453 lines
46 KiB
XML
453 lines
46 KiB
XML
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<mods:title id="12E58931A596B018D09A91B2433100A9">A taxonomic revision of the Palaearctic members of the subgenus Lasius s. str. (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="4DAC0F190B5B2CDEB3DE64FAE72F8FE1">Seifert, Bernhard</mods:namePart>
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<mods:title id="32A55BDBE06BD01D3E27A8445F0D909B">Soil Organisms</mods:title>
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<mods:part id="13BEBB647982B1DCEC4A183EF0455914">
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<mods:date id="7AB57C10D5E243BDE99F8E0192659F9D">2020</mods:date>
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<treatment id="153287B6FD0AFFFAFF71FBAE5826F8A4" LSID="urn:lsid:plazi:treatment:153287B6FD0AFFFAFF71FBAE5826F8A4" httpUri="http://treatment.plazi.org/id/153287B6FD0AFFFAFF71FBAE5826F8A4" lastPageId="56" lastPageNumber="71" pageId="55" pageNumber="70">
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<subSubSection id="D581652BFD0AFFF5FF71FBAE58A3FB93" box="[212,750,1094,1122]" pageId="55" pageNumber="70" type="nomenclature">
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<paragraph id="9D2436A0FD0AFFF5FF71FBAE58A3FB93" blockId="55.[211,802,1094,1975]" box="[212,750,1094,1122]" pageId="55" pageNumber="70">
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<emphasis id="AFEFEAB2FD0AFFF5FF71FBAE58A3FB93" bold="true" box="[212,750,1094,1122]" pageId="55" pageNumber="70">
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4.4.39
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<taxonomicName id="5A9B4D23FD0AFFF5FE91FBAE58A3FB93" ID-CoL="3SGWP" authority="(Olivier 1791)" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[308,750,1094,1122]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginatus">
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<emphasis id="AFEFEAB2FD0AFFF5FE91FBAE5874FB91" bold="true" box="[308,569,1094,1120]" italics="true" pageId="55" pageNumber="70">Lasius emarginatus</emphasis>
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(Olivier 1791)
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="D581652BFD0AFFF5FF76FB825881FB70" box="[211,716,1129,1153]" pageId="55" pageNumber="70" type="reference_group">
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<paragraph id="9D2436A0FD0AFFF5FF76FB825881FB70" blockId="55.[211,802,1094,1975]" box="[211,716,1129,1153]" pageId="55" pageNumber="70">
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<taxonomicName id="5A9B4D23FD0AFFF5FF76FB825873FB70" authority="Olivier 1791" authorityName="Olivier" authorityYear="1791" box="[211,574,1129,1153]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginata">
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<emphasis id="AFEFEAB2FD0AFFF5FF76FB825BFEFB70" box="[211,435,1130,1153]" italics="true" pageId="55" pageNumber="70">Formica emarginata</emphasis>
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Olivier 1791
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</taxonomicName>
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[description]
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</paragraph>
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</subSubSection>
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<subSubSection id="D581652BFD0AFFF5FF71FB635876F867" pageId="55" pageNumber="70" type="materials_examined">
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<paragraph id="9D2436A0FD0AFFF5FF71FB635876F867" blockId="55.[211,802,1094,1975]" pageId="55" pageNumber="70">
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Type specimens are assumed to be lost but Olivier‘s description of specimens found in the Provence allows the following conclusions. The absolute length of males (”deux lignes“ =
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<quantity id="5A639B45FD0AFFF5FE21FB055B9AFAF4" box="[388,471,1261,1285]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="55" pageNumber="70" unit="mm" value="4.5">4.5 mm</quantity>
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) and of gynes (”pres de quatre lignes“ = nearly 9.0 mm) and petiole shape indicate a
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<taxonomicName id="5A9B4D23FD0AFFF5FF71FAC75B56FAB7" authorityName="Fabricius" authorityYear="1804" box="[212,283,1327,1350]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="genus">
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<emphasis id="AFEFEAB2FD0AFFF5FF71FAC75B56FAB7" box="[212,283,1327,1350]" italics="true" pageId="55" pageNumber="70">Lasius</emphasis>
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</taxonomicName>
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and exclude other genera of Formicinae occurring in this particular region. The rectangular, emarginate petiole scale, the dark reddish brown head, the lighter reddish brown ventral part of the mesosoma compared to its dark reddish-brown dorsal part, and the light (not infuscated!) wings in the gyne exclude species of the subgenus
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<taxonomicName id="5A9B4D23FD0AFFF5FED5FA1C5841F9FD" box="[368,524,1524,1548]" pageId="55" pageNumber="70" rank="subGenus" subGenus="Chthonolasius">
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<emphasis id="AFEFEAB2FD0AFFF5FED5FA1C5841F9FD" box="[368,524,1524,1548]" italics="true" pageId="55" pageNumber="70">Chthonolasius</emphasis>
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</taxonomicName>
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with rectangular scales. The nuptial flight in the evening by the end of June is another reported trait. All these statements multiply to a fair probability that the interpretation of
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<taxonomicName id="5A9B4D23FD0AFFF5FD60F9BF5B1CF97E" authorityName="Olivier" authorityYear="1791" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginata">
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<emphasis id="AFEFEAB2FD0AFFF5FD60F9BF5B1CF97E" italics="true" pageId="55" pageNumber="70">Formica emarginata</emphasis>
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</taxonomicName>
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applied by the myrmecologists during the last 150 years was correct. As there are four cryptic species close to
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<taxonomicName id="5A9B4D23FD0AFFF5FE24F952581AF920" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[385,599,1722,1745]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginatus">
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<emphasis id="AFEFEAB2FD0AFFF5FE24F952581AF920" box="[385,599,1722,1745]" italics="true" pageId="55" pageNumber="70">Lasius emarginatus</emphasis>
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</taxonomicName>
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alone in the Westpalaearctic, a
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<typeStatus id="42208802FD0AFFF5FED2F9325B83F903" box="[375,462,1754,1778]" pageId="55" pageNumber="70" type="neotype">neotype</typeStatus>
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fixation in a specimen from
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<collectingCountry id="E58C7630FD0AFFF5FF71F9135B50F8E3" box="[212,285,1787,1810]" name="France" pageId="55" pageNumber="70">France</collectingCountry>
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is indicated. Herewith I fix a
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<typeStatus id="42208802FD0AFFF5FDDBF9135898F8E2" box="[638,725,1787,1811]" pageId="55" pageNumber="70" type="neotype">neotype</typeStatus>
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in the top worker on a pin with
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<specimenCount id="8B9DFD29FD0AFFF5FE47F8F4583BF8C5" box="[482,630,1820,1844]" count="3" pageId="55" pageNumber="70" type="worker">three workers</specimenCount>
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labelled ”FRA: Frankreich-06, Prov. Savoie,
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<quantity id="5A639B45FD0AFFF5FD8EF8D55835F8A4" box="[555,632,1853,1877]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="55" pageNumber="70" unit="km" value="12.0">12 km</quantity>
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E Belley, im Rhonetal, 250 mH,
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<date id="E9251060FD0AFFF5FE02F8B55850F884" box="[423,541,1885,1909]" pageId="55" pageNumber="70" value="1996-04-14">14.04.1996</date>
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, 133 Leg. A. Schulz, K. Vock“; depository:
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<collectionCode id="FB8AAE65FD0AFFF5FE03F8965BACF867" box="[422,481,1918,1942]" pageId="55" pageNumber="70">SMN</collectionCode>
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Görlitz.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D581652BFD0AFFF5FF71F8775E42FEDB" pageId="55" pageNumber="70" type="reference_group">
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||
<paragraph id="9D2436A0FD0AFFF5FF71F87758DEF846" blockId="55.[211,802,1094,1975]" box="[212,659,1951,1975]" pageId="55" pageNumber="70">
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<emphasis id="AFEFEAB2FD0AFFF5FF71F87758DEF846" bold="true" box="[212,659,1951,1975]" pageId="55" pageNumber="70">
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<taxonomicName id="5A9B4D23FD0AFFF5FF71F87758C2F846" authority="Forel 1874" authorityName="Forel" authorityYear="1874" box="[212,655,1951,1975]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="brunneoemarginatus">
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<emphasis id="AFEFEAB2FD0AFFF5FF71F8775842F846" bold="true" box="[212,527,1951,1975]" italics="true" pageId="55" pageNumber="70">Lasius brunneoemarginatus</emphasis>
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Forel 1874
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</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph id="9D2436A0FD0AFFF5FCEAFF1A5E42FEDB" blockId="55.[824,1437,241,1975]" pageId="55" pageNumber="70">
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||
<taxonomicName id="5A9B4D23FD0AFFF5FCEAFF1A5FD0FEF8" authority="Forel" authorityName="Forel" authorityYear="1874" box="[847,1437,241,265]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="variety" species="emarginatus" variety="brunneoemarginatus">
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<emphasis id="AFEFEAB2FD0AFFF5FCEAFF1A5E60FEF8" box="[847,1069,242,265]" italics="true" pageId="55" pageNumber="70">Lasius emarginatus</emphasis>
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var.
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<emphasis id="AFEFEAB2FD0AFFF5FBD6FF195F1BFEF8" box="[1139,1366,241,265]" italics="true" pageId="55" pageNumber="70">brunneoemarginatus</emphasis>
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Forel
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</taxonomicName>
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1874 [description]
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</paragraph>
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</subSubSection>
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<subSubSection id="D581652BFD0AFFF5FCEAFEDB5FD0FBCE" pageId="55" pageNumber="70" type="discussion">
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<paragraph id="9D2436A0FD0AFFF5FCEAFEDB5FD0FBCE" blockId="55.[824,1437,241,1975]" pageId="55" pageNumber="70">
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The full text of Forel‘s description is: ”
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<emphasis id="AFEFEAB2FD0AFFF5FAB3FEDC5FD1FEBA" box="[1302,1436,307,331]" italics="true" pageId="55" pageNumber="70">
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L.
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<taxonomicName id="5A9B4D23FD0AFFF5FCEAFEBD599AFE9D" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[847,983,341,364]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginatus">
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<emphasis id="AFEFEAB2FD0AFFF5FCEAFEBD599AFE9D" box="[847,983,341,364]" italics="true" pageId="55" pageNumber="70">brunneoemarginatus</emphasis>
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</taxonomicName>
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</emphasis>
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, ou plutot
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<emphasis id="AFEFEAB2FD0AFFF5FBD6FEBD5FDBFE9D" box="[1139,1430,340,364]" italics="true" pageId="55" pageNumber="70">
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<taxonomicName id="5A9B4D23FD0AFFF5FBD6FEBD5EB6FE9D" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[1139,1275,341,364]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginatus">emarginatus</taxonomicName>
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<taxonomicName id="5A9B4D23FD0AFFF5FAB5FEBC5FDBFE9D" authorityName="Forel" authorityYear="1874" box="[1296,1430,340,364]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="brunneoides">brunneoides</taxonomicName>
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</emphasis>
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. Formes de l‘
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<taxonomicName id="5A9B4D23FD0AFFF5FC71FE9E5E11FE7C" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[980,1116,374,397]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginatus">
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<emphasis id="AFEFEAB2FD0AFFF5FC71FE9E5E11FE7C" box="[980,1116,374,397]" italics="true" pageId="55" pageNumber="70">emarginatus</emphasis>
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</taxonomicName>
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plus claires et moins poilues, vivant sous l‘écorce des arbres.“ and later, on page 217, he stated as collecting localities Lugano and
