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<mods:title id="F685F70FC2E3F9ED0A68C8309292B5FE">Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<taxonomicName id="658FC8F84B0EFFC2E7B8FAFAFD5BFAED" ID-CoL="85ZWC" authority="Banner &amp; Banner, 1966" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[151,656,1367,1394]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="songkla" status="stat. nov.">
<emphasis id="90FB6F694B0EFFC2E7B8FAFAFEAAFAED" bold="true" box="[151,353,1367,1394]" italics="true" pageId="74" pageNumber="75">Alpheus songkla</emphasis>
<bibRefCitation id="C61ECE8A4B0EFFC2E647FAF5FD5BFAED" author="Banner, A. H. &amp; Banner, D. M." box="[360,656,1368,1394]" pageId="74" pageNumber="75" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner, 1966</bibRefCitation>
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, 
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(
<figureCitation id="3AB4AFFE4B0EFFC2E7B0FAD1FF36FA09" box="[159,253,1404,1430]" captionStart="FIGURE 41" captionStartId="77.[151,250,1873,1897]" captionTargetBox="[151,1435,594,1849]" captionTargetId="figure-243@77.[151,1435,594,1849]" captionTargetPageId="77" captionText="FIGURE 41. Alpheus songkla Banner &amp; Banner, 1966 stat. nov.: holotype, female (cl 6.1 mm) from Thale Sap (= Songkhla Lake), Thailand (USNM 120407); A—habitus (most appendages detached), dorsal view; B—major (left) cheliped, lateral view; C—same, mesial view; D—minor (right) cheliped, lateral view; E—third pereiopod, lateral view. Photographs by Robert M. Lasley Jr." figureDoi="http://doi.org/10.5281/zenodo.7912425" httpUri="https://zenodo.org/record/7912425/files/figure.png" pageId="74" pageNumber="75">Figs. 41</figureCitation>
, 
<figureCitation id="3AB4AFFE4B0EFFC2E626FAD1FEF2FA08" box="[265,313,1404,1431]" captionStart="FIGURE 52" captionStartId="106.[151,250,1909,1933]" captionTargetBox="[156,1430,181,1885]" captionTargetId="figure-21@106.[156,1430,181,1885]" captionTargetPageId="106" captionText="FIGURE 52. Distributional maps for species of Alpheus treated in this study:A—A. takla sp. nov.; B—A. mangalis sp. nov. and A. cf. mangalis sp. nov.; C—A. songkla Banner &amp; Banner, 1966 and A. cf. songkla (see text); D—A. paludicola Kemp, 1915, A. nipa Banner &amp; Banner, 1985 and A. bunburius Banner &amp; Banner, 1982; E—A. pontederiae de Rochebrune, 1883; F—A. firmus Kim &amp; Abele, 1988 and A. cf. firmus (see text)." figureDoi="http://doi.org/10.5281/zenodo.7912457" httpUri="https://zenodo.org/record/7912457/files/figure.png" pageId="74" pageNumber="75">52C</figureCitation>
)
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[see also 
<figureCitation id="3AB4AFFE4B0EFFC2E7D0FA0DFE84FA25" box="[255,335,1440,1466]" captionStart="FIGURE 42" captionStartId="78.[151,250,1821,1845]" captionTargetBox="[177,1407,193,1786]" captionTargetId="figure-21@78.[164,1422,181,1797]" captionTargetPageId="78" captionText="FIGURE 42. Alpheus cf. songkla stat. nov.: male (cl 8.1 mm) [A–I], male (cl 10.4 mm) [J–M], both from near Ho Chi Minh City, Vietnam (MNHN-IU-2018-5655); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—major (left) cheliped, chela, mesial; F—same, chela, carpus and distal part of merus, lateral; G—same, dactylus; H—minor (right) cheliped, chela, carpus and distal part of merus, lateral view; I—second pereiopod, lateral view; J—third pereiopod, lateral view; K—same, dactylus, dorsal view; L—fourth pereiopod, ischium, lateral view; M—possibly abnormal minor (right) cheliped, chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912427" httpUri="https://zenodo.org/record/7912427/files/figure.png" pageId="74" pageNumber="75">Fig. 42</figureCitation>
for 
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<emphasis id="90FB6F694B0EFFC2E652FA0DFE5DFA25" box="[381,406,1440,1466]" italics="true" pageId="74" pageNumber="75">A.</emphasis>
cf. 
<emphasis id="90FB6F694B0EFFC2E6EFFA0DFDDCFA25" box="[448,535,1440,1466]" italics="true" pageId="74" pageNumber="75">songkla</emphasis>
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]
</paragraph>
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<taxonomicName id="658FC8F84B0EFFC2E7B8FA72FD28FA68" ID-CoL="85ZWC" authority="Banner &amp; Banner 1966: 147" authorityName="Banner &amp; Banner" authorityPageNumber="147" authorityYear="1966" box="[151,739,1503,1527]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="74" pageNumber="75" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B0EFFC2E7B8FA72FE0AFA68" box="[151,449,1503,1527]" italics="true" pageId="74" pageNumber="75">Alpheus malabaricus songkla</emphasis>
<bibRefCitation id="C61ECE8A4B0EFFC2E6E9FA4DFD28FA68" author="Banner, A. H. &amp; Banner, D. M." box="[454,739,1503,1527]" pageId="74" pageNumber="75" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966: 147</bibRefCitation>
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, fig. 56; 
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<bibRefCitation id="C61ECE8A4B0EFFC2E416FA72FB53FA68" author="Angsupanich, S. &amp; Kuwabara, R." box="[825,1176,1503,1527]" pageId="74" pageNumber="75" pagination="1 - 13" refId="ref68563" refString="Angsupanich, S. &amp; Kuwabara, R. (1999) Distribution of macrobenthic fauna in Phawong and U-Taphao canals flowing into a lagoonal lake, Songkhla, Thailand. Lakes &amp; Reservoirs: Research and Management, 4, 1 - 13. https: // doi. org / 10.1046 / j. 1440 - 1770.1999.00080. x" type="journal article" year="1999">Angsupanich &amp; Kuwabara 1999: 6</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="232E956A4B0EFFC2E38EFA72FF3BF989" author="Angsupanich, S. &amp; Siripech, A. &amp; Charoenpornthip, M." page="376" pageId="74" pageNumber="75" year="2005">
<bibRefCitation id="C61ECE8A4B0EFFC2E38EFA72FF3BF989" author="Angsupanich, S. &amp; Siripech, A. &amp; Charoenpornthip, M." pageId="74" pageNumber="75" pagination="365 - 390" refId="ref68624" refString="Angsupanich, S., Siripech, A. &amp; Charoenpornthip, M. (2005) Macrobenthic fauna community in the Middle Songkhla Lake, southern Thailand. Songklanakarin Journal of Science and Technology, 27 (Supplement 1), 365 - 390." type="journal article" year="2005">
Angsupanich 
<emphasis id="90FB6F694B0EFFC2E203FA4DFA95FA68" box="[1324,1374,1503,1527]" italics="true" pageId="74" pageNumber="75">et al.</emphasis>
2005: 376
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; 
<treatmentCitation id="232E956A4B0EFFC2E7D4FA53FE05F989" author="Naiyanetr, P." box="[251,462,1534,1558]" page="173" pageId="74" pageNumber="75" year="2007">
<bibRefCitation id="C61ECE8A4B0EFFC2E7D4FA53FE05F989" author="Naiyanetr, P." box="[251,462,1534,1558]" pageId="74" pageNumber="75" refId="ref73355" refString="Naiyanetr, P. (2007) Checklist of crustacean fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr 2007: 173</bibRefCitation>
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.
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</paragraph>
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(?) 
<emphasis id="90FB6F694B0EFFC2E795F9B1FDC9F9AB" box="[186,514,1564,1588]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="658FC8F84B0EFFC2E795F9B1FE2EF9AB" ID-CoL="85ZWC" authorityName="Banner &amp; Banner" authorityYear="1966" box="[186,485,1564,1588]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="74" pageNumber="75" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">Alpheus malabaricus songkla</taxonomicName>
.—
</emphasis>
<bibRefCitation id="C61ECE8A4B0EFFC2E52DF9B1FD08F9AB" author="Thomas, M. M." box="[514,707,1564,1588]" pageId="74" pageNumber="75" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976: 668</bibRefCitation>
.
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<emphasis id="90FB6F694B0EFFC2E7B8F9F0FE89F9E8" bold="true" box="[151,322,1629,1655]" pageId="74" pageNumber="75">Type material</emphasis>
. 
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<typeStatus id="7D340DD94B0EFFC2E67AF9F0FE0BF9E8" box="[341,448,1629,1655]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
, female (cl 
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), 
<specimenCode id="F2291B004B0EFFC2E5E8F9F0FCB0F9E8" box="[711,891,1629,1655]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="75" type="Museum">USNM 120407</specimenCode>
, 
<collectingCountry id="DA98F3EB4B0EFFC2E4A5F9F0FC3BF9E8" box="[906,1008,1629,1655]" name="Thailand" pageId="74" pageNumber="75">Thailand</collectingCountry>
, 
<collectingMunicipality id="425429014B0EFFC2E32FF9F0FBBEF9E8" box="[1024,1141,1629,1655]" pageId="74" pageNumber="75">Thale Sap</collectingMunicipality>
(
<collectingRegion id="604B7D994B0EFFC2E3A6F9F0FB3EF9E8" box="[1161,1269,1629,1655]" country="Thailand" name="Songkhla" pageId="74" pageNumber="75">Songkhla</collectingRegion>
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), 
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, commercial shrimp trawl, sandy bottom, depth: approximately 
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, leg. 
<collectorName id="0F7AD6AD4B0EFFC2E493F92FFB87F903" box="[956,1100,1666,1692]" pageId="74" pageNumber="75">A.H. Banner</collectorName>
, 
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;
</materialsCitation>
<materialsCitation id="12E7B9264B0EFFC2E3CDF92FFC26F920" ID-GBIF-Occurrence="4104078358" collectingDate="1963-03-27" collectionCode="USNM, BR" collectorName="A. H. Banner" country="Thailand" elevation="1" location="Lake" municipality="Thale Sap" pageId="74" pageNumber="75" specimenCode="USNM 120407, BR44, USNM 120408" specimenCount="2" stateProvince="Songkhla" typeStatus="paratype">
<typeStatus id="7D340DD94B0EFFC2E3CDF92FFA8EF903" box="[1250,1349,1666,1692]" pageId="74" pageNumber="75" type="paratype">paratype</typeStatus>
, female (cl 
<quantity id="65771E9E4B0EFFC2E793F908FEDDF920" box="[188,278,1701,1727]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="74" pageNumber="75" unit="mm" value="6.4">6.4 mm</quantity>
), 
<specimenCode id="F2291B004B0EFFC2E606F908FE13F920" box="[297,472,1701,1727]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="75" type="Museum">USNM 120408</specimenCode>
, same collection data as for previous specimen.
</materialsCitation>
</paragraph>
<paragraph id="A230B37B4B0EFFC2E7E8F964FD3EF8CF" blockId="74.[151,1436,1629,2016]" pageId="74" pageNumber="75">
<emphasis id="90FB6F694B0EFFC2E7E8F964FE17F97C" bold="true" box="[199,476,1737,1763]" pageId="74" pageNumber="75">Tentative identification</emphasis>
. 
<taxonomicName id="658FC8F84B0EFFC2E6C5F964FD0CF97C" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[490,711,1737,1763]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0EFFC2E6C5F964FD8FF97C" box="[490,580,1737,1763]" italics="true" pageId="74" pageNumber="75">Alpheus</emphasis>
<materialsCitation id="12E7B9264B0EFFC2E563F964FD0CF97C" ID-GBIF-Occurrence="4104078308" box="[588,711,1737,1763]" collectionCode="ZRC" collectorName="Pulau Ubin &amp; P. K. L. Ng" country="Singapore" location="Singapore" pageId="74" pageNumber="75" specimenCode="ZRC 1992.11117, 11119" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
cf. 
<emphasis id="90FB6F694B0EFFC2E55FF964FD0CF97C" box="[624,711,1737,1763]" italics="true" pageId="74" pageNumber="75">songkla</emphasis>
</materialsCitation>
</taxonomicName>
. 
<emphasis id="90FB6F694B0EFFC2E5FAF967FC8CF97B" box="[725,839,1738,1764]" italics="true" pageId="74" pageNumber="75">
<collectingCountry id="DA98F3EB4B0EFFC2E5FAF967FC8CF97B" box="[725,839,1738,1764]" name="Singapore" pageId="74" pageNumber="75">Singapore</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0EFFC2E478F964FC66F97C" box="[855,941,1737,1763]" pageId="74" pageNumber="75" type="male">2 males</specimenCount>
(cl 8.8, 10.0 mm), 
<specimenCount id="B48978F24B0EFFC2E3A9F964FB2CF97C" box="[1158,1255,1737,1763]" pageId="74" pageNumber="75" type="female">1 female</specimenCount>
(
<quantity id="65771E9E4B0EFFC2E3D8F964FA98F97C" box="[1271,1363,1737,1763]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="74" pageNumber="75" unit="mm" value="9.6">9.6 mm</quantity>
), 
<specimenCode id="F2291B004B0EFFC2E249F964FED0F898" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="74" pageNumber="75">ZRC 1992.11117</specimenCode>
– 
<specimenCode id="F2291B004B0EFFC2E606F940FEA6F898" box="[297,365,1773,1799]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="74" pageNumber="75">11119</specimenCode>
, 
<collectorName id="0F7AD6AD4B0EFFC2E656F940FE37F898" box="[377,508,1773,1799]" pageId="74" pageNumber="75">Pulau Ubin</collectorName>
, mudflat, leg. 
<collectorName id="0F7AD6AD4B0EFFC2E58FF943FCDFF897" box="[672,788,1774,1800]" pageId="74" pageNumber="75">P.K.L. Ng</collectorName>
, 
<materialsCitation id="12E7B9264B0EFFC2E40EF940FDF1F8D0" ID-GBIF-Occurrence="4104078352" collectingDate="2012-02-14" collectionCode="ZRC" collectorName="H. H. Ng; H. Wong &amp; Y. L. Teo" country="Singapore" pageId="74" pageNumber="75" specimenCode="ZRC 2014.0665" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
08.1987; 
<specimenCount id="B48978F24B0EFFC2E4A3F940FC13F898" box="[908,984,1773,1799]" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0EFFC2E329F940FBABF897" box="[1030,1120,1773,1800]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="74" pageNumber="75" unit="mm" value="6.8">6.8 mm</quantity>
, missing minor cheliped), 
<specimenCount id="B48978F24B0EFFC2E2A1F940FF25F8B4" pageId="74" pageNumber="75" type="female">2 females</specimenCount>
(cl 5.6, 
<quantity id="65771E9E4B0EFFC2E664F8BCFE6EF8B4" box="[331,421,1809,1836]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="74" pageNumber="75" unit="mm" value="7.5">7.5 mm</quantity>
), 
<specimenCode id="F2291B004B0EFFC2E697F8BCFDBBF8B3" box="[440,624,1809,1836]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="74" pageNumber="75">ZRC 2014.0665</specimenCode>
, near 
<collectorName id="0F7AD6AD4B0EFFC2E59DF8BCFC70F8B4" box="[690,955,1809,1835]" pageId="74" pageNumber="75">Sarimbun Scouts Camp</collectorName>
(Jalan Bahtera), seining on mud, leg. 
<collectorName id="0F7AD6AD4B0EFFC2E249F8BFFF77F8CF" pageId="74" pageNumber="75">H.H. Ng</collectorName>
, H. Wong &amp; 
<collectorName id="0F7AD6AD4B0EFFC2E661F89BFE7BF8CF" box="[334,432,1846,1872]" pageId="74" pageNumber="75">Y.L. Teo</collectorName>
, 
<date id="D63195BB4B0EFFC2E693F898FDF1F8D0" box="[444,570,1845,1871]" pageId="74" pageNumber="75" value="2012-02-14">
<collectingDate id="C6756C534B0EFFC2E693F898FDF1F8D0" box="[444,570,1845,1871]" pageId="74" pageNumber="75" value="2012-02-14">14.02.2012</collectingDate>
</date>
</materialsCitation>
[48193–48195].
</paragraph>
<paragraph id="A230B37B4B0EFFC2E7E8F8F4FB92F807" blockId="74.[151,1436,1629,2016]" pageId="74" pageNumber="75">
<materialsCitation id="12E7B9264B0EFFC2E7E8F8F4FDA9F8EC" ID-GBIF-Occurrence="4104078387" box="[199,610,1881,1908]" country="Malaysia" location="Malaysia" pageId="74" pageNumber="75" specimenCount="3" specimenCount-male="3">
<emphasis id="90FB6F694B0EFFC2E7E8F8F4FEE4F8EC" box="[199,303,1881,1907]" italics="true" pageId="74" pageNumber="75">
<collectingCountry id="DA98F3EB4B0EFFC2E7E8F8F4FEE4F8EC" box="[199,303,1881,1907]" name="Malaysia" pageId="74" pageNumber="75">Malaysia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0EFFC2E66EF8F4FE51F8EC" box="[321,410,1881,1907]" pageId="74" pageNumber="75" type="male">3 males</specimenCount>
(cl 7.4–8.0 mm)
</materialsCitation>
, 
<materialsCitation id="12E7B9264B0EFFC2E55FF8F4FB9EF807" ID-GBIF-Occurrence="4104078562" collectingDate="2002-05-11" collectionCode="OUMNH, ZC" collectorName="Telok Tebrau &amp; Z. Jaafar" country="Malaysia" location="Terengganu" pageId="74" pageNumber="75" specimenCode="ZRC 2009.0306" specimenCount="1" specimenCount-female="1" stateProvince="Terengganu">
<specimenCount id="B48978F24B0EFFC2E55FF8F4FD1FF8EC" box="[624,724,1881,1907]" pageId="74" pageNumber="75" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B0EFFC2E429F8F4FCA8F8EC" box="[774,867,1881,1908]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="74" pageNumber="75" unit="mm" value="9.5">9.5 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0EFFC2E456F8F4FC22F8EC" box="[889,1001,1881,1907]" pageId="74" pageNumber="75">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0EFFC2E4C3F8F4FBDAF8EC" box="[1004,1041,1881,1907]" country="Austria" name="Zoological Collection, University of Vienna" pageId="74" pageNumber="75" type="University or college">ZC</collectionCode>
. 2019.06.50 [ex 
<specimenCode id="F2291B004B0EFFC2E3FBF8F4FA59F8EC" box="[1236,1426,1881,1907]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="74" pageNumber="75">ZRC 2009.0306</specimenCode>
], 
<collectingRegion id="604B7D994B0EFFC2E7B8F8D3FED5F807" box="[151,286,1918,1944]" country="Malaysia" name="Terengganu" pageId="74" pageNumber="75">Terengganu</collectingRegion>
, 
<collectorName id="0F7AD6AD4B0EFFC2E607F8D0FE75F808" box="[296,446,1917,1943]" pageId="74" pageNumber="75">Telok Tebrau</collectorName>
, intertidal mudflats, with gobies, leg. 
