
)
</heading>
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E7653FCFBB52A56" blockId="3.[317,1270,1236,1373]" pageId="3" pageNumber="48">
<materialsCitation id="6A8A3CD0FF9FFFA32E7653FCFBB52A56" collectingDate="2003-12-01" collectorName="Foldvari &amp; Peregovits &amp; Szappanos &amp; No" country="Thailand" location="Mae Ta Man" municipality="Mai" pageId="3" pageNumber="48" specimenCount="1" typeStatus="holotype">
<typeStatus id="0559882FFF9FFFA32E7653FCFE242BBB" box="[382,471,1236,1258]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
male: 
<collectingCountry id="A2F5761DFF9FFFA32D1353FCFD692BBB" box="[539,666,1236,1258]" name="Thailand" pageId="3" pageNumber="48">THAILAND</collectingCountry>
, 
<location id="DF3D6056FF9FFFA32DAC53FCFCD12BBB" LSID="urn:lsid:plazi:treatment:524B879BFF9FFFA72D605345FC682A16:DF3D6056FF9FFFA32DAC53FCFCD12BBB" box="[676,802,1236,1258]" country="Thailand" municipality="Mai" name="Mae Ta Man" pageId="3" pageNumber="48">Mae Ta Man</location>
elephant park, 
<quantity id="1D1A9B68FF9FFFA32CB253FCFC0B2BBB" box="[954,1016,1236,1258]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="48" unit="km" value="45.0">45 km</quantity>
N ofChiang 
<collectingMunicipality id="3A39ACF7FF9FFFA32B7F53FCFB532BBB" box="[1143,1184,1236,1258]" pageId="3" pageNumber="48">Mai</collectingMunicipality>
, 
<date id="AE5C104DFF9FFFA32BA353FCFE9E2A56" pageId="3" pageNumber="48" value="2003-12-01">
<collectingDate id="BE18E9A5FF9FFFA32BA353FCFE9E2A56" pageId="3" pageNumber="48" value="2003-12-01">01. 12. 2003</collectingDate>
</date>
– swept on elephant dung, leg. 
<collectorName id="7717535BFF9FFFA32DA053D9FD0F2A56" box="[680,764,1265,1287]" pageId="3" pageNumber="48">Földvári</collectorName>
, 
<collectorName id="7717535BFF9FFFA32C0F53D9FC9F2A56" box="[775,876,1265,1287]" pageId="3" pageNumber="48">Peregovits</collectorName>
&amp; 
<collectorName id="7717535BFF9FFFA32C8653D9FC042A56" box="[910,1015,1265,1287]" pageId="3" pageNumber="48">Szappanos</collectorName>
, 
<collectorName id="7717535BFF9FFFA32B0A53D9FBEC2A56" box="[1026,1055,1265,1287]" pageId="3" pageNumber="48">No</collectorName>
25.
</materialsCitation>
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E765226FE1B2A0C" blockId="3.[317,1270,1236,1373]" pageId="3" pageNumber="48">
<materialsCitation id="6A8A3CD0FF9FFFA32E765226FE172A0C" collectionCode="HNHM" pageId="3" pageNumber="48" specimenCount="37" specimenCount-female="21" specimenCount-male="16" typeStatus="paratype">
<typeStatus id="0559882FFF9FFFA32E765226FE2C2A75" box="[382,479,1294,1316]" pageId="3" pageNumber="48" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="CCE4FD04FF9FFFA32EEF5226FDCE2A75" box="[487,573,1294,1316]" count="11" pageId="3" pageNumber="48" type="male">11 males</specimenCount>
, 
<specimenCount id="CCE4FD04FF9FFFA32D4E5226FD5D2A75" box="[582,686,1294,1316]" count="17" pageId="3" pageNumber="48" type="female">17 females</specimenCount>
(
<collectionCode id="BCF3AE48FF9FFFA32DB05226FCF62A75" box="[696,773,1294,1316]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="3" pageNumber="48">HNHM</collectionCode>
, 
<specimenCount id="CCE4FD04FF9FFFA32C065226FCAB2A75" box="[782,856,1294,1316]" count="3" pageId="3" pageNumber="48" type="male">3 males</specimenCount>
, 
<specimenCount id="CCE4FD04FF9FFFA32C695226FC4E2A75" box="[865,957,1294,1316]" count="4" pageId="3" pageNumber="48" type="female">4 females</specimenCount>
in alcohol): same as for 
<typeStatus id="0559882FFF9FFFA32BAA5226FB052A75" box="[1186,1270,1294,1316]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
[genitalia of 
<specimenCount id="CCE4FD04FF9FFFA32EBA5202FE082A11" box="[434,507,1322,1344]" count="2" pageId="3" pageNumber="48" type="male">2 males</specimenCount>
and females each in a plastic microvial with glycerol, wings ofa specimen pre - pared on a slide]
</materialsCitation>
.
