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<document id="B4B5C51A0C4B4BA197425BD3338626CD" ID-DOI="10.3897/fr.27.133914" ID-ZooBank="A30F4FFF-547A-4323-80C0-771663DF6FB7" ID-publisher-id="133914" URI-arpha="66B94E5B-EC0A-579D-B7ED-F85ED4A51F96" URI-zoobank="http://zoobank.org/A30F4FFF-547A-4323-80C0-771663DF6FB7" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1730736172305" checkinUser="pensoft" docAuthor="Abbassi, Nasrollah, Salehi Tinooni, Mohammad, Ghorbani Dehnavi, Mahdi, Shakeri, Safoora &amp; Eshaghi, Ali" docDate="2024" docId="6424B77523EE56BD8EB6098F67E676D9" docLanguage="en" docName="Fossil Record 27(2): 265-287" docOrigin="Fossil Record 27 (2)" docSource="https://fr.pensoft.net/article/133914/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Lophiopus Ellenberger 1980" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="287" masterDocId="66B94E5BEC0A579DB7EDF85ED4A51F96" masterDocTitle="Oligocene vertebrate footprints from the Lower Red Formation, Central Iran" masterLastPageNumber="287" masterPageNumber="265" pageNumber="265" updateTime="1730752961050" updateUser="admin">
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<mods:title id="4703B28E8CD29F7056345186BF65220B">Oligocene vertebrate footprints from the Lower Red Formation, Central Iran</mods:title>
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<mods:name id="A06B678235BEC1A56FF47A6BBC6BCAB4" type="personal">
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<mods:namePart id="D7D83F200C62788BDD0D032FCC9CE4B7">Abbassi, Nasrollah</mods:namePart>
<mods:nameIdentifier id="9B18CE1D6E2ECBEC0ADF0B9E948F3D91" type="ORCID">0000-0002-9994-9122</mods:nameIdentifier>
<mods:affiliation id="8EB4DEA4ECFED47ADD6E4861DB3E6AC2">Department of Geology, Faculty of Sciences, University of Zanjan, Zanjan, 45371 - 38971, Iran</mods:affiliation>
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<mods:namePart id="A4147B296283505BB624C32AFC1EEBC2">Salehi Tinooni, Mohammad</mods:namePart>
<mods:affiliation id="A2B838AE613D26B17110235ED3A0515F">Geological Survey of Iran, South-East Territory (Kerman), No. 10, Bastani-ye Parizi Blvd. Kerman, 76157 - 36841, Iran</mods:affiliation>
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<mods:namePart id="2703B80B3EA205452A4627BACA0FDC30">Ghorbani Dehnavi, Mahdi</mods:namePart>
<mods:affiliation id="E7E5006A198FE9AD6DD6CAB89B703E06">Geological Survey of Iran, South-East Territory (Kerman), No. 10, Bastani-ye Parizi Blvd. Kerman, 76157 - 36841, Iran</mods:affiliation>
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<mods:namePart id="543CE4266B0E0F545753346CA048346B">Shakeri, Safoora</mods:namePart>
<mods:affiliation id="3187A28F6F46CB4E606AF5013D0E1D84">Department of Geology, Faculty of Sciences, University of Zanjan, Zanjan, 45371 - 38971, Iran &amp; Research Affairs Office, Faculty of Engineering, University of Zanjan, Zanjan 45371 - 38971, Iran</mods:affiliation>
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<mods:namePart id="6409A78F7F4D143E59F37092CC43F0B0">Eshaghi, Ali</mods:namePart>
<mods:affiliation id="21B5F20DD2584FB6D688C8FAD7E7D229">Research Affairs Office, Faculty of Engineering, University of Zanjan, Zanjan 45371 - 38971, Iran</mods:affiliation>
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<mods:title id="59717472833BBBD843D68A0FC8814A77">Fossil Record</mods:title>
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<mods:date id="C0656E982783C7985D54BA29DC719F16">2024</mods:date>
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<mods:number id="EAFD86CA7E1F57FF358F521781E2496C">2024-11-04</mods:number>
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<treatment id="6424B77523EE56BD8EB6098F67E676D9" ID-arpha="6424B775-23EE-56BD-8EB6-098F67E676D9" LSID="urn:lsid:plazi:treatment:6424B77523EE56BD8EB6098F67E676D9" httpUri="http://treatment.plazi.org/id/6424B77523EE56BD8EB6098F67E676D9">
<subSubSection id="0366815714E740F110D28C023DA67CE4" type="nomenclature">
<paragraph id="33112D5F2A7713133848B619F9961EC8">
<heading id="90BC99CE830C6B720C2C711B110CFB28" reason="label">Ichnospecies</heading>
<taxonomicName id="BAFAA2FA35C48083271D6C013E785E1C" authorityName="Ellenberger" authorityYear="1980" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
<emphasis id="F820D99E22F41F3376DFB939E0ED5703" italics="true">Lophiopus</emphasis>
</taxonomicName>
isp.
