324 lines
39 KiB
XML
324 lines
39 KiB
XML
<document id="AD9C7CA29D8D6B94B8D4B91300A5AA9A" ID-CLB-Dataset="54139" ID-DOI="10.11646/zootaxa.5243.1.1" ID-GBIF-Dataset="1476f0e1-3886-401f-bf56-c294a0a99962" ID-ISSN="1175-5326" ID-Zenodo-Dep="7642400" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1676448183279" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Sunbin & Jia, Xinyang" docDate="2023" docId="039587ADBF00FF95FF7F450726B4FE58" docLanguage="en" docName="zootaxa.5243.1.1.pdf" docOrigin="Zootaxa 5243 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5243.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Guizhaphaenops wanghaoi Tian & Huang & Jia 2023, n. sp." docType="treatment" docVersion="1" lastPageNumber="61" masterDocId="FFACFFD5BF38FFA9FFE841652260FFF5" masterDocTitle="A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="57" updateTime="1731937328377" updateUser="felipe">
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<mods:title id="7C247B35FB9FCDC7FC11AD9557C46701">A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:namePart id="9CFB99742D69EB64C9AA9298402A1881">Tian, Mingyi</mods:namePart>
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<mods:namePart id="B4EC4671538DEEF0E33407F03BA7C2DA">Huang, Sunbin</mods:namePart>
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<mods:namePart id="B3B9F9DAB8541A022FAAB0289B910197">Jia, Xinyang</mods:namePart>
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<mods:date id="0B4C8C5346CF64A77318759E86A2F76A">2023</mods:date>
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<mods:number id="C6D3AAAAC8222230862680DEF622A77F">2023-02-15</mods:number>
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<treatment id="039587ADBF00FF95FF7F450726B4FE58" ID-DOI="http://doi.org/10.5281/zenodo.7642394" ID-GBIF-Taxon="209296928" ID-Zenodo-Dep="7642394" LSID="urn:lsid:plazi:treatment:039587ADBF00FF95FF7F450726B4FE58" httpUri="http://treatment.plazi.org/id/039587ADBF00FF95FF7F450726B4FE58" lastPageId="60" lastPageNumber="61" pageId="56" pageNumber="57">
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<subSubSection id="C3266530BF00FF91FF7F450720E5FB88" box="[151,645,1122,1149]" pageId="56" pageNumber="57" type="nomenclature">
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<paragraph id="8B8336BBBF00FF91FF7F450720E5FB88" blockId="56.[151,645,1122,1185]" box="[151,645,1122,1149]" pageId="56" pageNumber="57">
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<heading id="D0CB81D7BF00FF91FF7F450720E5FB88" bold="true" box="[151,645,1122,1149]" fontSize="11" level="2" pageId="56" pageNumber="57" reason="4">
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<emphasis id="B948EAA9BF00FF91FF7F450720E5FB88" bold="true" box="[151,645,1122,1149]" pageId="56" pageNumber="57">
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<taxonomicName id="4C3C4D38BF00FF91FF7F4507205BFB88" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[151,571,1122,1149]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="wanghaoi" status="sp. nov.">
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<emphasis id="B948EAA9BF00FF91FF7F4507205BFB88" bold="true" box="[151,571,1122,1149]" italics="true" pageId="56" pageNumber="57">Guizhaphaenops (s. str.) wanghaoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B57D2BF00FF91FDAA450620E5FB88" box="[578,645,1123,1149]" pageId="56" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3266530BF00FF91FF7F45E22301FB54" box="[151,353,1159,1185]" pageId="56" pageNumber="57" type="description">
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<paragraph id="8B8336BBBF00FF91FF7F45E22301FB54" blockId="56.[151,645,1122,1185]" box="[151,353,1159,1185]" pageId="56" pageNumber="57">
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<figureCitation id="13072A3EBF00FF91FF7F45E2236EFB54" box="[151,270,1159,1185]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="56" pageNumber="57">Figures 1e</figureCitation>
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,
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<figureCitation id="13072A3EBF00FF91FEF345E22301FB54" box="[283,353,1159,1185]" captionStart-0="FIGURE 38" captionStart-1="FIGURE 39" captionStart-2="FIGURE 40" captionStartId-0="57.[151,250,1944,1968]" captionStartId-1="58.[151,250,1982,2006]" captionStartId-2="59.[151,250,1973,1997]" captionTargetBox-1="[168,1419,1030,1951]" captionTargetBox-2="[162,1426,345,1950]" captionTargetId-0="figure-20@57.[349,1239,181,1921]" captionTargetId-1="figure-447@58.[167,1419,1029,1959]" captionTargetId-2="figure-104@59.[162,1426,345,1950]" captionTargetPageId-0="57" captionTargetPageId-1="58" captionTargetPageId-2="59" captionText-0="FIGURE 38. Habitus of Guizhaphaenops (s. str.) wanghaoi n. sp., male, holotype." captionText-1="FIGURE 39. Guizhaphaenops (s. str.) wanghaoi n. sp. A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view." captionText-2="FIGURE 40. Weixiao Tiankeng, the type locality of Guizhaphaenops (s. str.) wanghaoi n. sp. A. environ of Weixiao Tiankeng, bird view; B. a descending caver; C. the main passage below the Tiankeng (photos courtesy of Ms. Rurui Ye)." figureDoi-0="http://doi.org/10.5281/zenodo.7647036" figureDoi-1="http://doi.org/10.5281/zenodo.7642466" figureDoi-2="http://doi.org/10.5281/zenodo.7642468" httpUri-0="https://zenodo.org/record/7647036/files/figure.png" httpUri-1="https://zenodo.org/record/7642466/files/figure.png" httpUri-2="https://zenodo.org/record/7642468/files/figure.png" pageId="56" pageNumber="57">38–40</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3266530BF00FF91FF7F45AB2167FAC4" pageId="56" pageNumber="57" type="materials_examined">
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<paragraph id="8B8336BBBF00FF91FF7F45AB2167FAC4" blockId="56.[151,1437,1230,2013]" pageId="56" pageNumber="57">
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<emphasis id="B948EAA9BF00FF91FF7F45AB2366FB1D" bold="true" box="[151,262,1230,1256]" pageId="56" pageNumber="57">Material.</emphasis>
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<materialsCitation id="3B543CE6BF00FF91FEF845AA2335FAC4" ID-GBIF-Occurrence="4030741309" collectingDate="2022-03-21" collectionCode="SCAU" collectorName="Hao Wang" country="China" county="Qinglong County" elevation="1345" latitude="23.16" location="Weixiao Tiankeng" longLatPrecision="754" longitude="113.36" municipality="Gaoling" pageId="56" pageNumber="57" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Guizhou Province" typeStatus="holotype">
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<typeStatus id="54878819BF00FF91FEF845AA2318FB1C" box="[272,376,1231,1257]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3AFD32BF00FF91FE6A45AA23DBFB1C" box="[386,443,1231,1257]" pageId="56" pageNumber="57" type="male">male</specimenCount>
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,
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<location id="8EE36060BF00FF91FE2145AA20FBFB1C" LSID="urn:lsid:plazi:treatment:039587ADBF00FF95FF7F450726B4FE58:8EE36060BF00FF91FE2145AA20FBFB1C" box="[457,667,1231,1257]" country="China" county="Qinglong County" latitude="23.16" longLatPrecision="754" longitude="113.36" municipality="Gaoling" name="Weixiao Tiankeng" pageId="56" pageNumber="57" stateProvince="Guizhou Province">Weixiao Tiankeng</location>
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,
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<collectingMunicipality id="6BE7ACC1BF00FF91FD4145AA2166FB1C" box="[681,774,1231,1257]" pageId="56" pageNumber="57">Gaoling</collectingMunicipality>
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,
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<collectingCounty id="62E24E37BF00FF91FCFC45AA21BCFB1C" box="[788,988,1231,1257]" pageId="56" pageNumber="57">Qinglong County</collectingCounty>
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,
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<collectingCounty id="62E24E37BF00FF91FC0145AA236DFAF8" pageId="56" pageNumber="57">Qianxinan Buyi & Miao Autonomous Prefecture</collectingCounty>
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, southwestern
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<collectingRegion id="49F8F859BF00FF91FE5A4596201FFAF8" box="[434,639,1267,1293]" country="China" name="Guizhou" pageId="56" pageNumber="57">Guizhou Province</collectingRegion>
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, southwestern
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<collectingCountry id="F32B762BBF00FF91FCCC4596210AFAF8" box="[804,874,1267,1293]" name="China" pageId="56" pageNumber="57">China</collectingCountry>
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,
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<geoCoordinate id="EE08507CBF00FF91FC9C459621B1FAF8" box="[884,977,1267,1293]" degrees="23.16" direction="north" orientation="latitude" pageId="56" pageNumber="57" precision="555" value="23.16">23.16 N</geoCoordinate>
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,
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<geoCoordinate id="EE08507CBF00FF91FC3345962621FAF8" box="[987,1089,1267,1293]" degrees="113.36" direction="east" orientation="longitude" pageId="56" pageNumber="57" precision="555" value="113.36">113.36 E</geoCoordinate>
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,
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<quantity id="4CC49B5EBF00FF91FBA4459726C3FAF8" box="[1100,1187,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="56" pageNumber="57" unit="m" value="1345.0">
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<elevation id="0011D188BF00FF91FBA4459726C3FAF8" box="[1100,1187,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="56" pageNumber="57" unit="m" value="1345.0">1345 m</elevation>
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</quantity>
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,
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<date id="FF82107BBF00FF91FB4645962754FAF8" box="[1198,1332,1267,1293]" pageId="56" pageNumber="57" value="2022-03-21">
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<collectingDate id="EFC6E993BF00FF91FB4645962754FAF8" box="[1198,1332,1267,1293]" pageId="56" pageNumber="57" value="2022-03-21">2022-III-21</collectingDate>
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</date>
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, leg.