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<collectingRegion id="5F5FF842FD0AFFF5FA99FE5F5FC5FE3E" box="[1340,1416,439,463]" country="Switzerland" name="Zuerich" pageId="55" pageNumber="70">Zurich</collectingRegion>
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. I investigated
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<specimenCount id="8B9DFD29FD0AFFF5FC72FE305E16FE01" box="[983,1115,472,496]" count="6" pageId="55" pageNumber="70" type="generic">6 specimens</specimenCount>
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in
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<collectionCode id="FB8AAE65FD0AFFF5FBD8FE305EF0FE1E" box="[1149,1213,472,495]" pageId="55" pageNumber="70">MHN</collectionCode>
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<collectingRegion id="5F5FF842FD0AFFF5FB66FE305F58FE01" box="[1219,1301,472,496]" country="Switzerland" name="Geneve" pageId="55" pageNumber="70">Genève</collectingRegion>
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which carry a printed ”
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<typeStatus id="42208802FD0AFFF5FC64FE115E4AFDE0" box="[961,1031,505,529]" pageId="55" pageNumber="70">Typus</typeStatus>
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“ label and ”L. brunneo emarginatus Forel
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<collectingRegion id="5F5FF842FD0AFFF5FC38FDF159A2FDC0" box="[925,1007,537,561]" country="Ukraine" name="Crimea" pageId="55" pageNumber="70">Crimée</collectingRegion>
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“ and ”ANTWEB CASENT 0911045“. According to disagreement with the published
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<typeStatus id="42208802FD0AFFF5FAC9FDD35FD1FDA2" box="[1388,1436,571,595]" pageId="55" pageNumber="70">type</typeStatus>
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localities these specimens cannot be considered as
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<typeStatus id="42208802FD0AFFF5FCEAFD9559C0FD64" box="[847,909,637,661]" pageId="55" pageNumber="70">types</typeStatus>
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. Furthermore, there is some doubt if the young Forel, who had little contacts to foreign collectors in 1873 has seen specimens from the
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<collectingRegion id="5F5FF842FD0AFFF5FB4FFD565F77FD27" box="[1258,1338,702,726]" country="Ukraine" name="Crimea" pageId="55" pageNumber="70">Crimea</collectingRegion>
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so early. According to Ivan Löbl (former curator in
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<collectionCode id="FB8AAE65FD0AFFF5FAF9FD375FD1FD07" box="[1372,1436,735,758]" pageId="55" pageNumber="70">MHN</collectionCode>
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||
<collectingRegion id="5F5FF842FD0AFFF5FCEAFCE859ECFCE9" box="[847,929,768,792]" country="Switzerland" name="Geneve" pageId="55" pageNumber="70">Genève</collectingRegion>
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||
) and the deceased former curator of
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<collectionCode id="FB8AAE65FD0AFFF5FAF9FCE85FD1FCE6" box="[1372,1436,768,791]" pageId="55" pageNumber="70">NHM</collectionCode>
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Basel, Walter Wittmer, Forel‘s wife placed printed
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<typeStatus id="42208802FD0AFFF5FCEAFCAA5932FCAB" box="[847,895,834,858]" pageId="55" pageNumber="70">type</typeStatus>
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labels to many specimens in the collection after Forel‘s death and probably Forel himself has made this error in his later life. The original handwritten label of Forel does not contain an indication of a
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<typeStatus id="42208802FD0AFFF5FAC9FC4C5FD1FC4D" box="[1388,1436,932,956]" pageId="55" pageNumber="70">type</typeStatus>
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||
status. The synonymization of
|
||
<taxonomicName id="5A9B4D23FD0AFFF5FB39FC2E5FD1FC2C" authorityName="Forel" authorityYear="1874" box="[1180,1436,965,989]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="brunneoemarginatus">
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||
<emphasis id="AFEFEAB2FD0AFFF5FB39FC2E5FD1FC2C" box="[1180,1436,965,989]" italics="true" pageId="55" pageNumber="70">L. brunneoemarginatus</emphasis>
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||
</taxonomicName>
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||
with
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<taxonomicName id="5A9B4D23FD0AFFF5FC2FFC0E5E7EFC0C" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[906,1075,998,1021]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="emarginatus">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FC2FFC0E5E7EFC0C" box="[906,1075,998,1021]" italics="true" pageId="55" pageNumber="70">L. emarginatus</emphasis>
|
||
</taxonomicName>
|
||
as it is stated here is based on the fact that the latter is the only species of the
|
||
<taxonomicName id="5A9B4D23FD0AFFF5FAF0FBEF5FD1FBEF" box="[1365,1436,1031,1054]" pageId="55" pageNumber="70">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FAF0FBEF5FD1FBEF" box="[1365,1436,1031,1054]" italics="true" pageId="55" pageNumber="70">Lasius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AFEFEAB2FD0AFFF5FCEAFBC0599AFBCE" box="[847,983,1064,1087]" italics="true" pageId="55" pageNumber="70">emarginatus</emphasis>
|
||
complex occurring in Lugano or
|
||
<collectingRegion id="5F5FF842FD0AFFF5FAE8FBCF5FD4FBCE" box="[1357,1433,1063,1087]" country="Switzerland" name="Zuerich" pageId="55" pageNumber="70">Zurich</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD0AFFF5FC9DFBAF599BFB53" pageId="55" pageNumber="70" type="reference_group">
|
||
<paragraph id="9D2436A0FD0AFFF5FC9DFBAF5EC3FBAE" blockId="55.[824,1437,241,1975]" box="[824,1166,1095,1120]" pageId="55" pageNumber="70">
|
||
<taxonomicName id="5A9B4D23FD0AFFF5FC9DFBAF5EC3FBAE" authority="Forel 1874" authorityName="Forel" authorityYear="1874" box="[824,1166,1095,1120]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="brunneoides">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FC9DFBAF5EC3FBAE" bold="true" box="[824,1166,1095,1120]" pageId="55" pageNumber="70">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FC9DFBAF5E42FBAE" bold="true" box="[824,1039,1095,1119]" italics="true" pageId="55" pageNumber="70">Lasius brunneoides</emphasis>
|
||
Forel 1874
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="9D2436A0FD0AFFF5FCEAFB825F1AFB70" blockId="55.[824,1437,241,1975]" box="[847,1367,1129,1153]" pageId="55" pageNumber="70">
|
||
<taxonomicName id="5A9B4D23FD0AFFF5FCEAFB825F1AFB70" authority="Forel 1874" authorityName="Forel" authorityYear="1874" box="[847,1367,1129,1153]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="variety" species="emarginatus" variety="brunneoides">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FCEAFB825E69FB70" box="[847,1060,1130,1153]" italics="true" pageId="55" pageNumber="70">Lasius emarginatus</emphasis>
|
||
var.
|
||
<emphasis id="AFEFEAB2FD0AFFF5FBFFFB815EACFB70" box="[1114,1249,1129,1153]" italics="true" pageId="55" pageNumber="70">brunneoides</emphasis>
|
||
Forel 1874
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="9D2436A0FD0AFFF5FCEAFB62599BFB53" blockId="55.[824,1437,241,1975]" box="[847,982,1162,1186]" pageId="55" pageNumber="70">[description]</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD0AFFF5FCEAFB445E48FAD7" pageId="55" pageNumber="70" type="discussion">
|
||
<paragraph id="9D2436A0FD0AFFF5FCEAFB445E48FAD7" blockId="55.[824,1437,241,1975]" pageId="55" pageNumber="70">
|
||
I agree with
|
||
<bibRefCitation id="F90A4B51FD0AFFF5FC43FB435ECFFB32" author="Wilson, E. O." box="[998,1154,1195,1219]" pageId="55" pageNumber="70" pagination="1 - 201" refId="ref53426" refString="Wilson, E. O. (1955): A monographic revision of the ant genus Lasius. - Bulletin of the Museum of Comparative Zoology 113: 1 - 201." type="journal article" year="1955">Wilson (1955)</bibRefCitation>
|
||
that
|
||
<taxonomicName id="5A9B4D23FD0AFFF5FB66FB445FD6FB32" authorityName="Forel" authorityYear="1874" box="[1219,1435,1195,1219]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="brunneoides">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FB66FB445FD6FB32" box="[1219,1435,1195,1219]" italics="true" pageId="55" pageNumber="70">Lasius brunneoides</emphasis>
|
||
</taxonomicName>
|
||
is an objective synonym of
|
||
<taxonomicName id="5A9B4D23FD0AFFF5FB33FB255FD1FB15" authorityName="Forel" authorityYear="1874" box="[1174,1436,1228,1252]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="70" phylum="Arthropoda" rank="species" species="brunneoemarginatus">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FB33FB255FD1FB15" box="[1174,1436,1228,1252]" italics="true" pageId="55" pageNumber="70">L. brunneoemarginatus</emphasis>
|
||
</taxonomicName>
|
||
because Forel applied two names on the same descriptive information.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD0AFFF5FCEAFAC65F48F9FD" pageId="55" pageNumber="70" type="materials_examined">
|
||
<paragraph id="9D2436A0FD0AFFF5FCEAFAC65F48F9FD" blockId="55.[824,1437,241,1975]" pageId="55" pageNumber="70">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FCEAFAC65E00FAB7" bold="true" box="[847,1101,1326,1350]" pageId="55" pageNumber="70">All material examined</emphasis>
|
||
. A total of 61 nest samples with
|
||
<specimenCount id="8B9DFD29FD0AFFF5FC9DFAA759F2FA96" box="[824,959,1359,1383]" count="154" pageId="55" pageNumber="70" type="worker">154 workers</specimenCount>
|
||
were subject to NUMOBAT investigation.
|
||
<location id="9844607BFD0AFFF5FC9DFA985934FA79" LSID="urn:lsid:plazi:treatment:153287B6FD0AFFFAFF71FBAE5826F8A4:9844607BFD0AFFF5FC9DFA985934FA79" box="[824,889,1392,1416]" country="Austria" municipality="For" name="These" pageId="55" pageNumber="70">These</location>
|
||
originated from
|
||
<collectingCountry id="E58C7630FD0AFFF5FB99FA985EC1FA79" box="[1084,1164,1392,1416]" name="Austria" pageId="55" pageNumber="70">Austria</collectingCountry>
|
||
(6 samples), Bosnia (1),
|
||
<collectingCountry id="E58C7630FD0AFFF5FC9DFA7959D8FA58" box="[824,917,1425,1449]" name="Bulgaria" pageId="55" pageNumber="70">Bulgaria</collectingCountry>
|
||
(3),
|
||
<collectingCountry id="E58C7630FD0AFFF5FC60FA795E59FA58" box="[965,1044,1425,1449]" name="Croatia" pageId="55" pageNumber="70">Croatia</collectingCountry>
|
||
(4), Czechia (2),
|
||
<collectingCountry id="E58C7630FD0AFFF5FB6EFA795F68FA58" box="[1227,1317,1425,1449]" name="United Kingdom" pageId="55" pageNumber="70">England</collectingCountry>
|
||
(1),
|
||
<collectingCountry id="E58C7630FD0AFFF5FAF6FA7A5FD1FA58" box="[1363,1436,1426,1449]" name="France" pageId="55" pageNumber="70">France</collectingCountry>
|
||
(3),
|
||
<collectingCountry id="E58C7630FD0AFFF5FCCFFA5A5982FA3B" box="[874,975,1458,1482]" name="Germany" pageId="55" pageNumber="70">Germany</collectingCountry>
|
||
(13),
|
||
<collectingCountry id="E58C7630FD0AFFF5FBBBFA5A5E26FA3B" box="[1054,1131,1458,1482]" name="Greece" pageId="55" pageNumber="70">Greece</collectingCountry>
|
||
(17),
|
||
<collectingCountry id="E58C7630FD0AFFF5FB1CFA5A5EB8FA3B" box="[1209,1269,1458,1482]" name="Israel" pageId="55" pageNumber="70">Israel</collectingCountry>
|
||
(1),
|
||
<collectingCountry id="E58C7630FD0AFFF5FA93FA5A5F2AFA3B" box="[1334,1383,1458,1482]" name="Italy" pageId="55" pageNumber="70">Italy</collectingCountry>
|
||
(7),
|
||
<collectingCountry id="E58C7630FD0AFFF5FC9DFA3B59D8FA1A" box="[824,917,1491,1515]" name="Slovakia" pageId="55" pageNumber="70">Slovakia</collectingCountry>
|
||
(1),
|
||
<collectingCountry id="E58C7630FD0AFFF5FC6BFA3B5E02FA1A" box="[974,1103,1491,1515]" name="Switzerland" pageId="55" pageNumber="70">Switzerland</collectingCountry>
|
||
(1), and the
|
||
<collectingCountry id="E58C7630FD0AFFF5FB43FA3B5F72FA1A" box="[1254,1343,1491,1515]" name="Ukraine" pageId="55" pageNumber="70">Ukraine</collectingCountry>
|
||
(1).
|
||
<collectingMunicipality id="7D40ACDAFD0AFFF5FADDFA3B5FD1FA1B" box="[1400,1436,1491,1514]" pageId="55" pageNumber="70">For</collectingMunicipality>
|
||
details see supplementary information SI1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD0AFFFAFCEAF9FC5845FE7C" lastPageId="56" lastPageNumber="71" pageId="55" pageNumber="70" type="distribution">
|
||
<paragraph id="9D2436A0FD0AFFFAFCEAF9FC5845FE7C" blockId="55.[824,1437,241,1975]" lastBlockId="56.[151,765,241,1877]" lastPageId="56" lastPageNumber="71" pageId="55" pageNumber="70">
|
||
<emphasis id="AFEFEAB2FD0AFFF5FCEAF9FC5E6AF9DC" bold="true" box="[847,1063,1556,1581]" pageId="55" pageNumber="70">Geographic range</emphasis>
|
||
.
|
||
<location id="9844607BFD0AFFF5FB98F9FD5EA3F9DD" LSID="urn:lsid:plazi:treatment:153287B6FD0AFFFAFF71FBAE5826F8A4:9844607BFD0AFFF5FB98F9FD5EA3F9DD" box="[1085,1262,1557,1581]" country="United Kingdom" name="Only European" pageId="55" pageNumber="70">Only European</location>
|
||
, meridional to south temperate.
|
||
<collectorName id="306E5376FD0AFFF5FBA5F9DE5E1BF9BF" box="[1024,1110,1590,1614]" pageId="55" pageNumber="70">From S</collectorName>
|
||
<collectingCountry id="E58C7630FD0AFFF5FBC1F9DE5EF3F9BF" box="[1124,1214,1590,1614]" name="United Kingdom" pageId="55" pageNumber="70">England</collectingCountry>
|
||
and
|
||
<collectingCountry id="E58C7630FD0AFFF5FAA4F9DE5F07F9BC" box="[1281,1354,1590,1613]" name="France" pageId="55" pageNumber="70">France</collectingCountry>
|
||
across
|
||
<location id="9844607BFD0AFFF5FC9DF9BE59AEF99F" LSID="urn:lsid:plazi:treatment:153287B6FD0AFFFAFF71FBAE5826F8A4:9844607BFD0AFFF5FC9DF9BE59AEF99F" box="[824,995,1622,1646]" country="Ukraine" name="Central Europe" pageId="55" pageNumber="70">Central Europe</location>
|
||
to the
|
||
<collectingCountry id="E58C7630FD0AFFF5FB9BF9BE5ED7F99F" box="[1086,1178,1622,1646]" name="Ukraine" pageId="55" pageNumber="70">Ukraine</collectingCountry>
|
||
; Iberia,
|
||
<location id="9844607BFD0AFFF5FB5BF9BE5F25F99F" LSID="urn:lsid:plazi:treatment:153287B6FD0AFFFAFF71FBAE5826F8A4:9844607BFD0AFFF5FB5BF9BE5F25F99F" box="[1278,1384,1622,1646]" country="Israel" latitude="31.798" longLatPrecision="72" longitude="35.146" municipality="Balkans" name="Apennine" pageId="55" pageNumber="70">Apennine</location>
|
||
and entire
|
||
<collectingMunicipality id="7D40ACDAFD0AFFF5FCDEF99F599EF97E" box="[891,979,1655,1679]" pageId="55" pageNumber="70">Balkans</collectingMunicipality>
|
||
south to
|
||
<geoCoordinate id="F8AF5067FD0AFFF5FB94F99F5E21F97E" box="[1073,1132,1655,1679]" degrees="37" direction="north" orientation="latitude" pageId="55" pageNumber="70" precision="55555" value="37.0">37°N</geoCoordinate>
|
||
. A single, extremely isolated finding from
|
||
<collectingCountry id="E58C7630FD0AFFF5FC6DF9705E49F941" box="[968,1028,1688,1712]" name="Israel" pageId="55" pageNumber="70">Israel</collectingCountry>
|
||
(
|
||
<geoCoordinate id="F8AF5067FD0AFFF5FBB6F9705E3AF941" box="[1043,1143,1688,1712]" degrees="31.798" direction="north" orientation="latitude" pageId="55" pageNumber="70" precision="55" value="31.798">31.798°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F8AF5067FD0AFFF5FB26F9705EA8F941" box="[1155,1253,1688,1712]" degrees="35.146" direction="east" orientation="longitude" pageId="55" pageNumber="70" precision="55" value="35.146">35.146°E</geoCoordinate>
|
||
, leg.
|
||
<collectorName id="306E5376FD0AFFF5FABBF9705FCDF941" box="[1310,1408,1688,1712]" pageId="55" pageNumber="70">Besuchet</collectorName>
|
||
&
|
||
<collectorName id="306E5376FD0AFFF5FC9DF9515927F920" box="[824,874,1721,1745]" pageId="55" pageNumber="70">Löbl</collectorName>
|
||
<date id="E9251060FD0AFFF5FCD1F95159A7F920" box="[884,1002,1721,1745]" pageId="55" pageNumber="70" value="1985-04-30">
|
||
<collectingDate id="F961E988FD0AFFF5FCD1F95159A7F920" box="[884,1002,1721,1745]" pageId="55" pageNumber="70" value="1985-04-30">1985.04.30</collectingDate>
|
||
</date>
|
||
) is interpreted here as anthropogenous introduction from
|
||
<location id="9844607BFD0AFFF5FBAFF9325ED8F903" LSID="urn:lsid:plazi:treatment:153287B6FD0AFFFAFF71FBAE5826F8A4:9844607BFD0AFFF5FBAFF9325ED8F903" box="[1034,1173,1754,1778]" country="Greece" latitude="52.5" municipality="Central Europe" name="Europe. The" pageId="55" pageNumber="70">Europe. The</location>
|
||
northern distributional limit in
|
||
<collectingMunicipality id="7D40ACDAFD0AFFF5FC3EF9135E0AF8E3" box="[923,1095,1787,1811]" pageId="55" pageNumber="70">Central Europe</collectingMunicipality>
|
||
was at
|
||
<geoCoordinate id="F8AF5067FD0AFFF5FB0EF9135EBAF8E2" box="[1195,1271,1787,1811]" degrees="52.5" direction="north" orientation="latitude" pageId="55" pageNumber="70" precision="5555" value="52.5">52.5°N</geoCoordinate>
|
||
in 1980, here planar to submontane, in
|
||
<collectorName id="306E5376FD0AFFF5FBF9F8F45E9DF8C5" box="[1116,1232,1820,1844]" pageId="55" pageNumber="70">Vorarlberg</collectorName>
|
||
up to
|
||
<quantity id="5A639B45FD0AFFF5FA84F8F45F27F8C2" box="[1313,1386,1820,1844]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="70" unit="m" value="600.0">
|
||
<elevation id="16B6D193FD0AFFF5FA84F8F45F27F8C2" box="[1313,1386,1820,1844]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="70" unit="m" value="600.0">600 m</elevation>
|
||
</quantity>
|
||
and in
|
||
<collectorName id="306E5376FD0AFFF5FCF3F8D559E1F8A4" box="[854,940,1853,1877]" pageId="55" pageNumber="70">N Tyrol</collectorName>
|
||
up to
|
||
<quantity id="5A639B45FD0AFFF5FC52F8D55E03F8A5" box="[1015,1102,1853,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="55" pageNumber="70" unit="m" value="1200.0">
|
||
<elevation id="16B6D193FD0AFFF5FC52F8D55E03F8A5" box="[1015,1102,1853,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="55" pageNumber="70" unit="m" value="1200.0">1200 m</elevation>
|
||
</quantity>
|
||
.
|
||
<collectorName id="306E5376FD0AFFF5FBFFF8D55E3CF8A5" box="[1114,1137,1853,1876]" pageId="55" pageNumber="70">In</collectorName>
|
||
<collectingCountry id="E58C7630FD0AFFF5FBDFF8D55E8AF8A4" box="[1146,1223,1853,1877]" name="Greece" pageId="55" pageNumber="70">Greece</collectingCountry>
|
||
at
|
||
<geoCoordinate id="F8AF5067FD0AFFF5FB48F8D55F6BF8A4" box="[1261,1318,1853,1877]" degrees="40" direction="north" orientation="latitude" pageId="55" pageNumber="70" precision="55555" value="40.0">40°N</geoCoordinate>
|
||
ascending to
|
||
<quantity id="5A639B45FD0AFFF5FCFDF8B659E3F884" box="[856,942,1886,1910]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="55" pageNumber="70" unit="m" value="1700.0">
|
||
<elevation id="16B6D193FD0AFFF5FCFDF8B659E3F884" box="[856,942,1886,1910]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="55" pageNumber="70" unit="m" value="1700.0">1700 m</elevation>
|
||
</quantity>
|
||
.
|
||
<collectorName id="306E5376FD0AFFF5FC18F8B55E7FF884" box="[957,1074,1885,1909]" pageId="55" pageNumber="70">Significant</collectorName>
|
||
northern range expansion since 1980 particularly in
|
||
<collectorName id="306E5376FD0AFFF5FBBCF8975EC0F867" box="[1049,1165,1919,1942]" pageId="55" pageNumber="70">W Europe</collectorName>
|
||
: after the first finding in 1983 it has colonized entire
|
||
<collectingCountry id="E58C7630FD0AFFF5FBCFF8775E84F846" box="[1130,1225,1951,1975]" name="Belgium" pageId="55" pageNumber="70">Belgium</collectingCountry>
|
||
, first records in
|
||
<collectingCountry id="E58C7630FD0AFFFAFADEF8775B54FEF8" lastPageId="56" lastPageNumber="71" name="Netherlands" pageId="55" pageNumber="70">the Netherlands</collectingCountry>
|
||
in 1996 and in S
|
||
<collectingCountry id="E58C7630FD05FFFAFE6BFF195865FEF8" box="[462,552,241,265]" name="United Kingdom" pageId="56" pageNumber="71">England</collectingCountry>
|
||
in 2006.
|
||
<location id="9844607BFD05FFFAFD2CFF1A58EDFEF8" LSID="urn:lsid:plazi:treatment:153287B6FD0AFFFAFF71FBAE5826F8A4:9844607BFD05FFFAFD2CFF1A58EDFEF8" box="[649,672,242,265]" country="United Kingdom" name="In" pageId="56" pageNumber="71">In</location>
|
||
2017, 13 sites were known in S
|
||
<collectingCountry id="E58C7630FD05FFFAFE2AFEFA5BA4FEDB" box="[399,489,274,298]" name="United Kingdom" pageId="56" pageNumber="71">England</collectingCountry>
|
||
and
|
||
<quantity id="5A639B45FD05FFFAFDBAFEFB5804FEDB" box="[543,585,274,298]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="56" pageNumber="71" unit="in" value="7.0">7 in</quantity>
|
||
<collectingCountry id="E58C7630FD05FFFAFDF4FEFA58B6FEDB" box="[593,763,274,298]" name="Netherlands" pageId="56" pageNumber="71">the Netherlands</collectingCountry>
|
||
with a northern range border at
|
||
<geoCoordinate id="F8AF5067FD05FFFAFE52FEDB5809FEBA" box="[503,580,307,331]" degrees="52.6" direction="north" orientation="latitude" pageId="56" pageNumber="71" precision="5555" value="52.6">52.6°N</geoCoordinate>
|
||
.