</paragraph>
</subSubSection>
<subSubSection id="75E44D5A3AA017D482140A13DA512D22" type="description">
<paragraph id="11C6624A78ED92FBA632C602921C2ACB">
<figureCitation id="2AF6FD37D499542D78E2C60B622712F0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A C. Lophiopus isp. preserved as convex hyporelief with its photogrammetry photo and outline sketch, sampled (IFMI - 684); D F. Moropopus elongatus preserved as convex hyporelief, including photogrammetric false-color depth map, sampled (IFMI - 685); Moropopus elongatus in lower bedding plane of large slab full of mud crack casts (G); H I. Close up view of behind footprints with outline sketch; J K. Front footprint with outline sketch." figureDoi="10.3897/fr.27.133914.figure7" httpUri="https://binary.pensoft.net/fig/1167588">Fig. 7 A C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2NAI" type="material">
<paragraph id="B3503255C95C7C1BBF7E9E532C5730AA">
<heading id="A8FBDD9E0E58D6BF69E7573ADF5AE27B" reason="title">Material.</heading>
</paragraph>
<paragraph id="D9B731A46F7D4685BD0CE0E4E61BBFCD">
<materialsCitation id="824C9FEA197D4409E45B9F04E0AB482D" collectionCode="IFMI" specimenCode="IFMI-684" specimenCount="1">
One specimen, collected (
<specimenCode id="79C2BE04D789FECFBD02886FB979818F">
<abbrev content-type="institution" id="ABBRID0EBOAI" xlink_title="Ichnofossil Museum of Iran">IFMI</abbrev>
-684
</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGOAI" type="description">
<paragraph id="9991A29159135B06690196E45FDEA9D2">
<heading id="94C8D9FABD15A65DDD1E2601406EB4FF" reason="title">Description.</heading>
</paragraph>
<paragraph id="D05CF2C62DE477818B7916BC18876F9D">
One tridactyl footprint preserved as convex hyporelief in the lower bedding plane of the fine-grained, dark-brown sandstone. This bedding plane comprises flute and groove casts. The surface of the footprint is not smooth, and erosional structures encompass the footprint. One of the lateral digits shows three digital pad imprints, and digit
<collectionCode id="685D5F516C41CD0C5B7DC952C6B47454">III</collectionCode>
and the other lateral digit imprint are thick and lacking digital pad. The tips of the digits are sharp, the proximal rim of the footprint is curved and the sole imprint is not developed.
</paragraph>
<caption id="A0811F94B70F2338DBE5787703EEA733" ID-DOI="10.3897/fr.27.133914.figure7" ID-arpha="6CACCE3E-6E8E-594C-BCE8-BDC8F68DB9F2" httpUri="https://binary.pensoft.net/fig/1167588" startId="F7">
<paragraph id="3BA885693138D84B50AF536B74FE3450">
<label id="0501292D231C870001876E9DD3E76264">Figure 7.</label>
</paragraph>
<paragraph id="24D613E51BE8C42402B7965C3E5573BD">
<emphasis id="34D2D0A47507AB1CB46264CB3094EFD2" bold="true">A C.</emphasis>
<taxonomicName id="E1318658105053EA46A45D0B33C0E0EE" authorityName="Ellenberger" authorityYear="1980" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
<emphasis id="8AC8C26D2D537C0ACA919056F49D85DC" italics="true">Lophiopus</emphasis>
</taxonomicName>
isp. preserved as convex hyporelief with its photogrammetry photo and outline sketch, sampled (
<abbrev content-type="institution" id="ABBRID0E3OAI" xlink_title="Ichnofossil Museum of Iran">IFMI</abbrev>
- 684);
<emphasis id="F47D21C77885E13A9CE07FE9769F0202" bold="true">D F.</emphasis>
<taxonomicName id="70E1A7470B30BDCEB5825063554BA19D" authorityName="Abbassi, Alinasiri &amp; Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="species" species="elongatus">
<emphasis id="668380AE3E0ECEE43DF1340C061B8D1E" italics="true">Moropopus elongatus</emphasis>
</taxonomicName>
preserved as convex hyporelief, including photogrammetric false-color depth map, sampled (
<abbrev content-type="institution" id="ABBRID0EOPAI" xlink_title="Ichnofossil Museum of Iran">IFMI</abbrev>
- 685);
