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<document id="983157935C8161AE28DEA028B1CCE252" ID-CLB-Dataset="27576" ID-DOI="10.11646/zootaxa.4554.1.7" ID-GBIF-Dataset="94cdd5b4-1428-46bc-ae1b-cea7e0976cfb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623528" ID-ZooBank="18AB16FA-399B-4A34-8811-B199C39F395D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554271133809" checkinUser="plazi" docAuthor="Li, Bing, Zhao, Zhe, Zhang, Chuntian &amp; Li, Shuqiang" docDate="2019" docId="03FEEF55FFF6FFCAFF391280C58720C6" docLanguage="en" docName="zootaxa.4554.1.7.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Troglocoelotes yosiianus Li &amp; Zhao &amp; Zhang &amp; Li 2019, comb. n." docType="treatment" docVersion="8" lastPageNumber="232" masterDocId="FFC7972DFFFBFFC7FFAE1779C713271F" masterDocTitle="Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China" masterLastPageNumber="238" masterPageNumber="219" pageNumber="232" updateTime="1698716248201" updateUser="ExternalLinkService">
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<mods:title id="3B18C4D5A56969FE74600A638CCBFC38">Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China</mods:title>
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<mods:namePart id="5DFE8519A4C8F7B9DD7ED2056F49FE80">Li, Bing</mods:namePart>
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<mods:namePart id="BF501983899E3AF1365C5C7CFFF541D1">Zhao, Zhe</mods:namePart>
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<mods:namePart id="42B0EB67627B4F817E8475FCCF905530">Zhang, Chuntian</mods:namePart>
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<mods:namePart id="5496B0602C33F174AA7120679B2C2142">Li, Shuqiang</mods:namePart>
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<treatment id="03FEEF55FFF6FFCAFF391280C58720C6" ID-DOI="http://doi.org/10.5281/zenodo.5943986" ID-GBIF-Taxon="154534946" ID-Zenodo-Dep="5943986" LSID="urn:lsid:plazi:treatment:03FEEF55FFF6FFCAFF391280C58720C6" httpUri="http://treatment.plazi.org/id/03FEEF55FFF6FFCAFF391280C58720C6" lastPageNumber="232" pageId="13" pageNumber="232">
<subSubSection id="C34D0DC8FFF6FFCAFF391280C408210C" box="[151,795,1529,1555]" pageId="13" pageNumber="232" type="nomenclature">
<paragraph id="8BE85E43FFF6FFCAFF391280C408210C" blockId="13.[151,795,1529,1555]" box="[151,795,1529,1555]" pageId="13" pageNumber="232">
<heading id="D0A0E92FFFF6FFCAFF391280C408210C" bold="true" box="[151,795,1529,1555]" fontSize="11" level="1" pageId="13" pageNumber="232" reason="1">
<emphasis id="B9238251FFF6FFCAFF391280C408210C" bold="true" box="[151,795,1529,1555]" pageId="13" pageNumber="232">
<taxonomicName id="4C5725C0FFF6FFCAFF391280C5BB210C" ID-CoL="598R8" authority="(Nishikawa, 1999)" authorityName="Li &amp; Zhao &amp; Zhang &amp; Li" authorityYear="2019" baseAuthorityName="Nishikawa" baseAuthorityYear="1999" box="[151,680,1529,1555]" class="Arachnida" family="Agelenidae" genus="Troglocoelotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="232" phylum="Arthropoda" rank="species" species="yosiianus" status="comb. nov.">
<emphasis id="B9238251FFF6FFCAFF391280C6D3210C" bold="true" box="[151,448,1529,1555]" italics="true" pageId="13" pageNumber="232">Troglocoelotes yosiianus</emphasis>
(
<bibRefCitation id="EFC623B2FFF6FFCAFE7E1280C5B3210C" author="Nishikawa, Y." box="[464,672,1529,1555]" pageId="13" pageNumber="232" pagination="23 - 26" refId="ref7015" refString="Nishikawa, Y. (1999) A new eyeless agelenid spider from a limestone cave in Guangxi, south China. Journal of the Speleological Society of Japan, 24, 23 - 26." type="journal article" year="1999">Nishikawa, 1999</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2103F2AFFF6FFCAFD1E1280C408210C" box="[688,795,1529,1555]" pageId="13" pageNumber="232" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D0DC8FFF6FFCAFF391146C201216B" pageId="13" pageNumber="232" type="reference_group">