<collectorName id="0F7AD6AD4B0EFFC2E449F8D3FC01F808" box="[870,970,1917,1944]" pageId="74" pageNumber="75">Z. Jaafar</collectorName>
, 
<date id="D63195BB4B0EFFC2E4FAF8D0FB9EF807" box="[981,1109,1917,1944]" pageId="74" pageNumber="75" value="2002-05-11">
<collectingDate id="C6756C534B0EFFC2E4FAF8D0FB9EF807" box="[981,1109,1917,1944]" pageId="74" pageNumber="75" value="2002-05-11">05.11.2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B0EFFC3E7E8F80CFE5BFF4A" blockId="74.[151,1436,1629,2016]" lastBlockId="75.[151,1437,150,2013]" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
<materialsCitation id="12E7B9264B0EFFC2E7E8F80CFED2F87F" ID-GBIF-Occurrence="4104078377" collectingDate="1925-06-20" collectionCode="USNM" collectorName="Sin Gora &amp; H. M. Smith" country="Thailand" location="Thailand" pageId="74" pageNumber="75" specimenCode="USNM 65561" specimenCount="1" specimenCount-male="1">
<emphasis id="90FB6F694B0EFFC2E7E8F80CFEE7F824" box="[199,300,1953,1979]" italics="true" pageId="74" pageNumber="75">
<collectingCountry id="DA98F3EB4B0EFFC2E7E8F80CFEE7F824" box="[199,300,1953,1979]" name="Thailand" pageId="74" pageNumber="75">Thailand</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0EFFC2E610F80CFE46F824" box="[319,397,1953,1979]" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0EFFC2E6EEF80CFDD5F824" box="[449,542,1953,1979]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="74" pageNumber="75" unit="mm" value="9.6">9.6 mm</quantity>
), 
<specimenCode id="F2291B004B0EFFC2E51AF80CFD12F823" box="[565,729,1953,1980]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="75" type="Museum">USNM 65561</specimenCode>
, “ 
<collectorName id="0F7AD6AD4B0EFFC2E5DDF80CFC97F824" box="[754,860,1953,1979]" pageId="74" pageNumber="75">Sin Gora</collectorName>
inland sea [locality not found], leg. 
<collectorName id="0F7AD6AD4B0EFFC2E222F80FFA53F824" box="[1293,1432,1953,1980]" pageId="74" pageNumber="75">H.M. Smith</collectorName>
, 
<date id="D63195BB4B0EFFC2E7B8F868FED2F87F" box="[151,281,1989,2016]" pageId="74" pageNumber="75" value="1925-06-20">
<collectingDate id="C6756C534B0EFFC2E7B8F868FED2F87F" box="[151,281,1989,2016]" pageId="74" pageNumber="75" value="1925-06-20">20.06.1925</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B0EFFC3E607F868FD05FF2E" ID-GBIF-Occurrence="4104078573" collectingDate="1925-05-21" collectionCode="USNM" collectorName="H. Smith" country="Thailand" lastPageId="75" lastPageNumber="76" location="Bangkok" pageId="74" pageNumber="75" specimenCode="USNM 65558" specimenCount="1" specimenCount-male="1" stateProvince="Krung Thep Mahanakhon">
<specimenCount id="B48978F24B0EFFC2E607F868FEB3F840" box="[296,376,1989,2015]" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0EFFC2E683F868FDDDF87F" box="[428,534,1989,2016]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="74" pageNumber="75" unit="mm" value="10.6">10.6 mm</quantity>
, missing both chelipeds), 
<specimenCode id="F2291B004B0EFFC2E464F868FC3BF87F" box="[843,1008,1989,2016]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="75" type="Museum">USNM 65558</specimenCode>
, 
<collectorName id="0F7AD6AD4B0EFFC2E4D0F868FB97F840" box="[1023,1116,1989,2015]" pageId="74" pageNumber="75">Paknam</collectorName>
, 
<collectorName id="0F7AD6AD4B0EFFC2E343F86BFAAAF840" box="[1132,1377,1989,2016]" pageId="74" pageNumber="75">Menam Chao Phraya</collectorName>
near 
<collectingRegion id="604B7D994B0FFFC3E7B8FF3AFF34FF2E" box="[151,255,151,177]" country="Thailand" name="Krung Thep Mahanakhon" pageId="75" pageNumber="76">Bangkok</collectingRegion>
, 
<collectorName id="0F7AD6AD4B0FFFC3E625FF3AFE55FF2E" box="[266,414,151,177]" pageId="75" pageNumber="76">Klang Krang</collectorName>
, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E6F6FF3AFD89FF2E" box="[473,578,151,177]" pageId="75" pageNumber="76">H. Smith</collectorName>
, 
<date id="D63195BB4B0FFFC3E563FF3BFD05FF2E" box="[588,718,150,177]" pageId="75" pageNumber="76" value="1925-05-21">
<collectingDate id="C6756C534B0FFFC3E563FF3BFD05FF2E" box="[588,718,150,177]" pageId="75" pageNumber="76" value="1925-05-21">21.05.1925</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B0FFFC3E5F6FF3AFE40FF4A" ID-GBIF-Occurrence="4104078511" collectingDate="1923-01-07" collectionCode="USNM" collectorName="H. Smith" country="Thailand" location="Bangpakong River" pageId="75" pageNumber="76" specimenCode="USNM 65477" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B0FFFC3E5F6FF3AFCF1FF2E" box="[729,826,151,177]" pageId="75" pageNumber="76" type="female">1 female</specimenCount>
(cl 9.0 mm), 
<specimenCode id="F2291B004B0FFFC3E4FDFF3AFBB9FF2E" box="[978,1138,150,177]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="75" pageNumber="76" type="Museum">USNM 65477</specimenCode>
/2, 
<location id="A750E5A04B0FFFC3E3BBFF3AFAA3FF2E" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E3BBFF3AFAA3FF2E" box="[1172,1384,151,177]" country="Thailand" name="Bangpakong River" pageId="75" pageNumber="76">Bangpakong River</location>
, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E7B8FF16FF34FF4A" box="[151,255,187,213]" pageId="75" pageNumber="76">H. Smith</collectorName>
, 
<date id="D63195BB4B0FFFC3E625FF16FE40FF4A" box="[266,395,187,213]" pageId="75" pageNumber="76" value="1923-01-07">
<collectingDate id="C6756C534B0FFFC3E625FF16FE40FF4A" box="[266,395,187,213]" pageId="75" pageNumber="76" value="1923-01-07">01.07.1923</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FF72FE27FEDE" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<materialsCitation id="12E7B9264B0FFFC3E7E8FF72FED2FE82" ID-GBIF-Occurrence="4104078330" collectingDate="1995-12-08" collectorName="Duyen-Hai &amp; H. Dung" country="Vietnam" location="Ho Chi Minh City" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Ho Chi Minh City">
<emphasis id="90FB6F694B0FFFC3E7E8FF72FEE7FF67" box="[199,300,223,248]" italics="true" pageId="75" pageNumber="76">
<collectingCountry id="DA98F3EB4B0FFFC3E7E8FF72FEEEFF67" box="[199,293,223,248]" name="Vietnam" pageId="75" pageNumber="76">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="B48978F24B0FFFC3E617FF72FE43FF66" box="[312,392,223,249]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E691FF72FDD5FF66" box="[446,542,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="75" pageNumber="76" unit="mm" value="8.1">8.1 mm</quantity>
), MNHN-IU-2018-5655, 
<collectorName id="0F7AD6AD4B0FFFC3E467FF72FC0FFF66" box="[840,964,223,249]" pageId="75" pageNumber="76">Duyen-Hai</collectorName>
near 
<collectingRegion id="604B7D994B0FFFC3E322FF72FB21FF66" box="[1037,1258,223,249]" country="Vietnam" name="Ho Chi Minh City" pageId="75" pageNumber="76">Ho Chi Minh City</collectingRegion>
, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E21FFF72FA53FF66" box="[1328,1432,223,249]" pageId="75" pageNumber="76">H. Dung</collectorName>
, 
<date id="D63195BB4B0FFFC3E7B8FEAFFED2FE82" box="[151,281,258,285]" pageId="75" pageNumber="76" value="1995-12-08">
<collectingDate id="C6756C534B0FFFC3E7B8FEAFFED2FE82" box="[151,281,258,285]" pageId="75" pageNumber="76" value="1995-12-08">12.08.1995</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B0FFFC3E608FEAEFE2CFEDE" ID-GBIF-Occurrence="4104078462" collectingDate="1995-12-08" collectorName="Duyen-Hai &amp; H. Dung" country="Vietnam" location="Ho Chi Minh City" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Ho Chi Minh City">
<specimenCount id="B48978F24B0FFFC3E608FEAEFEBEFE82" box="[295,373,259,285]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E688FEAEFDC4FE82" box="[423,527,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="75" pageNumber="76" unit="mm" value="10.4">10.4 mm</quantity>
, minor cheliped abnormal, see below), MNHN-IU-2019-2290, same collection data as for previous specimen
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FEE6FB7CFE16" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<materialsCitation id="12E7B9264B0FFFC3E7E8FEE6FB78FE16" ID-GBIF-Occurrence="4104078392" collectingDate="2004-04-12" collectionCode="NTM" collectorName="S. K. Horner &amp; G. M. Dally" country="Australia" county="Arnhem Land" elevation="2" latitude="-12.056483" location="Nilpaiwa Islands" longLatPrecision="1" longitude="134.88058" municipality="Milingimbi" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Northern Territory">
<emphasis id="90FB6F694B0FFFC3E7E8FEE6FEE4FEFA" box="[199,303,331,357]" italics="true" pageId="75" pageNumber="76">
<collectingCountry id="DA98F3EB4B0FFFC3E7E8FEE6FEE4FEFA" box="[199,303,331,357]" name="Australia" pageId="75" pageNumber="76">Australia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0FFFC3E612FEE6FE4DFEFA" box="[317,390,331,357]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E680FEE7FDCCFEFA" box="[431,519,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="75" pageNumber="76" unit="mm" value="6.5">6.5 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E536FEE6FD9CFEFA" box="[537,599,331,357]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="75" pageNumber="76">NTM</collectionCode>
Cr.015100, 
<collectingRegion id="604B7D994B0FFFC3E5F1FEE6FC67FEFA" box="[734,940,331,357]" country="Australia" name="Northern Territory" pageId="75" pageNumber="76">Northern Territory</collectingRegion>
, 
<collectingCounty id="4B51CBF74B0FFFC3E49AFEE6FB99FEFA" box="[949,1106,331,357]" pageId="75" pageNumber="76">Arnhem Land</collectingCounty>
, 
<collectingMunicipality id="425429014B0FFFC3E373FEE6FB14FEFA" box="[1116,1247,331,357]" pageId="75" pageNumber="76">Milingimbi</collectingMunicipality>
, east of 
<location id="A750E5A04B0FFFC3E219FEE6FF22FE16" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E219FEE6FF22FE16" country="Australia" county="Arnhem Land" latitude="-12.056483" longLatPrecision="1" longitude="134.88058" municipality="Milingimbi" name="Nilpaiwa Islands" pageId="75" pageNumber="76" stateProvince="Northern Territory">Nilpaiwa Islands</location>
, 
<geoCoordinate id="C7BBD5BC4B0FFFC3E7DAFEC2FEB4FE16" box="[245,383,367,393]" degrees="12" direction="south" minutes="03.389" orientation="latitude" pageId="75" pageNumber="76" precision="1" value="-12.056483">12º03.389’S</geoCoordinate>
<geoCoordinate id="C7BBD5BC4B0FFFC3E6A9FEC3FDE8FE16" box="[390,547,366,393]" degrees="134" direction="east" minutes="52.835" orientation="longitude" pageId="75" pageNumber="76" precision="1" value="134.88058">134º52.835’E</geoCoordinate>
, depth: 
<quantity id="65771E9E4B0FFFC3E554FEC2FD76FE16" box="[635,701,367,393]" metricMagnitude="0" metricUnit="m" metricValue="2.2" pageId="75" pageNumber="76" unit="m" value="2.2">
<elevation id="29A254484B0FFFC3E554FEC2FD76FE16" box="[635,701,367,393]" metricMagnitude="0" metricUnit="m" metricValue="2.2" pageId="75" pageNumber="76" unit="m" value="2.2">2.2 m</elevation>
</quantity>
, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E5D6FEC2FC4AFE16" box="[761,897,367,393]" pageId="75" pageNumber="76">S.K. Horner</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B0FFFC3E48AFEC2FBEDFE16" box="[933,1062,367,393]" pageId="75" pageNumber="76">G.M. Dally</collectorName>
, 
<date id="D63195BB4B0FFFC3E31DFEC2FB78FE16" box="[1074,1203,367,393]" pageId="75" pageNumber="76" value="2004-04-12">
<collectingDate id="C6756C534B0FFFC3E31DFEC2FB78FE16" box="[1074,1203,367,393]" pageId="75" pageNumber="76" value="2004-04-12">04.12.2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FE3FFE2DFE6A" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<emphasis id="90FB6F694B0FFFC3E7E8FE3FFE1FFE33" bold="true" box="[199,468,402,428]" pageId="75" pageNumber="76">Comparative material</emphasis>
. 
<taxonomicName id="658FC8F84B0FFFC3E6C9FE3EFC6CFE32" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[486,935,402,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" sensu="lato" species="malabaricus">
<emphasis id="90FB6F694B0FFFC3E6C9FE3EFD1CFE32" box="[486,727,403,429]" italics="true" pageId="75" pageNumber="76">Alpheus malabaricus</emphasis>
(
<bibRefCitation id="C61ECE8A4B0FFFC3E5C5FE3EFC54FE32" author="Fabricius, J. C." box="[746,927,402,429]" pageId="75" pageNumber="76" refId="ref71244" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens Insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Officina Libraria Kortii, Flensbergi et Lipsiae, xxx + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
)
</taxonomicName>
<emphasis id="90FB6F694B0FFFC3E49DFE3EFBECFE32" box="[946,1063,403,429]" italics="true" pageId="75" pageNumber="76">
<taxonomicNameLabel id="8BC8D2124B0FFFC3E49DFE3EFBECFE32" box="[946,1063,403,429]" pageId="75" pageNumber="76" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
(
<emphasis id="90FB6F694B0FFFC3E314FE3EFBB2FE33" box="[1083,1145,403,428]" italics="true" pageId="75" pageNumber="76">sensu</emphasis>
<bibRefCitation id="C61ECE8A4B0FFFC3E3ABFE3EFA5CFE32" author="Banner, D. M. &amp; Banner, A. H." box="[1156,1431,403,429]" pageId="75" pageNumber="76" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B0FFFC3E7B8FE1AFED5FE4E" author="Chace, F. A. Jr." box="[151,286,439,465]" pageId="75" pageNumber="76" pagination="1 - 99" refId="ref69928" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
; but see discussion below). 
<materialsCitation id="12E7B9264B0FFFC3E561FE1AFE28FE6A" ID-GBIF-Occurrence="4104078410" collectionCode="OUMNH, ZC, T" collectorName="Chang Hua &amp; Y. Chan" country="Taiwan" location="Taiwan" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1">
<emphasis id="90FB6F694B0FFFC3E561FE1AFD54FE4F" box="[590,671,439,464]" italics="true" pageId="75" pageNumber="76">
<collectingCountry id="DA98F3EB4B0FFFC3E561FE1AFD54FE4F" box="[590,671,439,464]" name="Taiwan" pageId="75" pageNumber="76">Taiwan</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0FFFC3E582FE1AFD3CFE4E" box="[685,759,439,465]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E430FE1AFCB3FE4E" box="[799,888,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="75" pageNumber="76" unit="mm" value="7.1">7.1 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E4A6FE1AFC32FE4E" box="[905,1017,439,465]" pageId="75" pageNumber="76">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0FFFC3E4D3FE1AFBEAFE4E" box="[1020,1057,439,465]" country="Austria" name="Zoological Collection, University of Vienna" pageId="75" pageNumber="76" type="University or college">ZC</collectionCode>
. 2019.06.51, 
<collectorName id="0F7AD6AD4B0FFFC3E39AFE1AFAFFFE4E" box="[1205,1332,439,465]" pageId="75" pageNumber="76">Chang Hua</collectorName>
, mudflat, leg. 
<collectionCode id="C49E2BBE4B0FFFC3E7E8FE76FF12FE6A" box="[199,217,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="75" pageNumber="76">T</collectionCode>
. 