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E7652A9FD702A81" blockId="3.[317,1270,1409,1805]" pageId="3" pageNumber="48">
Measurements in mm: body length 0.77 (
<typeStatus id="0559882FFF9FFFA32C1B52A9FC9F2AC6" box="[787,876,1409,1431]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
), 0.72–0.85 (
<typeStatus id="0559882FFF9FFFA32CE452A9FBB32AC6" box="[1004,1088,1409,1431]" pageId="3" pageNumber="48" type="paratype">paratype</typeStatus>
males), 0.75–0.95 (
<typeStatus id="0559882FFF9FFFA32E4B52B6FE642AE5" box="[323,407,1438,1460]" pageId="3" pageNumber="48" type="paratype">paratype</typeStatus>
females), wing length 0.68 (
<typeStatus id="0559882FFF9FFFA32DD152B6FCC12AE5" box="[729,818,1438,1460]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
), 0.65–0.75, 0.66–0.77, wing breadth 0.25 (
<typeStatus id="0559882FFF9FFFA32E4C5292FE6E2A81" box="[324,413,1466,1488]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
), 0.24–0.33, 0.26–0.34.
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E7652FFFB352ABC" blockId="3.[317,1270,1409,1805]" box="[382,1222,1495,1517]" pageId="3" pageNumber="48">General colour dark to black; some parts ofbody with brownish grey microtomentum.</paragraph>
<paragraph id="DA5D368DFF9FFFA42E7652DCFC4D2D71" blockId="3.[317,1270,1409,1805]" lastBlockId="4.[317,1270,436,544]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
Head much longer than high. Head with frons broad, frons mostly dark microtomentose. Anterior part covered with adpressed small scales; ifall the large central area offrons, which includes ocelli, back to occiput, is interpreted as frontal (ocellar) triangle, this triangle is bordered by a densely hatched stripe (
<figureCitation id="42D92A08FF9FFFA32EDD5162FDFC2931" box="[469,527,1610,1632]" captionStart="Figs 1–3" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 1–3. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 1–2, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
). Face (prefrons) black, with extremely deep and strongly microtomentose antennal concavities. Interantennal tubercle small. Three pairs ofshort lateroclinate fronto-orbitals; a row ofmedioclinate inner orbitals. No ocellar pair, a number ofsmall postocellar setulae, postocellar setae indistinct. A pair oflong outer verticals. Presence or absence ofinner verticals difficult to inter - pret, as longer, paired setae between outer verticals absent. Eye much reduced, slightly oval. Scape and pedicel black. Pedicel with a 
<quantity id="1D1A9B68FF9FFFA32DAE51F2FCB729A1" box="[678,836,1754,1776]" metricMagnitude="-4" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="3" pageNumber="48" unit="mm" value="0.1075" valueMax="0.11" valueMin="0.105">0.105–0.11 mm</quantity>
long dorsal sub-basal seta (
<figureCitation id="42D92A08FF9FFFA32B6751F2FB4229A1" box="[1135,1201,1754,1776]" captionStart="Figs 1–3" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 1–3. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 1–2, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
). First flagellomere rounded, ca. 