|
||
<collectorName id="306E5376FD05FFFAFDF5FEDB58C2FEBA" box="[592,655,307,331]" pageId="56" pageNumber="71">There</collectorName>
|
||
is a large geographic overlap area with the sister species
|
||
<taxonomicName id="5A9B4D23FD05FFFAFD22FEBD58B1FE9D" authorityName="Zimmerman" authorityYear="1934" box="[647,764,340,364]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="illyricus">
|
||
<emphasis id="AFEFEAB2FD05FFFAFD22FEBD58B1FE9D" box="[647,764,340,364]" italics="true" pageId="56" pageNumber="71">L. illyricus</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<collectorName id="306E5376FD05FFFAFF7EFE9D5B7EFE7C" box="[219,307,373,397]" pageId="56" pageNumber="71">Balkans</collectorName>
|
||
and south
|
||
<collectingCountry id="E58C7630FD05FFFAFE0DFE9D5849FE7C" box="[424,516,373,397]" name="Ukraine" pageId="56" pageNumber="71">Ukraine</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD05FFFAFF0BFE7E5B9BFE5F" box="[174,470,406,430]" pageId="56" pageNumber="71" type="biology_ecology">
|
||
<paragraph id="9D2436A0FD05FFFAFF0BFE7E5B9BFE5F" blockId="56.[151,765,241,1877]" box="[174,470,406,430]" pageId="56" pageNumber="71">
|
||
<emphasis id="AFEFEAB2FD05FFFAFF0BFE7E5B4EFE5F" bold="true" box="[174,259,406,430]" pageId="56" pageNumber="71">Biology</emphasis>
|
||
. See Seifert (2018).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD05FFFAFF0BFE5F5B93FAF4" pageId="56" pageNumber="71" type="diagnosis">
|
||
<paragraph id="9D2436A0FD05FFFAFF0BFE5F5877FDE1" blockId="56.[151,765,241,1877]" pageId="56" pageNumber="71">
|
||
<emphasis id="AFEFEAB2FD05FFFAFF0BFE5F5B50FE3E" bold="true" box="[174,285,439,463]" pageId="56" pageNumber="71">Diagnosis</emphasis>
|
||
(
|
||
<tableCitation id="D019031BFD05FFFAFE8BFE5F5B38FE3E" box="[302,373,439,463]" captionStart="Tab" captionStartId="76.[277,325,397,423]" captionTargetBox="[278,1394,469,1686]" captionTargetId="graphics-59@76.[273,1400,460,1690]" captionTargetPageId="76" captionText="Tab. 8: Westpalaearctic species related to Lasius emarginatus" pageId="56" pageNumber="71">Tab. 8</tableCitation>
|
||
,
|
||
<figureCitation id="05A02A25FD05FFFAFE26FE5F5B97FE3E" box="[387,474,439,463]" captionStart="Figs" captionStartId="26.[151,191,1192,1214]" captionTargetBox="[153,1318,748,1176]" captionTargetId="figure-44@26.[725,1321,748,1177]" captionTargetPageId="26" captionText="Figs. 75 and 76: Lasius emarginatus" figureDoi="http://doi.org/10.5281/zenodo.10832304" httpUri="https://zenodo.org/record/10832304/files/figure.png" pageId="56" pageNumber="71">Figs. 75</figureCitation>
|
||
–76; key; images in www. AntWeb.org with specimen identifiers CASENT0172762, CASENT0179933, CASENT0280445):
|
||
</paragraph>
|
||
<paragraph id="9D2436A0FD05FFFAFF0BFDF15B93FAF4" blockId="56.[151,765,241,1877]" pageId="56" pageNumber="71">
|
||
Absolute size rather large (CS 962 µm). Head and scape length indices large (CL/CW
|
||
<subScript id="011F34E5FD05FFFAFD8EFDA2580EFDA9" attach="left" box="[555,579,586,600]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
1.085, SL/CS
|
||
<subScript id="011F34E5FD05FFFAFD46FDA258B6FDA9" attach="left" box="[739,763,586,600]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
1.067); postocular distance small, eye and torulo-clypeal distance large (PoOc/CL
|
||
<subScript id="011F34E5FD05FFFAFE1CFD645B9CFD6B" attach="left" box="[441,465,652,666]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
0.217, EYE/CS
|
||
<subScript id="011F34E5FD05FFFAFD35FD6458E5FD6B" attach="left" box="[656,680,652,666]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
0.253, dClAn
|
||
<subScript id="011F34E5FD05FFFAFF45FD455AB5FD4A" attach="left" box="[224,248,685,699]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
5.38); terminal segment of maxillary palp long (MP6/CS
|
||
<subScript id="011F34E5FD05FFFAFF59FD265B59FD2D" attach="left" box="[252,276,718,732]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
0.221). Number of mandibular dents large (MaDe
|
||
<subScript id="011F34E5FD05FFFAFF41FD075AB1FD0C" attach="left" box="[228,252,751,765]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
8.76). Pubescence on clypeus dilute (sqPDCL
|
||
<subScript id="011F34E5FD05FFFAFD46FD0758B6FD0C" attach="left" box="[739,763,751,765]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
5.14); frontal pubescence short (PLF
|
||
<subScript id="011F34E5FD05FFFAFD8BFCF8580BFCEF" attach="left" box="[558,582,784,798]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
24.8). All body parts with standing setae of medium length, (PnHL/ CS
|
||
<subScript id="011F34E5FD05FFFAFF12FCB95A82FCAE" attach="left" box="[183,207,849,863]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
0.139, GuHL/CS
|
||
<subScript id="011F34E5FD05FFFAFE23FCB95BD3FCAE" attach="left" box="[390,414,849,863]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
0.126, nOcc
|
||
<subScript id="011F34E5FD05FFFAFD87FCB95877FCAE" attach="left" box="[546,570,849,863]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
12.2, nGen
|
||
<subScript id="011F34E5FD05FFFAFD13FCB95883FCAE" attach="left" box="[694,718,849,863]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
6.5, nGu
|
||
<subScript id="011F34E5FD05FFFAFF63FC9A5A93FC71" attach="left" box="[198,222,882,896]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
6.8, nSc
|
||
<subScript id="011F34E5FD05FFFAFE92FC9A5B02FC71" attach="left" box="[311,335,882,896]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
10.4, nHT
|
||
<subScript id="011F34E5FD05FFFAFE64FC9A5B94FC71" attach="left" box="[449,473,882,896]" fontSize="6" pageId="56" pageNumber="71">900</subScript>
|
||
18.1). The propodeal dome is (as in
|
||
<taxonomicName id="5A9B4D23FD05FFFAFF5AFC6C5BE7FC6A" authorityName="Zimmerman" authorityYear="1934" box="[255,426,899,923]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="illyricus">
|
||
<emphasis id="AFEFEAB2FD05FFFAFF5AFC6C5BE7FC6A" box="[255,426,899,923]" italics="true" pageId="56" pageNumber="71">Lasius illyricus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5A9B4D23FD05FFFAFE19FC6C5877FC6A" authorityName="Seifert" authorityYear="1992" box="[444,570,899,923]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="tebessae">
|
||
<emphasis id="AFEFEAB2FD05FFFAFE19FC6C5877FC6A" box="[444,570,899,923]" italics="true" pageId="56" pageNumber="71">L. tebessae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5A9B4D23FD05FFFAFDDDFC6C58B1FC6A" authority="Seifert, 2020" authorityName="Seifert" authorityYear="2020" box="[632,764,899,923]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="maltaeus" status="sp. nov.">
|
||
<emphasis id="AFEFEAB2FD05FFFAFDDDFC6C58B1FC6A" box="[632,764,899,923]" italics="true" pageId="56" pageNumber="71">L. maltaeus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4DC57C9FD05FFFAFF32FC4C5ABDFC4D" box="[151,240,932,956]" pageId="56" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) higher than in usually seen in
|
||
<taxonomicName id="5A9B4D23FD05FFFAFDD1FC4D58F6FC4D" authorityName="Fabricius" authorityYear="1804" box="[628,699,933,956]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="AFEFEAB2FD05FFFAFDD1FC4D58F6FC4D" box="[628,699,933,956]" italics="true" pageId="56" pageNumber="71">Lasius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4DC57C9FD05FFFAFD6CFC4C58B0FC4A" box="[713,765,932,955]" pageId="56" pageNumber="71" sensu="stricto">s.str.</taxonomicNameLabel>
|
||
Coloration: Two color morphs occur. The light morph has head, coxae, femora and tibiae medium reddish-brown to dark brown with a reddish tinge whereas mesosoma, anterior clypeal margin, scape, petiole and tarsae are orange. The dark morph is almost concolorous medium to dark brown, is restricted to the south Balkans and may occur sympatric with the light morph. Attempts to separate the light and dark color morph by exploratory or hypothesis-driven data analyses using the 16 standard NUMOBAT characters failed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD05FFFAFF0BFAE55826F8A4" pageId="56" pageNumber="71" type="discussion">
|
||
<paragraph id="9D2436A0FD05FFFAFF0BFAE55826F8A4" blockId="56.[151,765,241,1877]" pageId="56" pageNumber="71">
|
||
<emphasis id="AFEFEAB2FD05FFFAFF0BFAE55B68FAD4" bold="true" box="[174,293,1293,1317]" pageId="56" pageNumber="71">Comments</emphasis>
|
||
. The Westpalaearctic species of the
|
||
<taxonomicName id="5A9B4D23FD05FFFAFD43FAE65B57FAB7" baseAuthorityName="Olivier" baseAuthorityYear="1791" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="emarginatus">
|
||
<emphasis id="AFEFEAB2FD05FFFAFD43FAE65B57FAB7" italics="true" pageId="56" pageNumber="71">L. emarginatus</emphasis>
|
||
</taxonomicName>
|
||
complex are characterized by a combination of elongated head, long scape, long maxillary palps, large eyes, large torulo-clypeal distance, short frontal pubescence and the gular setae not being much shorter or equal in length to pronotal setae (
|
||
<tableCitation id="D019031BFD05FFFAFEF4FA5A5BDEFA3B" box="[337,403,1458,1482]" captionStart="Tab" captionStartId="76.[277,325,397,423]" captionTargetBox="[278,1394,469,1686]" captionTargetId="graphics-59@76.[273,1400,460,1690]" captionTargetPageId="76" captionText="Tab. 8: Westpalaearctic species related to Lasius emarginatus" pageId="56" pageNumber="71">Tab. 8</tableCitation>
|
||
). The only species of this complex occurring sympatric with
|
||
<taxonomicName id="5A9B4D23FD05FFFAFE11FA3C5815FA1A" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[436,600,1492,1515]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="emarginatus">
|
||
<emphasis id="AFEFEAB2FD05FFFAFE11FA3C5815FA1A" box="[436,600,1492,1515]" italics="true" pageId="56" pageNumber="71">L. emarginatus</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="5A9B4D23FD05FFFAFD25FA3C58B9FA1A" authorityName="Zimmerman" authorityYear="1934" box="[640,756,1491,1515]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="illyricus">
|
||
<emphasis id="AFEFEAB2FD05FFFAFD25FA3C58B9FA1A" box="[640,756,1491,1515]" italics="true" pageId="56" pageNumber="71">L. illyricus</emphasis>
|
||
</taxonomicName>
|
||
. The mean classification errors for 52 and 39 samples of
|
||
<taxonomicName id="5A9B4D23FD05FFFAFF32F9FD5B77F9DD" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[151,314,1557,1580]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="emarginatus">
|
||
<emphasis id="AFEFEAB2FD05FFFAFF32F9FD5B77F9DD" box="[151,314,1557,1580]" italics="true" pageId="56" pageNumber="71">L. emarginatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5A9B4D23FD05FFFAFED6F9FD5BAAF9DC" authorityName="Zimmerman" authorityYear="1934" box="[371,487,1557,1581]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="illyricus">
|
||
<emphasis id="AFEFEAB2FD05FFFAFED6F9FD5BAAF9DC" box="[371,487,1557,1581]" italics="true" pageId="56" pageNumber="71">L. illyricus</emphasis>
|
||
</taxonomicName>
|
||
respectively, in which all standard NUMOBAT characters have been recorded, were 3.3% in NC-Ward, 2.2% in NC-part.hclust, 0% in NC-part. kmeans and 0% in NC-NMDS.kmeans. A mean error rate of 1.4% relative to the controlling discriminant function is a good confirmation of heterospecificity according to the GAGE species concept. The classification error by the LDA on the individual level was 2.2% for
|
||
<specimenCount id="8B9DFD29FD05FFFAFDF8F9135893F8E2" box="[605,734,1787,1811]" count="232" pageId="56" pageNumber="71" type="worker">232 workers</specimenCount>
|
||
of both species. For separation from the allopatric species
|
||
<taxonomicName id="5A9B4D23FD05FFFAFD43F8F45AB4F8A4" authority="Seifert, 2020" authorityName="Seifert" authorityYear="2020" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="maltaeus" status="sp. nov.">
|
||
<emphasis id="AFEFEAB2FD05FFFAFD43F8F45AB4F8A4" italics="true" pageId="56" pageNumber="71">L. maltaeus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4DC57C9FD05FFFAFEA5F8D55B1FF8A5" box="[256,338,1853,1876]" pageId="56" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="5A9B4D23FD05FFFAFE20F8D55BB3F8A4" authorityName="Seifert" authorityYear="1992" box="[389,510,1853,1877]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="71" phylum="Arthropoda" rank="species" species="tebessae">
|
||
<emphasis id="AFEFEAB2FD05FFFAFE20F8D55BB3F8A4" box="[389,510,1853,1877]" italics="true" pageId="56" pageNumber="71">L. tebessae</emphasis>
|
||
</taxonomicName>
|
||
see there.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |