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<mods:title id="9EB94C24DAFE52235188688D6B7A4683">Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species</mods:title>
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<mods:namePart id="FC20B513648BC470071C322E2914DB79">Zhou, Hong-Zhang</mods:namePart>
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<treatment id="03EC87E6FF8C88119DBF1800FABF0DFC" ID-DOI="http://doi.org/10.5281/zenodo.5691662" ID-GBIF-Taxon="119606316" ID-Zenodo-Dep="5691662" LSID="urn:lsid:plazi:treatment:03EC87E6FF8C88119DBF1800FABF0DFC" httpUri="http://treatment.plazi.org/id/03EC87E6FF8C88119DBF1800FABF0DFC" lastPageId="11" lastPageNumber="312" pageId="9" pageNumber="310">
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<paragraph id="8BFA36F0FF8C88139DBF1800FDA80F0E" blockId="9.[151,605,295,355]" box="[151,605,295,322]" pageId="9" pageNumber="310">
<heading id="D0B2819CFF8C88139DBF1800FDA80F0E" bold="true" box="[151,605,295,322]" fontSize="11" level="1" pageId="9" pageNumber="310" reason="1">
<taxonomicName id="4C454D73FF8C88139DBF1800FDA80F0E" ID-CoL="4Q98M" authority="Perreau, 1993" authorityName="Perreau" authorityYear="1993" box="[151,605,295,322]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="franki">
<emphasis id="B931EAE2FF8C88139DBF1800FDA80F0E" bold="true" box="[151,605,295,322]" pageId="9" pageNumber="310">
<emphasis id="B931EAE2FF8C88139DBF1800FE5D0F0D" bold="true" box="[151,424,295,321]" italics="true" pageId="9" pageNumber="310">Ptomaphaginus franki</emphasis>
<bibRefCitation id="EFD44B01FF8C88139C87180FFDA80F0E" author="Perreau" box="[431,605,296,322]" pageId="9" pageNumber="310" refString="Perreau, M. (1993) Nouvelles especes de Ptomaphaginus de Thailande et de Taiwan (Coleoptera, Cholevidae). Bulletin de la Societe Entomologique de France, 97 (4), 343 - 347." type="journal article" year="1993">Perreau, 1993</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFA36F0FF8C88139DBF186DFEBC0F2F" blockId="9.[151,605,295,355]" box="[151,329,330,355]" pageId="9" pageNumber="310">
(
<figureCitation id="137E2A75FF8C88139DB7186DFF1B0F2F" box="[159,238,330,355]" captionStart="FIGURE 5" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[152,1436,990,1875]" captionTargetId="figure@9.[151,1436,985,1875]" captionTargetPageId="9" captionText="FIGURE 5. Ptomaphaginus franki Perreau, 1993 ♂: habitus (dorsal view; paratype). Scales: 1 mm." httpUri="https://zenodo.org/record/243865/files/figure.png" pageId="9" pageNumber="310">Figs. 5</figureCitation>
; 6AK)
</paragraph>
<paragraph id="8BFA36F0FF8C88139DBF18B4FDC00FAA" blockId="9.[151,1436,402,486]" pageId="9" pageNumber="310">
<bibRefCitation id="EFD44B01FF8C88139DBF18B4FED20FE4" author="Perreau" box="[151,295,402,424]" pageId="9" pageNumber="310" refString="Perreau, M. (1993) Nouvelles especes de Ptomaphaginus de Thailande et de Taiwan (Coleoptera, Cholevidae). Bulletin de la Societe Entomologique de France, 97 (4), 343 - 347." type="journal article" year="1993">Perreau, 1993</bibRefCitation>
: 343 (
<taxonomicName id="4C454D73FF8C88139C4E18B4FDF00FE4" box="[358,517,403,424]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8C88139C4E18B4FDF00FE4" box="[358,517,403,424]" italics="true" pageId="9" pageNumber="310">Ptomaphaginus</emphasis>
</taxonomicName>
;
<typeStatus id="54FE8852FF8C88139F3A18B4FDB10FE4" box="[530,580,403,424]" pageId="9" pageNumber="310">Type</typeStatus>
locality:
<collectingCountry id="F3527660FF8C88139F8C18B5FD020FE4" box="[676,759,402,424]" name="China" pageId="9" pageNumber="310">CHINA</collectingCountry>
,
<collectingCountry id="F3527660FF8C88139E2918B5FCBA0FE4" box="[769,847,402,424]" name="Taiwan" pageId="9" pageNumber="310">Taiwan</collectingCountry>
: Fenchihu,
<quantity id="4CBD9B15FF8C88139EEC18B5FBE10FE4" box="[964,1044,402,424]" metricMagnitude="3" metricUnit="m" metricValue="1.409" pageId="9" pageNumber="310" unit="m" value="1409.0">1409 m</quantity>
);
<bibRefCitation id="EFD44B01FF8C8813990F18B4FB420FE4" author="Perreau" box="[1063,1207,402,424]" pageId="9" pageNumber="310" refString="Perreau, M. (1996) Contribution a la connaissance des Cholevidae du Japon et de Taiwan (Coleoptera). Revue Suisse de Zoologie, 103 (1), 283 - 297." type="journal article" year="1996">Perreau, 1996</bibRefCitation>
: 290 (
<taxonomicName id="4C454D73FF8C881399DE18B4FA610FE4" box="[1270,1428,403,424]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8C881399DE18B4FA610FE4" box="[1270,1428,403,424]" italics="true" pageId="9" pageNumber="310">Ptomaphaginus</emphasis>
</taxonomicName>
; new distributional locations);
<bibRefCitation id="EFD44B01FF8C88139F341895FD4F0F8B" author="Perreau" box="[540,698,433,455]" pageId="9" pageNumber="310" refString="Perreau, M. (2000) Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae. Memories de la Societe entomologique de France, 4, 1 - 460." type="journal article" year="2000">Perreau, 2000</bibRefCitation>
: 371 (
<taxonomicName id="4C454D73FF8C88139E3A1896FC440F8A" box="[786,945,433,454]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8C88139E3A1896FC440F8A" box="[786,945,433,454]" italics="true" pageId="9" pageNumber="310">Ptomaphaginus</emphasis>
</taxonomicName>
; catalog);
<bibRefCitation id="EFD44B01FF8C8813991F1896FAAA0F8B" author="Hoshina" box="[1079,1375,433,455]" pageId="9" pageNumber="310" refString="Hoshina, H. &amp; Sugaya, H. (2003) A taxonomic study of the genus Ptomaphaginus (Coleoptera: Leiodidae: Cholevinae) from the Ryukyu Islands, Japan. Entomological Review of Japan, 58 (2), 121 - 131." type="journal article" year="2003">Hoshina &amp; Sugaya, 2003</bibRefCitation>
: 127 (
<taxonomicName id="4C454D73FF8C88139DE718F7FE980FA9" box="[207,365,464,485]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8C88139DE718F7FE980FA9" box="[207,365,464,485]" italics="true" pageId="9" pageNumber="310">Ptomaphaginus</emphasis>
</taxonomicName>
; aedeagus figures).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF8C88139DBF1B35FB960C03" pageId="9" pageNumber="310" type="materials_examined">
<paragraph id="8BFA36F0FF8C88139DBF1B35FB960C03" blockId="9.[151,1437,530,915]" pageId="9" pageNumber="310">
<emphasis id="B931EAE2FF8C88139DBF1B35FD0D0C67" bold="true" box="[151,760,530,555]" pageId="9" pageNumber="310">
Material examined.
<typeStatus id="54FE8852FF8C88139CB31B35FDF90C67" box="[411,524,530,555]" pageId="9" pageNumber="310" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F3527660FF8C88139F0A1B35FD740C67" box="[546,641,530,555]" name="China" pageId="9" pageNumber="310">CHINA</collectingCountry>
,
<collectingCountry id="F3527660FF8C88139FBF1B35FD060C67" box="[663,755,530,555]" name="Taiwan" pageId="9" pageNumber="310">Taiwan</collectingCountry>
:
</emphasis>
<specimenCount id="9D43FD79FF8C88139E211B34FCD80C67" box="[777,813,531,555]" pageId="9" pageNumber="310" type="generic">1♂</specimenCount>
, Fenchihu,
<quantity id="4CBD9B15FF8C88139EEC1B34FBD00C66" box="[964,1061,531,555]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="310" unit="m" value="1400.0">1400 m</quantity>
,
<date id="FFFB1030FF8C881399131B34FB450C67" box="[1083,1200,531,555]" pageId="9" pageNumber="310" value="1977-05-10">10.V.1977</date>
, Klapperich leg. //
<typeStatus id="54FE8852FF8C88139DBF1B10FED00C03" box="[151,293,567,591]" pageId="9" pageNumber="310" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C454D73FF8C88139C181B11FD590C02" authority="M. Perreau" authorityName="M. Perreau" box="[304,684,566,591]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="franki">
<emphasis id="B931EAE2FF8C88139C181B11FDDC0C03" box="[304,553,566,591]" italics="true" pageId="9" pageNumber="310">Ptomaphaginus franki</emphasis>
M. Perreau
</taxonomicName>
det. 1992 // Coll. M. Perreau (
<collectionCode id="ED54AE35FF8C8813992B1B10FBA20C03" box="[1027,1111,567,591]" httpUri="http://grbio.org/cool/5fiw-1xnw" name="Centre for Medicinal Plants Research" pageId="9" pageNumber="310">CMPR</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF8C88119DEF1B7DFBD10D0F" lastPageId="11" lastPageNumber="312" pageId="9" pageNumber="310" type="description">
<paragraph id="8BFA36F0FF8C88139DEF1B7DFEF70C93" blockId="9.[151,1437,530,915]" pageId="9" pageNumber="310">
<emphasis id="B931EAE2FF8C88139DEF1B7DFE810C3F" bold="true" box="[199,372,602,627]" pageId="9" pageNumber="310">Redescription.</emphasis>
<emphasis id="B931EAE2FF8C88139C531B7DFE400C3F" box="[379,437,602,627]" italics="true" pageId="9" pageNumber="310">Male</emphasis>
. EBL:
<quantity id="4CBD9B15FF8C88139F2D1B7DFD990C3E" box="[517,620,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="2.53" pageId="9" pageNumber="310" unit="mm" value="2.53">2.53 mm</quantity>
. Length of different body parts: HL: AL: PL: ELL = 0.41: 0.78: 0.66:
<quantity id="4CBD9B15FF8C88139DBF1B58FF080CDA" box="[151,253,639,663]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="9" pageNumber="310" unit="mm" value="1.37">1.37 mm</quantity>
; width: HW: EW: PW: ELW = 0.70: 0.08: 1.09:
<quantity id="4CBD9B15FF8C88139E681B58FC530CDA" box="[832,934,639,663]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="9" pageNumber="310" unit="mm" value="1.11">1.11 mm</quantity>
. Proportion of antennomeres from base to tip in µm (length × width): 124 × 41, 107 × 47, 63 × 42, 40 × 45, 44 × 48, 30 × 54, 57 × 71, 25 × 70, 63 × 79, 61 × 78, 109 × 79.
</paragraph>
<paragraph id="8BFA36F0FF8C88139DEF1BCDFF070DDF" blockId="9.[151,1437,530,915]" pageId="9" pageNumber="310">
Habitus elongated oval, relatively convex and lustreless. Moderately pigmented: mostly chestnut brown: only mouthparts, basal five antennomeres and apical half of ultimate antennomere, elytral apex, and apical part of tarsi more or less yellowish. Dorsum continually clothed with fine, recumbent and sallow pubescence. Insertions of pubescence on dorsal surfaces of head, pronotum, scutellum, elytra and femora align along transverse striolations (
<figureCitation id="137E2A75FF8C88139DB71A5DFF110DDF" box="[159,228,890,915]" captionStart="FIGURE 5" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[152,1436,990,1875]" captionTargetId="figure@9.[151,1436,985,1875]" captionTargetPageId="9" captionText="FIGURE 5. Ptomaphaginus franki Perreau, 1993 ♂: habitus (dorsal view; paratype). Scales: 1 mm." httpUri="https://zenodo.org/record/243865/files/figure.png" pageId="9" pageNumber="310">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF3A6678FF8C88139DBF1E58FB6A09D9" box="[151,1183,1918,1941]" httpUri="https://zenodo.org/record/243865/files/figure.png" pageId="9" pageNumber="310" targetBox="[152,1436,990,1875]" targetPageId="9">
<paragraph id="8BFA36F0FF8C88139DBF1E58FB6A09D9" blockId="9.[151,1183,1918,1941]" box="[151,1183,1918,1941]" pageId="9" pageNumber="310">
<emphasis id="B931EAE2FF8C88139DBF1E58FD3709D9" bold="true" box="[151,706,1918,1941]" pageId="9" pageNumber="310">
FIGURE 5.
<taxonomicName id="4C454D73FF8C88139C311E59FD5409D9" authority="Perreau, 1993" authorityName="Perreau" authorityYear="1993" box="[281,673,1918,1941]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="franki">
<emphasis id="B931EAE2FF8C88139C311E59FDF009D8" bold="true" box="[281,517,1918,1940]" italics="true" pageId="9" pageNumber="310">Ptomaphaginus franki</emphasis>
Perreau, 1993
</taxonomicName>
♂:
</emphasis>
habitus (dorsal view; paratype). Scales: 1 mm.
</paragraph>
</caption>
<paragraph id="8BFA36F0FF8F88109DEF19B0FE520F6B" blockId="10.[151,1436,151,295]" pageId="10" pageNumber="311">
Head short and convex, transversely and superficially striolated, interspaces narrower than that on pronotum, HW
<emphasis id="B931EAE2FF8F88109DED1998FF240E94" bold="true" box="[197,209,191,216]" pageId="10" pageNumber="311">/</emphasis>
HL = 1.72; anterior margin round. Compound eyes normally developed, EW
<emphasis id="B931EAE2FF8F881099141998FBBD0E94" bold="true" box="[1084,1096,191,216]" pageId="10" pageNumber="311">/</emphasis>
HW = 0.11. Antennae slender but not very long (
<figureCitation id="137E2A75FF8F88109C4C19C0FE530F4C" box="[356,422,231,256]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="10" pageNumber="311">Fig. 6</figureCitation>
A), AL
<emphasis id="B931EAE2FF8F88109CDE19C0FDF70F4C" bold="true" box="[502,514,231,256]" pageId="10" pageNumber="311">/</emphasis>
HW = 1.11; 5th slightly longer than 4th; 6th wider than long; 9th and 10th a little wider than long; 11th pear-like.
</paragraph>
<caption id="DF3A6678FF8F88109DBF1E43FD68099A" httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="10" pageNumber="311" targetBox="[167,1416,362,1856]" targetPageId="10">
<paragraph id="8BFA36F0FF8F88109DBF1E43FD68099A" blockId="10.[151,1436,1892,2006]" pageId="10" pageNumber="311">
<emphasis id="B931EAE2FF8F88109DBF1E43FD3C0936" bold="true" box="[151,713,1892,1914]" pageId="10" pageNumber="311">
FIGURE 6.
<taxonomicName id="4C454D73FF8F88109C331E43FD5F0936" authority="Perreau, 1993" authorityName="Perreau" authorityYear="1993" box="[283,682,1892,1914]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="franki">
<emphasis id="B931EAE2FF8F88109C331E43FDFD0936" bold="true" box="[283,520,1892,1914]" italics="true" pageId="10" pageNumber="311">Ptomaphaginus franki</emphasis>
Perreau, 1993
</taxonomicName>
: A
</emphasis>
, antenna ♂ (dorsal view);
<emphasis id="B931EAE2FF8F88109EF61E43FC1A0936" bold="true" box="[990,1007,1892,1914]" pageId="10" pageNumber="311">B</emphasis>
, proleg ♂ (ventral view);
<emphasis id="B931EAE2FF8F881099D71E43FAE40936" bold="true" box="[1279,1297,1892,1914]" pageId="10" pageNumber="311">C</emphasis>
, pronotum ♂ (dorsal view);
<emphasis id="B931EAE2FF8F88109C181EA4FEB709D5" bold="true" box="[304,322,1923,1945]" pageId="10" pageNumber="311">D</emphasis>
, elytral apex ♂ (dorsoapical view);
<emphasis id="B931EAE2FF8F88109FE81EA4FD2409D5" bold="true" box="[704,721,1923,1945]" pageId="10" pageNumber="311">E</emphasis>
, ventrite VII ♂ (ventral view);
<emphasis id="B931EAE2FF8F881099081EA4FBD809D5" bold="true" box="[1056,1069,1923,1945]" pageId="10" pageNumber="311">F</emphasis>
, ventrite VIII ♂ (ventral view);
<emphasis id="B931EAE2FF8F881098AD1EA4FA6309D5" bold="true" box="[1413,1430,1923,1945]" pageId="10" pageNumber="311">G</emphasis>
, abdominal segment IX ♂ (ventral view);
<emphasis id="B931EAE2FF8F88109F141E85FDA509F4" bold="true" box="[572,592,1954,1976]" pageId="10" pageNumber="311">H</emphasis>
, aedeagus (dorsal view);
<emphasis id="B931EAE2FF8F88109E7E1E85FC9509F4" bold="true" box="[854,864,1954,1976]" pageId="10" pageNumber="311">I</emphasis>
, aedeagus (ventral view);
<emphasis id="B931EAE2FF8F881099461E85FB8F09F4" bold="true" box="[1134,1146,1954,1976]" pageId="10" pageNumber="311">J</emphasis>
, aedeagus (lateral view);
<emphasis id="B931EAE2FF8F881098AA1E85FA6309F4" bold="true" box="[1410,1430,1954,1976]" pageId="10" pageNumber="311">K</emphasis>
, spermatheca (after Perreau, 1993). Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BFA36F0FF8E88119DEF19BBFA730EB0" blockId="11.[151,1436,155,944]" pageId="11" pageNumber="312">
Pronotum transverse and convex (
<figureCitation id="137E2A75FF8E88119F7019BCFD550EF8" box="[600,672,155,180]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
C), widest directly before corners, PW
<emphasis id="B931EAE2FF8E8811994F19BCFB860EF8" bold="true" box="[1127,1139,155,180]" pageId="11" pageNumber="312">/</emphasis>
PL = 1.66. Sides slightly curved, narrowing from posterior to anterior; hind corners drawn out and blunt. Posterior margin with distinct postero-lateral emargination. Surface transversely striolated, SP: 4649, interspaces narrower than that on elytra.
</paragraph>
<paragraph id="8BFA36F0FF8E88119DEF182CFD730F20" blockId="11.[151,1436,155,944]" pageId="11" pageNumber="312">
Elytra of medium length and weakly convex, widest directly after base, ELL
<emphasis id="B931EAE2FF8E8811990B182DFBDA0F6F" bold="true" box="[1059,1071,266,291]" pageId="11" pageNumber="312">/</emphasis>
EW = 1.24. Sides feebly curved, gradually narrowing from base to apices; apices obliquely truncated (
<figureCitation id="137E2A75FF8E88119E8A1808FC130F04" box="[930,998,303,328]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
D). Surface transversely striolated, SE: 8992. Metathoracic wings fully developed.
</paragraph>
<paragraph id="8BFA36F0FF8E88119DEF185CFD690F97" blockId="11.[151,1436,155,944]" pageId="11" pageNumber="312">
Prolegs robust, with basal three protarsomeres moderately expanded (
<figureCitation id="137E2A75FF8E88119ED4185CFBB00FD8" box="[1020,1093,379,404]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
B): TW
<emphasis id="B931EAE2FF8E881199B6185DFB5F0FDF" bold="true" box="[1182,1194,378,403]" pageId="11" pageNumber="312">/</emphasis>
BTW = 1.35. Spinal arrangement on ventral side of protibia as shown in
<figureCitation id="137E2A75FF8E88119FF018B8FCEF0FF4" box="[728,794,415,440]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
B. Profemora wider than protibiae, and carried strong and long hairs on ventral side. Mesotibiae arcuate.
</paragraph>
<paragraph id="8BFA36F0FF8E88119DEF18C0FAFC0C04" blockId="11.[151,1436,155,944]" pageId="11" pageNumber="312">
Ventrite VII with two large teeth at posterior margin, and many small spines located around middle emargination (
<figureCitation id="137E2A75FF8E88119C131B2CFE750C68" box="[315,384,523,548]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
E). Ventrite VIII with a large fossa subround and moderately shallow, almost not emarginate posteriorly (
<figureCitation id="137E2A75FF8E88119C371B08FE960C04" box="[287,355,559,584]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
F). Segment IX: spiculum gastrale wide, widely rounded at posterior end (
<figureCitation id="137E2A75FF8E8811998A1B08FB100C04" box="[1186,1253,559,584]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
G).
</paragraph>
<paragraph id="8BFA36F0FF8E88119DEF1B74FDF60D53" blockId="11.[151,1436,155,944]" pageId="11" pageNumber="312">
Aedeagus stout and distinctly incurved around middle in dorsal view (
<figureCitation id="137E2A75FF8E88119EDE1B74FBC80C20" box="[1014,1085,595,620]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
H): median lobe widest before base; parameres narrow, firmly attached to median lobe, strongly and sinuately curved, totally invisible before middle. Ventral operculum distinctly divided in apical part, each lobe narrowly rounded apically (
<figureCitation id="137E2A75FF8E881199FE1BBCFAEF0CF8" box="[1238,1306,667,692]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">
Fig.
<date id="FFFB1030FF8E881198221BBCFAEF0CF8" box="[1290,1306,667,692]" pageId="11" pageNumber="312">6</date>
</figureCitation>
I). In lateral view, median lobe very thick and strongly bent ventrad at about apical half, apex almost not thinned, and four ventrally-oriented setae inserted below apex as well as two just before middle (
<figureCitation id="137E2A75FF8E8811996A1BC4FB7F0CB0" box="[1090,1162,739,764]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
J). Internal stylus wide, multiannulate at about basal 1/3.
</paragraph>
<paragraph id="8BFA36F0FF8E88119DEF1A0CFBD10D0F" blockId="11.[151,1436,155,944]" box="[199,1060,811,836]" pageId="11" pageNumber="312">
<emphasis id="B931EAE2FF8E88119DEF1A0CFEEE0D08" box="[199,283,811,836]" italics="true" pageId="11" pageNumber="312">Female</emphasis>
. Unexamined. Spermatheca C shape (
<figureCitation id="137E2A75FF8E88119FF01A0CFCEE0D08" box="[728,795,811,836]" captionStart="FIGURE 6" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[167,1416,362,1856]" captionTargetId="figure@10.[151,1436,347,1871]" captionTargetPageId="10" captionText="FIGURE 6. Ptomaphaginus franki Perreau, 1993: A, antenna ♂ (dorsal view); B, proleg ♂ (ventral view); C, pronotum ♂ (dorsal view); D, elytral apex ♂ (dorsoapical view); E, ventrite VII ♂ (ventral view); F, ventrite VIII ♂ (ventral view); G, abdominal segment IX ♂ (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view); K, spermatheca (after Perreau, 1993). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243866/files/figure.png" pageId="11" pageNumber="312">Fig. 6</figureCitation>
K; after
<bibRefCitation id="EFD44B01FF8E88119E501A0BFBED0D0F" author="Perreau" box="[888,1048,811,835]" pageId="11" pageNumber="312" refString="Perreau, M. (1993) Nouvelles especes de Ptomaphaginus de Thailande et de Taiwan (Coleoptera, Cholevidae). Bulletin de la Societe Entomologique de France, 97 (4), 343 - 347." type="journal article" year="1993">Perreau, 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF8E88119DEF1A68FDD50D24" box="[199,544,847,872]" pageId="11" pageNumber="312" type="distribution">
<paragraph id="8BFA36F0FF8E88119DEF1A68FDD50D24" blockId="11.[151,1436,155,944]" box="[199,544,847,872]" pageId="11" pageNumber="312">
<emphasis id="B931EAE2FF8E88119DEF1A68FE970D24" bold="true" box="[199,354,847,872]" pageId="11" pageNumber="312">Distribution:</emphasis>
<collectingCountry id="F3527660FF8E88119C411A68FE590D24" box="[361,428,847,872]" name="China" pageId="11" pageNumber="312">China</collectingCountry>
(
<collectingCountry id="F3527660FF8E88119C931A68FDE10D24" box="[443,532,847,872]" name="Taiwan" pageId="11" pageNumber="312">Taiwan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF8E88119DEF1A54FABF0DFC" pageId="11" pageNumber="312" type="discussion">
<paragraph id="8BFA36F0FF8E88119DEF1A54FABF0DFC" blockId="11.[151,1436,155,944]" pageId="11" pageNumber="312">
<emphasis id="B931EAE2FF8E88119DEF1A54FECE0DC0" bold="true" box="[199,315,883,908]" pageId="11" pageNumber="312">Remarks.</emphasis>
For
<taxonomicName id="4C454D73FF8E88119C501A54FDDD0DC0" box="[376,552,883,908]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8E88119C501A54FDDD0DC0" box="[376,552,883,908]" italics="true" pageId="11" pageNumber="312">Ptomaphaginus</emphasis>
</taxonomicName>
species from
<collectingCountry id="F3527660FF8E88119FF91A54FCE20DC0" box="[721,791,883,908]" name="China" pageId="11" pageNumber="312">China</collectingCountry>
, only this one has aedeagus distinctly incurved around middle when viewed dorsally, and parameres strongly and sinuately curved, totally invisible before middle.
</paragraph>
</subSubSection>
</treatment>
</document>