<taxonomicName id="72947E9AFAA3D4E1C0EE8FADD19C3219" authorityName="Abbassi, Alinasiri &amp; Lucas" authorityYear="2017" genus="Moropopus" higherTaxonomySource="GBIF" kingdom="Animalia" rank="species" species="elongatus">
<emphasis id="3635F850B7194F7920BC551E675A4998" italics="true">Moropopus elongatus</emphasis>
</taxonomicName>
in lower bedding plane of large slab full of mud crack casts (
<emphasis id="49E2B9130B61BFFCD95480203F43B1EA" bold="true">G</emphasis>
);
<emphasis id="767DB43CBCE05B51B6569CAB0E0F850C" bold="true">H I.</emphasis>
Close up view of behind footprints with outline sketch;
<emphasis id="DF89B97B3175F0171502222622A0A82C" bold="true">J K.</emphasis>
Front footprint with outline sketch.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EJQAI" type="discussion">
<paragraph id="CCA7AA7596B6F793F68587A6C17E15F9">
<heading id="EF996B49AA543EBB98F34DA7F1B23E22" reason="title">Discussion.</heading>
</paragraph>
<paragraph id="24877EA0F83644DCC7CCD6E028521EB1">
<bibRefCitation id="86D36432267BC91FD6A9656338DEBB7E" author="Ellenberger P" journalOrPublisher="Lavocat" pagination="37-78" refId="B31" refString="Ellenberger P (1980) Sur les empreintes de pas de gros mammiferes de l'Eocene superieur de Garrigues-Ste-Eulalie (Gard). Paleovertebrata, Memoire Jubile R. Lavocat 9: 37 - 78." title="Sur les empreintes de pas de gros mammiféres de lEocène supérieur de Garrigues-Ste-Eulalie (Gard). Paleovertebrata, Mémoire Jubilé R." volume="9" year="1980">Ellenberger, 1980</bibRefCitation>
considered two ichnospecies for
<taxonomicName id="68DFDFAA0F09118244CAC1A093C10ACB" authorityName="Ellenberger" authorityYear="1980" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
<emphasis id="A6A5C66BC87E2A340B1F518D4F548D3E" italics="true">Lophiopus</emphasis>
</taxonomicName>
,
<taxonomicName id="1DAF99E958548D5780E56B1D89009D4A" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="species" species="lalus">
<emphasis id="312C04384DC918D0186C653BB6FCEA31" italics="true">L. lalus</emphasis>
</taxonomicName>
and
<taxonomicName id="B4795D3B157023CDA80143623263C29C" authorityName="Ellenberger" authorityYear="1980" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="species" species="rapidus">
<emphasis id="B51AC92537E2CEA458063CA39606E83A" italics="true">L. rapidus</emphasis>
</taxonomicName>
.
<taxonomicName id="92EC57DEBF340E1EB3F364012FB21D9F" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="species" species="lalus">
<emphasis id="627FBD0347D16F24E8E363F28E5D0635" italics="true">L. lalus</emphasis>
</taxonomicName>
has a marked “ heel mark ” and an elongate digit
<collectionCode id="C7B4437C6315A5A2E7C8599EF1516240">III</collectionCode>
, whereas
<taxonomicName id="787508A90ABA54D911B72BCF070FED5C" authorityName="Ellenberger" authorityYear="1980" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="species" species="rapidus">
<emphasis id="ED3BE8FF3ACB42CA7C92B53111633A13" italics="true">L. rapidus</emphasis>
</taxonomicName>
is wider, and its lateral or central digits are more rounded along the tips. No digital pads were mentioned for ichnospecies of
<taxonomicName id="F813D77D15EE6DB5CDC97474EA2F8F34" authorityName="Ellenberger" authorityYear="1980" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
<emphasis id="DC033BEFCB7B17B4012D2A6F24DE720F" italics="true">Lophiopus</emphasis>
</taxonomicName>
. The studied footprints of the Deh Nar area show similar morphology to
<taxonomicName id="B0C5519E95F5A4E4BDCA77B3CC80B3AA" authorityName="Ellenberger" authorityYear="1980" class="Mammalia" family="Equidae" genus="Lophiopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" phylum="Chordata" rank="genus">
<emphasis id="53EB942FF9591D1EE0BAC1BB894E9C64" italics="true">Lophiopus</emphasis>
</taxonomicName>
in their round proximal outline, undeveloped sole imprint and distinct lateral digit imprints.
</paragraph>
</subSubSection>
</treatment>
</document>