<collectorName id="0F7AD6AD4B0FFFC3E7F3FE76FEFCFE6A" box="[220,311,475,501]" pageId="75" pageNumber="76">Y. Chan</collectorName>
<emphasis id="90FB6F694B0FFFC3E611FE76FEBDFE6A" box="[318,374,475,501]" italics="true" pageId="75" pageNumber="76">et al.</emphasis>
, 07.1996
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FE52FBEEFD1A" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<materialsCitation id="12E7B9264B0FFFC3E7E8FE52FED2FDA2" ID-GBIF-Occurrence="4104078417" collectingDate="2013-08-17" collectionCode="OUMNH, ZC" collectorName="S. Hajisamae" country="Thailand" location="Lake Thale Sap" pageId="75" pageNumber="76" specimenCount="1" specimenCount-female="1" stateProvince="Songkhla">
<emphasis id="90FB6F694B0FFFC3E7E8FE52FEE7FD86" box="[199,300,511,537]" italics="true" pageId="75" pageNumber="76">
<collectingCountry id="DA98F3EB4B0FFFC3E7E8FE52FEE7FD86" box="[199,300,511,537]" name="Thailand" pageId="75" pageNumber="76">Thailand</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0FFFC3E613FE52FE56FD86" box="[316,413,511,537]" pageId="75" pageNumber="76" type="female">1 female</specimenCount>
(cl 12.0 mm), 
<collectionCode id="C49E2BBE4B0FFFC3E569FE52FD7DFD86" box="[582,694,511,537]" pageId="75" pageNumber="76">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0FFFC3E596FE52FD15FD86" box="[697,734,511,537]" country="Austria" name="Zoological Collection, University of Vienna" pageId="75" pageNumber="76" type="University or college">ZC</collectionCode>
. 2019.06.52, 
<location id="A750E5A04B0FFFC3E459FE52FBEDFD86" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E459FE52FBEDFD86" box="[886,1062,511,537]" country="Thailand" name="Lake Thale Sap" pageId="75" pageNumber="76" stateProvince="Songkhla">Lake Thale Sap</location>
(= 
<collectingRegion id="604B7D994B0FFFC3E361FE52FB77FD86" box="[1102,1212,511,537]" country="Thailand" name="Songkhla" pageId="75" pageNumber="76">Songkhla</collectingRegion>
), leg. 
<collectorName id="0F7AD6AD4B0FFFC3E22EFE52FA53FD86" box="[1281,1432,511,537]" pageId="75" pageNumber="76">S. Hajisamae</collectorName>
, 
<date id="D63195BB4B0FFFC3E7B8FD8EFED2FDA2" box="[151,281,547,573]" pageId="75" pageNumber="76" value="2013-08-17">
<collectingDate id="C6756C534B0FFFC3E7B8FD8EFED2FDA2" box="[151,281,547,573]" pageId="75" pageNumber="76" value="2013-08-17">17.08.2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B0FFFC3E60BFD8EFA94FDA2" ID-GBIF-Occurrence="4104078519" box="[292,1375,546,573]" collectingDate="2008-04-03" collectionCode="OUMNH, ZC" collectorName="S. Hajisamae" country="Thailand" elevation="5" location="Bay" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Pattani">
<specimenCount id="B48978F24B0FFFC3E60BFD8EFEA4FDA2" box="[292,367,547,573]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E6B5FD8EFDCAFDA2" box="[410,513,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="75" pageNumber="76" unit="mm" value="11.7">11.7 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E53BFD8EFD4FFDA2" box="[532,644,547,573]" pageId="75" pageNumber="76">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0FFFC3E5A8FD8EFD67FDA2" box="[647,684,547,573]" country="Austria" name="Zoological Collection, University of Vienna" pageId="75" pageNumber="76" type="University or college">ZC</collectionCode>
. 2019.06.53, 
<collectingRegion id="604B7D994B0FFFC3E46DFD8EFC5BFDA2" box="[834,912,547,573]" country="Thailand" name="Pattani" pageId="75" pageNumber="76">Pattani</collectingRegion>
<location id="A750E5A04B0FFFC3E4B9FD8EFC0FFDA2" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E4B9FD8EFC0FFDA2" box="[918,964,547,573]" country="Thailand" name="Bay" pageId="75" pageNumber="76" stateProvince="Pattani">Bay</location>
, Pattani, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E374FD8EFB39FDA2" box="[1115,1266,547,573]" pageId="75" pageNumber="76">S. Hajisamae</collectorName>
, 01.2013;
</materialsCitation>
<materialsCitation id="12E7B9264B0FFFC3E249FD8EFAA9FDFE" ID-GBIF-Occurrence="4104078433" collectingDate="2008-04-03" collectionCode="OUMNH, ZC" collectorName="S. Hajisamae" country="Thailand" elevation="5" location="Bay" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Pattani">
1 ov. female (cl 
<quantity id="65771E9E4B0FFFC3E63FFDEBFEA0FDFE" box="[272,363,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="75" pageNumber="76" unit="mm" value="9.5">9.5 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E651FDEAFE25FDFE" box="[382,494,583,609]" pageId="75" pageNumber="76">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0FFFC3E6DEFDEAFDDDFDFE" box="[497,534,583,609]" country="Austria" name="Zoological Collection, University of Vienna" pageId="75" pageNumber="76" type="University or college">ZC</collectionCode>
. 2019.06.54, 
<collectingRegion id="604B7D994B0FFFC3E581FDEAFD34FDFE" box="[686,767,583,609]" country="Thailand" name="Phuket" pageId="75" pageNumber="76">Phuket</collectingRegion>
, E 
<location id="A750E5A04B0FFFC3E40CFDEAFC73FDFE" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E40CFDEAFC73FDFE" box="[803,952,583,609]" country="Thailand" name="Chalong Bay" pageId="75" pageNumber="76" stateProvince="Phuket">Chalong Bay</location>
, mudflat in front of mangrove, depth:
</materialsCitation>
<materialsCitation id="12E7B9264B0FFFC3E246FDEBFBEEFD1A" ID-GBIF-Occurrence="4104078555" collectingDate="2008-04-03" collectionCode="OUMNH, ZC" collectorName="S. Hajisamae" country="Thailand" elevation="5" location="Bay" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Pattani">
&lt;
<quantity id="65771E9E4B0FFFC3E256FDEBFF64FD1A" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="75" pageNumber="76" unit="m" value="0.5">
<elevation id="29A254484B0FFFC3E256FDEBFF64FD1A" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="75" pageNumber="76" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, suction pump, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E6A0FDC6FE33FD1A" box="[399,504,619,645]" pageId="75" pageNumber="76">A. Anker</collectorName>
, 
<collectorName id="0F7AD6AD4B0FFFC3E52CFDC6FDB9FD1A" box="[515,626,619,645]" pageId="75" pageNumber="76">J.C.Y. Lai</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B0FFFC3E5B9FDC6FD29FD1A" box="[662,738,619,645]" pageId="75" pageNumber="76">M. Ng</collectorName>
, 
<date id="D63195BB4B0FFFC3E5C2FDC6FCA7FD1A" box="[749,876,619,645]" pageId="75" pageNumber="76" value="2008-04-03">
<collectingDate id="C6756C534B0FFFC3E5C2FDC6FCA7FD1A" box="[749,876,619,645]" pageId="75" pageNumber="76" value="2008-04-03">04.03.2008</collectingDate>
</date>
[
<collectingCountry id="DA98F3EB4B0FFFC3E456FDC6FC69FD1A" box="[889,930,619,645]" name="Philippines" pageId="75" pageNumber="76">PH</collectingCountry>
12-08-036].
</materialsCitation>
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FD22FAE6FB8E" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<emphasis id="90FB6F694B0FFFC3E7E8FD22FEFEFD36" box="[199,309,655,681]" italics="true" pageId="75" pageNumber="76">
<collectingCountry id="DA98F3EB4B0FFFC3E7E8FD22FEFEFD36" box="[199,309,655,681]" name="Indonesia" pageId="75" pageNumber="76">Indonesia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0FFFC3E66BFD22FE45FD36" box="[324,398,655,681]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E695FD22FDEBFD36" box="[442,544,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="75" pageNumber="76" unit="mm" value="11.7">11.7 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E51CFD22FD68FD36" box="[563,675,655,681]" pageId="75" pageNumber="76">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0FFFC3E58AFD22FD01FD36" box="[677,714,655,681]" country="Austria" name="Zoological Collection, University of Vienna" pageId="75" pageNumber="76" type="University or college">ZC</collectionCode>
. 2019.06.55, 
<collectingRegion id="604B7D994B0FFFC3E44FFD22FC6CFD36" box="[864,935,655,681]" country="Indonesia" name="Papua Barat" pageId="75" pageNumber="76">Papua</collectingRegion>
, Ajkwa Island, mangrove, leg. A. Darmawan 
<emphasis id="90FB6F694B0FFFC3E7B8FD1EFF07FD52" box="[151,204,691,717]" italics="true" pageId="75" pageNumber="76">et al</emphasis>
., 
<date id="D63195BB4B0FFFC3E7CAFD1EFEACFD52" box="[229,359,691,717]" pageId="75" pageNumber="76" value="2012-07-20">20.07.2012</date>
; 1 ov. female (cl 
<quantity id="65771E9E4B0FFFC3E516FD1EFD6EFD52" box="[569,677,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="75" pageNumber="76" unit="mm" value="12.4">12.4 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E593FD1EFD22FD52" box="[700,745,691,717]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="75" pageNumber="76" type="Museum">QM</collectionCode>
W25803-1, 
<collectingRegion id="604B7D994B0FFFC3E452FD1EFC0FFD52" box="[893,964,691,717]" country="Indonesia" name="Papua Barat" pageId="75" pageNumber="76">Papua</collectingRegion>
, Ajkwa River estuary, 
<geoCoordinate id="C7BBD5BC4B0FFFC3E3FEFD1FFAD5FD52" box="[1233,1310,690,717]" degrees="4" direction="south" minutes="50" orientation="latitude" pageId="75" pageNumber="76" precision="925" value="-4.8333335">4°50’S</geoCoordinate>
<geoCoordinate id="C7BBD5BC4B0FFFC3E205FD1FFA5CFD52" box="[1322,1431,690,717]" degrees="136" direction="east" minutes="50" orientation="longitude" pageId="75" pageNumber="76" precision="925" value="136.83333">136°50’E</geoCoordinate>
, Environmental Laboratory, PT Freeport, Sta. Ajk-36, estuarine habitat, 
<date id="D63195BB4B0FFFC3E4E3FD7AFB85FD6E" box="[972,1102,727,753]" pageId="75" pageNumber="76" value="2000-03-30">30.03.2000</date>
; 
<specimenCount id="B48978F24B0FFFC3E373FD7AFB7EFD6E" box="[1116,1205,727,753]" pageId="75" pageNumber="76" type="male">3 males</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E3C9FD7AFA5BFD6E" box="[1254,1424,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.2399999999999998" metricValueMax="1.46" metricValueMin="1.02" pageId="75" pageNumber="76" unit="mm" value="12.399999999999999" valueMax="14.6" valueMin="10.2">10.2–14.6 mm</quantity>
), 
<specimenCount id="B48978F24B0FFFC3E7B8FD56FF37FC8A" box="[151,252,763,789]" pageId="75" pageNumber="76" type="female">1 female</specimenCount>
(cl 11.0 mm), 
<collectionCode id="C49E2BBE4B0FFFC3E681FD56FE10FC8A" box="[430,475,763,789]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="75" pageNumber="76" type="Museum">QM</collectionCode>
W25802, 
<collectingRegion id="604B7D994B0FFFC3E578FD56FD55FC8A" box="[599,670,763,789]" country="Indonesia" name="Papua Barat" pageId="75" pageNumber="76">Papua</collectingRegion>
, West Ajkwa River, river mouth at Lanal Base, 
<geoCoordinate id="C7BBD5BC4B0FFFC3E3FEFD57FAD4FC8A" box="[1233,1311,762,789]" degrees="4" direction="south" minutes="50" orientation="latitude" pageId="75" pageNumber="76" precision="925" value="-4.8333335">4°50’S</geoCoordinate>
<geoCoordinate id="C7BBD5BC4B0FFFC3E205FD57FA5CFC8A" box="[1322,1431,762,789]" degrees="136" direction="east" minutes="50" orientation="longitude" pageId="75" pageNumber="76" precision="925" value="136.83333">136°50’E</geoCoordinate>
, Environmental Laboratory, PT Freeport, Sta. Ajk-36, estuarine habitat, 
<date id="D63195BB4B0FFFC3E4EAFCB2FB8CFCA6" box="[965,1095,799,825]" pageId="75" pageNumber="76" value="1999-07-29">29.07.1999</date>
; 1 ov. female (cl 
<quantity id="65771E9E4B0FFFC3E23FFCB2FAB2FCA6" box="[1296,1401,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="75" pageNumber="76" unit="mm" value="11.6">11.6 mm</quantity>
), 
<specimenCount id="B48978F24B0FFFC3E2A1FCB2FF06FCC2" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
major cheliped, 
<collectionCode id="C49E2BBE4B0FFFC3E6A5FCEEFE31FCC2" box="[394,506,835,861]" pageId="75" pageNumber="76">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0FFFC3E6D2FCEEFDE9FCC2" box="[509,546,835,861]" country="Austria" name="Zoological Collection, University of Vienna" pageId="75" pageNumber="76" type="University or college">ZC</collectionCode>
. 2019.06.56, Lombok, Lembar, prawn ponds, muddy banks of brackish stream, low tide, suction pump, leg. A. Anker, I.S. Pratama, M. Firdaus &amp; D.L. Rahayu, 
<date id="D63195BB4B0FFFC3E33BFCCBFB59FC1E" box="[1044,1170,870,897]" pageId="75" pageNumber="76" value="2014-05-14">14.05.2014</date>
[St5-14]; 
<specimenCount id="B48978F24B0FFFC3E22BFCCAFA85FC1E" box="[1284,1358,871,897]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E256FCCAFF0CFC3A" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="75" pageNumber="76" unit="mm" value="9.4">9.4 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B0FFFC3E6BEFC26FE20FC3A" box="[401,491,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="75" pageNumber="76" unit="mm" value="7.8">7.8 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E6D0FC26FDA4FC3A" box="[511,623,907,933]" pageId="75" pageNumber="76">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B0FFFC3E55EFC26FD5DFC3A" box="[625,662,907,933]" country="Austria" name="Zoological Collection, University of Vienna" pageId="75" pageNumber="76" type="University or college">ZC</collectionCode>
. 2019.06.57, Lombok, Teluk Sekotong, near mangrove, 
<quantity id="65771E9E4B0FFFC3E220FC26FA4AFC3A" box="[1295,1409,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="75" pageNumber="76" unit="m" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 m</quantity>
at low tide, suction pump, leg. A. Anker, I.S. Pratama, M. Firdaus &amp; D.L. Rahayu, 
<date id="D63195BB4B0FFFC3E30EFC03FB55FC56" box="[1057,1182,942,969]" pageId="75" pageNumber="76" value="2014-05-14">14.05.2014</date>
[St6-03]; 1 ov. female (cl indet.), 
<collectionCode id="C49E2BBE4B0FFFC3E639FC7EFE87FC72" box="[278,332,979,1005]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="75" pageNumber="76">ZRC</collectionCode>
2014.0682, Anambas, sta. 
<collectionCode id="C49E2BBE4B0FFFC3E5ABFC7EFD02FC72" box="[644,713,979,1005]" pageId="75" pageNumber="76">EAJL</collectionCode>
03, Pulau Jemaja Teluk, deep sheltered sandy bay, northern and eastern sides with fringing 
<taxonomicName id="658FC8F84B0FFFC3E6E9FC5AFD82FB8E" box="[454,585,1015,1041]" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" kingdom="Plantae" order="Malpighiales" pageId="75" pageNumber="76" phylum="Tracheophyta" rank="genus">
<emphasis id="90FB6F694B0FFFC3E6E9FC5AFD82FB8E" box="[454,585,1015,1041]" italics="true" pageId="75" pageNumber="76">Rhizophora</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B0FFFC3E5AFFC5AFD38FB8F" box="[640,755,1015,1040]" class="Magnoliopsida" family="Rhizophoraceae" genus="Bruguiera" kingdom="Plantae" order="Malpighiales" pageId="75" pageNumber="76" phylum="Tracheophyta" rank="genus">
<emphasis id="90FB6F694B0FFFC3E5AFFC5AFD38FB8F" box="[640,755,1015,1040]" italics="true" pageId="75" pageNumber="76">Bruguiera</emphasis>
</taxonomicName>
mangrove inlets, leg. J.C.Y. Lai 
<emphasis id="90FB6F694B0FFFC3E34DFC5AFB58FB8E" box="[1122,1171,1015,1041]" italics="true" pageId="75" pageNumber="76">et al</emphasis>
., 
<date id="D63195BB4B0FFFC3E387FC5AFAE2FB8E" box="[1192,1321,1015,1041]" pageId="75" pageNumber="76" value="2002-03-13">13.03.2002</date>
.