<quantity id="1D1A9B68FF9FFFA32D3D51DFFD66285C" box="[565,661,1783,1805]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="48" unit="mm" value="0.06">0.060 mm</quantity>
long with dorsal arista. First flagellomere with 5 extremely long dorsal setae, not much shorter than arista (longest 
<quantity id="1D1A9B68FF98FFA42C50569CFBF52E9B" box="[856,1030,436,458]" metricMagnitude="-4" metricUnit="m" metricValue="1.125" metricValueMax="1.15" metricValueMin="1.1" pageId="4" pageNumber="49" unit="mm" value="0.1125" valueMax="0.115" valueMin="0.11">
<geoCoordinate id="BFD6504AFF98FFA42C50569CFC7D2E9B" box="[856,910,436,458]" degrees="0.110" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="55" value="0.11">0.110</geoCoordinate>
<geoCoordinate id="BFD6504AFF98FFA42C87569CFC232E9B" box="[911,976,436,458]" degrees="0.115" direction="west" orientation="longitude" pageId="4" pageNumber="49" precision="55" value="-0.115">–0.115</geoCoordinate>
mm
</quantity>
), in addition with 3–4 
<quantity id="1D1A9B68FF98FFA42E3556F9FE132EB6" box="[317,480,465,487]" metricMagnitude="-5" metricUnit="m" metricValue="4.1" metricValueMax="4.5" metricValueMin="3.7" pageId="4" pageNumber="49" unit="mm" value="0.040999999999999995" valueMax="0.045" valueMin="0.037">
<geoCoordinate id="BFD6504AFF98FFA42E3556F9FE802EB6" box="[317,371,465,487]" degrees="0.037" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="55" value="0.037">0.037</geoCoordinate>
<geoCoordinate id="BFD6504AFF98FFA42E7B56F9FE462EB6" box="[371,437,465,487]" degrees="0.045" direction="west" orientation="longitude" pageId="4" pageNumber="49" precision="55" value="-0.045">–0.045</geoCoordinate>
mm
</quantity>
long apical and ventral setae and with shorter cilia. Arista 
<quantity id="1D1A9B68FF98FFA42B1356F9FB4D2EB6" box="[1051,1214,465,487]" metricMagnitude="-4" metricUnit="m" metricValue="1.385" metricValueMax="1.42" metricValueMin="1.35" pageId="4" pageNumber="49" unit="mm" value="0.1385" valueMax="0.142" valueMin="0.135">
<geoCoordinate id="BFD6504AFF98FFA42B1356F9FBA32EB6" box="[1051,1104,465,487]" degrees="0.135" direction="north" orientation="latitude" pageId="4" pageNumber="49" precision="55" value="0.135">0.135</geoCoordinate>
<geoCoordinate id="BFD6504AFF98FFA42B5856F9FB612EB6" box="[1104,1170,465,487]" degrees="0.142" direction="west" orientation="longitude" pageId="4" pageNumber="49" precision="55" value="-0.142">–0.142</geoCoordinate>
mm
</quantity>
long, i.e. a little more than 2 times as long as 1st flagellomere, practically bare. Vibrissa rather long, about as long as antenna. Gena narrow. 5–6 peristomals. No genal seta.
</paragraph>
<caption id="8E9D6605FF98FFA42E3551FBFB062857" ID-DOI="http://doi.org/10.5281/zenodo.12586912" ID-Zenodo-Dep="12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="4" pageNumber="49" startId="4.[317,360,1747,1769]" targetBox="[342,1247,578,1717]" targetPageId="4" targetType="figure">
<paragraph id="DA5D368DFF98FFA42E3551FBFB062857" blockId="4.[317,1270,1747,1826]" pageId="4" pageNumber="49">
<emphasis id="E896EA9FFF98FFA42E3551FBFE6429B8" bold="true" box="[317,407,1747,1769]" pageId="4" pageNumber="49">Figs 4–8.</emphasis>
<taxonomicName id="1DE24D0EFF98FFA42E9351FBFD8129B8" authorityName="Papp" authorityYear="2004" box="[411,626,1747,1769]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF98FFA42E9351FBFD8129B8" box="[411,626,1747,1769]" italics="true" pageId="4" pageNumber="49">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF98FFA42D7E51FBFD5A29B8" box="[630,681,1747,1769]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view: 5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-
</paragraph>
</caption>
<paragraph id="DA5D368DFF98FFA42E955024FB652873" blockId="4.[317,1270,1747,1826]" box="[413,1174,1804,1826]" pageId="4" pageNumber="49">
nification. Scales: 
<quantity id="1D1A9B68FF98FFA42D475024FD6B2873" box="[591,664,1804,1826]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="49" unit="mm" value="0.2">0.2 mm</quantity>
for 
<figureCitation id="42D92A08FF98FFA42DC85024FCDF2873" box="[704,812,1804,1826]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="4" pageNumber="49">Figs 4–5, 8</figureCitation>
, 
<quantity id="1D1A9B68FF98FFA42C3F5024FC732873" box="[823,896,1804,1826]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="49" unit="mm" value="0.1">0.1 mm</quantity>
for 
<figureCitation id="42D92A08FF98FFA42CA15024FC102873" box="[937,995,1804,1826]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="4" pageNumber="49">Fig. 6</figureCitation>
, 
<quantity id="1D1A9B68FF98FFA42CE55024FBC52873" box="[1005,1078,1804,1826]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="49" unit="mm" value="0.4">0.4 mm</quantity>
for 
<figureCitation id="42D92A08FF98FFA42B565024FB652873" box="[1118,1174,1804,1826]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="4" pageNumber="49">Fig. 7</figureCitation>
</paragraph>
<caption id="8E9D6605FF99FFA52E355192FB052858" ID-DOI="http://doi.org/10.5281/zenodo.12586914" ID-Zenodo-Dep="12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="5" pageNumber="50" startId="5.[317,360,1722,1744]" targetBox="[346,1235,435,1687]" targetPageId="5" targetType="figure">
<paragraph id="DA5D368DFF99FFA52E355192FB052858" blockId="5.[317,1270,1722,1830]" pageId="5" pageNumber="50">
<emphasis id="E896EA9FFF99FFA52E355192FE562981" bold="true" box="[317,421,1722,1744]" pageId="5" pageNumber="50">Figs 9–15.</emphasis>
<taxonomicName id="1DE24D0EFF99FFA52EA65192FD7B2981" authorityName="Papp" authorityYear="2004" box="[430,648,1722,1744]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF99FFA52EA65192FD7B2981" box="[430,648,1722,1744]" italics="true" pageId="5" pageNumber="50">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF99FFA52D985192FD342981" box="[656,711,1722,1744]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme,
</paragraph>
</caption>
<paragraph id="DA5D368DFF99FFA52E4F5038FB1F2877" blockId="5.[317,1270,1722,1830]" box="[327,1260,1808,1830]" pageId="5" pageNumber="50">
sublateral view, 15 = genital complex, lateral view. Scale: 
<quantity id="1D1A9B68FF99FFA52C625038FC422877" box="[874,945,1808,1830]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="50" unit="mm" value="0.2">0.2 mm</quantity>
for 
<figureCitation id="42D92A08FF99FFA52CDE5038FBFD2877" box="[982,1038,1808,1830]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="5" pageNumber="50">Fig. 9</figureCitation>
, 
<quantity id="1D1A9B68FF99FFA52B1E5038FBAE2877" box="[1046,1117,1808,1830]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="50" unit="mm" value="0.1">0.1 mm</quantity>
for 
<figureCitation id="42D92A08FF99FFA52B8A5038FB1F2877" box="[1154,1260,1808,1830]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="5" pageNumber="50">Figs 10–15</figureCitation>
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E76569CFB452D55" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Thorax black, dark microtomentose. Prothorax (
<figureCitation id="42D92A08FF9AFFA62C4F569CFC8C2E9B" box="[839,895,436,458]" captionStart="Figs 1–3" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 1–3. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 1–2, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
) consisting ofextremely reduced scler - ites. Anepisternum present as a narrow oblique sclerite, medially only 
<quantity id="1D1A9B68FF9AFFA62CDF56F9FBD82EB6" box="[983,1067,465,487]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="51" unit="mm" value="0.04">0.04 mm</quantity>
broad, without setae. Anepimeron membranous. Katepisternum with 1 comparatively long seta at dorsal margin.
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E765522FB1E2DBB" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Legs short and thick, black, but trochanters, knees, apices oftibiae, and tarsi somewhat lighter. Fore femur thickened, with longer setae on dorsal side and sub-basally on ventral side. Mid femur with 1 anterior apical seta and with a number ofmedium-long setae. Hind femur very thick with a dorsal seta at apical 1/4. Fore tibia with 3 long dorsal setae: at apical 1/4, medially and subapically. Mid tibia without the usual subapical crown ofsetae, but with a strong ventroapical spur and with a shorter, though rather thick, anterodorsal spur (
<figureCitation id="42D92A08FF9AFFA62C0855B2FCC92DE1" box="[768,826,666,688]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 5</figureCitation>
). Hind tibia extremely thick, only 3 times longer than thick (
<figureCitation id="42D92A08FF9AFFA62EC7559FFDF92D9C" box="[463,522,695,717]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 7</figureCitation>
), with an extremely long and thick ventroapical spur, with a long (
<quantity id="1D1A9B68FF9AFFA62B83559FFB1C2D9C" box="[1163,1263,695,717]" metricMagnitude="-5" metricUnit="m" metricValue="5.7" pageId="6" pageNumber="51" unit="mm" value="0.057">0.057 mm</quantity>
) dorsal seta opposite the base ofventroapical spur and with 3 pairs ofantero- and postero-dorsal setae.
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E3555D9FE592CC6" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Fore basitarsus large and thick, about as long as tarsomeres 2 and 3 combined. Mid basitarsus (
<figureCitation id="42D92A08FF9AFFA62BBE55D9FB1C2C56" box="[1206,1263,753,775]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 6</figureCitation>
) with an extremely thick and long ventroapical thorn and 3 other ventral thorns. Mid tarsomere 2 and 3 each with a strong apical thorn. Hind basitarsus (
<figureCitation id="42D92A08FF9AFFA62C285402FCA92C11" box="[800,858,810,832]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 8</figureCitation>
) with three anterior (anteroventral) thorn. Hind tarsomeres 2 and 3 each with an apical (anterior to ventral) thorn, which is longer and setiform on tarsomere 3. All tarsomeres with a pair ofdorsal (i.e. antero- and postero-dorsal) long thin setae (
<figureCitation id="42D92A08FF9AFFA62E4C54A9FE6F2CC6" box="[324,412,897,919]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Figs 6, 8</figureCitation>
).
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E7654B6FB582B76" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Wing peculiar (
<figureCitation id="42D92A08FF9AFFA62D1A54B6FDB92CE5" box="[530,586,926,948]" captionStart="Figs 4–8" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 4–8. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 5–8 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 4</figureCitation>
) both in form and venation. Wing membrane and veins unicolorous yellowish. R-M at distal part ofmid third ofdiscal cell. Costal vein with 2 long setae proximally to R 
<subScript id="466634C8FF9AFFA62BE754EFFB052C82" attach="right" box="[1263,1270,967,979]" fontSize="5" pageId="6" pageNumber="51">1</subScript>
break, 13 to 15 long setae on 2nd and 3rd costal section. Wing vein ratios: C-sections 1–3 (
<typeStatus id="0559882FFF9AFFA62B9B54FFFB1F2CBC" box="[1171,1260,983,1005]" pageId="6" pageNumber="51" type="paratype">paratype</typeStatus>
): ca. 
<quantity id="1D1A9B68FF9AFFA62E5654DCFE312B5B" box="[350,450,1012,1034]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="51" unit="mm" value="0.075">0.075 mm</quantity>
, 
<quantity id="1D1A9B68FF9AFFA62EC454DCFDC52B5B" box="[460,566,1012,1034]" metricMagnitude="-4" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="51" unit="mm" value="0.105">0. 105 mm</quantity>
, 
<quantity id="1D1A9B68FF9AFFA62D3754DCFD502B5B" box="[575,675,1012,1034]" metricMagnitude="-4" metricUnit="m" metricValue="3.42" pageId="6" pageNumber="51" unit="mm" value="0.342">0.342 mm</quantity>
, M vein index = 
<quantity id="1D1A9B68FF9AFFA62C4E54DCFC5B2B5B" box="[838,936,1012,1034]" metricMagnitude="-4" metricUnit="m" metricValue="3.36" pageId="6" pageNumber="51" unit="mm" value="0.336">0.336 mm</quantity>
/ 
<quantity id="1D1A9B68FF9AFFA62CA754DCFBFC2B5B" box="[943,1039,1012,1034]" metricMagnitude="-4" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="51" unit="mm" value="0.105">0.105 mm</quantity>
= 3.2, dM-Cu ca. 
<quantity id="1D1A9B68FF9AFFA62BB754DCFE962B76" metricMagnitude="-5" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="51" unit="mm" value="0.033">0.033 mm</quantity>
, R-M-dM-Cu / dM-Cu ratio ca. 3.5. Halter dark grey, knob usually darker than stem.