<paragraph id="8BE85E43FFF6FFCAFF391146C5A5214A" blockId="13.[151,1298,1599,1653]" box="[151,694,1599,1622]" pageId="13" pageNumber="232">
<treatmentCitationGroup id="AB47796DFFF6FFCAFF391146C5A5214A" box="[151,694,1599,1622]" pageId="13" pageNumber="232">
<taxonomicName id="4C5725C0FFF6FFCAFF391146C53A214A" authority="Nishikawa 1999: 23" authorityName="Nishikawa" authorityPageNumber="23" authorityYear="1999" box="[151,553,1599,1622]" class="Arachnida" family="Agelenidae" genus="Coelotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="232" phylum="Arthropoda" rank="species" species="yosiianus">
<emphasis id="B9238251FFF6FFCAFF391146C6452149" box="[151,342,1599,1622]" italics="true" pageId="13" pageNumber="232">Coelotes yosiianus</emphasis>
<treatmentCitation id="0AF67852FFF6FFCAFEF51146C53A214A" author="Nishikawa, Y." box="[347,553,1599,1621]" page="23" pageId="13" pageNumber="232" year="1999">
<bibRefCitation id="EFC623B2FFF6FFCAFEF51146C516214A" author="Nishikawa, Y." box="[347,517,1599,1621]" pageId="13" pageNumber="232" pagination="23 - 26" refId="ref7015" refString="Nishikawa, Y. (1999) A new eyeless agelenid spider from a limestone cave in Guangxi, south China. Journal of the Speleological Society of Japan, 24, 23 - 26." type="journal article" year="1999">Nishikawa 1999</bibRefCitation>
: 23
</treatmentCitation>
</taxonomicName>
, figs 15 (
<specimenCount id="9D5195CAFFF6FFCAFD391146C5BB214A" box="[663,680,1599,1621]" pageId="13" pageNumber="232" type="female"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE85E43FFF6FFCAFF391126C201216B" blockId="13.[151,1298,1599,1653]" box="[151,1298,1630,1653]" pageId="13" pageNumber="232">
<treatmentCitationGroup id="AB47796DFFF6FFCAFF391126C201216B" box="[151,1298,1630,1653]" pageId="13" pageNumber="232">
<taxonomicName id="4C5725C0FFF6FFCAFF391126C515216B" ID-CoL="92ZRS" authority="(Nishikawa)" authorityName="Li &amp; Zhao &amp; Zhang &amp; Li" authorityYear="2019" baseAuthorityName="Li &amp; Zhao &amp; Zhang &amp; Li" baseAuthorityYear="1999" box="[151,518,1630,1652]" class="Arachnida" family="Agelenidae" genus="Draconarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="232" phylum="Arthropoda" rank="species" species="yosiianus">
<emphasis id="B9238251FFF6FFCAFF391126C66D216B" box="[151,382,1631,1652]" italics="true" pageId="13" pageNumber="232">Draconarius yosiianus</emphasis>
(Nishikawa)
</taxonomicName>
:
<treatmentCitation id="0AF67852FFF6FFCAFDA11126C5BA216B" author="Wang, X. P." box="[527,681,1630,1652]" page="69" pageId="13" pageNumber="232" year="2002">
<bibRefCitation id="EFC623B2FFF6FFCAFDA11126C594216B" author="Wang, X. P." box="[527,647,1631,1652]" pageId="13" pageNumber="232" pagination="1 - 150" refId="ref7176" refString="Wang, X. P. (2002) A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History, 269, 1 - 150. https: // doi. org / 10.1206 / 0003 - 0090 (2002) 269 &lt;0001: AGLROT&gt; 2.0. CO; 2" type="journal article" year="2002">Wang 2002</bibRefCitation>
: 69
</treatmentCitation>
(
<specimenCount id="9D5195CAFFF6FFCAFD171126C5D9216A" box="[697,714,1631,1653]" pageId="13" pageNumber="232" type="female"></specimenCount>
);
<treatmentCitation id="0AF67852FFF6FFCAFD4E1126C494216B" author="Wang, X. P." box="[736,903,1630,1652]" page="556" pageId="13" pageNumber="232" year="2003">
<bibRefCitation id="EFC623B2FFF6FFCAFD4E1126C444216B" author="Wang, X. P." box="[736,855,1631,1652]" pageId="13" pageNumber="232" pagination="499 - 662" refId="ref7240" refString="Wang, X. P. (2003) Species revision of the coelotine spider genera Bifidocoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae). Proceedings of the California Academy of Sciences, 54, 499 - 662." type="journal article" year="2003">Wang 2003</bibRefCitation>
: 556
</treatmentCitation>
(
<specimenCount id="9D5195CAFFF6FFCAFC391126C4BB216A" box="[919,936,1631,1653]" pageId="13" pageNumber="232" type="female"></specimenCount>
);
<treatmentCitation id="0AF67852FFF6FFCAFC111127C39C216B" author="Zhu, M. S. &amp; Wang, X. P. &amp; Zhang, Z. S." box="[959,1167,1630,1652]" page="415" pageId="13" pageNumber="232" year="2017">
<bibRefCitation id="EFC623B2FFF6FFCAFC111127C34E216B" author="Zhu, M. S. &amp; Wang, X. P. &amp; Zhang, Z. S." box="[959,1117,1630,1652]" pageId="13" pageNumber="232" refId="ref7659" refString="Zhu, M. S., Wang, X. P. &amp; Zhang, Z. S. (2017) Fauna Sinica: Invertebrata Uol. 59: Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp." type="book" year="2017">
Zhu
<emphasis id="B9238251FFF6FFCAFC401126C332216B" box="[1006,1057,1630,1652]" italics="true" pageId="13" pageNumber="232">et al.</emphasis>
2017
</bibRefCitation>
: 415
</treatmentCitation>
, fig.277 (
<specimenCount id="9D5195CAFFF6FFCAFB5C1127C210216B" box="[1266,1283,1630,1652]" pageId="13" pageNumber="232" type="female"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D0DC8FFF6FFCAFF3911D9C3092039" pageId="13" pageNumber="232" type="diagnosis">
<paragraph id="8BE85E43FFF6FFCAFF3911D9C3092039" blockId="13.[151,1436,1696,2009]" pageId="13" pageNumber="232">
<emphasis id="B9238251FFF6FFCAFF3911D9C60221A6" bold="true" box="[151,273,1696,1721]" pageId="13" pageNumber="232">Diagnosis.</emphasis>
Females are similar to
<taxonomicName id="4C5725C0FFF6FFCAFDB611DBC5D521A6" authority="Li &amp; Zhao &amp; Zhang &amp; Li, 2019" authorityName="Li &amp; Zhao &amp; Zhang &amp; Li" authorityYear="2019" box="[536,710,1696,1721]" class="Arachnida" family="Agelenidae" genus="Troglocoelotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="232" phylum="Arthropoda" rank="species" species="nongchiensis" status="sp. nov.">
<emphasis id="B9238251FFF6FFCAFDB611DBC53521A6" box="[536,550,1698,1721]" italics="true" pageId="13" pageNumber="232">T</emphasis>
.
<emphasis id="B9238251FFF6FFCAFD9A11D9C5D521A6" box="[564,710,1696,1721]" italics="true" pageId="13" pageNumber="232">nongchiensis</emphasis>
</taxonomicName>
<emphasis id="B9238251FFF6FFCAFD6311D9C41E21A6" bold="true" box="[717,781,1696,1721]" pageId="13" pageNumber="232">
<taxonomicNameLabel id="A2103F2AFFF6FFCAFD6311D9C41E21A6" box="[717,781,1696,1721]" pageId="13" pageNumber="232" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the epigynal teeth pointed medially and with a tip bent upwards, but can be distinguished from by the copulatory ducts having more coils. Females are also similar to
<taxonomicName id="4C5725C0FFF6FFCAFA2911BFC7CA201E" authorityName="Li &amp; Zhao &amp; Zhang &amp; Li" authorityYear="2019" baseAuthorityName="Chen, Zhu &amp; Kim" baseAuthorityYear="2008" class="Arachnida" family="Agelenidae" genus="Troglocoelotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="232" phylum="Arthropoda" rank="species" species="tortus">
<emphasis id="B9238251FFF6FFCAFA2911BFC28721C2" box="[1415,1428,1734,1757]" italics="true" pageId="13" pageNumber="232">T</emphasis>
.
<emphasis id="B9238251FFF6FFCAFF391193C7CA201E" box="[151,217,1770,1793]" italics="true" pageId="13" pageNumber="232">tortus</emphasis>
</taxonomicName>
by the rectangular posterior margin of epigynal and long copulatory ducts, but can be distinguished by the epigynal teeth pointed medially and the middle parts of copulatory ducts widest.