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FBB6FEA3FB3E" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<materialsCitation id="12E7B9264B0FFFC3E7E8FBB6FE6BFBC6" ID-GBIF-Occurrence="4104078471" collectionCode="ZRC" collectorName="Pulau Ubin &amp; P. K. L. Ng" country="Singapore" location="Singapore" pageId="75" pageNumber="76" specimenCode="ZRC 1992.11124, 11126" specimenCount="2" specimenCount-male="2">
<emphasis id="90FB6F694B0FFFC3E7E8FBB6FEF2FBAA" box="[199,313,1051,1077]" italics="true" pageId="75" pageNumber="76">
<collectingCountry id="DA98F3EB4B0FFFC3E7E8FBB6FEF2FBAA" box="[199,313,1051,1077]" name="Singapore" pageId="75" pageNumber="76">Singapore</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0FFFC3E663FBB6FE6DFBAA" box="[332,422,1051,1077]" pageId="75" pageNumber="76" type="male">2 males</specimenCount>
(cl 15.4, 
<quantity id="65771E9E4B0FFFC3E531FBB7FD41FBAA" box="[542,650,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="75" pageNumber="76" unit="mm" value="15.9">15.9 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B0FFFC3E449FBB6FC18FBAA" box="[870,979,1051,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="75" pageNumber="76" unit="mm" value="16.7">16.7 mm</quantity>
), 
<specimenCode id="F2291B004B0FFFC3E4C5FBB6FB7BFBAA" box="[1002,1200,1051,1077]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="75" pageNumber="76">ZRC 1992.11124</specimenCode>
– 
<specimenCode id="F2291B004B0FFFC3E390FBB6FAC8FBAA" box="[1215,1283,1051,1077]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="75" pageNumber="76">11126</specimenCode>
, 
<collectorName id="0F7AD6AD4B0FFFC3E23DFBB6FA53FBAA" box="[1298,1432,1051,1077]" pageId="75" pageNumber="76">Pulau Ubin</collectorName>
, mudflat, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E602FB92FE6BFBC6" box="[301,416,1087,1113]" pageId="75" pageNumber="76">P.K.L. Ng</collectorName>
</materialsCitation>
, 08.1987; 
<materialsCitation id="12E7B9264B0FFFC3E53AFB92FEB3FBE2" ID-GBIF-Occurrence="4104078539" collectionCode="ZRC" collectorName="Pulau Ubin &amp; P. K. L. Ng" country="Singapore" location="Singapore" pageId="75" pageNumber="76" specimenCode="ZRC 1992.11122, 11123" specimenCount="2" specimenCount-male="2">
<specimenCount id="B48978F24B0FFFC3E53AFB92FDA0FBC6" box="[533,619,1087,1113]" pageId="75" pageNumber="76" type="male">2 males</specimenCount>
(cl 12.4, 
<quantity id="65771E9E4B0FFFC3E5F9FB93FCF5FBC6" box="[726,830,1086,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="75" pageNumber="76" unit="mm" value="12.5">12.5 mm</quantity>
), 
<specimenCode id="F2291B004B0FFFC3E47EFB92FBDFFBC6" box="[849,1044,1087,1113]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="75" pageNumber="76">ZRC 1992.11122</specimenCode>
– 
<specimenCode id="F2291B004B0FFFC3E30CFB92FBADFBC6" box="[1059,1126,1087,1113]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="75" pageNumber="76">11123</specimenCode>
, same collection data as for previous specimens
</materialsCitation>
; 
<materialsCitation id="12E7B9264B0FFFC3E6ABFBCEFEAFFB3E" ID-GBIF-Occurrence="4104078393" collectingDate="1994-12-16" collectionCode="ZRC" collectorName="Reef Ecology Study Team" country="Singapore" location="Kallang Basin" pageId="75" pageNumber="76" specimenCode="ZRC 1994.4392" specimenCount="1" specimenCount-male="1">
<specimenCount id="B48978F24B0FFFC3E6ABFBCEFE1BFBE2" box="[388,464,1123,1149]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 9.0 mm), 
<specimenCode id="F2291B004B0FFFC3E542FBCEFCEDFBE2" box="[621,806,1123,1149]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="75" pageNumber="76">ZRC 1994.4392</specimenCode>
, 
<location id="A750E5A04B0FFFC3E41DFBCEFC1EFBE2" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E41DFBCEFC1EFBE2" box="[818,981,1123,1149]" country="Singapore" name="Kallang Basin" pageId="75" pageNumber="76">Kallang Basin</location>
, sta. 5, dredge, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E396FBCEFF13FB3E" pageId="75" pageNumber="76">Reef Ecology Study Team</collectorName>
, 
<date id="D63195BB4B0FFFC3E7CCFB2AFEAFFB3E" box="[227,356,1159,1185]" pageId="75" pageNumber="76" value="1994-12-16">
<collectingDate id="C6756C534B0FFFC3E7CCFB2AFEAFFB3E" box="[227,356,1159,1185]" pageId="75" pageNumber="76" value="1994-12-16">16.12.1994</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FB06FE5DFAAE" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<materialsCitation id="12E7B9264B0FFFC3E7E8FB06FD45FB76" ID-GBIF-Occurrence="4104078390" collectingDate="2000-12-05" collectionCode="QM" collectorName="J. Johnson &amp; A. Gill" country="Australia" elevation="300" location="Queensland" pageId="75" pageNumber="76" specimenCode="W25812" specimenCount="1" specimenCount-male="1" stateProvince="Queensland">
<emphasis id="90FB6F694B0FFFC3E7E8FB06FEE4FB5A" box="[199,303,1195,1221]" italics="true" pageId="75" pageNumber="76">
<collectingCountry id="DA98F3EB4B0FFFC3E7E8FB06FEE4FB5A" box="[199,303,1195,1221]" name="Australia" pageId="75" pageNumber="76">Australia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B0FFFC3E611FB06FE41FB5A" box="[318,394,1195,1221]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B0FFFC3E523FB06FDF2FB5A" box="[524,569,1195,1221]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="75" pageNumber="76" type="Museum">QM</collectionCode>
<specimenCode id="F2291B004B0FFFC3E56FFB07FD68FB5A" box="[576,675,1194,1221]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="75" pageNumber="76" type="Museum">W25812</specimenCode>
, SE 
<collectingRegion id="604B7D994B0FFFC3E5F9FB06FC96FB5A" box="[726,861,1195,1221]" country="Australia" name="Queensland" pageId="75" pageNumber="76">Queensland</collectingRegion>
, 
<collectorName id="0F7AD6AD4B0FFFC3E446FB06FC28FB5A" box="[873,995,1195,1221]" pageId="75" pageNumber="76">Innes Park</collectorName>
, creek, 
<quantity id="65771E9E4B0FFFC3E316FB06FB4BFB5A" box="[1081,1152,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="75" pageNumber="76" unit="m" value="300.0">
<elevation id="29A254484B0FFFC3E316FB06FB4BFB5A" box="[1081,1152,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="75" pageNumber="76" unit="m" value="300.0">300 m</elevation>
</quantity>
from mouth, mangroves, sand, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E621FB62FE4EFB76" box="[270,389,1231,1257]" pageId="75" pageNumber="76">J. Johnson</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B0FFFC3E683FB62FE35FB76" box="[428,510,1231,1257]" pageId="75" pageNumber="76">A. Gill</collectorName>
, 
<date id="D63195BB4B0FFFC3E523FB63FD45FB76" box="[524,654,1230,1257]" pageId="75" pageNumber="76" value="2000-12-05">
<collectingDate id="C6756C534B0FFFC3E523FB63FD45FB76" box="[524,654,1230,1257]" pageId="75" pageNumber="76" value="2000-12-05">12.05.2000</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B0FFFC3E5B3FB62FE59FAAE" ID-GBIF-Occurrence="4104078343" collectingDate="1991-04-06" collectionCode="NTM" collectorName="M. Burke" country="Australia" latitude="-12.413333" location="Ludmilla Creek" longLatPrecision="129" longitude="130.845" pageId="75" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Northern Territory">
<specimenCount id="B48978F24B0FFFC3E5B3FB62FD22FB76" box="[668,745,1231,1257]" pageId="75" pageNumber="76" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B0FFFC3E435FB62FC4FFB76" box="[794,900,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="75" pageNumber="76" unit="mm" value="13.4">13.4 mm</quantity>
), 1 ov. female (cl 
<quantity id="65771E9E4B0FFFC3E377FB62FB08FB76" box="[1112,1219,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="75" pageNumber="76" unit="mm" value="14.9">14.9 mm</quantity>
), 
<collectionCode id="C49E2BBE4B0FFFC3E3F7FB62FADDFB76" box="[1240,1302,1231,1257]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="75" pageNumber="76">NTM</collectionCode>
<collectorName id="0F7AD6AD4B0FFFC3E20FFB62FA8AFB76" box="[1312,1345,1231,1257]" pageId="75" pageNumber="76">Cr.</collectorName>
008307, 
<collectingRegion id="604B7D994B0FFFC3E7B8FB5EFEA3FA92" box="[151,360,1267,1293]" country="Australia" name="Northern Territory" pageId="75" pageNumber="76">Northern Territory</collectingRegion>
, 
<collectorName id="0F7AD6AD4B0FFFC3E65AFB5EFE07FA92" box="[373,460,1267,1293]" pageId="75" pageNumber="76">Darwin</collectorName>
, 
<location id="A750E5A04B0FFFC3E6F7FB5EFD41FA92" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E6F7FB5EFD41FA92" box="[472,650,1267,1293]" country="Australia" latitude="-12.413333" longLatPrecision="129" longitude="130.845" name="Ludmilla Creek" pageId="75" pageNumber="76" stateProvince="Northern Territory">Ludmilla Creek</location>
mouth, 
<geoCoordinate id="C7BBD5BC4B0FFFC3E5C6FB5EFC90FA92" box="[745,859,1267,1293]" degrees="12" direction="south" minutes="24.8" orientation="latitude" pageId="75" pageNumber="76" precision="92" value="-12.413333">12º24.8’S</geoCoordinate>
, 
<geoCoordinate id="C7BBD5BC4B0FFFC3E448FB5FFC25FA92" box="[871,1006,1266,1293]" degrees="130" direction="east" minutes="50.7" orientation="longitude" pageId="75" pageNumber="76" precision="92" value="130.845">130º50.7’ E</geoCoordinate>
, undisturbed mangrove, low tide, leg. 
<collectorName id="0F7AD6AD4B0FFFC3E7B8FABAFECEFAAE" box="[151,261,1303,1329]" pageId="75" pageNumber="76">M. Burke</collectorName>
, 
<date id="D63195BB4B0FFFC3E63EFABAFE59FAAE" box="[273,402,1303,1329]" pageId="75" pageNumber="76" value="1991-04-06">
<collectingDate id="C6756C534B0FFFC3E63EFABAFE59FAAE" box="[273,402,1303,1329]" pageId="75" pageNumber="76" value="1991-04-06">04.06.1991</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B0FFFC3E7E8FA97FA5CF9EA" pageId="75" pageNumber="76" type="description">
<paragraph id="A230B37B4B0FFFC3E7E8FA97FE77FAE6" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<emphasis id="90FB6F694B0FFFC3E7E8FA97FE9AFACB" bold="true" box="[199,337,1338,1364]" pageId="75" pageNumber="76">Description</emphasis>
. See 
<bibRefCitation id="C61ECE8A4B0FFFC3E6A2FA96FD69FACA" author="Banner, A. H. &amp; Banner, D. M." box="[397,674,1339,1365]" pageId="75" pageNumber="76" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
for original description and illustrations, as 
<taxonomicName id="658FC8F84B0FFFC3E3A2FA96FA5FFACA" authorityName="Banner &amp; Banner" authorityYear="1966" box="[1165,1428,1339,1365]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B0FFFC3E3A2FA96FA5FFACA" box="[1165,1428,1339,1365]" italics="true" pageId="75" pageNumber="76">A. malabaricus songkla</emphasis>
</taxonomicName>
; see also discussion below.
</paragraph>
<paragraph id="A230B37B4B0FFFC3E7E8FA2FFA5CF9EA" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<emphasis id="90FB6F694B0FFFC3E7E8FA2FFEB1FA02" bold="true" box="[199,378,1410,1437]" pageId="75" pageNumber="76">Colour pattern</emphasis>
. The colour pattern of the 
<typeStatus id="7D340DD94B0FFFC3E58DFA2EFD19FA02" box="[674,722,1411,1437]" pageId="75" pageNumber="76">type</typeStatus>
specimens from 
<collectingRegion id="604B7D994B0FFFC3E4BEFA2EFC30FA02" box="[913,1019,1411,1437]" country="Thailand" name="Songkhla" pageId="75" pageNumber="76">Songkhla</collectingRegion>
Lake is unknown. The colour pattern of the Indian specimens identified by 
<bibRefCitation id="C61ECE8A4B0FFFC3E576FA0AFCC1FA5E" author="Thomas, M. M." box="[601,778,1447,1473]" pageId="75" pageNumber="76" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas (1976)</bibRefCitation>
as 
<taxonomicName id="658FC8F84B0FFFC3E417FA0AFB80FA5E" authorityName="Banner &amp; Banner" authorityYear="1966" box="[824,1099,1447,1473]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B0FFFC3E417FA0AFB80FA5E" box="[824,1099,1447,1473]" italics="true" pageId="75" pageNumber="76">A. malabaricus songkla</emphasis>
</taxonomicName>
was described as following: general body colour cream with dark brown cross bands along posterior margins of pleonites and carapace; tips of uropods and anterior border of carapace between orbital hood and lateral angle dark brown; antennal flagella bluish violet, antennular flagella with brownish tinge; chelipeds grey with violet inner [mesial?] depression of large chela; remaining portion of chela and legs pinkish; exopods and endopods of pleopods [erroneously called chelipeds] bright red with paler bases; underside of pleon and chela white (adapted from 
<bibRefCitation id="C61ECE8A4B0FFFC3E32EF9F6FB55F9EA" author="Thomas, M. M." box="[1025,1182,1627,1653]" pageId="75" pageNumber="76" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976</bibRefCitation>
; however, see below).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B0FFFC3E7E8F9D2FCD2F906" box="[199,793,1662,1689]" pageId="75" pageNumber="76" type="materials_examined">
<paragraph id="A230B37B4B0FFFC3E7E8F9D2FCD2F906" blockId="75.[151,1437,150,2013]" box="[199,793,1662,1689]" pageId="75" pageNumber="76">
<materialsCitation id="12E7B9264B0FFFC3E7E8F9D2FCDEF906" ID-GBIF-Occurrence="4104078507" box="[199,789,1662,1689]" country="Thailand" location="Thale Sap" municipality="Lake" pageId="75" pageNumber="76" specimenCount="1" stateProvince="Songkhla" typeStatus="holotype">
<emphasis id="90FB6F694B0FFFC3E7E8F9D2FEABF907" bold="true" box="[199,352,1662,1689]" pageId="75" pageNumber="76">
<typeStatus id="7D340DD94B0FFFC3E7E8F9D2FEC9F906" box="[199,258,1663,1689]" pageId="75" pageNumber="76">Type</typeStatus>
locality
</emphasis>
. 
<location id="A750E5A04B0FFFC3E642F9D2FE15F906" LSID="urn:lsid:plazi:treatment:2A26026D4B0EFFD8E7B8FAFAFBB6FE6A:A750E5A04B0FFFC3E642F9D2FE15F906" box="[365,478,1663,1689]" country="Thailand" municipality="Lake" name="Thale Sap" pageId="75" pageNumber="76" stateProvince="Songkhla">Thale Sap</location>
(
<collectingRegion id="604B7D994B0FFFC3E6C2F9D2FD92F906" box="[493,601,1663,1689]" country="Thailand" name="Songkhla" pageId="75" pageNumber="76">Songkhla</collectingRegion>
<collectingMunicipality id="425429014B0FFFC3E570F9D2FD50F906" box="[607,667,1663,1689]" pageId="75" pageNumber="76">Lake</collectingMunicipality>
), 
<collectingCountry id="DA98F3EB4B0FFFC3E581F9D2FCDEF906" box="[686,789,1663,1689]" name="Thailand" pageId="75" pageNumber="76">Thailand</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B0FFFC3E7E8F90FFE57F8B6" pageId="75" pageNumber="76" type="distribution">
<paragraph id="A230B37B4B0FFFC3E7E8F90FFE57F8B6" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<emphasis id="90FB6F694B0FFFC3E7E8F90FFE92F923" bold="true" box="[199,345,1698,1724]" pageId="75" pageNumber="76">Distribution</emphasis>
. Indo-West Pacific: with certainty known only from the 
<typeStatus id="7D340DD94B0FFFC3E4D7F90EFBE3F922" box="[1016,1064,1699,1725]" pageId="75" pageNumber="76">type</typeStatus>
locality in 
<collectingCountry id="DA98F3EB4B0FFFC3E39DF90EFAD3F922" box="[1202,1304,1699,1725]" name="Thailand" pageId="75" pageNumber="76">Thailand</collectingCountry>
, Thale Sap = 
<collectingRegion id="604B7D994B0FFFC3E79EF96AFED0F97E" box="[177,283,1735,1761]" country="Thailand" name="Songkhla" pageId="75" pageNumber="76">Songkhla</collectingRegion>
Lake (
<bibRefCitation id="C61ECE8A4B0FFFC3E641F96AFD4AF97E" author="Banner, A. H. &amp; Banner, D. M." box="[366,641,1735,1761]" pageId="75" pageNumber="76" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
); record from 
<collectingCountry id="DA98F3EB4B0FFFC3E405F96AFCAFF97E" box="[810,868,1735,1761]" name="India" pageId="75" pageNumber="76">India</collectingCountry>
(
<bibRefCitation id="C61ECE8A4B0FFFC3E45AF96AFBDFF97E" author="Thomas, M. M." box="[885,1044,1735,1761]" pageId="75" pageNumber="76" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976</bibRefCitation>
) requires confirmation; additional records of 
<taxonomicName id="658FC8F84B0FFFC3E621F946FE6CF89A" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[270,423,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0FFFC3E621F946FEEDF89A" box="[270,294,1771,1797]" italics="true" pageId="75" pageNumber="76">A.</emphasis>
cf. 
<emphasis id="90FB6F694B0FFFC3E67FF946FE6CF89A" box="[336,423,1771,1797]" italics="true" pageId="75" pageNumber="76">songkla</emphasis>
</taxonomicName>
from 
<collectingCountry id="DA98F3EB4B0FFFC3E6C5F946FD9BF89A" box="[490,592,1771,1797]" name="Thailand" pageId="75" pageNumber="76">Thailand</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0FFFC3E574F946FD76F89A" box="[603,701,1771,1797]" name="Vietnam" pageId="75" pageNumber="76">Vietnam</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0FFFC3E5E8F946FCFBF89A" box="[711,816,1771,1797]" name="Malaysia" pageId="75" pageNumber="76">Malaysia</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0FFFC3E414F946FC66F89A" box="[827,941,1771,1797]" name="Singapore" pageId="75" pageNumber="76">Singapore</collectingCountry>
and 
<collectingRegion id="604B7D994B0FFFC3E4CDF946FB7AF89A" box="[994,1201,1771,1797]" country="Australia" name="Northern Territory" pageId="75" pageNumber="76">Northern Territory</collectingRegion>
, 
<collectingCountry id="DA98F3EB4B0FFFC3E394F946FAE9F89A" box="[1211,1314,1771,1797]" name="Australia" pageId="75" pageNumber="76">Australia</collectingCountry>
(
<figureCitation id="3AB4AFFE4B0FFFC3E21FF946FA5FF89A" box="[1328,1428,1770,1797]" captionStart="FIGURE 52" captionStartId="106.[151,250,1909,1933]" captionTargetBox="[156,1430,181,1885]" captionTargetId="figure-21@106.[156,1430,181,1885]" captionTargetPageId="106" captionText="FIGURE 52. Distributional maps for species of Alpheus treated in this study:A—A. takla sp. nov.; B—A. mangalis sp. nov. and A. cf. mangalis sp. nov.; C—A. songkla Banner &amp; Banner, 1966 and A. cf. songkla (see text); D—A. paludicola Kemp, 1915, A. nipa Banner &amp; Banner, 1985 and A. bunburius Banner &amp; Banner, 1982; E—A. pontederiae de Rochebrune, 1883; F—A. firmus Kim &amp; Abele, 1988 and A. cf. firmus (see text)." figureDoi="http://doi.org/10.5281/zenodo.7912457" httpUri="https://zenodo.org/record/7912457/files/figure.png" pageId="75" pageNumber="76">Fig. 52C</figureCitation>
) (see discussion below).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B0FFFC3E7E8F89FFCF8F8D2" box="[199,819,1842,1869]" pageId="75" pageNumber="76" type="vernacular_names">
<paragraph id="A230B37B4B0FFFC3E7E8F89FFCF8F8D2" blockId="75.[151,1437,150,2013]" box="[199,819,1842,1869]" pageId="75" pageNumber="76">
<emphasis id="90FB6F694B0FFFC3E7E8F89FFE38F8D2" bold="true" box="[199,499,1842,1869]" pageId="75" pageNumber="76">Common name proposed</emphasis>
. Songhkla snapping shrimp.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B0FFFC3E7E8F8FAFD6BF842" pageId="75" pageNumber="76" type="biology_ecology">
<paragraph id="A230B37B4B0FFFC3E7E8F8FAFD6BF842" blockId="75.[151,1437,150,2013]" pageId="75" pageNumber="76">
<emphasis id="90FB6F694B0FFFC3E7E8F8FAFE7EF8EF" bold="true" box="[199,437,1878,1905]" pageId="75" pageNumber="76">Ecology and biology</emphasis>
. Shallow subtidal species found on mud and sand bottoms usually off mangroves or inside brackish and saltwater lagoons; the South-East Asian and Australian specimens were collected in 
<quantity id="65771E9E4B0FFFC3E3F9F8D6FAD7F80A" box="[1238,1308,1915,1941]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="75" pageNumber="76" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1–3 m</quantity>
deep water (
<bibRefCitation id="C61ECE8A4B0FFFC3E7B0F832FE6CF826" author="Banner, A. H. &amp; Banner, D. M." box="[159,423,1951,1977]" pageId="75" pageNumber="76" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
; present study), whereas the Indian specimens (which may or may not be 
<taxonomicName id="658FC8F84B0FFFC3E3F3F832FA9AF826" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[1244,1361,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0FFFC3E3F3F832FA9AF826" box="[1244,1361,1951,1977]" italics="true" pageId="75" pageNumber="76">A. songkla</emphasis>
</taxonomicName>
) came from a depth range of 
<quantity id="65771E9E4B0FFFC3E6BEF86EFE2DF842" box="[401,486,1987,2013]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="75" pageNumber="76" unit="m" value="6.5" valueMax="10.0" valueMin="3.0">3–10 m</quantity>
(
<bibRefCitation id="C61ECE8A4B0FFFC3E6DBF86EFD58F842" author="Thomas, M. M." box="[500,659,1987,2013]" pageId="75" pageNumber="76" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B08FFD8E7E8FF3BFBB6FE6A" lastPageId="80" lastPageNumber="81" pageId="76" pageNumber="77" type="discussion">
<paragraph id="A230B37B4B08FFC4E7E8FF3BFBC6FEFA" blockId="76.[151,1437,150,2039]" pageId="76" pageNumber="77">
<emphasis id="90FB6F694B08FFC4E7E8FF3BFE73FF2E" bold="true" box="[199,440,150,177]" pageId="76" pageNumber="77">Taxonomic remarks</emphasis>
. 
<bibRefCitation id="C61ECE8A4B08FFC4E6E8FF3AFD28FF2E" author="Banner, A. H. &amp; Banner, D. M." box="[455,739,151,177]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
separated their new subspecies 
<taxonomicName id="658FC8F84B08FFC4E37EFF3AFA96FF2E" authorityName="Banner &amp; Banner" authorityYear="1966" box="[1105,1373,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E37EFF3AFA96FF2E" box="[1105,1373,151,177]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
from 
<taxonomicName id="658FC8F84B08FFC4E7B8FF16FE89FF4A" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[151,322,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E7B8FF16FE89FF4A" box="[151,322,187,213]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
, including the forms known as 
<taxonomicName id="658FC8F84B08FFC4E58AFF16FB71FF4A" authority="Ortmann, 1890" authorityName="Ortmann" authorityYear="1890" box="[677,1210,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="dolichodactylus">
<emphasis id="90FB6F694B08FFC4E58AFF16FBC2FF4A" box="[677,1033,187,213]" italics="true" pageId="76" pageNumber="77">A. malabaricus dolichodactylus</emphasis>
<bibRefCitation id="C61ECE8A4B08FFC4E33FFF16FB71FF4A" author="Ortmann, A." box="[1040,1210,187,213]" pageId="76" pageNumber="77" pagination="437 - 542" refId="ref73624" refString="Ortmann, A. (1890) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beru ¨ cksichtigung der von Herrn Dr. D ˆ derlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. I. Theil. Die Unterordnung Natantia (Boas) (Abtheilungen: Penaeidae und Eucyphidea = Caridae der Autoren). Zoologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 5, 437 - 542, pls. 36, 37." type="book chapter" year="1890">Ortmann, 1890</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B08FFC4E3DEFF16FE6EFF66" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="leptopus">
<emphasis id="90FB6F694B08FFC4E3DEFF16FF3DFF66" italics="true" pageId="76" pageNumber="77">A. malabaricus leptopus</emphasis>
<bibRefCitation id="C61ECE8A4B08FFC4E7D0FF72FE6EFF66" author="De Man, J. G." box="[255,421,223,249]" pageId="76" pageNumber="77" pagination="287 - 319" refId="ref70953" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeniging, (2) 11, 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
(both now in the synonymy of 
<taxonomicName id="658FC8F84B08FFC4E43BFF72FC09FF66" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[788,962,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E43BFF72FC09FF66" box="[788,962,223,249]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
, see below), by the relative length of the fingers of the minor chela, which are 1.5 times as long as the palm (
<emphasis id="90FB6F694B08FFC4E4A7FEAEFC54FE83" box="[904,927,259,284]" italics="true" pageId="76" pageNumber="77">vs</emphasis>
. at least three times as long as the palm in the other three forms). In addition, in 
<taxonomicName id="658FC8F84B08FFC4E521FE8AFCDDFEDE" authorityName="Banner &amp; Banner" authorityYear="1966" box="[526,790,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E521FE8AFCDDFEDE" box="[526,790,295,321]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
, the distal parts of the fingers are strongly crossing, which is not the case of 
<taxonomicName id="658FC8F84B08FFC4E675FEE6FD76FEFA" authorityName="Ortmann" authorityYear="1890" box="[346,701,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="dolichodactylus">
<emphasis id="90FB6F694B08FFC4E675FEE6FD76FEFA" box="[346,701,331,357]" italics="true" pageId="76" pageNumber="77">A. malabaricus dolichodactylus</emphasis>
</taxonomicName>
and 
<emphasis id="90FB6F694B08FFC4E5DBFEE6FBC6FEFA" box="[756,1037,331,357]" italics="true" pageId="76" pageNumber="77">
<taxonomicName id="658FC8F84B08FFC4E5DBFEE6FBC3FEFA" authorityName="De Man" authorityYear="1910" box="[756,1032,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="leptopus">A. malabaricus leptopus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="A230B37B4B08FFC4E7E8FEC2FD1BFC3D" blockId="76.[151,1437,150,2039]" pageId="76" pageNumber="77">
In the general shape and proportions of the minor chela, 
<taxonomicName id="658FC8F84B08FFC4E47CFEC2FBABFE16" authorityName="Banner &amp; Banner" authorityYear="1966" box="[851,1120,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E47CFEC2FBABFE16" box="[851,1120,367,393]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
approaches 
<taxonomicName id="658FC8F84B08FFC4E3C0FEC2FE5EFE32" authority="Banner, 1959" authorityName="Banner" authorityYear="1959" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="mackayi">
<emphasis id="90FB6F694B08FFC4E3C0FEC2FF3FFE32" italics="true" pageId="76" pageNumber="77">A. malabaricus mackayi</emphasis>
<bibRefCitation id="C61ECE8A4B08FFC4E7D3FE3EFE5EFE32" author="Banner, A. H." box="[252,405,402,429]" pageId="76" pageNumber="77" pagination="130 - 155" refId="ref69125" refString="Banner, A. H. (1959) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part IV. Various small collections from the Central Pacific area, including supplementary notes on alpheids from Hawaii. Pacific Science, 13, 130 - 155." type="journal article" year="1959">Banner, 1959</bibRefCitation>
</taxonomicName>
, which was elevated to species rank, as 
<taxonomicName id="658FC8F84B08FFC4E476FE3EFC1EFE32" authorityName="A. H. Banner" authorityYear="1959" box="[857,981,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="90FB6F694B08FFC4E476FE3EFC1EFE32" box="[857,981,403,429]" italics="true" pageId="76" pageNumber="77">A. mackayi</emphasis>
</taxonomicName>
(
<bibRefCitation id="C61ECE8A4B08FFC4E4CAFE3EFBB1FE32" author="Banner, A. H." box="[997,1146,402,429]" pageId="76" pageNumber="77" pagination="130 - 155" refId="ref69125" refString="Banner, A. H. (1959) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part IV. Various small collections from the Central Pacific area, including supplementary notes on alpheids from Hawaii. Pacific Science, 13, 130 - 155." type="journal article" year="1959">Banner 1959</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B08FFC4E3A9FE3EFA5BFE32" author="Banner, A. H. &amp; Banner, D. M." box="[1158,1424,403,429]" pageId="76" pageNumber="77" pagination="423 - 437" refId="ref69210" refString="Banner, A. H. &amp; Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part XVII. Additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner &amp; Banner 1974</bibRefCitation>
). However, 
<bibRefCitation id="C61ECE8A4B08FFC4E622FE1AFE6CFE4E" author="Chace, F. A. Jr." box="[269,423,439,465]" pageId="76" pageNumber="77" pagination="1 - 99" refId="ref69928" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace (1988)</bibRefCitation>
placed all the above-mentioned species and varieties, including 
<taxonomicName id="658FC8F84B08FFC4E3A1FE1AFA57FE4E" authorityName="Banner &amp; Banner" authorityYear="1966" box="[1166,1436,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E3A1FE1AFA57FE4E" box="[1166,1436,439,465]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B08FFC4E7E9FE76FE89FE6A" authorityName="A. H. Banner" authorityYear="1959" box="[198,322,475,501]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="90FB6F694B08FFC4E7E9FE76FE89FE6A" box="[198,322,475,501]" italics="true" pageId="76" pageNumber="77">A. mackayi</emphasis>
</taxonomicName>
, as well as 
<taxonomicName id="658FC8F84B08FFC4E6EDFE76FCF3FE6A" authority="Ortmann, 1890" authorityName="Ortmann" authorityYear="1890" box="[450,824,475,501]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="90FB6F694B08FFC4E6EDFE76FD4EFE6A" box="[450,645,475,501]" italics="true" pageId="76" pageNumber="77">A. macrodactylus</emphasis>
<bibRefCitation id="C61ECE8A4B08FFC4E5A3FE76FCF3FE6A" author="Ortmann, A." box="[652,824,475,501]" pageId="76" pageNumber="77" pagination="437 - 542" refId="ref73624" refString="Ortmann, A. (1890) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beru ¨ cksichtigung der von Herrn Dr. D ˆ derlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. I. Theil. Die Unterordnung Natantia (Boas) (Abtheilungen: Penaeidae und Eucyphidea = Caridae der Autoren). Zoologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 5, 437 - 542, pls. 36, 37." type="book chapter" year="1890">Ortmann, 1890</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B08FFC4E46DFE76FA84FE6A" authority="Banner &amp; Banner, 1982" authorityName="Banner &amp; Banner" authorityYear="1982" box="[834,1359,475,501]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="trefzae">
<emphasis id="90FB6F694B08FFC4E46DFE76FBF4FE6A" box="[834,1087,475,501]" italics="true" pageId="76" pageNumber="77">A. malabaricus trefzae</emphasis>
<bibRefCitation id="C61ECE8A4B08FFC4E36AFE76FA84FE6A" author="Banner, D. M. &amp; Banner, A. H." box="[1093,1359,475,501]" pageId="76" pageNumber="77" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner, 1982</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B08FFC4E2ABFE76FE2AFD86" authority="Wicksten, 1983" authorityName="Wicksten" authorityYear="1983" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mazatlanicus">
<emphasis id="90FB6F694B08FFC4E2ABFE76FEE0FD86" italics="true" pageId="76" pageNumber="77">A. mazatlanicus</emphasis>
<bibRefCitation id="C61ECE8A4B08FFC4E61FFE52FE2AFD86" author="Wicksten, M. K." box="[304,481,511,537]" pageId="76" pageNumber="77" pagination="1 - 59" refId="ref74826" refString="Wicksten, M. K. (1983) A monograph on the shallow water caridean shrimps of the Gulf of California, Mexico. Allan Hancock Monographs in Marine Biology, 13, 1 - 59." type="journal article" year="1983">Wicksten, 1983</bibRefCitation>
</taxonomicName>
, in the synonymy of 
<taxonomicName id="658FC8F84B08FFC4E5E9FE52FCBBFD86" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[710,880,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E5E9FE52FCBBFD86" box="[710,880,511,537]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
. The clearly premature synonymisations of 
<bibRefCitation id="C61ECE8A4B08FFC4E279FE52FF29FDA2" author="Chace, F. A. Jr." pageId="76" pageNumber="77" pagination="1 - 99" refId="ref69928" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace (1988)</bibRefCitation>
resulted in the morphologically and ecologically highly variable 
<emphasis id="90FB6F694B08FFC4E4F2FD8EFAC0FDA2" box="[989,1291,547,573]" italics="true" pageId="76" pageNumber="77">
<taxonomicName id="658FC8F84B08FFC4E4F2FD8EFB1FFDA3" box="[989,1236,547,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="sensu">
A. malabaricus 
<taxonomicNameLabel id="8BC8D2124B08FFC4E3B9FD8EFB1FFDA3" box="[1174,1236,547,572]" pageId="76" pageNumber="77" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
, a “species” with a vast geographic range, intertidal and deep-water populations, specimens with relatively short to extremely elongated fingers of the minor cheliped, specimens with the dactylar plunger of the major cheliped ranging from very large and stout to greatly reduced, and a great deal of other “intraspecific variation”. Although the revision of the entire 
<taxonomicName id="658FC8F84B08FFC4E62AFD1FFE64FD53" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[261,431,690,716]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E62AFD1FFE64FD53" box="[261,431,690,716]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
complex (
<taxonomicName id="658FC8F84B08FFC4E50BFD1FFC57FD53" authority="sensu Chace 1988" authorityName="sensu Chace" authorityYear="1988" box="[548,924,690,716]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E50BFD1FFCD9FD53" box="[548,786,690,716]" italics="true" pageId="76" pageNumber="77">A. malabaricus sensu</emphasis>
<bibRefCitation id="C61ECE8A4B08FFC4E437FD1FFC57FD53" author="Chace, F. A. Jr." box="[792,924,690,716]" pageId="76" pageNumber="77" pagination="1 - 99" refId="ref69928" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
</taxonomicName>
) is well beyond the scope of the present study, the author sees no reason for treating 
<taxonomicName id="658FC8F84B08FFC4E518FD7BFD32FD6F" authorityName="Ortmann" authorityYear="1890" box="[567,761,726,752]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="90FB6F694B08FFC4E518FD7BFD32FD6F" box="[567,761,726,752]" italics="true" pageId="76" pageNumber="77">A. macrodactylus</emphasis>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B08FFC4E428FD7BFC49FD6F" authorityName="A. H. Banner" authorityYear="1959" box="[775,898,726,752]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="90FB6F694B08FFC4E428FD7BFC49FD6F" box="[775,898,726,752]" italics="true" pageId="76" pageNumber="77">A. mackayi</emphasis>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B08FFC4E4BFFD7BFB89FD6F" authorityName="Wicksten" authorityYear="1983" box="[912,1090,726,752]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mazatlanicus">
<emphasis id="90FB6F694B08FFC4E4BFFD7BFB89FD6F" box="[912,1090,726,752]" italics="true" pageId="76" pageNumber="77">A. mazatlanicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B08FFC4E358FD7BFAB5FD6F" authorityName="Banner &amp; Banner" authorityYear="1966" box="[1143,1406,726,752]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E358FD7BFAB5FD6F" box="[1143,1406,726,752]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
as junior synonyms of 
<taxonomicName id="658FC8F84B08FFC4E656FD57FDEFFC8C" authorityName="sensu Chace" authorityYear="1988" box="[377,548,761,788]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E656FD57FDEFFC8C" box="[377,548,761,788]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="C61ECE8A4B08FFC4E576FD57FCAAFC8C" author="Banner, A. H. &amp; Banner, D. M." box="[601,865,761,788]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B08FFC4E443FD54FC6CFC8C" author="Banner, A. H. &amp; Banner, D. M." box="[876,935,761,787]" pageId="76" pageNumber="77" pagination="423 - 437" refId="ref69210" refString="Banner, A. H. &amp; Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part XVII. Additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">1974</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B08FFC4E49CFD54FC25FC8C" author="Banner, D. M. &amp; Banner, A. H." box="[947,1006,761,787]" pageId="76" pageNumber="77" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B08FFC4E4D6FD54FBFFFC8C" author="Banner, A. H. &amp; Banner, D. M." box="[1017,1076,761,787]" pageId="76" pageNumber="77" pagination="1 - 164" refId="ref69267" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B08FFC4E36FFD54FAD0FC8C" author="Kim, W. &amp; Abele, L. G." box="[1088,1307,761,787]" pageId="76" pageNumber="77" pagination="1 - 119" refId="ref72505" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
), whilst the taxonomic status of 
<taxonomicName id="658FC8F84B08FFC4E650FCB0FD2EFCA8" authorityName="Ortmann" authorityYear="1890" box="[383,741,797,823]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="dolichodactylus">
<emphasis id="90FB6F694B08FFC4E650FCB0FD2EFCA8" box="[383,741,797,823]" italics="true" pageId="76" pageNumber="77">A. malabaricus dolichodactylus</emphasis>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B08FFC4E5DAFCB0FBC1FCA8" authorityName="De Man" authorityYear="1910" box="[757,1034,797,823]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="leptopus">
<emphasis id="90FB6F694B08FFC4E5DAFCB0FBC1FCA8" box="[757,1034,797,823]" italics="true" pageId="76" pageNumber="77">A. malabaricus leptopus</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B08FFC4E36CFCB0FA8FFCA8" authorityName="Banner &amp; Banner" authorityYear="1982" box="[1091,1348,797,823]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="trefzae">
<emphasis id="90FB6F694B08FFC4E36CFCB0FA8FFCA8" box="[1091,1348,797,823]" italics="true" pageId="76" pageNumber="77">A. malabaricus trefzae</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="C61ECE8A4B08FFC4E253FCB3FEC6FCC4" author="De Man, J. G." pageId="76" pageNumber="77" pagination="133 - 465" refId="ref70992" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga-Expeditie, 39 a 1 (2), 133 - 465; (1915) Supplement - Explanations of plates of Alpheidae, pls. 1 - 23. E. J. Brill, Leiden. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article" year="1911">De Man 1911</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B08FFC4E635FCECFDEFFCC4" author="Banner, D. M. &amp; Banner, A. H." box="[282,548,833,859]" pageId="76" pageNumber="77" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
) will need further clarification. Nevertheless, awaiting a long-needed revision of 
<taxonomicName id="658FC8F84B08FFC4E799FCC8FEAAFCE1" authorityName="sensu Chace" authorityYear="1988" box="[182,353,868,895]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E799FCC8FEAAFCE1" box="[182,353,868,895]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
, the comparative material examined in this study is listed under 
<emphasis id="90FB6F694B08FFC4E319FCC8FA91FCE1" box="[1078,1370,868,895]" italics="true" pageId="76" pageNumber="77">
<taxonomicName id="658FC8F84B08FFC4E319FCC8FAEDFCE1" box="[1078,1318,868,895]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="sensu">
A. malabaricus 
<taxonomicNameLabel id="8BC8D2124B08FFC4E3C7FCC8FAEDFCE1" box="[1256,1318,869,894]" pageId="76" pageNumber="77" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
, even though it certainly contains more than one species.