</paragraph>
<caption id="8E9D6605FF9AFFA62E3551C7FB932873" ID-DOI="http://doi.org/10.5281/zenodo.12586916" ID-Zenodo-Dep="12586916" httpUri="https://zenodo.org/record/12586916/files/figure.png" pageId="6" pageNumber="51" startId="6.[317,360,1775,1797]" targetBox="[321,1265,1159,1725]" targetPageId="6" targetType="figure">
<paragraph id="DA5D368DFF9AFFA62E3551C7FB932873" blockId="6.[317,1270,1775,1826]" pageId="6" pageNumber="51">
<emphasis id="E896EA9FFF9AFFA62E3551C7FE5C2854" bold="true" box="[317,431,1775,1797]" pageId="6" pageNumber="51">Figs 16–17.</emphasis>
<taxonomicName id="1DE24D0EFF9AFFA62EBE51C7FD7D2854" authorityName="Papp" authorityYear="2004" box="[438,654,1775,1797]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF9AFFA62EBE51C7FD7D2854" box="[438,654,1775,1797]" italics="true" pageId="6" pageNumber="51">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF9AFFA62D9B51C7FD3B2854" box="[659,712,1775,1797]" pageId="6" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
, paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm
</paragraph>
</caption>
<paragraph id="DA5D368DFF9BFFA72E76569CFC8B2D71" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">
Abdomen with well-sclerotized preabdominal sclerites. Terga black, broad, with fine setae also on lateral margins. Preabdominal sterna broad but less sclerotized, 5th sternum broad, unmodified. Postabdominal sclerites well-sclerotized, darker than preabdomen; 6th to 8th sterna strongly asymmetrical as in other copromyzine genera (
<figureCitation id="42D92A08FF9BFFA72C0F5522FC9F2D71" box="[775,876,522,544]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Figs 9–11</figureCitation>
).
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E76550FFB8F2C75" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">
Male genital morphology peculiar, as described above. Epandrium strongly convex (globose), without ventral lateral cleft but fused and closed ventrally as in some genera of 
<taxonomicName id="1DE24D0EFF9BFFA72B70556CFB002D0B" box="[1144,1267,580,602]" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="subFamily" subFamily="Limosininae">Limosininae</taxonomicName>
. Epandrium with a pair ofcomparatively long subdorsal (caudal) setae. Cerci distinct but small, not fused with epandrium (
<figureCitation id="42D92A08FF9BFFA72D115556FDAC2DC5" box="[537,607,638,660]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 10</figureCitation>
), without setae. Surstylus (
<figureCitation id="42D92A08FF9BFFA72C685556FC572DC5" box="[864,932,638,660]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 11</figureCitation>
) long thin, digitiform with a longer medial subapical seta and with 2 small apical setae. Hypandrium small. Phallic complex (
<figureCitation id="42D92A08FF9BFFA72BAF55B2FB1D2DE1" box="[1191,1262,666,688]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 15</figureCitation>
) rather compact. Phallus with rather normal distiphallus, basiphallus with a thick, intricately sclerotized dorsal projection, which is almost as large as distiphallus (
<figureCitation id="42D92A08FF9BFFA72CD755FCFBD52DBB" box="[991,1062,724,746]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 15</figureCitation>
). Phallapodeme slen- der, comparatively short, strongly curved in its distal third. Epiphallus seemingly absent. Ejaculatory apodeme (
<figureCitation id="42D92A08FF9BFFA72EAA5426FDE22C75" box="[418,529,782,804]" captionStart="Figs 9–15" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 9–15. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Figs 14–15</figureCitation>
) mushroom-shaped with an additional sclerite at its distal part.
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E765402FC022CBC" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">
Female tergite 7 with a pair ofextremely long setae. Cercus with a long apical and 2 thick dor - sal setae (
<figureCitation id="42D92A08FF9BFFA72E91546FFE132C0C" box="[409,480,839,861]" captionStart="Figs 16–17" captionStartId="6.[317,360,1775,1797]" captionTargetBox="[321,1265,1159,1725]" captionTargetId="figure-362@6.[317,1269,1154,1726]" captionTargetPageId="6" captionText="Figs 16–17. Immoderatus foldvarii sp. n., paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.12586916" httpUri="https://zenodo.org/record/12586916/files/figure.png" pageId="7" pageNumber="52">Fig. 16</figureCitation>
). Two spermathecae (
<figureCitation id="42D92A08FF9BFFA72DA7546FFD062C0C" box="[687,757,839,861]" captionStart="Figs 16–17" captionStartId="6.[317,360,1775,1797]" captionTargetBox="[321,1265,1159,1725]" captionTargetId="figure-362@6.[317,1269,1154,1726]" captionTargetPageId="6" captionText="Figs 16–17. Immoderatus foldvarii sp. n., paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.12586916" httpUri="https://zenodo.org/record/12586916/files/figure.png" pageId="7" pageNumber="52">Fig. 17</figureCitation>
), similar in shape but uneven in size (
<specimenCount id="CCE4FD04FF9BFFA72B50546FFB3E2C0C" box="[1112,1229,839,861]" count="2" pageId="7" pageNumber="52" type="female">two females</specimenCount>
dissected), spermathecae not globular as in several genera ofCopromyzinae (
<taxonomicName id="1DE24D0EFF9BFFA72B1C544CFB822C2B" authorityName="Lioy" authorityYear="1864" box="[1044,1137,868,890]" class="Insecta" family="Sphaeroceridae" genus="Lotophila" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72B1C544CFB822C2B" box="[1044,1137,868,890]" italics="true" pageId="7" pageNumber="52">Lotophila</emphasis>
</taxonomicName>
, 
<taxonomicName id="1DE24D0EFF9BFFA72B76544CFB1C2C2B" authorityName="Macquart" authorityYear="1835" box="[1150,1263,868,890]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72B76544CFB1C2C2B" box="[1150,1263,868,890]" italics="true" pageId="7" pageNumber="52">Crumomyia</emphasis>
</taxonomicName>
, 
<taxonomicName id="1DE24D0EFF9BFFA72E3554A9FE392CC6" box="[317,458,897,919]" class="Insecta" family="Sphaeroceridae" genus="Achaetothorax" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72E3554A9FE392CC6" box="[317,458,897,919]" italics="true" pageId="7" pageNumber="52">Achaetothorax</emphasis>
</taxonomicName>
, 
<taxonomicName id="1DE24D0EFF9BFFA72ED454A9FD882CC6" authorityName="Hedicke" authorityYear="1923" box="[476,635,897,919]" class="Insecta" family="Sphaeroceridae" genus="Gymnometopina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72ED454A9FD882CC6" box="[476,635,897,919]" italics="true" pageId="7" pageNumber="52">Gymnometopina</emphasis>
</taxonomicName>
, 
<taxonomicName id="1DE24D0EFF9BFFA72D8554A9FCE52CC6" authorityName="Vanschuytbroeck" authorityYear="1948" box="[653,790,897,919]" class="Insecta" family="Sphaeroceridae" genus="Metaborborus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72D8554A9FCE52CC6" box="[653,790,897,919]" italics="true" pageId="7" pageNumber="52">Metaborborus</emphasis>
</taxonomicName>
, etc., see 
<bibRefCitation id="BE734B7CFF9BFFA72C8F54A9FB772CC6" author="NORRBOM, A. L. &amp; KIM, K. C." box="[903,1156,897,919]" pageId="7" pageNumber="52" pagination="167 - 225" refId="ref3210" refString="NORRBOM, A. L. &amp; KIM, K. C. (1985 a) Systematics ofCrumomyia Macquart and Alloborborus (Diptera: Sphaeroceridae). Systematic Entomology 10: 167 - 225." type="journal article" year="1985">
<smallCapsWord id="DCBBA051FF9BFFA72C8F54A9FC1A2CC4" baselines="913,913" box="[903,1001,897,919]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norrbom" pageId="7" pageNumber="52">NORRBOM</smallCapsWord>
&amp; 
<smallCapsWord id="DCBBA051FF9BFFA72B1B54A9FBCF2CC4" baselines="913,913" box="[1043,1084,897,919]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kim" pageId="7" pageNumber="52">KIM</smallCapsWord>
1985 
<emphasis id="E896EA9FFF9BFFA72B7F54A9FB772CC6" box="[1143,1156,897,919]" italics="true" pageId="7" pageNumber="52">a</emphasis>
</bibRefCitation>
,b, 
<bibRefCitation id="BE734B7CFF9BFFA72BAF54A9FE282CE5" author="PAPP, L. &amp; NORRBOM, A. L." pageId="7" pageNumber="52" pagination="321 - 344" refId="ref3314" refString="PAPP, L. &amp; NORRBOM, A. L. (1992) Taxonomy ofthe Palaeotropical genus Achaetothorax Hedicke (Diptera, Sphaeroceridae). Acta zool. hung. 38: 321 - 344." type="journal article" year="1992">
<smallCapsWord id="DCBBA051FF9BFFA72BAF54A9FB242CC4" baselines="913,913" box="[1191,1239,897,919]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Papp" pageId="7" pageNumber="52">PAPP</smallCapsWord>
&amp; 
<smallCapsWord id="DCBBA051FF9BFFA72E3554B6FE6C2CE3" baselines="942,942" box="[317,415,926,948]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norrbom" pageId="7" pageNumber="52">NORRBOM</smallCapsWord>
1992
</bibRefCitation>
). Ducts laterally directed, which again is peculiar (similar lateroclinate ducts are known in 
<taxonomicName id="1DE24D0EFF9BFFA72EAD5492FC952C81" authority="ROHACEK &amp; MARSHALL, 1998" authorityName="ROHACEK &amp; MARSHALL" authorityYear="1998" box="[421,870,954,976]" class="Insecta" family="Sphaeroceridae" genus="Sphaeromitra" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72EAD5492FDDA2C81" box="[421,553,954,976]" italics="true" pageId="7" pageNumber="52">Sphaeromitra</emphasis>
<bibRefCitation id="BE734B7CFF9BFFA72D3B5492FC952C81" author="ROHACEK, J. &amp; MARSHALL, S. A." box="[563,870,954,976]" pageId="7" pageNumber="52" pagination="455 - 491" refId="ref3406" refString="ROHACEK, J. &amp; MARSHALL, S. A. (1998) Revision ofHomalomitrinae subfam. n. (Diptera: Sphaero - ceridae), with description ofa new genus and three new species. European Journal ofEntomol - ogy 95: 455 - 491." type="journal article" year="1998">
<smallCapsWord id="DCBBA051FF9BFFA72D3B5492FD622C9E" baselines="970,971" box="[563,657,954,976]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Rohacek" pageId="7" pageNumber="52">ROHÁČEK</smallCapsWord>
&amp; 
<smallCapsWord id="DCBBA051FF9BFFA72DB05492FCD02C9E" baselines="970,971" box="[696,803,954,976]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Marshall" pageId="7" pageNumber="52">MARSHALL</smallCapsWord>
, 1998
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="1DE24D0EFF9BFFA72C7C5492FBE62C81" box="[884,1045,954,976]" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="subFamily" subFamily="Homalomitrinae">Homalomitrinae</taxonomicName>
), but the shape ofthe spermatheca ofthis species is otherwise unique amongst sphaerocerids.
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E7654DCFCA62B15" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">This new species is so peculiar that I think a new genus deserves to be erected for it. However, I have no objective criteria to separate generic or specific characters. For this reason, some characters are given in both the generic and specific descriptions.</paragraph>
<paragraph id="DA5D368DFF9BFFA72E76534FFCCC2BE6" blockId="7.[317,1265,1127,1207]" pageId="7" pageNumber="52">
<materialsCitation id="6A8A3CD0FF9BFFA72E76534FFCC92BE6" country="Vietnam" location="Viet Nam" pageId="7" pageNumber="52" specimenCount="1">
I name this new species to honour my young colleague, Dr 
<smallCapsWord id="DCBBA051FF9BFFA72CA5534FFC0F2B2D" baselines="1143,1144" box="[941,1020,1127,1149]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Mihaly" pageId="7" pageNumber="52">MIHÁLY</smallCapsWord>
<smallCapsWord id="DCBBA051FF9BFFA72B09534FFB902B2D" baselines="1143,1144" box="[1025,1123,1127,1149]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Foldvari" pageId="7" pageNumber="52">FÖLDVÁRI</smallCapsWord>
, one ofthe collectors ofthe 
<typeStatus id="0559882FFF9BFFA72EB053ACFE122BCB" box="[440,481,1156,1178]" pageId="7" pageNumber="52">type</typeStatus>
series, and who also collected much other invaluable dipterous material during his joint 2003 collection trip to 
<collectingCountry id="A2F5761DFF9BFFA72D595389FD422BE6" box="[593,689,1185,1207]" name="Vietnam" pageId="7" pageNumber="52">Viet Nam</collectingCountry>
and 
<collectingCountry id="A2F5761DFF9BFFA72DEA5389FCC92BE6" box="[738,826,1185,1207]" name="Thailand" pageId="7" pageNumber="52">Thailand</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="DA5D368DFF9BFFA72C1B53C1FCEC2BAE" blockId="7.[787,799,1257,1279]" box="[787,799,1257,1279]" pageId="7" pageNumber="52">*</paragraph>
<paragraph id="DA5D368DFF9BFFA72E76523CFB052A7B" blockId="7.[317,1270,1300,1351]" box="[382,1270,1300,1322]" pageId="7" pageNumber="52">
<emphasis id="E896EA9FFF9BFFA72E76523CFDDC2A7B" box="[382,559,1300,1322]" italics="true" pageId="7" pageNumber="52">Acknowledgements</emphasis>
– This study was supported by the National Scientific Research Fund (OTKA,
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E355219FC682A16" blockId="7.[317,1270,1300,1351]" box="[317,923,1329,1351]" pageId="7" pageNumber="52">No. T 042540). I am grateful to the two reviewers for advice.</paragraph>
</subSubSection>
</treatment>
</document>