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<collectorName id="26C9536DBF00FF91FA86459622BBFAC4" pageId="56" pageNumber="57">Hao Wang</collectorName>
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, in
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<collectionCode id="ED2DAE7EBF00FF91FEEB44722331FAC4" box="[259,337,1303,1329]" pageId="56" pageNumber="57">SCAU</collectionCode>
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;
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</materialsCitation>
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<materialsCitation id="3B543CE6BF00FF91FEB444722167FAC4" ID-GBIF-Occurrence="4030741308" box="[348,775,1303,1329]" collectingDate="2022-03-21" collectionCode="SCAU" collectorName="Hao Wang" country="China" county="Qinglong County" elevation="1345" latitude="23.16" location="Weixiao Tiankeng" longLatPrecision="754" longitude="113.36" municipality="Gaoling" pageId="56" pageNumber="57" specimenCount="2" specimenCount-female="2" stateProvince="Guizhou Province" typeStatus="paratype">
|
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<typeStatus id="54878819BF00FF91FEB4447223ACFAC4" box="[348,460,1303,1329]" pageId="56" pageNumber="57" type="paratype">paratypes</typeStatus>
|
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:
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<specimenCount id="9D3AFD32BF00FF91FE3E44722025FAC4" box="[470,581,1303,1329]" pageId="56" pageNumber="57" type="female">2 females</specimenCount>
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, IBID, in
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<collectionCode id="ED2DAE7EBF00FF91FD5D44722163FAC4" box="[693,771,1303,1329]" collectionName="SCAU" pageId="56" pageNumber="57">SCAU</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3266530BF00FF91FF2F445F238AFA8C" pageId="56" pageNumber="57" type="diagnosis">
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<paragraph id="8B8336BBBF00FF91FF2F445F238AFA8C" blockId="56.[151,1437,1230,2013]" pageId="56" pageNumber="57">
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<emphasis id="B948EAA9BF00FF91FF2F445F2322FAA1" bold="true" box="[199,322,1338,1364]" pageId="56" pageNumber="57">Diagnosis.</emphasis>
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Medium-sized trechine, semi-aphaenopsian, eyeless and depigmented, body glabrous and concolorous yellowish brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3266530BF00FF93FF2F44E7271FFC18" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57" type="description">
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<paragraph id="8B8336BBBF00FF91FF2F44E72623FA68" blockId="56.[151,1437,1230,2013]" box="[199,1091,1410,1437]" pageId="56" pageNumber="57">
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||
<emphasis id="B948EAA9BF00FF91FF2F44E72339FA69" bold="true" box="[199,345,1410,1436]" pageId="56" pageNumber="57">Description.</emphasis>
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||
Length:
|
||
<quantity id="4CC49B5EBF00FF91FE5644E62028FA68" box="[446,584,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="6.699999999999999" metricValueMax="6.8" metricValueMin="6.6" pageId="56" pageNumber="57" unit="mm" value="6.699999999999999" valueMax="6.8" valueMin="6.6">6.6–6.8 mm</quantity>
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; width:
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<quantity id="4CC49B5EBF00FF91FD4A44E6214CFA68" box="[674,812,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="56" pageNumber="57" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.3–2.4 mm</quantity>
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||
. Habitus as in
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||
<figureCitation id="13072A3EBF00FF91FC2744E6265FFA68" box="[975,1087,1411,1437]" captionStart="FIGURE 38" captionStartId="57.[151,250,1944,1968]" captionTargetId="figure-20@57.[349,1239,181,1921]" captionTargetPageId="57" captionText="FIGURE 38. Habitus of Guizhaphaenops (s. str.) wanghaoi n. sp., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7647036" httpUri="https://zenodo.org/record/7647036/files/figure.png" pageId="56" pageNumber="57">Figure 38</figureCitation>
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.
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</paragraph>
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<paragraph id="8B8336BBBF00FF91FF2F44C226BAF9FC" blockId="56.[151,1437,1230,2013]" pageId="56" pageNumber="57">Body yellowish brown, mouthparts, palps and antennae pale. Head, pronotum, elytra and underside of head and thorax smooth and glabrous. Microsculptural engraved meshes more or less isometric on head, while transversely striate on pronotum and elytra. Fore body much shorter than elytra, (HLm+PL) / EL = 0.86–0.87.</paragraph>
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<paragraph id="8B8336BBBF00FF91FF2F4776263BF828" blockId="56.[151,1437,1230,2013]" pageId="56" pageNumber="57">
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Head elongate, longer than wide, HLm/HW = 1.95–1.96, HLl/HW = 1.45–1.57; genae feebly expanded, widest at about middle of head excluding mandibles; frons and vertex moderately convex; clypeus transverse, 4-setose; labrum shallowly emarginate at frontal margin, 6-setose, median two smaller and shorter; frontal furrows subparallel-sided in basal half, strongly divergent posteriorly; mandibles moderately developed, feebly curved apically, right mandibular tooth bidentate; only anterior supraorbital setiferous pores present, located at about the middle of the head respectively; labial suture absent, mentum bisetose, tooth short and bifid at tip, base largely concave; submentum 10-setose; ligula adnated with paraglossae, 8-setose at apex; palpomeres thin and elongate, the 2
|
||
<superScript id="7C499BF3BF00FF91FAA0478F2738F90D" attach="left" box="[1352,1368,1770,1784]" fontSize="6" pageId="56" pageNumber="57">nd</superScript>
|
||
labial palpomere bisetose on inner margin, with an additional seta on outer margin, the 3
|
||
<superScript id="7C499BF3BF00FF91FBC2466B2658F8E9" attach="left" box="[1066,1080,1806,1820]" fontSize="6" pageId="56" pageNumber="57">rd</superScript>
|
||
maxillary palpomere with three setae at apex; 3
|
||
<superScript id="7C499BF3BF00FF91FEAD46572333F8B5" attach="left" box="[325,339,1842,1856]" fontSize="6" pageId="56" pageNumber="57">rd</superScript>
|
||
labial and 4
|
||
<superScript id="7C499BF3BF00FF91FE364657238BF8B5" attach="left" box="[478,491,1842,1856]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
palpomere maxillary palpomeres smooth and glabrous; the 2
|
||
<superScript id="7C499BF3BF00FF91FB76465726CEF8B5" attach="left" box="[1182,1198,1842,1856]" fontSize="6" pageId="56" pageNumber="57">nd</superScript>
|
||
labial palpomere 1.3 times longer than 3
|
||
<superScript id="7C499BF3BF00FF91FE9F463323E5F891" attach="none" box="[375,389,1878,1892]" fontSize="6" pageId="56" pageNumber="57">rd</superScript>
|
||
, the 3
|
||
<superScript id="7C499BF3BF00FF91FE27463323BDF891" attach="left" box="[463,477,1878,1892]" fontSize="6" pageId="56" pageNumber="57">rd</superScript>
|
||
maxillary palpomere 1.2 times longer than 4
|
||
<superScript id="7C499BF3BF00FF91FC0F46332194F891" attach="none" box="[999,1012,1878,1892]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
; antennae pubescent from pedicel to 11
|
||
<superScript id="7C499BF3BF00FF91FF5A461F22DFF87D" attach="left" box="[178,191,1914,1928]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
antennomere; scape with several long setae, smooth and slightly stouter than other, the 10
|
||
<superScript id="7C499BF3BF00FF91FB39461F26BEF87D" attach="left" box="[1233,1246,1914,1928]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
the shortest; the comparative length of each antennomere from the 1
|
||
<superScript id="7C499BF3BF00FF91FD0046FA2093F858" attach="left" box="[744,755,1951,1965]" fontSize="6" pageId="56" pageNumber="57">st</superScript>
|
||
to 11
|
||
<superScript id="7C499BF3BF00FF91FCDD46FB2122F859" attach="left" box="[821,834,1950,1964]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
in
|
||
<typeStatus id="54878819BF00FF91FC8146FA21ABF84C" box="[873,971,1951,1977]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
as: 1
|
||
<superScript id="7C499BF3BF00FF91FBE246FA2675F858" attach="left" box="[1034,1045,1951,1965]" fontSize="6" pageId="56" pageNumber="57">st</superScript>
|
||
(1.15) / 2
|
||
<superScript id="7C499BF3BF00FF91FB6146FB26F9F859" attach="left" box="[1161,1177,1950,1964]" fontSize="6" pageId="56" pageNumber="57">nd</superScript>
|
||
(1.25) / 3
|
||
<superScript id="7C499BF3BF00FF91FAE646FB277CF859" attach="left" box="[1294,1308,1950,1964]" fontSize="6" pageId="56" pageNumber="57">rd</superScript>
|
||
(1.55) / 4
|
||
<superScript id="7C499BF3BF00FF91FA7846FB27FDF859" attach="none" box="[1424,1437,1950,1964]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.50) / 5
|
||
<superScript id="7C499BF3BF00FF91FF1746A7236CF825" attach="left" box="[255,268,1986,2000]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.40) / 6
|
||
<superScript id="7C499BF3BF00FF91FE9246A723E7F825" attach="left" box="[378,391,1986,2000]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.30) / 7
|
||
<superScript id="7C499BF3BF00FF91FE1D46A72062F825" attach="left" box="[501,514,1986,2000]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.25) / 8
|
||
<superScript id="7C499BF3BF00FF91FD9846A7201DF825" attach="left" box="[624,637,1986,2000]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.15) / 9
|
||
<superScript id="7C499BF3BF00FF91FD0346A72098F825" attach="left" box="[747,760,1986,2000]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.10) / 10
|
||
<superScript id="7C499BF3BF00FF91FC9C46A721E1F825" attach="left" box="[884,897,1986,2000]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.00) / 11
|
||
<superScript id="7C499BF3BF00FF91FC1446A72669F825" attach="left" box="[1020,1033,1986,2000]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
(1.35).
|
||
</paragraph>
|
||
<caption id="DF436633BF01FF90FF7F46FD2181F844" ID-DOI="http://doi.org/10.5281/zenodo.7647036" ID-Zenodo-Dep="7647036" box="[151,993,1944,1969]" httpUri="https://zenodo.org/record/7647036/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1944,1968]" targetBox="[350,1222,197,1901]" targetPageId="57">
|
||
<paragraph id="8B8336BBBF01FF90FF7F46FD2181F844" blockId="57.[151,993,1944,1969]" box="[151,993,1944,1969]" pageId="57" pageNumber="58">
|
||
<emphasis id="B948EAA9BF01FF90FF7F46FD2340F845" bold="true" box="[151,288,1944,1968]" pageId="57" pageNumber="58">FIGURE 38.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C3C4D38BF01FF90FE7F46FC2097F844" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[407,759,1945,1969]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="wanghaoi" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF01FF90FE7F46FC2097F844" box="[407,759,1945,1969]" italics="true" pageId="57" pageNumber="58">Guizhaphaenops (s. str.) wanghaoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF01FF90FD1646FC2157F844" bold="true" box="[766,823,1945,1969]" pageId="57" pageNumber="58">
|
||
<taxonomicNameLabel id="A27B57D2BF01FF90FD1646FC2157F844" box="[766,823,1945,1969]" pageId="57" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8336BBBF02FF93FF2F41F22186FE7C" blockId="58.[151,1437,151,1005]" pageId="58" pageNumber="59">Propleura invisible from above. Pronotum sub-orbicular, as long as wide; much wider but shorter than head, PnW/HW = 1.24–1.33, PnL/HLm = 0.71–0.74, PnL/HLl = 0.84–0.86; lateral margins finely bordered and reflexed upwardly throughout, fore angles slightly protruding and obtuse, hind angles widely obtuse; widest at about apical 3/7; only the anterior marginal seta present, located at about apical 1/6 of pronotum, the posterior absent; base slightly narrower than front, PbW/PfW = 0.73–0.78, both unbordered; front slightly emarginate, while base nearly straight, but shortly curved near hind angles; disc moderately convex; fore and posterior transverse impressions weakly-marked, basal foveae indistinct; median line clear. Scutellum small.</paragraph>
|
||
<paragraph id="8B8336BBBF02FF93FF2F40F62645FD5C" blockId="58.[151,1437,151,1005]" pageId="58" pageNumber="59">
|
||
Elytra elongate ovate, much longer than wide, EL/EW = 1.58–1.60; slightly widened apically; much wider than pronotum, EW/PnW = 1.83–1.87; lateral margins finely bordered and smooth throughout, but ciliate near humeral portion, widest at about middle; disc strongly convex, striae shallow, punctures visible, intervals flat, apical striole not well-marked; chaetotaxy (
|
||
<figureCitation id="13072A3EBF02FF93FE0F409A200FFDEC" box="[487,623,511,537]" captionStart="FIGURE 39" captionStartId="58.[151,250,1982,2006]" captionTargetBox="[168,1419,1030,1951]" captionTargetId="figure-447@58.[167,1419,1029,1959]" captionTargetPageId="58" captionText="FIGURE 39. Guizhaphaenops (s. str.) wanghaoi n. sp. A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642466" httpUri="https://zenodo.org/record/7642466/files/figure.png" pageId="58" pageNumber="59">Figure 39A</figureCitation>
|
||
): basal pore present; only a dorsal setiferous pore present on the 3
|
||
<superScript id="7C499BF3BF02FF93FABF409B2705FDF9" attach="left" box="[1367,1381,510,524]" fontSize="6" pageId="58" pageNumber="59">rd</superScript>
|
||
stria on each elytron, located at a little before middle; preapical pore present at about apical 2/13 of elytra, located in the anastomosis of 3
|
||
<superScript id="7C499BF3BF02FF93FEB94323233FFDA1" attach="left" box="[337,351,582,596]" fontSize="6" pageId="58" pageNumber="59">rd</superScript>
|
||
and 4
|
||
<superScript id="7C499BF3BF02FF93FE49432323CEFDA1" attach="left" box="[417,430,582,596]" fontSize="6" pageId="58" pageNumber="59">th</superScript>
|
||
striae, closer to elytral suture than to margin; the humeral group of the marginal umbilicate pores not aggregated, the 2
|
||
<superScript id="7C499BF3BF02FF93FE24430F23BCFD8D" attach="left" box="[460,476,618,632]" fontSize="6" pageId="58" pageNumber="59">nd</superScript>