</paragraph>
</subSubSection>
<subSubSection id="C34D0DC8FFF6FFCAFF691049C55E2056" box="[199,589,1840,1865]" pageId="13" pageNumber="232" type="description">
<paragraph id="8BE85E43FFF6FFCAFF691049C55E2056" blockId="13.[151,1436,1696,2009]" box="[199,589,1840,1865]" pageId="13" pageNumber="232">
<emphasis id="B9238251FFF6FFCAFF691049C64A2056" bold="true" box="[199,345,1840,1865]" pageId="13" pageNumber="232">Description.</emphasis>
See
<bibRefCitation id="EFC623B2FFF6FFCAFE201049C55B2056" author="Nishikawa, Y." box="[398,584,1840,1865]" pageId="13" pageNumber="232" pagination="23 - 26" refId="ref7015" refString="Nishikawa, Y. (1999) A new eyeless agelenid spider from a limestone cave in Guangxi, south China. Journal of the Speleological Society of Japan, 24, 23 - 26." type="journal article" year="1999">Nishikawa 1999</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D0DC8FFF6FFCAFF69102CC6EF20AA" pageId="13" pageNumber="232" type="materials_examined">
<paragraph id="8BE85E43FFF6FFCAFF69102CC28F208E" blockId="13.[151,1436,1696,2009]" pageId="13" pageNumber="232">
<emphasis id="B9238251FFF6FFCAFF69102CC6FD2071" bold="true" box="[199,494,1877,1902]" pageId="13" pageNumber="232">
Type material.
<materialsCitation id="3B3F541EFFF6FFCAFE2F102CC6FD2071" ID-GBIF-Occurrence="2234227911" box="[385,494,1877,1902]" collectingDate="1998-04-21" collectionCode="NSMT" collectorName="Y. Nishikawa" country="China" county="Bama County" elevation="280" location="Jiabao Cave" municipality="Xingren Village" pageId="13" pageNumber="232" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="54ECE0E1FFF6FFCAFE2F102CC6FD2071" box="[385,494,1877,1902]" pageId="13" pageNumber="232" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
<specimenCount id="9D5195CAFFF6FFCAFE59102FC51F2072" box="[503,524,1878,1901]" pageId="13" pageNumber="232" type="female"></specimenCount>
(NSMT, not examined):
<location id="8E880898FFF6FFCAFC9F102CC4AF2072" LSID="urn:lsid:plazi:treatment:03FEEF55FFF6FFCAFF391280C58720C6:8E880898FFF6FFCAFC9F102CC4AF2072" box="[817,956,1877,1902]" country="China" county="Bama County" municipality="Xingren Village" name="Jiabao Cave" pageId="13" pageNumber="232" stateProvince="Guangxi">Jiabao Cave</location>
(
<quantity id="4CAFF3A6FFF6FFCAFC63102CC30F2072" box="[973,1052,1877,1901]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="232" unit="m" value="280.0">
<elevation id="007AB970FFF6FFCAFC63102CC30F2072" box="[973,1052,1877,1901]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="232" unit="m" value="280.0">280 m</elevation>
</quantity>
),
<collectingMunicipality id="6B8CC439FFF6FFCAFB9C102CC3FF2071" box="[1074,1260,1877,1902]" pageId="13" pageNumber="232">Xingren Village</collectingMunicipality>
,
<collectingCounty id="628926CFFFF6FFCAFB57102FC28B2072" box="[1273,1432,1877,1901]" pageId="13" pageNumber="232">Bama County</collectingCounty>
,
<collectingRegion id="499390A1FFF6FFCAFF391001C7E9208E" box="[151,250,1912,1937]" country="China" name="Guangxi" pageId="13" pageNumber="232">Guangxi</collectingRegion>
,
<collectingCountry id="F3401ED3FFF6FFCAFEAB1001C658208E" box="[261,331,1912,1937]" name="China" pageId="13" pageNumber="232">China</collectingCountry>
,
<date id="FFE97883FFF6FFCAFEF81000C6F2208E" box="[342,481,1913,1937]" pageId="13" pageNumber="232" value="1998-04-21">
<collectingDate id="EFAD816BFFF6FFCAFEF81000C6F2208E" box="[342,481,1913,1937]" pageId="13" pageNumber="232" value="1998-04-21">21. IV. 1998</collectingDate>
</date>
,
<collectorName id="26A23B95FFF6FFCAFE421000C591208E" box="[492,642,1912,1937]" pageId="13" pageNumber="232">Y. Nishikawa</collectorName>
leg.
<materialsCitation id="3B3F541EFFF6FFCAFD161001C28F208E" ID-GBIF-Occurrence="2234227968" box="[696,1436,1912,1937]" collectingDate="1997-04-06" collectionCode="NSMT" collectorName="Y. Nishikawa" country="China" pageId="13" pageNumber="232" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="B9238251FFF6FFCAFD161001C437208E" bold="true" box="[696,804,1912,1937]" pageId="13" pageNumber="232">
<typeStatus id="54ECE0E1FFF6FFCAFD161001C437208E" box="[696,804,1912,1937]" pageId="13" pageNumber="232" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="9D5195CAFFF6FFCAFC9A1000C444208E" box="[820,855,1913,1937]" pageId="13" pageNumber="232" type="female">1♀</specimenCount>
(
<collectionCode id="ED46C686FFF6FFCAFCCB1000C4A9208E" box="[869,954,1913,1937]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="13" pageNumber="232">NSMT</collectionCode>
): same cave,
<date id="FFE97883FFF6FFCAFBE31001C3D8208E" box="[1101,1227,1912,1937]" pageId="13" pageNumber="232" value="1997-04-06">
<collectingDate id="EFAD816BFFF6FFCAFBE31001C3D8208E" box="[1101,1227,1912,1937]" pageId="13" pageNumber="232" value="1997-04-06">6. IV. 1997</collectingDate>
</date>
,
<collectorName id="26A23B95FFF6FFCAFB781000C27F208E" box="[1238,1388,1912,1937]" pageId="13" pageNumber="232">Y. Nishikawa</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BE85E43FFF6FFCAFF6910E4C6EF20AA" blockId="13.[151,1436,1696,2009]" box="[199,508,1949,1974]" pageId="13" pageNumber="232">
<emphasis id="B9238251FFF6FFCAFF6910E4C6A320A9" bold="true" box="[199,432,1949,1974]" pageId="13" pageNumber="232">Material examined.</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="C34D0DC8FFF6FFCAFF6910B9C58720C6" box="[199,660,1984,2009]" pageId="13" pageNumber="232" type="distribution">
<paragraph id="8BE85E43FFF6FFCAFF6910B9C58720C6" blockId="13.[151,1436,1696,2009]" box="[199,660,1984,2009]" pageId="13" pageNumber="232">
<emphasis id="B9238251FFF6FFCAFF6910B9C67320C6" bold="true" box="[199,352,1984,2009]" pageId="13" pageNumber="232">Distribution.</emphasis>
<collectingCountry id="F3401ED3FFF6FFCAFEC910B9C6B920C6" box="[359,426,1984,2009]" name="China" pageId="13" pageNumber="232">China</collectingCountry>
(
<collectingRegion id="499390A1FFF6FFCAFE1710B9C50F20C6" box="[441,540,1984,2009]" country="China" name="Guangxi" pageId="13" pageNumber="232">Guangxi</collectingRegion>
) (
<figureCitation id="136C42C6FFF6FFCAFD9D10B9C59420C6" box="[563,647,1984,2009]" captionStart="FIGURE 14" captionStartId="17.[151,250,1139,1161]" captionTargetBox="[151,1436,195,1117]" captionTargetId="figure@17.[151,1436,195,1117]" captionTargetPageId="17" captionText="FIGURE 14. Distribution records of Troglocoelotes species from South China. 1. T. bailongensis Z. Zhao &amp; S. Li sp. n.; 2. T. banmenensis Z. Zhao &amp; S. Li sp. n.; 3. T. liangensis Z. Zhao &amp; S. Li sp. n.; 4. T. nongchiensis Z. Zhao &amp; S. Li sp. n.; 5. T. proximus (Chen, Zhu &amp; Kim, 2008) comb. n.; 6. T. qixianensis Z. Zhao &amp; S. Li sp. n.; 7. T. tortus (Chen, Zhu &amp; Kim, 2008) comb. n.; 8. T. yosiianus (Nishikawa, 1999) comb. n.; 9. T. yumiganensis Z. Zhao &amp; S. Li sp. n." httpUri="https://zenodo.org/record/2623558/files/figure.png" pageId="13" pageNumber="232">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>