</paragraph>
<paragraph id="A230B37B4B08FFC4E7E8FC00FBF7F99D" blockId="76.[151,1437,150,2039]" pageId="76" pageNumber="77">
<bibRefCitation id="C61ECE8A4B08FFC4E7E8FC00FE15FC59" author="Banner, A. H. &amp; Banner, D. M." box="[199,478,940,967]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
noted that 
<taxonomicName id="658FC8F84B08FFC4E574FC01FCA8FC59" authorityName="Banner &amp; Banner" authorityYear="1966" box="[603,867,940,966]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E574FC01FCA8FC59" box="[603,867,940,966]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="658FC8F84B08FFC4E4D8FC01FBB8FC59" authorityName="A. H. Banner" authorityYear="1959" box="[1015,1139,940,966]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="90FB6F694B08FFC4E4D8FC01FBB8FC59" box="[1015,1139,940,966]" italics="true" pageId="76" pageNumber="77">A. mackayi</emphasis>
</taxonomicName>
by the longer rostrum; the lack of rostro-orbital furrows; and the fingers of the minor chela slenderer and with longer, distally stronger crossing tips. In 
<taxonomicName id="658FC8F84B08FFC4E7C4FC59FE38FB92" authorityName="Banner &amp; Banner" authorityYear="1966" box="[235,499,1011,1038]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E7C4FC59FE38FB92" box="[235,499,1011,1038]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
, the orbital hoods seem to be less projecting than in 
<taxonomicName id="658FC8F84B08FFC4E315FC59FB7DFB92" authorityName="A. H. Banner" authorityYear="1959" box="[1082,1206,1011,1038]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="90FB6F694B08FFC4E315FC59FB7DFB92" box="[1082,1206,1011,1038]" italics="true" pageId="76" pageNumber="77">A. mackayi</emphasis>
</taxonomicName>
, whereas the second article of the antennular peduncle is noticeably shorter than in 
<taxonomicName id="658FC8F84B08FFC4E44AFBB5FC28FBAE" authorityName="A. H. Banner" authorityYear="1959" box="[869,995,1047,1074]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="90FB6F694B08FFC4E44AFBB5FC28FBAE" box="[869,995,1047,1074]" italics="true" pageId="76" pageNumber="77">A. mackayi</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="C61ECE8A4B08FFC4E334FBB5FAE2FBAE" author="Banner, A. H. &amp; Banner, D. M." box="[1051,1321,1047,1074]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
: fig. 56A; 
<bibRefCitation id="C61ECE8A4B08FFC4E7B8FB96FEE0FBC9" author="Banner, A. H." box="[151,299,1083,1110]" pageId="76" pageNumber="77" pagination="130 - 155" refId="ref69125" refString="Banner, A. H. (1959) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part IV. Various small collections from the Central Pacific area, including supplementary notes on alpheids from Hawaii. Pacific Science, 13, 130 - 155." type="journal article" year="1959">Banner 1959</bibRefCitation>
: fig. 12a, b). Furthermore, the telson of 
<taxonomicName id="658FC8F84B08FFC4E5C4FB96FC3EFBCA" authorityName="Banner &amp; Banner" authorityYear="1966" box="[747,1013,1083,1109]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E5C4FB96FC3EFBCA" box="[747,1013,1083,1109]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
has straight lateral margins; these are strongly convex in 
<taxonomicName id="658FC8F84B08FFC4E642FBF2FE23FBE6" authorityName="A. H. Banner" authorityYear="1959" box="[365,488,1119,1145]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="90FB6F694B08FFC4E642FBF2FE23FBE6" box="[365,488,1119,1145]" italics="true" pageId="76" pageNumber="77">A. mackayi</emphasis>
</taxonomicName>
(cf. 
<emphasis id="90FB6F694B08FFC4E534FBF2FD9AFBE6" box="[539,593,1119,1145]" italics="true" pageId="76" pageNumber="77">idem</emphasis>
: fig. 56H; fig. 
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). The main differences between 
<taxonomicName id="658FC8F84B08FFC4E3BBFBF2FA57FBE6" authorityName="Banner &amp; Banner" authorityYear="1966" box="[1172,1436,1119,1145]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E3BBFBF2FA57FBE6" box="[1172,1436,1119,1145]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B08FFC4E7E7FB2EFE47FB03" authorityName="Ortmann" authorityYear="1890" box="[200,396,1154,1181]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="90FB6F694B08FFC4E7E7FB2EFE47FB03" box="[200,396,1154,1181]" italics="true" pageId="76" pageNumber="77">A. macrodactylus</emphasis>
</taxonomicName>
consist in the less projecting orbital hoods (cf. 
<bibRefCitation id="C61ECE8A4B08FFC4E487FB2EFB78FB03" author="Banner, A. H. &amp; Banner, D. M." box="[936,1203,1154,1181]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
: fig. 56A; 
<bibRefCitation id="C61ECE8A4B08FFC4E202FB2EFEE7FB5F" author="Banner, D. M. &amp; Banner, A. H." pageId="76" pageNumber="77" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
: fig. 65a); the ratio of the fingers to the palm in the major chela being around 
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<emphasis id="90FB6F694B08FFC4E3DFFB0AFEC4FB7B" italics="true" pageId="76" pageNumber="77">
<taxonomicName id="658FC8F84B08FFC4E3DFFB0AFF25FB7B" authorityName="Banner &amp; Banner" authorityYear="1966" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">A. malabaricus songkla</taxonomicName>
vs
</emphasis>
. closer to 1.0 in 
<taxonomicName id="658FC8F84B08FFC4E6F9FB67FD50FB7B" authorityName="Ortmann" authorityYear="1890" box="[470,667,1226,1252]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="90FB6F694B08FFC4E6F9FB67FD50FB7B" box="[470,667,1226,1252]" italics="true" pageId="76" pageNumber="77">A. macrodactylus</emphasis>
</taxonomicName>
(cf. 
<emphasis id="90FB6F694B08FFC4E5FCFB67FCC2FB7B" box="[723,777,1226,1252]" italics="true" pageId="76" pageNumber="77">idem</emphasis>
: fig. 56B, C; fig. 65b, c); and the minor cheliped fingers somewhat shorter relative to the palm (1.5 times as long as the palm), without armature on the cutting edges and with the fingertips strongly crossing in 
<emphasis id="90FB6F694B08FFC4E54AFABFFC53FAB4" box="[613,920,1297,1324]" italics="true" pageId="76" pageNumber="77">
<taxonomicName id="658FC8F84B08FFC4E54AFABFFCBDFAB4" authorityName="Banner &amp; Banner" authorityYear="1966" box="[613,886,1297,1324]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">A. malabaricus songkla</taxonomicName>
vs
</emphasis>
. noticeably longer (1.8 times as long as the palm), with small teeth in the proximal portion of the cutting edges and with the fingertips not strongly crossing in 
<taxonomicName id="658FC8F84B08FFC4E7B8FAF4FE90FAEC" authorityName="Ortmann" authorityYear="1890" box="[151,347,1369,1395]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="90FB6F694B08FFC4E7B8FAF4FE90FAEC" box="[151,347,1369,1395]" italics="true" pageId="76" pageNumber="77">A. macrodactylus</emphasis>
</taxonomicName>
(cf. 
<emphasis id="90FB6F694B08FFC4E6BEFAF4FE0CFAEC" box="[401,455,1369,1395]" italics="true" pageId="76" pageNumber="77">idem</emphasis>
: fig. 56D, E; fig. 65e, f). Finally, 
<taxonomicName id="658FC8F84B08FFC4E468FAF4FB99FAEC" authorityName="Banner &amp; Banner" authorityYear="1966" box="[839,1106,1369,1395]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E468FAF4FB99FAEC" box="[839,1106,1369,1395]" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
can be easily separated from the eastern Pacific 
<taxonomicName id="658FC8F84B08FFC4E65EFAD0FDEDFA09" authorityName="Wicksten" authorityYear="1983" box="[369,550,1404,1431]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="mazatlanicus">
<emphasis id="90FB6F694B08FFC4E65EFAD0FDEDFA09" box="[369,550,1404,1431]" italics="true" pageId="76" pageNumber="77">A. mazatlanicus</emphasis>
</taxonomicName>
, for instance, by the shape of the rostro-orbital area; the length of the second article of the antennular peduncle; the shape of the major chela dactylus; and the relative proportions of the carpal subarticles in the second pereiopod (cf. 
<bibRefCitation id="C61ECE8A4B08FFC4E579FA68FC94FA41" author="Banner, A. H. &amp; Banner, D. M." box="[598,863,1476,1503]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
: fig. 56; 
<bibRefCitation id="C61ECE8A4B08FFC4E4EDFA69FB57FA41" author="Kim, W. &amp; Abele, L. G." box="[962,1180,1476,1502]" pageId="76" pageNumber="77" pagination="1 - 119" refId="ref72505" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
: fig. 36). Therefore, 
<taxonomicName id="658FC8F84B08FFC4E2ABFA69FE4AF99D" authorityName="Banner &amp; Banner" authorityYear="1966" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B08FFC4E2ABFA69FE4AF99D" italics="true" pageId="76" pageNumber="77">A. malabaricus songkla</emphasis>
</taxonomicName>
is herein elevated to full species rank, as 
<taxonomicName id="658FC8F84B08FFC4E47BFA45FC01F99D" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[852,970,1512,1538]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="songkla" status="stat. nov.">
<emphasis id="90FB6F694B08FFC4E47BFA45FC01F99D" box="[852,970,1512,1538]" italics="true" pageId="76" pageNumber="77">A. songkla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B08FFC4E4FEFA45FBF7F99D" bold="true" box="[977,1084,1512,1538]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="8BC8D2124B08FFC4E4FEFA45FBF7F99D" box="[977,1084,1512,1538]" pageId="76" pageNumber="77">stat. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="A230B37B4B08FFC4E7E8F9A6FE51F963" blockId="76.[151,1437,150,2039]" pageId="76" pageNumber="77">
Despite the fact that 
<taxonomicName id="658FC8F84B08FFC4E691F9A1FDF3F9BA" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[446,568,1547,1574]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B08FFC4E691F9A1FDF3F9BA" box="[446,568,1547,1574]" italics="true" pageId="76" pageNumber="77">A. songkla</emphasis>
</taxonomicName>
is morphologically different from all species and subspecies (or varieties) currently assigned to the 
<taxonomicName id="658FC8F84B08FFC4E696F99DFDACF9D6" authorityName="sensu Chace" authorityYear="1988" box="[441,615,1583,1610]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E696F99DFDACF9D6" box="[441,615,1583,1610]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
complex, it remains a problematic taxon. The description by 
<bibRefCitation id="C61ECE8A4B08FFC4E203F99DFEF3F9F2" author="Banner, A. H. &amp; Banner, D. M." pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
was based exclusively on females, i.e., the morphology of the male minor chela, of critical importance for taxonomy of 
<taxonomicName id="658FC8F84B08FFC4E677F9DAFE79F90E" authorityName="Fabricius" authorityYear="1798" box="[344,434,1655,1681]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="90FB6F694B08FFC4E677F9DAFE79F90E" box="[344,434,1655,1681]" italics="true" pageId="76" pageNumber="77">Alpheus</emphasis>
</taxonomicName>
, currently remains unknown. Therefore, at this stage, a redescription or a new diagnosis for 
<taxonomicName id="658FC8F84B08FFC4E790F936FEFDF92B" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[191,310,1690,1717]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B08FFC4E790F936FEFDF92B" box="[191,310,1690,1717]" italics="true" pageId="76" pageNumber="77">A. songkla</emphasis>
</taxonomicName>
would not be very useful. However, an opportunity is taken to correct a number of small errors and inaccuracies in the original description of 
<taxonomicName id="658FC8F84B08FFC4E55CF912FD21F947" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[627,746,1726,1753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B08FFC4E55CF912FD21F947" box="[627,746,1726,1753]" italics="true" pageId="76" pageNumber="77">A. songkla</emphasis>
</taxonomicName>
by 
<bibRefCitation id="C61ECE8A4B08FFC4E439F912FBF9F947" author="Banner, A. H. &amp; Banner, D. M." box="[790,1074,1726,1753]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
, based on re-examination of the 
<typeStatus id="7D340DD94B08FFC4E7B8F94FFF0CF963" box="[151,199,1762,1788]" pageId="76" pageNumber="77">type</typeStatus>
material (
<figureCitation id="3AB4AFFE4B08FFC4E616F94FFE46F963" box="[313,397,1762,1788]" captionStart="FIGURE 41" captionStartId="77.[151,250,1873,1897]" captionTargetBox="[151,1435,594,1849]" captionTargetId="figure-243@77.[151,1435,594,1849]" captionTargetPageId="77" captionText="FIGURE 41. Alpheus songkla Banner &amp; Banner, 1966 stat. nov.: holotype, female (cl 6.1 mm) from Thale Sap (= Songkhla Lake), Thailand (USNM 120407); A—habitus (most appendages detached), dorsal view; B—major (left) cheliped, lateral view; C—same, mesial view; D—minor (right) cheliped, lateral view; E—third pereiopod, lateral view. Photographs by Robert M. Lasley Jr." figureDoi="http://doi.org/10.5281/zenodo.7912425" httpUri="https://zenodo.org/record/7912425/files/figure.png" pageId="76" pageNumber="77">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="A230B37B4B08FFC5E7E8F8ABFB74FDA5" blockId="76.[151,1437,150,2039]" lastBlockId="77.[151,1436,150,571]" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
One of the most important features of 
<taxonomicName id="658FC8F84B08FFC4E55FF8ABFD2EF8BF" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[624,741,1798,1824]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B08FFC4E55FF8ABFD2EF8BF" box="[624,741,1798,1824]" italics="true" pageId="76" pageNumber="77">A. songkla</emphasis>
</taxonomicName>
setting it clearly apart from the 
<taxonomicName id="658FC8F84B08FFC4E366F8ABFB3FF8BF" authorityName="sensu Chace" authorityYear="1988" box="[1097,1268,1798,1824]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B08FFC4E366F8ABFB3FF8BF" box="[1097,1268,1798,1824]" italics="true" pageId="76" pageNumber="77">A. malabaricus</emphasis>
</taxonomicName>
complex is the relatively weak anterior projection of the orbital hoods (
<bibRefCitation id="C61ECE8A4B08FFC4E426F884FBD8F8DC" author="Banner, A. H. &amp; Banner, D. M." box="[777,1043,1833,1860]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
: fig. 56A; see also 
<figureCitation id="3AB4AFFE4B08FFC4E3C7F884FA92F8DC" box="[1256,1369,1833,1859]" captionStart="FIGURE 41" captionStartId="77.[151,250,1873,1897]" captionTargetBox="[151,1435,594,1849]" captionTargetId="figure-243@77.[151,1435,594,1849]" captionTargetPageId="77" captionText="FIGURE 41. Alpheus songkla Banner &amp; Banner, 1966 stat. nov.: holotype, female (cl 6.1 mm) from Thale Sap (= Songkhla Lake), Thailand (USNM 120407); A—habitus (most appendages detached), dorsal view; B—major (left) cheliped, lateral view; C—same, mesial view; D—minor (right) cheliped, lateral view; E—third pereiopod, lateral view. Photographs by Robert M. Lasley Jr." figureDoi="http://doi.org/10.5281/zenodo.7912425" httpUri="https://zenodo.org/record/7912425/files/figure.png" pageId="76" pageNumber="77">Figs. 41A</figureCitation>
, 
<figureCitation id="3AB4AFFE4B08FFC4E24BF884FA53F8DC" box="[1380,1432,1833,1859]" captionStart="FIGURE 42" captionStartId="78.[151,250,1821,1845]" captionTargetBox="[177,1407,193,1786]" captionTargetId="figure-21@78.[164,1422,181,1797]" captionTargetPageId="78" captionText="FIGURE 42. Alpheus cf. songkla stat. nov.: male (cl 8.1 mm) [A–I], male (cl 10.4 mm) [J–M], both from near Ho Chi Minh City, Vietnam (MNHN-IU-2018-5655); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—major (left) cheliped, chela, mesial; F—same, chela, carpus and distal part of merus, lateral; G—same, dactylus; H—minor (right) cheliped, chela, carpus and distal part of merus, lateral view; I—second pereiopod, lateral view; J—third pereiopod, lateral view; K—same, dactylus, dorsal view; L—fourth pereiopod, ischium, lateral view; M—possibly abnormal minor (right) cheliped, chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912427" httpUri="https://zenodo.org/record/7912427/files/figure.png" pageId="76" pageNumber="77">42A</figureCitation>
; cf. 