|
||
pore closer to marginal gutter than other, the 4
|
||
<superScript id="7C499BF3BF02FF93FC10430F2665FD8D" attach="left" box="[1016,1029,618,632]" fontSize="6" pageId="58" pageNumber="59">th</superScript>
|
||
pore widely separated from 3
|
||
<superScript id="7C499BF3BF02FF93FAB4430F270AFD8D" attach="none" box="[1372,1386,618,632]" fontSize="6" pageId="58" pageNumber="59">rd</superScript>
|
||
; the middle group (5
|
||
<superScript id="7C499BF3BF02FF93FEA443EB2339FD69" attach="left" box="[332,345,654,668]" fontSize="6" pageId="58" pageNumber="59">th</superScript>
|
||
and 6
|
||
<superScript id="7C499BF3BF02FF93FE7543EB23CAFD69" attach="left" box="[413,426,654,668]" fontSize="6" pageId="58" pageNumber="59">th</superScript>
|
||
pores) close to each other; the anguloapical pore absent.
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF02FF93FF2F43D626EFFCE0" blockId="58.[151,1437,151,1005]" pageId="58" pageNumber="59">
|
||
Legs moderately developed; fore tibia with a longitudinal sulcus on dorsal surface; the 1
|
||
<superScript id="7C499BF3BF02FF93FB3843D626BBFD34" attach="left" box="[1232,1243,691,705]" fontSize="6" pageId="58" pageNumber="59">st</superScript>
|
||
tarsomere much shorter than 2
|
||
<superScript id="7C499BF3BF02FF93FEDB43B32323FD11" attach="none" box="[307,323,726,740]" fontSize="6" pageId="58" pageNumber="59">nd</superScript>
|
||
–4
|
||
<superScript id="7C499BF3BF02FF93FE8843B3230DFD11" attach="left" box="[352,365,726,740]" fontSize="6" pageId="58" pageNumber="59">th</superScript>
|
||
tarsomeres together in fore leg, whereas as long as those in middle and hind legs respectively. The 1
|
||
<superScript id="7C499BF3BF02FF93FF30439E2283FCFC" attach="left" box="[216,227,763,777]" fontSize="6" pageId="58" pageNumber="59">st</superScript>
|
||
protarsomere elongated and slightly enlarged, denticulate inwardly at apex in male.
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF02FF93FF2F427A2162FCA8" blockId="58.[151,1437,151,1005]" pageId="58" pageNumber="59">Ventrites smooth at sides, with some short pubescence medially, IV-VI each with a pair of paramedial setae, VII bisetose apically in male, while quadrisetose in female.</paragraph>
|
||
<paragraph id="8B8336BBBF02FF93FF2F4202271FFC18" blockId="58.[151,1437,151,1005]" pageId="58" pageNumber="59">
|
||
Male genitalia (
|
||
<figureCitation id="13072A3EBF02FF93FE984202206EFC74" box="[368,526,871,897]" captionStart="FIGURE 39" captionStartId="58.[151,250,1982,2006]" captionTargetBox="[168,1419,1030,1951]" captionTargetId="figure-447@58.[167,1419,1029,1959]" captionTargetPageId="58" captionText="FIGURE 39. Guizhaphaenops (s. str.) wanghaoi n. sp. A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642466" httpUri="https://zenodo.org/record/7642466/files/figure.png" pageId="58" pageNumber="59">Figure 39B,C</figureCitation>
|
||
):The median lobe moderately sclerotized, slender and elongate, widely emarginate; base small, with a large sagittal aileron; copulatory piece not well-marked, about 1/4 as long as the median lobe; apical lobe gradually tapered, apex blunt; in dorsal view, apical lobe narrowly rounded at the extremity; parameres moderately developed, similar each other in shape, the right and left each with four and three long setae at apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF436633BF02FF93FF7F46DB204FF80F" ID-DOI="http://doi.org/10.5281/zenodo.7642466" ID-Zenodo-Dep="7642466" httpUri="https://zenodo.org/record/7642466/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,250,1982,2006]" targetBox="[168,1419,1030,1951]" targetPageId="58">
|
||
<paragraph id="8B8336BBBF02FF93FF7F46DB204FF80F" blockId="58.[151,1436,1982,2042]" pageId="58" pageNumber="59">
|
||
<emphasis id="B948EAA9BF02FF93FF7F46DB2341F823" bold="true" box="[151,289,1982,2006]" pageId="58" pageNumber="59">FIGURE 39.</emphasis>
|
||
<taxonomicName id="4C3C4D38BF02FF93FEC046DB20EBF823" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[296,651,1982,2006]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="wanghaoi" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF02FF93FEC046DB20EBF823" box="[296,651,1982,2006]" italics="true" pageId="58" pageNumber="59">Guizhaphaenops (s. str.) wanghaoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF02FF93FD7A46DB20ACF823" bold="true" box="[658,716,1982,2006]" pageId="58" pageNumber="59">
|
||
<taxonomicNameLabel id="A27B57D2BF02FF93FD7A46DB20ACF823" box="[658,716,1982,2006]" pageId="58" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3266530BF03FF92FF2F41F22090FEB4" pageId="59" pageNumber="60" type="discussion">
|
||
<paragraph id="8B8336BBBF03FF92FF2F41F22090FEB4" blockId="59.[151,1436,151,321]" pageId="59" pageNumber="60">
|
||
<emphasis id="B948EAA9BF03FF92FF2F41F2235BFF44" bold="true" box="[199,315,151,177]" pageId="59" pageNumber="60">Remarks.</emphasis>
|
||
This new species is similar to
|
||
<taxonomicName id="4C3C4D38BF03FF92FD7241F22752FF44" authority="Deuve & Queinnec, 2014" authorityName="Deuve & Queinnec" authorityYear="2014" box="[666,1330,151,177]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="pouillyi">
|
||
<emphasis id="B948EAA9BF03FF92FD7241F2266AFF44" box="[666,1034,151,177]" italics="true" pageId="59" pageNumber="60">Guizhaphaenops (s. str.) pouillyi</emphasis>
|
||
<bibRefCitation id="EFAD4B4ABF03FF92FBF941F22752FF44" author="Deuve, T. & Queinnec, E." box="[1041,1330,151,177]" pageId="59" pageNumber="60" pagination="467 - 471" refId="ref27756" refString="Deuve, T. & Queinnec, E. (2014) Deux nouveaux Trechini de la Chine du Sud-Ouest (Coleoptera, Caraboidea, Trechinae). Bulletin de la Societe entomologique de France, 119, 467 - 471. https: // doi. org / 10.3406 / bsef. 2014.2427" type="journal article" year="2014">Deuve & Quéinnec, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
recorded from Yin Dong (also known as Luoshui Dong II) along Gesuo River in Panzhou Qu of Liupanshui Shi, about 34 kilometres in linear distance from Weixiao Tiankeng (
|
||
<bibRefCitation id="EFAD4B4ABF03FF92FD0541BA2669FF0C" author="Deuve, T. & Queinnec, E." box="[749,1033,223,249]" pageId="59" pageNumber="60" pagination="467 - 471" refId="ref27756" refString="Deuve, T. & Queinnec, E. (2014) Deux nouveaux Trechini de la Chine du Sud-Ouest (Coleoptera, Caraboidea, Trechinae). Bulletin de la Societe entomologique de France, 119, 467 - 471. https: // doi. org / 10.3406 / bsef. 2014.2427" type="journal article" year="2014">Deuve & Quéinnec 2014</bibRefCitation>
|
||
). However,
|
||
<taxonomicName id="4C3C4D38BF03FF92FB6641BA2710FF0C" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[1166,1392,223,249]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="wanghaoi">
|
||
<emphasis id="B948EAA9BF03FF92FB6641BA2710FF0C" box="[1166,1392,223,249]" italics="true" pageId="59" pageNumber="60">G. (s. str.) wanghaoi</emphasis>
|
||
</taxonomicName>
|
||
has thinner head and pronotum, presence of posterior dorsal pore and the preapical pore on elytra vs. both absent but presence of anterior dorsal pore in
|
||
<taxonomicName id="4C3C4D38BF03FF92FDF54042208AFEB4" authorityName="Deuve & Queinnec" authorityYear="2014" box="[541,746,295,321]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="pouillyi">
|
||
<emphasis id="B948EAA9BF03FF92FDF54042208AFEB4" box="[541,746,295,321]" italics="true" pageId="59" pageNumber="60">G. (s. str.) pouillyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF436633BF03FF92FF7F46D02690F807" ID-DOI="http://doi.org/10.5281/zenodo.7642468" ID-Zenodo-Dep="7642468" httpUri="https://zenodo.org/record/7642468/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,250,1973,1997]" targetBox="[162,1426,345,1950]" targetPageId="59">
|
||
<paragraph id="8B8336BBBF03FF92FF7F46D02690F807" blockId="59.[151,1437,1973,2034]" pageId="59" pageNumber="60">
|
||
<emphasis id="B948EAA9BF03FF92FF7F46D02340F838" bold="true" box="[151,288,1973,1997]" pageId="59" pageNumber="60">FIGURE 40.</emphasis>
|
||
Weixiao Tiankeng, the type locality of
|
||
<taxonomicName id="4C3C4D38BF03FF92FD4546D0266DF838" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[685,1037,1973,1997]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="wanghaoi" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF03FF92FD4546D0266DF838" box="[685,1037,1973,1997]" italics="true" pageId="59" pageNumber="60">Guizhaphaenops (s. str.) wanghaoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF03FF92FBFB46D3262BF83B" bold="true" box="[1043,1099,1974,1998]" pageId="59" pageNumber="60">
|
||
<taxonomicNameLabel id="A27B57D2BF03FF92FBFB46D3262BF83B" box="[1043,1099,1974,1998]" pageId="59" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. environ of Weixiao Tiankeng, bird view; B. a descending caver; C. the main passage below the Tiankeng (photos courtesy of Ms. Rurui Ye).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3266530BF04FF95FF2F41F22376FF20" pageId="60" pageNumber="61" type="etymology">
|
||
<paragraph id="8B8336BBBF04FF95FF2F41F22376FF20" blockId="60.[151,1437,151,429]" pageId="60" pageNumber="61">
|
||
<emphasis id="B948EAA9BF04FF95FF2F41F2232EFF44" bold="true" box="[199,334,151,177]" pageId="60" pageNumber="61">Etymology.</emphasis>
|
||
In honour of Mr. Hao Wang (an member of “Explore
|
||
<collectingCountry id="F32B762BBF04FF95FC5D41F2219AFF44" box="[949,1018,151,177]" name="China" pageId="60" pageNumber="61">China</collectingCountry>
|
||
” team), who found and collected the
|
||
<typeStatus id="54878819BF04FF95FF7F41DE22A7FF20" box="[151,199,187,213]" pageId="60" pageNumber="61">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3266530BF04FF95FF2F41BB26B4FE58" pageId="60" pageNumber="61" type="distribution">
|
||
<paragraph id="8B8336BBBF04FF95FF2F41BB2720FF0C" blockId="60.[151,1437,151,429]" box="[199,1344,222,249]" pageId="60" pageNumber="61">
|
||
<emphasis id="B948EAA9BF04FF95FF2F41BB2301FF0D" bold="true" box="[199,353,222,248]" pageId="60" pageNumber="61">Distribution.</emphasis>
|
||
<collectingCountry id="F32B762BBF04FF95FE8F41BA23CAFF0C" box="[359,426,223,249]" name="China" pageId="60" pageNumber="61">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49F8F859BF04FF95FE5141BA2040FF0C" box="[441,544,223,249]" country="China" name="Guizhou" pageId="60" pageNumber="61">Guizhou</collectingRegion>
|
||
). Known only from Weixiao Tiankeng in Qinglong County (
|
||
<figureCitation id="13072A3EBF04FF95FB2B41BA2750FF0C" box="[1219,1328,223,249]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="60" pageNumber="61">Figure 1e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF04FF95FF2F406626B4FE58" blockId="60.[151,1437,151,429]" pageId="60" pageNumber="61">
|
||
Weixiao (which means “smiling” in Chinese) Tiankeng is located near Gaoling Village in southwestern Qinglong County. It has a wide and smiling-like opening and a depth of about 210 metres. The bottom connects to an underground creek (
|
||
<figureCitation id="13072A3EBF04FF95FE90402E238AFE90" box="[376,490,331,357]" captionStart="FIGURE 40" captionStartId="59.[151,250,1973,1997]" captionTargetBox="[162,1426,345,1950]" captionTargetId="figure-104@59.[162,1426,345,1950]" captionTargetPageId="59" captionText="FIGURE 40. Weixiao Tiankeng, the type locality of Guizhaphaenops (s. str.) wanghaoi n. sp. A. environ of Weixiao Tiankeng, bird view; B. a descending caver; C. the main passage below the Tiankeng (photos courtesy of Ms. Rurui Ye)." figureDoi="http://doi.org/10.5281/zenodo.7642468" httpUri="https://zenodo.org/record/7642468/files/figure.png" pageId="60" pageNumber="61">Figure 40</figureCitation>
|
||
). Along the main passage, there are several chambers and branches with high degree of humidity (Rurui Ye, personal communication). The
|
||
<typeStatus id="54878819BF04FF95FCE4400A215CFE7C" box="[780,828,367,393]" pageId="60" pageNumber="61">type</typeStatus>
|
||
specimens were collected under stone in a chamber. Other cave animals also observed inside this Tiankeng were crickets, pseudoscorpions and frogs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |