347 lines
43 KiB
XML
347 lines
43 KiB
XML
<document id="5EED6989E3D79739E0B25BE168B177ED" ID-CLB-Dataset="29008" ID-DOI="10.11646/zootaxa.4459.3.9" ID-GBIF-Dataset="ad9c1f20-5e3c-400f-9ca8-7a8c46347063" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458925" ID-ZooBank="79B91BA4-C861-4EA1-85D9-7AE039D950B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539290441872" checkinUser="plazi" docAuthor="Gill, Anthony C., Russell, Barry C. & Nelson, Gary" docDate="2018" docId="03FE1B36474CD44DFF25FB3BFF61B5DA" docLanguage="en" docName="zootaxa.4459.3.9.pdf" docOrigin="Zootaxa 4459 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Engraulis nasutus Castelnau 1878" docType="treatment" docVersion="7" lastPageNumber="572" masterDocId="FFC7634E474AD44AFFB2FF94FFBAB67F" masterDocTitle="F. L. de Castelnau’s Norman River fishes housed in the Macleay Museum, University of Sydney" masterLastPageNumber="574" masterPageNumber="565" pageNumber="571" updateTime="1698578944772" updateUser="ExternalLinkService">
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<mods:title id="97CA345585B694B45B0AAE335F7A8777">F. L. de Castelnau’s Norman River fishes housed in the Macleay Museum, University of Sydney</mods:title>
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<mods:namePart id="3D2BE32F698ED67DC4DF927E6E53A22C">Gill, Anthony C.</mods:namePart>
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<mods:namePart id="0F98FE1C30185F051E5EE794DF7C5C09">Russell, Barry C.</mods:namePart>
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<mods:namePart id="1D42B84CAC748DFF4BDE8719C962DF48">Nelson, Gary</mods:namePart>
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<mods:title id="86AD6976E64A4CB2AAF10326B4592E35">Zootaxa</mods:title>
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<mods:date id="CC321194A7B77F41334F619C7DC836D6">2018</mods:date>
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<mods:number id="AD98C3717BC5616B05EA540FE22FFBC6">2018-08-16</mods:number>
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<treatment id="03FE1B36474CD44DFF25FB3BFF61B5DA" ID-DOI="http://doi.org/10.5281/zenodo.5961905" ID-GBIF-Taxon="148692885" ID-Zenodo-Dep="5961905" LSID="urn:lsid:plazi:treatment:03FE1B36474CD44DFF25FB3BFF61B5DA" httpUri="http://treatment.plazi.org/id/03FE1B36474CD44DFF25FB3BFF61B5DA" lastPageId="7" lastPageNumber="572" pageId="6" pageNumber="571">
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<subSubSection id="C34DF9AB474CD44CFF25FB3BFEA6B294" pageId="6" pageNumber="571" type="nomenclature">
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<paragraph id="8BE8AA20474CD44CFF25FB3BFC9DB2B6" blockId="6.[151,807,1199,1259]" box="[151,807,1199,1225]" pageId="6" pageNumber="571">
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<heading id="D0A01D4C474CD44CFF25FB3BFC9DB2B6" bold="true" box="[151,807,1199,1225]" fontSize="11" level="1" pageId="6" pageNumber="571" reason="1">
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<emphasis id="B9237632474CD44CFF25FB3BFC9DB2B6" bold="true" box="[151,807,1199,1225]" pageId="6" pageNumber="571">
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<taxonomicName id="4C57D1A3474CD44CFF25FB3BFD86B2B6" ID-CoL="6F8X7" authority="Castelnau 1878" authorityName="Castelnau" authorityYear="1878" box="[151,572,1199,1225]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
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<emphasis id="B9237632474CD44CFF25FB3BFECFB2B6" bold="true" box="[151,373,1199,1225]" italics="true" pageId="6" pageNumber="571">Engraulis nasutus</emphasis>
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<bibRefCitation id="EFC6D7D1474CD44CFECEFB3BFD86B2B6" box="[380,572,1199,1225]" pageId="6" pageNumber="571" refId="ref4823" refString="Castelnau, F. L. de (1878 a) Australian fishes. New or little known species. Proceedings of the Linnean Society of New South Wales, 2, 225 - 248, pls. 2 - 3." type="journal article">Castelnau 1878</bibRefCitation>
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</taxonomicName>
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[=
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<taxonomicName id="4C57D1A3474CD44CFDD7FB3BFCA7B2B6" box="[613,797,1199,1225]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasuta">
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<emphasis id="B9237632474CD44CFDD7FB3BFCA7B2B6" bold="true" box="[613,797,1199,1225]" italics="true" pageId="6" pageNumber="571">Thryssa nasuta</emphasis>
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</taxonomicName>
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]
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE8AA20474CD44CFF25FB46FEA6B294" blockId="6.[151,807,1199,1259]" box="[151,284,1234,1259]" pageId="6" pageNumber="571">
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<figureCitation id="136CB6A5474CD44CFF25FB46FEA6B294" box="[151,284,1234,1259]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="7.[151,250,1844,1866]" captionStartId-1="8.[151,250,1151,1173]" captionTargetBox-0="[151,1436,988,1823]" captionTargetBox-1="[151,1436,195,1127]" captionTargetId-0="figure@7.[151,1436,988,1823]" captionTargetId-1="figure@8.[151,1436,195,1127]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 6. Engraulis nasutus Castlenau, syntypes, MAMU F.1194, 99.5 (upper) and 104.5 (lower) mm SL. Photo by Y.K. Tea." captionText-1="FIGURE 7. Engraulis nasutus Castlenau, syntypes, MAMU F.1194, 99.5 (upper) and 104.5 (lower) mm SL. Radiograph by J. King." httpUri-0="https://zenodo.org/record/1458937/files/figure.png" httpUri-1="https://zenodo.org/record/1458939/files/figure.png" pageId="6" pageNumber="571">Figures 6–7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34DF9AB474CD44CFF25FA8EFEC5B39E" pageId="6" pageNumber="571" type="reference_group">
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<paragraph id="8BE8AA20474CD44CFF25FA8EFCE4B34F" blockId="6.[151,862,1306,1328]" box="[151,862,1306,1328]" pageId="6" pageNumber="571">
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<materialsCitation id="3B3FA07D474CD44CFF25FA8EFCE4B34F" ID-GBIF-Occurrence="1917413287" box="[151,862,1306,1328]" location="Norman River" pageId="6" pageNumber="571" specimenCount="1" typeStatus="holotype">
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<treatmentCitationGroup id="AB478D0E474CD44CFF25FA8EFCE4B34F" box="[151,862,1306,1328]" pageId="6" pageNumber="571">
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<taxonomicName id="4C57D1A3474CD44CFF25FA8EFD9FB34F" ID-CoL="6F8X7" authority="Castelnau 1878 b: 51" authorityName="Castelnau" authorityPageNumber="51" authorityYear="1878" box="[151,549,1306,1328]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
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<emphasis id="B9237632474CD44CFF25FA8EFEF4B34F" box="[151,334,1306,1328]" italics="true" pageId="6" pageNumber="571">Engraulis nasutus</emphasis>
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<treatmentCitation id="0AF68C31474CD44CFEE6FA8EFD9FB34F" author="" box="[340,549,1306,1328]" page="51" pageId="6" pageNumber="571">
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<bibRefCitation id="EFC6D7D1474CD44CFEE6FA8EFDB8B34F" box="[340,514,1306,1328]" pageId="6" pageNumber="571" refId="ref4823" refString="Castelnau, F. L. de (1878 a) Australian fishes. New or little known species. Proceedings of the Linnean Society of New South Wales, 2, 225 - 248, pls. 2 - 3." type="journal article">Castelnau 1878b</bibRefCitation>
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: 51
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</treatmentCitation>
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||
</taxonomicName>
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||
(
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<typeStatus id="54EC1482474CD44CFD80FA8EFDE5B34F" box="[562,607,1306,1328]" pageId="6" pageNumber="571">type</typeStatus>
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locality,
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||
<location id="8E88FCFB474CD44CFD0EFA8FFCE9B34F" LSID="urn:lsid:plazi:treatment:03FE1B36474CD44DFF25FB3BFF61B5DA:8E88FCFB474CD44CFD0EFA8FFCE9B34F" box="[700,851,1306,1328]" name="Norman River" pageId="6" pageNumber="571">Norman River</location>
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).
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</treatmentCitationGroup>
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE8AA20474CD44CFF25FAC8FEC5B39E" blockId="6.[151,1437,1372,2010]" pageId="6" pageNumber="571">
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<bibRefCitation id="EFC6D7D1474CD44CFF25FAC8FED3B30A" box="[151,361,1372,1397]" pageId="6" pageNumber="571" refId="ref4823" refString="Castelnau, F. L. de (1878 a) Australian fishes. New or little known species. Proceedings of the Linnean Society of New South Wales, 2, 225 - 248, pls. 2 - 3." type="journal article">Castelnau (1878b)</bibRefCitation>
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described
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<taxonomicName id="4C57D1A3474CD44CFE5EFAC8FD00B30A" ID-CoL="6F8X7" authorityName="Castelnau" authorityYear="1878" box="[492,698,1372,1397]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
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<emphasis id="B9237632474CD44CFE5EFAC8FD00B30A" box="[492,698,1372,1397]" italics="true" pageId="6" pageNumber="571">Engraulis nasutus</emphasis>
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</taxonomicName>
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on the basis of an unspecified number of specimens from the Norman River. He noted that “I have seen only one adult specimen seven inches long, but I have a small specimen preserved in spirits, which is silvery with the upper parts of a light brown, fins yellow”. This suggests he had at least two specimens.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34DF9AB474CD44DFF75FA79FF61B5DA" lastPageId="7" lastPageNumber="572" pageId="6" pageNumber="571" type="materials_examined">
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||
<paragraph id="8BE8AA20474CD44CFF75FA79FCB3B1C9" blockId="6.[151,1437,1372,2010]" pageId="6" pageNumber="571">
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Identification of the species has been problematic. Macleay (1879) considered it to be a valid species of
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<taxonomicName id="4C57D1A3474CD44CFF25F984FEBFB056" box="[151,261,1552,1577]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="genus">
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<emphasis id="B9237632474CD44CFF25F984FEBFB056" box="[151,261,1552,1577]" italics="true" pageId="6" pageNumber="571">Engraulis</emphasis>
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</taxonomicName>
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, stating (p. 367): “This species is described by Count Castelnau […] from one adult specimen,
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<quantity id="4CAF07C5474CD44CFA8DF985FA26B056" box="[1343,1436,1552,1577]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="6" pageNumber="571" unit="in" value="7.0">7 inches</quantity>
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long, sent to him from the Norman River, Gulf of Carpentaria. Its special distinguishing character seems to be a strong longitudinal ridge along the top of the head.” Macleay’s mention of a single specimen is curious, given that there are two specimens in
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<collectionCode id="ED4632E5474CD44CFE68F9E9FD8EB0EA" box="[474,564,1661,1685]" country="Australia" httpUri="http://grbio.org/cool/6adq-425v" name="University of Sydney, Macleay Museum" pageId="6" pageNumber="571">MAMU</collectionCode>
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(see below).
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<bibRefCitation id="EFC6D7D1474CD44CFD64F9E9FCC3B0EA" box="[726,889,1661,1686]" pageId="6" pageNumber="571" refId="ref5553" refString="Ogilby, J. D. (1910) On new or insufficiently described fishes. Proceedings of the Royal Society of Queensland, 23, 1 - 55." type="journal article">Ogilby (1910)</bibRefCitation>
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considered
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<taxonomicName id="4C57D1A3474CD44CFBB5F9EAFB3AB0EA" authorityName="Castelnau" authorityYear="1878" box="[1031,1152,1662,1685]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
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<emphasis id="B9237632474CD44CFBB5F9EAFB3AB0EA" box="[1031,1152,1662,1685]" italics="true" pageId="6" pageNumber="571">E. nasutus</emphasis>
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</taxonomicName>
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to be a valid species of
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<taxonomicName id="4C57D1A3474CD44CFF25F934FEBBB0C6" authorityName="Jordan & Evermann" authorityYear="1895" box="[151,257,1696,1721]" class="Actinopterygii" family="Engraulidae" genus="Anchovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="genus">
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<emphasis id="B9237632474CD44CFF25F934FEBBB0C6" box="[151,257,1696,1721]" italics="true" pageId="6" pageNumber="571">Anchovia</emphasis>
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</taxonomicName>
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<collectingCountry id="F340EAB0474CD44CFEB5F934FEE8B0C6" box="[263,338,1696,1721]" name="Jordan" pageId="6" pageNumber="571">Jordan</collectingCountry>
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& Evermann in
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<bibRefCitation id="EFC6D7D1474CD44CFDBBF934FD1DB0C6" box="[521,679,1696,1721]" pageId="6" pageNumber="571" refId="ref5241" refString="Jordan, D. S. (1895) The fishes of Sinaloa. Proceedings of the California Academy of Sciences, Series 2, 5, 377 - 514, pls 26 - 55." type="journal article">
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<collectingCountry id="F340EAB0474CD44CFDBBF934FDEEB0C6" box="[521,596,1696,1721]" name="Jordan" pageId="6" pageNumber="571">Jordan</collectingCountry>
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(1895)
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</bibRefCitation>
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, and compared it with his new species
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<taxonomicName id="4C57D1A3474CD44CFBEAF935FB68B0C6" box="[1112,1234,1697,1721]" class="Actinopterygii" family="Engraulidae" genus="Anchovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="aesturia">
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<emphasis id="B9237632474CD44CFBEAF935FB68B0C6" box="[1112,1234,1697,1721]" italics="true" pageId="6" pageNumber="571">A. aesturia</emphasis>
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</taxonomicName>
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. It is unlikely that Ogilby saw
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<typeStatus id="54EC1482474CD44CFE9AF952FEE2B0A1" box="[296,344,1734,1758]" pageId="6" pageNumber="571">type</typeStatus>
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||
material of
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<taxonomicName id="4C57D1A3474CD44CFE42F952FDD3B0A2" authorityName="Castelnau" authorityYear="1878" box="[496,617,1734,1757]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
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<emphasis id="B9237632474CD44CFE42F952FDD3B0A2" box="[496,617,1734,1757]" italics="true" pageId="6" pageNumber="571">E. nasutus</emphasis>
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</taxonomicName>
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, as all of his comparative data for the species is identical to that in Castelnau’s original description.