<bibRefCitation id="C61ECE8A4B08FFC4E793F8E3FE0CF8F8" author="Banner, A. H. &amp; Banner, D. M." box="[188,455,1869,1896]" pageId="76" pageNumber="77" pagination="423 - 437" refId="ref69210" refString="Banner, A. H. &amp; Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part XVII. Additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner &amp; Banner 1974</bibRefCitation>
: fig. 12a; 
<bibRefCitation id="C61ECE8A4B08FFC4E517F8E3FC88F8F8" author="Banner, D. M. &amp; Banner, A. H." box="[568,835,1869,1896]" pageId="76" pageNumber="77" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
: fig. 64a, 65a; 
<bibRefCitation id="C61ECE8A4B08FFC4E4C3F8E0FB03F8F8" author="Kim, W. &amp; Abele, L. G." box="[1004,1224,1869,1895]" pageId="76" pageNumber="77" pagination="1 - 119" refId="ref72505" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119. https: // doi. org / 10.5479 / si. 00810282.454" type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
: fig. 36a, b; 
<bibRefCitation id="C61ECE8A4B08FFC4E279F8E0FF18F814" author="Chace, F. A. Jr." pageId="76" pageNumber="77" pagination="1 - 99" refId="ref69928" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
: fig. 9a). The antennal basicerite of the 
<typeStatus id="7D340DD94B08FFC4E5BAF8DCFD3CF814" box="[661,759,1905,1931]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
(USNM 120407) has a small, curved tooth in the middle of the distolateral margin (apparently broken on the right side), which seems to correspond to “minute lateral spine” in the description of 
<bibRefCitation id="C61ECE8A4B08FFC4E6AEF814FD57F84D" author="Banner, A. H. &amp; Banner, D. M." box="[385,668,1976,2003]" pageId="76" pageNumber="77" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
. However, the antennal basicerite of the examined 
<typeStatus id="7D340DD94B08FFC4E3F5F814FAF1F84C" box="[1242,1338,1977,2003]" pageId="76" pageNumber="77" type="paratype">paratype</typeStatus>
(USNM 120408) has a similar curved tooth only on the right side, being unarmed on the left side. The ischium of the third pereiopod has a small spiniform seta, as correctly figured by 
<bibRefCitation id="C61ECE8A4B09FFC5E464FF3AFBABFF2E" author="Banner, A. H. &amp; Banner, D. M." box="[843,1120,151,177]" pageId="77" pageNumber="78" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
: fig. 56G). The propodus of the third pereiopod has a row of tightly appressed spiniform setae between the long stiff setae; only the latter were shown by 
<bibRefCitation id="C61ECE8A4B09FFC5E627FF72FDDCFF67" author="Banner, A. H. &amp; Banner, D. M." box="[264,535,222,249]" pageId="77" pageNumber="78" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
: fig. 56G), who stated that the propodus is “bearing strong setae but no spinules”, which is incorrect. Furthermore, the mesial face of the major chela is not perfectly smooth, but has a large field of fine granules distally, i.e., on the distal portion of the palm and most of the pollex, reminiscent of the granulation of 
<taxonomicName id="658FC8F84B09FFC5E7B8FEE7FE8DFEFC" authorityName="De Man" authorityYear="1920" box="[151,326,329,356]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B09FFC5E7B8FEE7FE8DFEFC" box="[151,326,329,356]" italics="true" pageId="77" pageNumber="78">A. eurydactylus</emphasis>
</taxonomicName>
(although weaker). The mesial subdistal ridge of the pollex is long and sharp, and departs from a conspicuous proximal bump, as in 
<taxonomicName id="658FC8F84B09FFC5E536FEC0FD7CFE18" authorityName="sensu De Man" authorityYear="1897" box="[537,695,365,391]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B09FFC5E536FEC0FD7CFE18" box="[537,695,365,391]" italics="true" pageId="77" pageNumber="78">A. euphrosyne</emphasis>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B09FFC5E5EBFEC0FCBAFE18" authorityName="De Man" authorityYear="1920" box="[708,881,365,391]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B09FFC5E5EBFEC0FCBAFE18" box="[708,881,365,391]" italics="true" pageId="77" pageNumber="78">A. eurydactylus</emphasis>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B09FFC5E451FEC0FC1EFE18" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[894,981,365,391]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B09FFC5E451FEC0FC1EFE18" box="[894,981,365,391]" italics="true" pageId="77" pageNumber="78">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B09FFC5E4F4FEC0FBF8FE18" bold="true" box="[987,1075,365,391]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="8BC8D2124B09FFC5E4F4FEC0FBF8FE18" box="[987,1075,365,391]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, etc. The last two taxonomically important features of the major chela (granulation and mesial subdistal ridge) were not mentioned nor illustrated by 
<bibRefCitation id="C61ECE8A4B09FFC5E795FE18FE18FE51" author="Banner, A. H. &amp; Banner, D. M." box="[186,467,436,463]" pageId="77" pageNumber="78" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
. In addition, the minor chela fingers of both examined type specimens are quite setose, covered by tufts of numerous stiff setae; these setae were omitted in the original illustrations of the minor cheliped by 
<bibRefCitation id="C61ECE8A4B09FFC5E795FE50FE01FD89" author="Banner, A. H. &amp; Banner, D. M." box="[186,458,508,535]" pageId="77" pageNumber="78" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
: fig. 56D, E). The ventromesial carina of the first article of the antennular peduncle (not illustrated in the original description) is broadly rounded-triangular and without an acute point.
</paragraph>
<caption id="F6F0E3F34B09FFC5E7B8F8FCFF32F84A" ID-DOI="http://doi.org/10.5281/zenodo.7912425" ID-Zenodo-Dep="7912425" httpUri="https://zenodo.org/record/7912425/files/figure.png" pageId="77" pageNumber="78" startId="77.[151,250,1873,1897]" targetBox="[151,1435,594,1849]" targetPageId="77" targetType="figure">
<paragraph id="A230B37B4B09FFC5E7B8F8FCFF32F84A" blockId="77.[151,1437,1873,2005]" pageId="77" pageNumber="78">
<emphasis id="90FB6F694B09FFC5E7B8F8FCFED1F8F6" bold="true" box="[151,282,1873,1897]" pageId="77" pageNumber="78">FIGURE 41</emphasis>
. 
<taxonomicName id="658FC8F84B09FFC5E607F8FCFD06F8F6" authority="Banner &amp; Banner, 1966" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[296,717,1873,1897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="songkla" status="stat. nov.">
<emphasis id="90FB6F694B09FFC5E607F8FCFE1BF8F6" box="[296,464,1873,1897]" italics="true" pageId="77" pageNumber="78">Alpheus songkla</emphasis>
<bibRefCitation id="C61ECE8A4B09FFC5E6F7F8FFFD06F8F6" author="Banner, A. H. &amp; Banner, D. M." box="[472,717,1873,1897]" pageId="77" pageNumber="78" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner, 1966</bibRefCitation>
</taxonomicName>
<emphasis id="90FB6F694B09FFC5E5FBF8FCFCFEF8F6" bold="true" box="[724,821,1873,1897]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="8BC8D2124B09FFC5E5FBF8FCFCFEF8F6" box="[724,821,1873,1897]" pageId="77" pageNumber="78">stat. nov.</taxonomicNameLabel>
</emphasis>
: holotype, female (cl 6.1 mm) from Thale Sap (= Songkhla Lake), Thailand (USNM 120407); A—habitus (most appendages detached), dorsal view; B—major (left) cheliped, lateral view; C—same, mesial view; D—minor (right) cheliped, lateral view; E—third pereiopod, lateral view. Photographs by Robert M. Lasley Jr.
</paragraph>
</caption>
<caption id="F6F0E3F34B0AFFC6E7B8F8B0FB38F876" ID-DOI="http://doi.org/10.5281/zenodo.7912427" ID-Zenodo-Dep="7912427" httpUri="https://zenodo.org/record/7912427/files/figure.png" pageId="78" pageNumber="79" startId="78.[151,250,1821,1845]" targetBox="[177,1407,193,1786]" targetPageId="78" targetType="figure">
<paragraph id="A230B37B4B0AFFC6E7B8F8B0FB38F876" blockId="78.[151,1436,1821,2025]" pageId="78" pageNumber="79">
<emphasis id="90FB6F694B0AFFC6E7B8F8B0FED1F8AA" bold="true" box="[151,282,1821,1845]" pageId="78" pageNumber="79">FIGURE 42</emphasis>
. 
<taxonomicName id="658FC8F84B0AFFC6E608F8B0FE3BF8AA" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[295,496,1821,1845]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="songkla" status="stat. nov.">
<emphasis id="90FB6F694B0AFFC6E608F8B0FEB2F8AA" box="[295,377,1821,1845]" italics="true" pageId="78" pageNumber="79">Alpheus</emphasis>
cf. 
<emphasis id="90FB6F694B0AFFC6E68EF8B0FE3BF8AA" box="[417,496,1821,1845]" italics="true" pageId="78" pageNumber="79">songkla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B0AFFC6E6D8F8B0FD93F8AA" bold="true" box="[503,600,1821,1845]" pageId="78" pageNumber="79">
<taxonomicNameLabel id="8BC8D2124B0AFFC6E6D8F8B0FD93F8AA" box="[503,600,1821,1845]" pageId="78" pageNumber="79">stat. nov.</taxonomicNameLabel>
</emphasis>
: male (cl 8.1 mm) [A–I], male (cl 10.4 mm) [J–M], both from near Ho Chi Minh City, Vietnam (MNHN-IU-2018-5655); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—major (left) cheliped, chela, mesial; F—same, chela, carpus and distal part of merus, lateral; G—same, dactylus; H—minor (right) cheliped, chela, carpus and distal part of merus, lateral view; I—second pereiopod, lateral view; J—third pereiopod, lateral view; K—same, dactylus, dorsal view; L—fourth pereiopod, ischium, lateral view; M—possibly abnormal minor (right) cheliped, chela, lateral view.
</paragraph>
</caption>
<paragraph id="A230B37B4B0BFFC7E7E8FF3AFEB1FE32" blockId="79.[151,1437,151,537]" pageId="79" pageNumber="80">
The taxonomic identity of 
<specimenCount id="B48978F24B0BFFC7E529FF3AFDB6FF2E" box="[518,637,151,177]" pageId="79" pageNumber="80" type="male">two males</specimenCount>
and 
<specimenCount id="B48978F24B0BFFC7E591FF3AFC9BFF2E" box="[702,848,151,177]" pageId="79" pageNumber="80" type="female">four females</specimenCount>
from southern 
<collectingCountry id="DA98F3EB4B0BFFC7E324FF3AFB8EFF2E" box="[1035,1093,151,177]" name="India" pageId="79" pageNumber="80">India</collectingCountry>
(Korapuzha estuary north of 
<collectingCountry id="DA98F3EB4B0BFFC7E7B8FF16FED9FF4A" box="[151,274,187,213]" name="India" pageId="79" pageNumber="80">Kozhikode</collectingCountry>
on the Malabar Coast) reported as 
<taxonomicName id="658FC8F84B0BFFC7E58EFF16FC25FF4A" authorityName="Banner &amp; Banner" authorityYear="1966" box="[673,1006,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B0BFFC7E58EFF16FC25FF4A" box="[673,1006,187,213]" italics="true" pageId="79" pageNumber="80">Alpheus malabaricus songkla</emphasis>
</taxonomicName>
by 
<bibRefCitation id="C61ECE8A4B0BFFC7E336FF16FB0EFF4A" author="Thomas, M. M." box="[1049,1221,187,213]" pageId="79" pageNumber="80" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas (1976)</bibRefCitation>
remains uncertain. According to 
<bibRefCitation id="C61ECE8A4B0BFFC7E601FF72FE10FF66" author="Thomas, M. M." box="[302,475,223,249]" pageId="79" pageNumber="80" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas (1976)</bibRefCitation>
, both male and female specimens had “dense setae” on the lateral margins of the minor chela fingers, which were not referred to as “balaeniceps setae” (the same author reported a balaeniceps minor chela in 
<taxonomicName id="658FC8F84B0BFFC7E79CFE8AFE9AFEDE" authorityName="sensu De Man" authorityYear="1897" box="[179,337,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B0BFFC7E79CFE8AFE9AFEDE" box="[179,337,295,321]" italics="true" pageId="79" pageNumber="80">A. euphrosyne</emphasis>
</taxonomicName>
). 
<bibRefCitation id="C61ECE8A4B0BFFC7E648FE8AFDB1FEDE" author="Banner, A. H. &amp; Banner, D. M." box="[359,634,295,321]" pageId="79" pageNumber="80" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
did not mention nor illustrated dense setae on the female minor cheliped of 
<taxonomicName id="658FC8F84B0BFFC7E79AFEE6FEE0FEFA" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[181,299,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E79AFEE6FEE0FEFA" box="[181,299,331,357]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
, but the re-examination of the 
<typeStatus id="7D340DD94B0BFFC7E5ACFEE6FD78FEFA" box="[643,691,331,357]" pageId="79" pageNumber="80">type</typeStatus>
material confirmed that the minor chela is indeed quite setose (see above). Noteworhy is that Thomas’ (1976) descriptions of the colour patterns of 
<taxonomicName id="658FC8F84B0BFFC7E306FEC2FB08FE16" authorityName="Banner &amp; Banner" authorityYear="1966" box="[1065,1219,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="songkla">
<emphasis id="90FB6F694B0BFFC7E306FEC2FB08FE16" box="[1065,1219,367,393]" italics="true" pageId="79" pageNumber="80">A. m. songkla</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B0BFFC7E3D3FEC2FA57FE16" authorityName="sensu De Man" authorityYear="1897" box="[1276,1436,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B0BFFC7E3D3FEC2FA57FE16" box="[1276,1436,367,393]" italics="true" pageId="79" pageNumber="80">A. euphrosyne</emphasis>
</taxonomicName>
are almost identical.
</paragraph>
<paragraph id="A230B37B4B0BFFC7E7E8FE1AFCC4FD86" blockId="79.[151,1437,151,537]" pageId="79" pageNumber="80">
The material from 
<collectingCountry id="DA98F3EB4B0BFFC7E68CFE1AFDDCFE4E" box="[419,535,439,465]" name="Singapore" pageId="79" pageNumber="80">Singapore</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0BFFC7E509FE1AFD44FE4E" box="[550,655,439,465]" name="Malaysia" pageId="79" pageNumber="80">Malaysia</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0BFFC7E5B1FE1AFCCBFE4E" box="[670,768,439,465]" name="Vietnam" pageId="79" pageNumber="80">Vietnam</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0BFFC7E422FE1AFCBAFE4E" box="[781,881,439,465]" name="Thailand" pageId="79" pageNumber="80">Thailand</collectingCountry>
and 
<collectingCountry id="DA98F3EB4B0BFFC7E483FE1AFBD8FE4E" box="[940,1043,439,465]" name="Australia" pageId="79" pageNumber="80">Australia</collectingCountry>
herein tentatively identified as 
<taxonomicName id="658FC8F84B0BFFC7E2ABFE1AFEDEFE6A" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E2ABFE1AFA56FE4E" box="[1412,1437,439,465]" italics="true" pageId="79" pageNumber="80">A.</emphasis>
cf. 
<emphasis id="90FB6F694B0BFFC7E791FE76FEDEFE6A" box="[190,277,475,501]" italics="true" pageId="79" pageNumber="80">songkla</emphasis>
</taxonomicName>
is morphologically somewhat heterogeneous and needs further study. The material from 
<collectingCountry id="DA98F3EB4B0BFFC7E20BFE76FA53FE6A" box="[1316,1432,475,501]" name="Singapore" pageId="79" pageNumber="80">Singapore</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0BFFC7E7B8FE52FF30FD86" box="[151,251,511,537]" name="Thailand" pageId="79" pageNumber="80">Thailand</collectingCountry>
and 
<collectingCountry id="DA98F3EB4B0BFFC7E61EFE52FE5BFD86" box="[305,400,511,537]" name="Vietnam" pageId="79" pageNumber="80">Vietnam</collectingCountry>
is discussed in more detail below.
</paragraph>
<paragraph id="A230B37B4B0BFFC7E7B8FDEAFD0BFC1E" blockId="79.[151,1437,582,1797]" pageId="79" pageNumber="80">
(1) The specimens from Pulau Ubin and Sarimbun, 
<collectingCountry id="DA98F3EB4B0BFFC7E5D3FDEAFCA5FDFE" box="[764,878,583,609]" name="Singapore" pageId="79" pageNumber="80">Singapore</collectingCountry>
(ZRC 1992.11117–11119, ZRC 2014.0665), are morphologically most similar to 
<taxonomicName id="658FC8F84B0BFFC7E51AFDC6FD67FD1A" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[565,684,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E51AFDC6FD67FD1A" box="[565,684,619,645]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
, e.g., in the configuration of the rostro-orbital area and antennules; the presence of a small tooth on the antennal basicerite; the relatively slender major chela, with a moderately developed plunger on the dactylus; and the general proportions of the female chela. The general characteristics of the male minor cheliped place these specimens close to 
<taxonomicName id="658FC8F84B0BFFC7E4AEFD7AFBFFFD6E" authorityName="De Man" authorityYear="1920" box="[897,1076,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E4AEFD7AFBFFFD6E" box="[897,1076,727,753]" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
. However, they differ from 
<taxonomicName id="658FC8F84B0BFFC7E2ABFD7AFE9EFC8A" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E2ABFD7AFE9EFC8A" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
in the longer distolateral tooth of the scaphocerite, exceeding the distal margin of the blade (
<emphasis id="90FB6F694B0BFFC7E27BFD56FAA0FC8B" box="[1364,1387,763,788]" italics="true" pageId="79" pageNumber="80">vs</emphasis>
. not exceeding it in 
<taxonomicName id="658FC8F84B0BFFC7E65FFCB2FDD5FCA6" authorityName="De Man" authorityYear="1920" box="[368,542,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E65FFCB2FDD5FCA6" box="[368,542,799,825]" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
), and in the dorsal surface of the telson without longitudinal depression (present in 
<taxonomicName id="658FC8F84B0BFFC7E7C9FCEEFE5CFCC2" authorityName="De Man" authorityYear="1920" box="[230,407,835,861]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E7C9FCEEFE5CFCC2" box="[230,407,835,861]" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
). In addition, the Singaporean specimens have slenderer chelipeds compared to those of similarly sized specimens of 
<taxonomicName id="658FC8F84B0BFFC7E524FCCAFD72FC1E" authorityName="De Man" authorityYear="1920" box="[523,697,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E524FCCAFD72FC1E" box="[523,697,871,897]" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A230B37B4B0BFFC7E7B8FC26FDF7FBC6" blockId="79.[151,1437,582,1797]" pageId="79" pageNumber="80">
(2) The male from “Sin Gora inland sea” in 
<collectingCountry id="DA98F3EB4B0BFFC7E5AEFC26FD2EFC3A" box="[641,741,907,933]" name="Thailand" pageId="79" pageNumber="80">Thailand</collectingCountry>
(USNM 65561) generally agrees with 
<taxonomicName id="658FC8F84B0BFFC7E3BDFC26FACDFC3A" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[1170,1286,907,933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E3BDFC26FACDFC3A" box="[1170,1286,907,933]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
, for instance, in the similar frontal area; the third pereiopod ischium armed with a spiniform seta; the third pereiopod propodus bearing stout appressed spines and long stiff setae; and the spatulate third pereiopod dactylus. However, the distolateral margin of the antennal basicerite of this specimen has no trace of tooth on both sides. The minor chela of this male specimen, which cannot be directly compared with that of the females of 
<taxonomicName id="658FC8F84B0BFFC7E3E3FBB6FA89FBAA" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[1228,1346,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E3E3FBB6FA89FBAA" box="[1228,1346,1051,1077]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
, is very similar to that of 
<taxonomicName id="658FC8F84B0BFFC7E6A8FB92FDFEFBC6" authorityName="De Man" authorityYear="1920" box="[391,565,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E6A8FB92FDFEFBC6" box="[391,565,1087,1113]" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A230B37B4B0BFFC7E7B8FBCEFCECFA7A" blockId="79.[151,1437,582,1797]" pageId="79" pageNumber="80">
(3) The incomplete male from Menam Chao Phraya near 
<collectingRegion id="604B7D994B0BFFC7E40AFBCEFC46FBE2" box="[805,909,1123,1149]" country="Thailand" name="Krung Thep Mahanakhon" pageId="79" pageNumber="80">Bangkok</collectingRegion>
, 
<collectingCountry id="DA98F3EB4B0BFFC7E4B7FBCEFC37FBE2" box="[920,1020,1123,1149]" name="Thailand" pageId="79" pageNumber="80">Thailand</collectingCountry>
(USNM 65558), matches 
<taxonomicName id="658FC8F84B0BFFC7E209FBCEFA57FBE2" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[1318,1436,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E209FBCEFA57FBE2" box="[1318,1436,1123,1149]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
in most characters, including the very similar rostro-orbital area; the presence of a small, sharp tooth on the distolateral margin of the antennal basicerite (on both sides); the major chela with a gently sloping dorsal shoulder and granulated distomesially (with minute granules on the distal portion of the palm, pollex and dactylus); the major chela pollex with a distinct mesial subdistal ridge; the second pereiopod with the first carpal article much longer than the second; and the propodus of the third and fourth pereiopods furnished with stout appressed spines and long stiff setae. However, the ischium of the third and fourth pereiopods of this specimen is unarmed (
<emphasis id="90FB6F694B0BFFC7E6ECFAF2FE11FAE7" box="[451,474,1375,1400]" italics="true" pageId="79" pageNumber="80">vs</emphasis>
. armed with a spiniform seta in 
<taxonomicName id="658FC8F84B0BFFC7E46AFAF2FC70FAE6" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[837,955,1375,1401]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E46AFAF2FC70FAE6" box="[837,955,1375,1401]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
), and the third pereiopod merus appears to be somewhat broader than in 
<taxonomicName id="658FC8F84B0BFFC7E53EFA2EFD4CFA02" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[529,647,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E53EFA2EFD4CFA02" box="[529,647,1411,1437]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
, as illustrated by 
<bibRefCitation id="C61ECE8A4B0BFFC7E461FA2EFBAAFA02" author="Banner, A. H. &amp; Banner, D. M." box="[846,1121,1411,1437]" pageId="79" pageNumber="80" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
: fig. 56G). The female from Bangpakong River, 
<collectingCountry id="DA98F3EB4B0BFFC7E68DFA0AFDCDFA5E" box="[418,518,1447,1473]" name="Thailand" pageId="79" pageNumber="80">Thailand</collectingCountry>
(USNM 65477/2), has essentially the same characterstics as the male from Menam Chao Phraya and seems to belong to the same species.
</paragraph>
<paragraph id="A230B37B4B0BFFC7E7B8FA42FB98F89A" blockId="79.[151,1437,582,1797]" pageId="79" pageNumber="80">
(4) The 
<specimenCount id="B48978F24B0BFFC7E7D5FA42FEA5F996" box="[250,366,1519,1545]" pageId="79" pageNumber="80" type="male">two males</specimenCount>
from Duyen-Hai near 
<collectingRegion id="604B7D994B0BFFC7E55AFA42FC83F996" box="[629,840,1519,1545]" country="Vietnam" name="Ho Chi Minh City" pageId="79" pageNumber="80">Ho Chi Minh City</collectingRegion>
, 
<collectingCountry id="DA98F3EB4B0BFFC7E47AFA42FC7FF996" box="[853,948,1519,1545]" name="Vietnam" pageId="79" pageNumber="80">Vietnam</collectingCountry>
(MNHN-IU-2018-5655, MNHN-IU-2019- 2290, see 
<figureCitation id="3AB4AFFE4B0BFFC7E618F9BEFE42F9B2" box="[311,393,1555,1581]" captionStart="FIGURE 42" captionStartId="78.[151,250,1821,1845]" captionTargetBox="[177,1407,193,1786]" captionTargetId="figure-21@78.[164,1422,181,1797]" captionTargetPageId="78" captionText="FIGURE 42. Alpheus cf. songkla stat. nov.: male (cl 8.1 mm) [A–I], male (cl 10.4 mm) [J–M], both from near Ho Chi Minh City, Vietnam (MNHN-IU-2018-5655); A—frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—major (left) cheliped, chela, mesial; F—same, chela, carpus and distal part of merus, lateral; G—same, dactylus; H—minor (right) cheliped, chela, carpus and distal part of merus, lateral view; I—second pereiopod, lateral view; J—third pereiopod, lateral view; K—same, dactylus, dorsal view; L—fourth pereiopod, ischium, lateral view; M—possibly abnormal minor (right) cheliped, chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912427" httpUri="https://zenodo.org/record/7912427/files/figure.png" pageId="79" pageNumber="80">Fig. 42</figureCitation>
), were initially identified by the author as 
<taxonomicName id="658FC8F84B0BFFC7E44AF9BEFBFCF9B2" authorityName="De Man" authorityYear="1920" box="[869,1079,1555,1581]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E44AF9BEFCB6F9B2" box="[869,893,1555,1581]" italics="true" pageId="79" pageNumber="80">A.</emphasis>
cf. 
<emphasis id="90FB6F694B0BFFC7E487F9BEFBFCF9B2" box="[936,1079,1555,1581]" italics="true" pageId="79" pageNumber="80">eurydactylus</emphasis>
</taxonomicName>
. They differ from the typical 
<taxonomicName id="658FC8F84B0BFFC7E2ABF9BEFE9EF9CE" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E2ABF9BEFE9EF9CE" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
, however, by the slenderer second pereiopod, with the first article much longer than the second; the third pereiopod ischium armed with a spiniform seta (as in 
<taxonomicName id="658FC8F84B0BFFC7E456F9F6FC25F9EA" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[889,1006,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E456F9F6FC25F9EA" box="[889,1006,1627,1653]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
); and the slightly different proportions of the minor chela fingers to the palm in the male with the normally developed minor cheliped (MNHN-IU-2018-5655). In the male with a somewhat abnormal minor cheliped (MNHN-IU-2019-2290), the palm has a distinct dorsal notch, whilst the fingers possess a weak balaeniceps crest. On the other hand, they appear to be more similar to the afore-mentioned specimens of 
<taxonomicName id="658FC8F84B0BFFC7E5D4F946FC5EF89A" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[763,917,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E5D4F946FCC7F89A" box="[763,780,1771,1797]" italics="true" pageId="79" pageNumber="80">A</emphasis>
. cf. 
<emphasis id="90FB6F694B0BFFC7E411F946FC5EF89A" box="[830,917,1771,1797]" italics="true" pageId="79" pageNumber="80">songkla</emphasis>
</taxonomicName>
from 
<collectingCountry id="DA98F3EB4B0BFFC7E4F5F946FB84F89A" box="[986,1103,1771,1797]" name="Singapore" pageId="79" pageNumber="80">Singapore</collectingCountry>
.
</paragraph>
<paragraph id="A230B37B4B0BFFD8E7E8F89EFDE0FF66" blockId="79.[151,1437,1843,2013]" lastBlockId="80.[151,1437,151,501]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
Thus, the above character combinations do not allow identifying the material from 
<collectingCountry id="DA98F3EB4B0BFFC7E3A5F89EFB35F8D2" box="[1162,1278,1843,1869]" name="Singapore" pageId="79" pageNumber="80">Singapore</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B0BFFC7E224F89EFAA1F8D2" box="[1291,1386,1843,1869]" name="Vietnam" pageId="79" pageNumber="80">Vietnam</collectingCountry>
and 
<collectingCountry id="DA98F3EB4B0BFFC7E7B8F8FAFF30F8EE" box="[151,251,1879,1905]" name="Thailand" pageId="79" pageNumber="80">Thailand</collectingCountry>
reliably as 
<taxonomicName id="658FC8F84B0BFFC7E6AFF8FAFE3CF8EE" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[384,503,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E6AFF8FAFE3CF8EE" box="[384,503,1879,1905]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
(or as 
<taxonomicName id="658FC8F84B0BFFC7E567F8FAFD3CF8EE" authorityName="De Man" authorityYear="1920" box="[584,759,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E567F8FAFD3CF8EE" box="[584,759,1879,1905]" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
) and the situation is further complicated by the variation in the armature of the antennal basicerite in the 
<typeStatus id="7D340DD94B0BFFC7E5A5F8D6FD71F80A" box="[650,698,1915,1941]" pageId="79" pageNumber="80">type</typeStatus>
material of 
<taxonomicName id="658FC8F84B0BFFC7E46FF8D6FC7EF80A" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[832,949,1915,1941]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E46FF8D6FC7EF80A" box="[832,949,1915,1941]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
and of the ischium of the third pereiopod in 
<taxonomicName id="658FC8F84B0BFFC7E7B8F832FE8DF826" authorityName="De Man" authorityYear="1920" box="[151,326,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B0BFFC7E7B8F832FE8DF826" box="[151,326,1951,1977]" italics="true" pageId="79" pageNumber="80">A. eurydactylus</emphasis>
</taxonomicName>
. Since 
<taxonomicName id="658FC8F84B0BFFC7E6B5F832FDDAF826" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[410,529,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E6B5F832FDDAF826" box="[410,529,1951,1977]" italics="true" pageId="79" pageNumber="80">A. songkla</emphasis>
</taxonomicName>
is presently known only from females, there is at least a possibility that some of the above-listed material of 
<taxonomicName id="658FC8F84B0BFFC7E6E1F86EFDADF842" authorityName="Anker" authorityYear="2023" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[462,614,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E6E1F86EFE2DF842" box="[462,486,1987,2013]" italics="true" pageId="79" pageNumber="80">A.</emphasis>
cf. 
<emphasis id="90FB6F694B0BFFC7E520F86EFDADF842" box="[527,614,1987,2013]" italics="true" pageId="79" pageNumber="80">songkla</emphasis>
</taxonomicName>
material may indeed represent 
<taxonomicName id="658FC8F84B0BFFC7E4EEF86EFA53F842" authority="sensu Banner &amp; Banner (1966)" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[961,1432,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B0BFFC7E4EEF86EFBB0F843" box="[961,1147,1987,2013]" italics="true" pageId="79" pageNumber="80">A. songkla sensu</emphasis>
<bibRefCitation id="C61ECE8A4B0BFFC7E3AEF86EFA53F842" author="Banner, A. H. &amp; Banner, D. M." box="[1153,1432,1987,2013]" pageId="79" pageNumber="80" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
</taxonomicName>
. Whatever the case might be, it seems most reasonable to treat 
<taxonomicName id="658FC8F84B14FFD8E47DFF3AFC03FF2E" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[850,968,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B14FFD8E47DFF3AFC03FF2E" box="[850,968,151,177]" italics="true" pageId="80" pageNumber="81">A. songkla</emphasis>
</taxonomicName>
as a species distinct from 
<taxonomicName id="658FC8F84B14FFD8E3DEFF3AFA57FF2E" authorityName="sensu Chace" authorityYear="1988" box="[1265,1436,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B14FFD8E3DEFF3AFA57FF2E" box="[1265,1436,151,177]" italics="true" pageId="80" pageNumber="81">A. malabaricus</emphasis>
</taxonomicName>
and allied forms, as well as from 
<taxonomicName id="658FC8F84B14FFD8E53AFF16FD0FFF4A" authorityName="De Man" authorityYear="1920" box="[533,708,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B14FFD8E53AFF16FD0FFF4A" box="[533,708,187,213]" italics="true" pageId="80" pageNumber="81">A. eurydactylus</emphasis>
</taxonomicName>
, although its taxonomic identity remains problematic due to the lack of males from the 
<typeStatus id="7D340DD94B14FFD8E6B5FF72FE01FF66" box="[410,458,223,249]" pageId="80" pageNumber="81">type</typeStatus>
locality.
</paragraph>
<paragraph id="A230B37B4B14FFD8E7E8FEAEFBB6FE6A" blockId="80.[151,1437,151,501]" pageId="80" pageNumber="81">
The presence of a mesial subdistal ridge on the major chela pollex in 
<taxonomicName id="658FC8F84B14FFD8E4E1FEAEFB8FFE82" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[974,1092,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B14FFD8E4E1FEAEFB8FFE82" box="[974,1092,259,285]" italics="true" pageId="80" pageNumber="81">A. songkla</emphasis>
</taxonomicName>
and 
<emphasis id="90FB6F694B14FFD8E355FEAEFA56FE82" box="[1146,1437,259,285]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="658FC8F84B14FFD8E355FEAEFAA1FE83" box="[1146,1386,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="malabaricus" subSpecies="sensu">
A. malabaricus 
<taxonomicNameLabel id="8BC8D2124B14FFD8E203FEAEFAA1FE83" box="[1324,1386,259,284]" pageId="80" pageNumber="81" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
(based on comparative material) may represent a phylogenetic link between the 
<taxonomicName id="658FC8F84B14FFD8E337FE8AFB08FEDE" authorityName="sensu Chace" authorityYear="1988" box="[1048,1219,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="90FB6F694B14FFD8E337FE8AFB08FEDE" box="[1048,1219,295,321]" italics="true" pageId="80" pageNumber="81">A. malabaricus</emphasis>
</taxonomicName>
complex and some species of the 
<emphasis id="90FB6F694B14FFD8E61BFEE6FD7BFEFA" box="[308,688,331,357]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="658FC8F84B14FFD8E61BFEE6FE19FEFA" authorityName="sensu De Man" authorityYear="1897" box="[308,466,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
— 
<taxonomicName id="658FC8F84B14FFD8E6C0FEE6FD7BFEFA" authorityName="sensu De Man" authorityYear="1897" box="[495,688,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="microrhynchus">A. microrhynchus</taxonomicName>
</emphasis>
complex. However, this hypothesis, and the phylogenetic affinities of 
<taxonomicName id="658FC8F84B14FFD8E797FEC2FEFAFE16" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[184,305,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B14FFD8E797FEC2FEFAFE16" box="[184,305,367,393]" italics="true" pageId="80" pageNumber="81">A. songkla</emphasis>
</taxonomicName>
, can be tested only by a recollection of both male and female specimens of 
<taxonomicName id="658FC8F84B14FFD8E384FEC2FAE8FE16" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[1195,1315,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B14FFD8E384FEC2FAE8FE16" box="[1195,1315,367,393]" italics="true" pageId="80" pageNumber="81">A. songkla</emphasis>
</taxonomicName>
at or near its 
<typeStatus id="7D340DD94B14FFD8E795FE3EFF21FE32" box="[186,234,403,429]" pageId="80" pageNumber="81">type</typeStatus>
locality; a full redescription of 
<taxonomicName id="658FC8F84B14FFD8E579FE3EFD05FE32" authorityName="sensu Banner &amp; Banner" authorityYear="1966" baseAuthorityName="Banner &amp; Banner" baseAuthorityYear="1966" box="[598,718,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="songkla">
<emphasis id="90FB6F694B14FFD8E579FE3EFD05FE32" box="[598,718,403,429]" italics="true" pageId="80" pageNumber="81">A. songkla</emphasis>
</taxonomicName>
, including taxonomically important features of the male minor chela (balaeniceps or not) and colour pattern; and a comprehensive molecular analysis of the 
<taxonomicName id="658FC8F84B14FFD8E391FE1AFA80FE4E" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[1214,1355,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="90FB6F694B14FFD8E391FE1AFA80FE4E" box="[1214,1355,439,465]" italics="true" pageId="80" pageNumber="81">A. edwardsii</emphasis>
</taxonomicName>
group, including numerous specimens from as many localities as possible from both complexes.
</paragraph>
</subSubSection>
</treatment>
</document>