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<bibRefCitation id="EFC6D7D1474CD44CFDB8F97CFD5CB17E" box="[522,742,1768,1793]" pageId="6" pageNumber="571" refId="ref5489" refString="McCulloch, A. R. (1929 a) A check-list of the fishes recorded from Australia. Part I. Australian Museum Memoirs, 5, 1 - 144." type="journal article">McCulloch (1929a)</bibRefCitation>
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||
regarded
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||
<taxonomicName id="4C57D1A3474CD44CFCEAF97EFC76B17E" authorityName="Castelnau" authorityYear="1878" box="[856,972,1770,1793]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
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||
<emphasis id="B9237632474CD44CFCEAF97EFC76B17E" box="[856,972,1770,1793]" italics="true" pageId="6" pageNumber="571">E. nasutus</emphasis>
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||
</taxonomicName>
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as a valid species of
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<taxonomicName id="4C57D1A3474CD44CFB0DF97CFF58B15A" authority="Fowler (1911)" authorityName="Fowler" authorityYear="1911" class="Actinopterygii" family="Engraulidae" genus="Anchoviella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="genus">
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<emphasis id="B9237632474CD44CFB0DF97CFAFFB17E" box="[1215,1349,1768,1793]" italics="true" pageId="6" pageNumber="571">Anchoviella</emphasis>
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||
<bibRefCitation id="EFC6D7D1474CD44CFAFEF97CFF58B15A" pageId="6" pageNumber="571" refId="ref5046" refString="Fowler, H. w. (1911) Notes on clupeoid fishes. Proceedings of the Academy of Natural Sciences of Philadelphia, 63, 204 - 221." type="journal article">Fowler (1911)</bibRefCitation>
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</taxonomicName>
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, but did not indicate whether he had examined the
|
||
<typeStatus id="54EC1482474CD44CFCAFF89AFC39B159" box="[797,899,1806,1830]" pageId="6" pageNumber="571" type="syntype">syntypes</typeStatus>
|
||
. Whitley (1964) considered both
|
||
<taxonomicName id="4C57D1A3474CD44CFB4AF89AFAD6B15A" authorityName="Castelnau" authorityYear="1878" box="[1272,1388,1806,1829]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9237632474CD44CFB4AF89AFAD6B15A" box="[1272,1388,1806,1829]" italics="true" pageId="6" pageNumber="571">E. nasutus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C57D1A3474CD44CFF25F8A5FE9BB136" authorityName="Ogibly" authorityYear="1910" box="[151,289,1841,1865]" class="Actinopterygii" family="Engraulidae" genus="Anchovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="aestuaria">
|
||
<emphasis id="B9237632474CD44CFF25F8A5FE9BB136" box="[151,289,1841,1865]" italics="true" pageId="6" pageNumber="571">A. aestuaria</emphasis>
|
||
</taxonomicName>
|
||
to be valid species of
|
||
<taxonomicName id="4C57D1A3474CD44CFD92F8A4FD30B136" authorityName="Jordan & Seale" authorityYear="1925" box="[544,650,1840,1865]" class="Actinopterygii" family="Engraulidae" genus="Thrissina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9237632474CD44CFD92F8A4FD30B136" box="[544,650,1840,1865]" italics="true" pageId="6" pageNumber="571">Thrissina</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFC6D7D1474CD44CFD20F8A4FC2FB136" box="[658,917,1840,1865]" pageId="6" pageNumber="571" refId="ref5277" refString="Jordan, D. S. & Seale, A. (1925) Analysis of the genera of anchovies or Engraulidae. Copeia, 141, 27 - 32. https: // doi. org / 10.2307 / 1437308" type="journal article">
|
||
<collectingCountry id="F340EAB0474CD44CFD20F8A4FD67B136" box="[658,733,1840,1865]" name="Jordan" pageId="6" pageNumber="571">Jordan</collectingCountry>
|
||
& Seale (1925)
|
||
</bibRefCitation>
|
||
. Whitehead
|
||
<emphasis id="B9237632474CD44CFB93F8A6FBE1B136" box="[1057,1115,1840,1865]" italics="true" pageId="6" pageNumber="571">et al.</emphasis>
|
||
(1988) tentatively placed
|
||
<taxonomicName id="4C57D1A3474CD44CFA36F8A6FF57B112" authorityName="Castelnau" authorityYear="1878" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9237632474CD44CFA36F8A6FF57B112" italics="true" pageId="6" pageNumber="571">E. nasutus</emphasis>
|
||
</taxonomicName>
|
||
in the synonymy of
|
||
<taxonomicName id="4C57D1A3474CD44CFE66F8C1FC88B111" authority="(Gray 1835)" authorityName="Gray" authorityYear="1835" baseAuthorityName="Gray" baseAuthorityYear="1835" box="[468,818,1877,1902]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="hamiltoni">
|
||
<emphasis id="B9237632474CD44CFE66F8C1FD25B111" box="[468,671,1877,1902]" italics="true" pageId="6" pageNumber="571">Thryssa hamiltoni</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC6D7D1474CD44CFD1FF8C1FC93B111" box="[685,809,1877,1902]" pageId="6" pageNumber="571" refId="ref5131" refString="Gray, J. E. (1835) Illustrations of Indian Zoology; Chiefly Selected From the Collection of Major-General Hardwicke, F. R. S. Fol. 2. Treuttel, wurtZ, Treuttel, Jun. and Richter, London, 15 pp. [pp. 88 - 102]" type="book chapter">Gray 1835</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which was followed also by Wongratana
|
||
<emphasis id="B9237632474CD44CFAB9F8C2FAFEB111" box="[1291,1348,1877,1902]" italics="true" pageId="6" pageNumber="571">et al.</emphasis>
|
||
(1999). Paxton
|
||
<emphasis id="B9237632474CD44CFF5EF8EEFE9FB1EE" box="[236,293,1912,1937]" italics="true" pageId="6" pageNumber="571">et al.</emphasis>
|
||
(2006) listed it as
|
||
<emphasis id="B9237632474CD44CFE4AF8EEFD29B1EE" box="[504,659,1912,1937]" italics="true" pageId="6" pageNumber="571">incertae sedis</emphasis>
|
||
in the
|
||
<taxonomicName id="4C57D1A3474CD44CFD53F8ECFCD1B1EE" authorityName="Gill" authorityYear="1861" box="[737,875,1912,1937]" class="Actinopterygii" family="Engraulidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="family">Engraulidae</taxonomicName>
|
||
, and as a possible synonym of
|
||
<emphasis id="B9237632474CD44CFB7BF8ECFA26B1EE" box="[1225,1436,1912,1937]" italics="true" pageId="6" pageNumber="571">
|
||
<taxonomicName id="4C57D1A3474CD44CFB7BF8ECFA22B1EE" baseAuthorityName="Gray" baseAuthorityYear="1835" box="[1225,1432,1912,1937]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="6" pageNumber="571" phylum="Chordata" rank="species" species="hamiltoni">Thryssa hamiltoni</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
They noted (p. 317): “
|
||
<typeStatus id="54EC1482474CD44CFE3CF809FE48B1CA" box="[398,498,1949,1973]" pageId="6" pageNumber="571" type="syntype">syntypes</typeStatus>
|
||
whereabouts unknown.”
|
||
</paragraph>
|
||
<paragraph id="8BE8AA20474CD44DFF75F855FBABB4FB" blockId="6.[151,1437,1372,2010]" lastBlockId="7.[151,1436,151,933]" lastPageId="7" lastPageNumber="572" pageId="6" pageNumber="571">
|
||
There are two specimens in the Macleay Museum (
|
||
<collectionCode id="ED4632E5474CD44CFCA1F855FCD5B1A6" box="[787,879,1985,2009]" country="Australia" httpUri="http://grbio.org/cool/6adq-425v" name="University of Sydney, Macleay Museum" pageId="6" pageNumber="571">MAMU</collectionCode>
|
||
F.1194;
|
||
<figureCitation id="136CB6A5474CD44CFC66F855FBE4B1A5" box="[980,1118,1985,2010]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="7.[151,250,1844,1866]" captionStartId-1="8.[151,250,1151,1173]" captionTargetBox-0="[151,1436,988,1823]" captionTargetBox-1="[151,1436,195,1127]" captionTargetId-0="figure@7.[151,1436,988,1823]" captionTargetId-1="figure@8.[151,1436,195,1127]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 6. Engraulis nasutus Castlenau, syntypes, MAMU F.1194, 99.5 (upper) and 104.5 (lower) mm SL. Photo by Y.K. Tea." captionText-1="FIGURE 7. Engraulis nasutus Castlenau, syntypes, MAMU F.1194, 99.5 (upper) and 104.5 (lower) mm SL. Radiograph by J. King." httpUri-0="https://zenodo.org/record/1458937/files/figure.png" httpUri-1="https://zenodo.org/record/1458939/files/figure.png" pageId="6" pageNumber="571">Figures 6–7</figureCitation>
|
||
), measuring 99.5 and
|
||
<quantity id="4CAF07C5474CD44DFAEFF855FF79B6D0" lastPageId="7" lastPageNumber="572" metricMagnitude="-1" metricUnit="m" metricValue="1.045" pageId="6" pageNumber="571" unit="mm" value="104.5">104.5 mm</quantity>
|
||
SL (TL not determinable owing to caudal-fin damage). They correspond to an index card stating “
|
||
<taxonomicName id="4C57D1A3474DD44DFA9FFF03FE8FB6AB" authority=", Cast." authorityName="Cast." class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9237632474DD44DFA9FFF03FF56B6AB" italics="true" pageId="7" pageNumber="572">Engraulis nasutus</emphasis>
|
||
, Cast.
|
||
</taxonomicName>
|
||
[…]
|
||
<specimenCount id="9D5161A9474DD44DFEC4FF28FE17B6AB" box="[374,429,188,212]" pageId="7" pageNumber="572" type="generic">2 sp.</specimenCount>
|
||
6″ Norman R., N.
|
||
<collectingCountry id="F340EAB0474DD44DFD3EFF28FD42B6AA" box="[652,760,188,213]" name="Australia" pageId="7" pageNumber="572">Australia</collectingCountry>
|
||
”. The specimens have the following characters (where two counts are presented, the first is from the
|
||
<quantity id="4CAF07C5474DD44DFDCFFF4BFD5EB688" box="[637,740,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="9.95" pageId="7" pageNumber="572" unit="mm" value="99.5">99.5 mm</quantity>
|
||
SL specimen): predorsal scutes 1; abdominal scutes sharply keeled, 14 prepelvic + 9 postpelvic = 23 total; maxilla relatively short, reaching to posterior border of preopercle; anal-fin rays 34 (iv,30; anterior ray tips damaged in
|
||
<quantity id="4CAF07C5474DD44DFD55FEB3FCE0B740" box="[743,858,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="1.045" pageId="7" pageNumber="572" unit="mm" value="104.5">104.5 mm</quantity>
|
||
SL specimen); total dorsal-fin rays 13; uppermost pectoral-fin ray not filamentous; teeth in jaws fine and conical, not canine-like; lower gill rakers 25 (checked in
|
||
<quantity id="4CAF07C5474DD44DFF25FEFBFEB3B7F8" box="[151,265,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="1.045" pageId="7" pageNumber="572" unit="mm" value="104.5">104.5 mm</quantity>
|
||
SL specimen only). This combination of characters is unique among engraulids to the species currently called
|
||
<taxonomicName id="4C57D1A3474DD44DFF53FE00FDE9B7D3" authority="(Ogilby 1910)" authorityName="Ogilby" authorityYear="1910" baseAuthorityName="Ogilby" baseAuthorityYear="1910" box="[225,595,404,429]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="aestuaria">
|
||
<emphasis id="B9237632474DD44DFF53FE00FE10B7D3" box="[225,426,404,429]" italics="true" pageId="7" pageNumber="572">Thryssa aestuaria</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC6D7D1474DD44DFE0BFE00FDF1B7D3" box="[441,587,404,429]" pageId="7" pageNumber="572" refId="ref5553" refString="Ogilby, J. D. (1910) On new or insufficiently described fishes. Proceedings of the Royal Society of Queensland, 23, 1 - 55." type="journal article">Ogilby 1910</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. According to Paxton
|
||
<emphasis id="B9237632474DD44DFCFFFE01FC3CB7D2" box="[845,902,404,429]" italics="true" pageId="7" pageNumber="572">et al.</emphasis>
|
||
(2006), there are three other species of
|
||
<taxonomicName id="4C57D1A3474DD44DFAF1FE00FA26B7D2" authorityName="Cuvier" authorityYear="1829" box="[1347,1436,404,429]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9237632474DD44DFAF1FE00FA26B7D2" box="[1347,1436,404,429]" italics="true" pageId="7" pageNumber="572">Thryssa</emphasis>
|
||
</taxonomicName>
|
||
known from the Gulf of Carpentaria:
|
||
<taxonomicName id="4C57D1A3474DD44DFDEBFE2DFD5FB7AF" baseAuthorityName="Gray" baseAuthorityYear="1835" box="[601,741,439,464]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="hamiltoni">
|
||
<emphasis id="B9237632474DD44DFDEBFE2DFD5FB7AF" box="[601,741,439,464]" italics="true" pageId="7" pageNumber="572">T. hamiltoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C57D1A3474DD44DFD4AFE2DFB76B7AF" authority="(Ramsay & Ogilby 1886)" authorityName="Ramsay & Ogilby" authorityYear="1886" baseAuthorityName="Ramsay & Ogilby" baseAuthorityYear="1886" box="[760,1228,439,464]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="scratchleyi">
|
||
<emphasis id="B9237632474DD44DFD4AFE2DFC29B7AF" box="[760,915,439,464]" italics="true" pageId="7" pageNumber="572">T. scratchleyi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC6D7D1474DD44DFC15FE2CFB7EB7AF" box="[935,1220,439,464]" pageId="7" pageNumber="572" refId="ref5752" refString="Ramsay, E. P. & Ogilby, J. D. (1886) A contribution to the knowledge of the fish-fauna of New Guinea. Proceedings of the Linnean Society of New South Wales, Series 2, 1, 8 - 20." type="journal article">Ramsay & Ogilby 1886</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C57D1A3474DD44DFABEFE2DFED3B78B" authority="(Broussonet 1782)" authorityName="Broussonet" authorityYear="1782" baseAuthorityName="Broussonet" baseAuthorityYear="1782" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="setirostris">
|
||
<emphasis id="B9237632474DD44DFABEFE2DFA21B7AF" box="[1292,1435,441,464]" italics="true" pageId="7" pageNumber="572">T. setirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC6D7D1474DD44DFF2CFE48FEDBB78B" box="[158,353,476,500]" pageId="7" pageNumber="572" refId="ref4780" refString="Broussonet, P. M. A. (1782) Ichthyologia, sistens piscium descriptiones et icones. Decas I. apud P. Elmsly, London, 49 pp. (unnumbered), 11 pls. (unnumbered)." type="book">Broussonet 1782</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The
|
||
<collectionCode id="ED4632E5474DD44DFE14FE48FDBAB78B" box="[422,512,476,500]" country="Australia" httpUri="http://grbio.org/cool/6adq-425v" name="University of Sydney, Macleay Museum" pageId="7" pageNumber="572">MAMU</collectionCode>
|
||
specimens differ from
|
||
<taxonomicName id="4C57D1A3474DD44DFCB6FE49FC30B78A" baseAuthorityName="Gray" baseAuthorityYear="1835" box="[772,906,476,501]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="hamiltoni">
|
||
<emphasis id="B9237632474DD44DFCB6FE49FC30B78A" box="[772,906,476,501]" italics="true" pageId="7" pageNumber="572">T. hamiltoni</emphasis>
|
||
</taxonomicName>
|
||
in having fewer prepelvic scutes (14 vs 15–20) and more lower gill rakers (25 vs 11–15); from
|
||
<taxonomicName id="4C57D1A3474DD44DFD07FD95FCF6B467" baseAuthorityName="Ramsay & Ogilby" baseAuthorityYear="1886" box="[693,844,511,536]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="scratchleyi">
|
||
<emphasis id="B9237632474DD44DFD07FD95FCF6B467" box="[693,844,511,536]" italics="true" pageId="7" pageNumber="572">T. scratchleyi</emphasis>
|
||
</taxonomicName>
|
||
in having fewer abdominal scutes (14 prepelvic + 9 postpelvic vs 19 + 12) and more lower gill rakers (
|
||
<quantity id="4CAF07C5474DD44DFD7AFDB0FC96B442" box="[712,812,548,573]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.842" metricValueMin="4.572" pageId="7" pageNumber="572" unit="in" value="20.5" valueMax="23.0" valueMin="18.0">18–23 in</quantity>
|
||
<taxonomicName id="4C57D1A3474DD44DFC80FDB1FC72B442" baseAuthorityName="Ramsay & Ogilby" baseAuthorityYear="1886" box="[818,968,548,573]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="scratchleyi">
|
||
<emphasis id="B9237632474DD44DFC80FDB1FC72B442" box="[818,968,548,573]" italics="true" pageId="7" pageNumber="572">T. scratchleyi</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="4C57D1A3474DD44DFBF9FDB1FB6FB443" baseAuthorityName="Broussonet" baseAuthorityYear="1782" box="[1099,1237,549,572]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="setirostris">
|
||
<emphasis id="B9237632474DD44DFBF9FDB1FB6FB443" box="[1099,1237,549,572]" italics="true" pageId="7" pageNumber="572">T. setirostris</emphasis>
|
||
</taxonomicName>
|
||
in having a much shorter maxilla (reaching to preopercle edge vs to at least tip of pectoral fins), fewer abdominal scutes (16–18 + 9– 10 =
|
||
<quantity id="4CAF07C5474DD44DFF63FDF8FE89B4FA" box="[209,307,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="6.731" metricValueMax="7.112" metricValueMin="6.35" pageId="7" pageNumber="572" unit="in" value="26.5" valueMax="28.0" valueMin="25.0">25–28 in</quantity>
|
||
<taxonomicName id="4C57D1A3474DD44DFE88FDF9FE7FB4FB" baseAuthorityName="Broussonet" baseAuthorityYear="1782" box="[314,453,621,644]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="setirostris">
|
||
<emphasis id="B9237632474DD44DFE88FDF9FE7FB4FB" box="[314,453,621,644]" italics="true" pageId="7" pageNumber="572">T. setirostris</emphasis>
|
||
</taxonomicName>
|
||
) and more lower gill rakers (
|
||
<quantity id="4CAF07C5474DD44DFCBEFDF8FCCAB4FA" box="[780,880,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="7" pageNumber="572" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
|
||
<taxonomicName id="4C57D1A3474DD44DFCC5FDF9FBBBB4FB" baseAuthorityName="Broussonet" baseAuthorityYear="1782" box="[887,1025,621,644]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="setirostris">
|
||
<emphasis id="B9237632474DD44DFCC5FDF9FBBBB4FB" box="[887,1025,621,644]" italics="true" pageId="7" pageNumber="572">T. setirostris</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE8AA20474DD44DFF75FD04FBD1B56B" blockId="7.[151,1436,151,933]" pageId="7" pageNumber="572">
|
||
We regard the specimens in MAMU F.1194 as syntypes of
|
||
<taxonomicName id="4C57D1A3474DD44DFCE9FD1BFB5EB4D7" authority="Castelnau, 1878" authorityName="Castelnau" authorityYear="1878" box="[859,1252,655,680]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9237632474DD44DFCE9FD1BFB9CB4D7" box="[859,1062,655,680]" italics="true" pageId="7" pageNumber="572">Engraulis nasutus</emphasis>
|
||
<bibRefCitation id="EFC6D7D1474DD44DFB9FFD1BFB5EB4D7" box="[1069,1252,655,680]" pageId="7" pageNumber="572" refId="ref4823" refString="Castelnau, F. L. de (1878 a) Australian fishes. New or little known species. Proceedings of the Linnean Society of New South Wales, 2, 225 - 248, pls. 2 - 3." type="journal article">Castelnau, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
, and identical to
|
||
<taxonomicName id="4C57D1A3474DD44DFF25FD20FDABB4B3" authority="Ogilby, 1910" authorityName="Ogilby" authorityYear="1910" box="[151,529,692,717]" class="Actinopterygii" family="Engraulidae" genus="Anchovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="aestuaria">
|
||
<emphasis id="B9237632474DD44DFF25FD20FEC8B4B3" box="[151,370,692,717]" italics="true" pageId="7" pageNumber="572">Anchovia aestuaria</emphasis>
|
||
<bibRefCitation id="EFC6D7D1474DD44DFEC8FD20FDABB4B3" box="[378,529,692,717]" pageId="7" pageNumber="572" refId="ref5553" refString="Ogilby, J. D. (1910) On new or insufficiently described fishes. Proceedings of the Royal Society of Queensland, 23, 1 - 55." type="journal article">Ogilby, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C57D1A3474DD44DFDA9FD20FD5DB4B3" authorityName="Castelnau" authorityYear="1878" box="[539,743,692,717]" class="Actinopterygii" family="Engraulidae" genus="Engraulis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9237632474DD44DFDA9FD20FD5DB4B3" box="[539,743,692,717]" italics="true" pageId="7" pageNumber="572">Engraulis nasutus</emphasis>
|
||
</taxonomicName>
|
||
thus is a subjective senior synonym of
|
||
<taxonomicName id="4C57D1A3474DD44DFB15FD20FA8AB4B3" authorityName="Ogilby" authorityYear="1910" box="[1191,1328,692,716]" class="Actinopterygii" family="Engraulidae" genus="Anchovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="aestuaria">
|
||
<emphasis id="B9237632474DD44DFB15FD20FA8AB4B3" box="[1191,1328,692,716]" italics="true" pageId="7" pageNumber="572">A. aestuaria</emphasis>
|
||
</taxonomicName>
|
||
. We here follow Eschmeyer
|
||
<emphasis id="B9237632474DD44DFED9FD4DFE1EB48F" box="[363,420,727,752]" italics="true" pageId="7" pageNumber="572">et al.</emphasis>
|
||
(2018, and references therein) in recognising
|
||
<taxonomicName id="4C57D1A3474DD44DFC1EFD43FB13B48F" authority="Cuvier (1829)" authorityName="Cuvier" authorityYear="1829" box="[940,1193,727,752]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9237632474DD44DFC1EFD43FBBFB48F" box="[940,1029,727,752]" italics="true" pageId="7" pageNumber="572">Thryssa</emphasis>
|
||
<bibRefCitation id="EFC6D7D1474DD44DFBBEFD43FB13B48F" box="[1036,1193,727,752]" pageId="7" pageNumber="572" refId="ref4901" refString="Cuvier, G. (1829) Le Regne Animal, distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee. Fol. 2. 2 nd Edition, CheZ Deterville, Paris, Xv + 406 pp." type="book">Cuvier (1829)</bibRefCitation>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="4C57D1A3474DD44DFA80FD43FA26B48F" authorityName="Jordan & Seale" authorityYear="1925" box="[1330,1436,727,752]" class="Actinopterygii" family="Engraulidae" genus="Thrissina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9237632474DD44DFA80FD43FA26B48F" box="[1330,1436,727,752]" italics="true" pageId="7" pageNumber="572">Thrissina</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFC6D7D1474DD44DFF25FD68FE29B56A" box="[151,403,764,789]" pageId="7" pageNumber="572" refId="ref5277" refString="Jordan, D. S. & Seale, A. (1925) Analysis of the genera of anchovies or Engraulidae. Copeia, 141, 27 - 32. https: // doi. org / 10.2307 / 1437308" type="journal article">Jordan & Seale (1925)</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFC6D7D1474DD44DFE10FD68FD85B56B" box="[418,575,764,789]" pageId="7" pageNumber="572" refId="ref5318" refString="Kottelat, M. (2013) The fishes of the inland waters of southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology Supplement, 27, 1 - 663." type="journal article">Kottelat 2013</bibRefCitation>
|
||
) as the correct generic name for
|
||
<taxonomicName id="4C57D1A3474DD44DFC1CFD68FBDEB56B" box="[942,1124,764,789]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9237632474DD44DFC1CFD68FBDEB56B" box="[942,1124,764,789]" italics="true" pageId="7" pageNumber="572">Thryssa nasutus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE8AA20474DD44DFF75FCB4FF61B5DA" blockId="7.[151,1436,151,933]" pageId="7" pageNumber="572">
|
||
<bibRefCitation id="EFC6D7D1474DD44DFF75FCB4FE09B547" box="[199,435,799,824]" pageId="7" pageNumber="572" refId="ref5674" refString="Pusey, B. J., Burrows, D. w., Kennard, M. J., Perna, C. N., Unmack, P. J., Allsop, Q. & Hammer, M. P. (2017) Freshwater fishes of northern Australia. Zootaxa, 4253 (1), 1 - 104. https: // doi. org / 10.11646 / ZootaXa. 4253.1.1" type="journal article">
|
||
Pusey
|
||
<emphasis id="B9237632474DD44DFEABFCB5FEE0B547" box="[281,346,799,824]" italics="true" pageId="7" pageNumber="572">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
recorded only a single engraulid from freshwaters of northern Australia,
|
||
<taxonomicName id="4C57D1A3474DD44DFAF1FC8BFEA8B522" baseAuthorityName="Ramsay & Ogilby" baseAuthorityYear="1886" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="scratchleyi">
|
||
<emphasis id="B9237632474DD44DFAF1FC8BFEA8B522" italics="true" pageId="7" pageNumber="572">Thryssa scratchleyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C57D1A3474DD44DFE92FCD0FE71B523" box="[288,459,836,861]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="nasuta">
|
||
<emphasis id="B9237632474DD44DFE92FCD0FE71B523" box="[288,459,836,861]" italics="true" pageId="7" pageNumber="572">Thryssa nasuta</emphasis>
|
||
</taxonomicName>
|
||
is known only from estuarine and marine areas (Whitehead
|
||
<emphasis id="B9237632474DD44DFBC6FCD1FB16B522" box="[1140,1196,836,861]" italics="true" pageId="7" pageNumber="572">et al.</emphasis>
|
||
1988, Wongratana
|
||
<emphasis id="B9237632474DD44DFA3AFCD1FF0EB5FF" italics="true" pageId="7" pageNumber="572">et al.</emphasis>
|
||
1999); presumably the syntypes of
|
||
<taxonomicName id="4C57D1A3474DD44DFDFEFCFDFD7BB5FF" box="[588,705,873,896]" class="Actinopterygii" family="Engraulidae" genus="Thryssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clupeiformes" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9237632474DD44DFDFEFCFDFD7BB5FF" box="[588,705,873,896]" italics="true" pageId="7" pageNumber="572">E. nasutus</emphasis>
|
||
</taxonomicName>
|
||
were collected from the lower, estuarine reaches of the Norman River.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |