439 lines
51 KiB
XML
439 lines
51 KiB
XML
<document id="DD0664315FCB4F53A3A4CB91DB1684B6" ID-DOI="10.35249/rche.48.2.22.13" ID-ISSN="0718-8994" ID-Zenodo-Dep="13205757" ID-ZooBank="6B82B548-73E8-4501-9121-AEEF94C1FEBB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722662833282" checkinUser="felipe" docAuthor="Barria, Manuel" docDate="2022" docId="03EC87AEFFE6FFAC1AA531E728E8FA4E" docLanguage="en" docName="RevChilEntomol.48.2.313-319.pdf" docOrigin="Revista Chilena de Entomología (Rev. Chil. Entomol.) 48 (2)" docSource="http://dx.doi.org/10.35249/rche.48.2.22.13" docStyle="DocumentStyle:451321A1F3358405F3D5797D4FC35865.3:RevChilEntomol.2020-.journal_article" docStyleId="451321A1F3358405F3D5797D4FC35865" docStyleName="RevChilEntomol.2020-.journal_article" docStyleVersion="3" docTitle="Chrysina mercedesae Barria 2022, sp. nov." docType="treatment" docUuid="DAC26EE3-320E-466F-A902-5EE3465D60FC" docUuidSource="ZooBank" docVersion="2" lastPageNumber="318" masterDocId="FFD5FFD6FFE7FFA91B3A321A2911FF80" masterDocTitle="A new species of Chrysina Kirby, 1828 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini) from Ecuador and inclusion into proposed veraguana species-group" masterLastPageNumber="319" masterPageNumber="313" pageNumber="314" updateTime="1722702160900" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="137905408EF97EFAEF5A7834E1A2C591">A new species of Chrysina Kirby, 1828 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini) from Ecuador and inclusion into proposed veraguana species-group</mods:title>
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<mods:roleTerm id="6AD082BCE5BC8C83D80161F0C8C6FE8A">Author</mods:roleTerm>
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<mods:namePart id="D533877E1576D14BFDFA9644EDD34169">Barria, Manuel</mods:namePart>
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<mods:typeOfResource id="38ECA212EA59FC9851C2D064AF27F467">text</mods:typeOfResource>
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<mods:title id="DAB6C13F1BE97810C9665A1237EE5699">Revista Chilena de Entomología</mods:title>
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</mods:titleInfo>
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<mods:part id="BE5655506A94F00EC7236FB24F4D4A18">
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<mods:date id="C2E4E923970359E6ECBE879046AC9AEF">2022</mods:date>
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<mods:detail id="439F7C8843FE14BD19186977B9BED869" type="series">
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<mods:title id="820F79B39E8A673D3885D738FD99768E">Rev. Chil. Entomol.</mods:title>
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<mods:number id="60258DD3F46A568FDB37EC123A7C9DBD">2022-05-27</mods:number>
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<mods:number id="1CE28001B9D01E6B6DF672399E17E0FB">48</mods:number>
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<mods:number id="417A79EB4476C3A573C38E960F335C5B">2</mods:number>
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<mods:url id="C58436A3B7D681E0C0FFB6398036DE9A">http://dx.doi.org/10.35249/rche.48.2.22.13</mods:url>
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<mods:classification id="03C7C6B2AFBA3FD7AA05582ACFC58796">journal article</mods:classification>
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</mods:mods>
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<treatment id="03EC87AEFFE6FFAC1AA531E728E8FA4E" LSID="urn:lsid:zoobank.org:act:DAC26EE3-320E-466F-A902-5EE3465D60FC" httpUri="http://treatment.plazi.org/id/03EC87AEFFE6FFAC1AA531E728E8FA4E" lastPageId="5" lastPageNumber="318" pageId="1" pageNumber="314">
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<subSubSection id="C35F6533FFE6FFA81AA531E72A28FB95" box="[415,825,1021,1045]" pageId="1" pageNumber="314" type="nomenclature">
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<paragraph id="8BFA36B8FFE6FFA81AA531E72A28FB95" blockId="1.[415,825,1021,1077]" box="[415,825,1021,1045]" pageId="1" pageNumber="314">
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<heading id="D0B281D4FFE6FFA81AA531E72A28FB95" box="[415,825,1021,1045]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="314" reason="4">
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<taxonomicName id="4C454D3BFFE6FFA81AA531E72BC1FB95" ID-CoL="999KM" authority="Barria" authorityName="Barria" authorityYear="2022" box="[415,720,1021,1045]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="mercedesae" status="sp. nov.">
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<emphasis id="B931EAAAFFE6FFA81AA531E72B91FB95" box="[415,640,1021,1045]" italics="true" pageId="1" pageNumber="314">Chrysina mercedesae</emphasis>
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Barria
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</taxonomicName>
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,
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<emphasis id="B931EAAAFFE6FFA819E131E72A28FB95" bold="true" box="[731,825,1021,1045]" pageId="1" pageNumber="314">
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<taxonomicNameLabel id="A20257D1FFE6FFA819E131E72A28FB95" box="[731,825,1021,1045]" pageId="1" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6533FFE6FFA8190836072BB4FBB5" box="[562,677,1053,1077]" pageId="1" pageNumber="314" type="description">
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<paragraph id="8BFA36B8FFE6FFA8190836072BB4FBB5" blockId="1.[415,825,1021,1077]" box="[562,677,1053,1077]" pageId="1" pageNumber="314">
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(
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<figureCitation id="137E2A3DFFE6FFA8190036072B8DFBB5" box="[570,668,1053,1077]" captionStart-0="Figures 1" captionStart-1="Figure 2" captionStartId-0="3.[91,172,1297,1319]" captionStartId-1="4.[151,222,697,719]" captionTargetBox-0="[104,1107,130,1279]" captionTargetBox-1="[200,1161,123,682]" captionTargetId-0="figure-60@3.[91,1149,121,1286]" captionTargetId-1="figure-55@4.[200,1161,120,685]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figures 1a-f. Chrysina mercedesae sp. nov. a-c) Male holotype, dorsal, ventral and lateral views. 1d-e) Female allotype, dorsal and ventral views. f) Female genital plates. / a-c) Holotipo macho, vistas dorsal, ventral y lateral. 1d-e) Alotipo hembra, vistas dorsal y ventral. f) Placas genitales de la hembra." captionText-1="Figure 2. Chrysina mercedesae sp. nov. a-c) Aedeagus holotype, lateral, caudal and dorsal views. / a-c) Edeago del holotipo, vistas lateral, caudal y dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.13205759" figureDoi-1="http://doi.org/10.5281/zenodo.13205761" httpUri-0="https://zenodo.org/record/13205759/files/figure.png" httpUri-1="https://zenodo.org/record/13205761/files/figure.png" pageId="1" pageNumber="314">Figs. 1-2</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35F6533FFE6FFA81B61364628E2FA15" pageId="1" pageNumber="314" type="materials_examined">
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<paragraph id="8BFA36B8FFE6FFA81B61364628E2FA15" blockId="1.[90,1150,1116,1429]" pageId="1" pageNumber="314">
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<emphasis id="B931EAAAFFE6FFA81B6136462841FBF4" bold="true" box="[91,336,1116,1140]" pageId="1" pageNumber="314">Material examined.</emphasis>
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<materialsCitation id="3B2D3CE5FFE6FFA81A5B364728FEFA15" collectingDate="2022-04-03" collectingDateMax="2022-04-07" collectingDateMin="2022-04-03" collectionCode="LOUNAZ, UV, MDBC" collectorName="Robacker D. C." country="Ecuador" elevation="400" latitude="0.5238889" location="Male" longLatPrecision="1" longitude="-79.214165" pageId="1" pageNumber="314" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Esmeraldas" typeStatus="holotype">
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<typeStatus id="54FE881AFFE6FFA81A9736472B07FBF5" box="[429,534,1117,1141]" pageId="1" pageNumber="314" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED54AE7DFFE6FFA8191436472BB8FBF5" box="[558,681,1117,1141]" pageId="1" pageNumber="314">LOUNAZ</collectionCode>
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), labeled: “
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<collectingCountry id="F3527628FFE6FFA8180636472ADAFBF5" box="[828,971,1117,1141]" name="Ecuador" pageId="1" pageNumber="314">ECUADOR</collectingCountry>
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:
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<collectingRegion id="4981F85AFFE6FFA818E5364729F7FB15" country="Ecuador" name="Esmeraldas" pageId="1" pageNumber="314">Provincia de Esmeraldas</collectingRegion>
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, Reserva Canandé,
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</date>
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, Barria M.D. &
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<collectorName id="26B0536EFFE6FFA81B4336872834FB35" box="[121,293,1181,1205]" pageId="1" pageNumber="314">Robacker D.C.</collectorName>
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coll., Metal-halide Hg lamp &
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||
<collectionCode id="ED54AE7DFFE6FFA819A636872BD5FB34" box="[668,708,1181,1204]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="1" pageNumber="314">UV</collectionCode>
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light trap // (red label)
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||
<typeStatus id="54FE881AFFE6FFA818D536872D6FFB35" box="[1007,1150,1181,1205]" pageId="1" pageNumber="314" type="holotype">HOLOTIPO</typeStatus>
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<specimenCount id="9D43FD31FFE6FFA81B6736DA2961FB54" box="[93,112,1216,1236]" count="1" pageId="1" pageNumber="314" type="male">♂</specimenCount>
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<taxonomicName id="4C454D3BFFE6FFA81B4336A728E8FB55" authority="Barria 2022" authorityName="Barria" authorityYear="2022" box="[121,505,1213,1237]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="mercedesae">
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||
<emphasis id="B931EAAAFFE6FFA81B4336A7284EFB55" box="[121,351,1213,1237]" italics="true" pageId="1" pageNumber="314">Chrysina mercedesae</emphasis>
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||
Barria 2022
|
||
</taxonomicName>
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||
”. Female
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||
<typeStatus id="54FE881AFFE6FFA8194936A72BC2FB55" box="[627,723,1213,1237]" pageId="1" pageNumber="314" type="allotype">allotype</typeStatus>
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(
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||
<collectionCode id="ED54AE7DFFE6FFA819DC36A72A70FB55" box="[742,865,1213,1237]" pageId="1" pageNumber="314">LOUNAZ</collectionCode>
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), labeled: “
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<collectingCountry id="F3527628FFE6FFA818D136A72D6BFB55" box="[1003,1146,1213,1237]" name="Ecuador" pageId="1" pageNumber="314">ECUADOR</collectingCountry>
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:
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<collectingRegion id="4981F85AFFE6FFA81B6136C72891FB75" box="[91,384,1245,1269]" country="Ecuador" name="Esmeraldas" pageId="1" pageNumber="314">Provincia de Esmeraldas</collectingRegion>
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, Reserva Canandé,
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,
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,
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<collectingDate id="EFBFE990FFE6FFA818CD36C72D69FB75" box="[1015,1144,1245,1269]" pageId="1" pageNumber="314" value="2022-04-03">3/IV/2022</collectingDate>
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</date>
|
||
, Barria M.D. &
|
||
<collectorName id="26B0536EFFE6FFA81A3D36E728A0FA95" box="[263,433,1277,1301]" pageId="1" pageNumber="314">Robacker D.C.</collectorName>
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coll., Hg &
|
||
<collectionCode id="ED54AE7DFFE6FFA8190736E72B74FA94" box="[573,613,1277,1300]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="1" pageNumber="314">UV</collectionCode>
|
||
light trap // (red label)
|
||
<typeStatus id="54FE881AFFE6FFA818BF36E72AECFA95" box="[901,1021,1277,1301]" pageId="1" pageNumber="314" type="alotype">ALOTIPO</typeStatus>
|
||
<specimenCount id="9D43FD31FFE6FFA81F3E371A2D02FA94" box="[1028,1043,1280,1300]" count="1" pageId="1" pageNumber="314" type="female">♀</specimenCount>
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<taxonomicName id="4C454D3BFFE6FFA81F2036E7287DFAB5" authority="Barria 2022" authorityName="Barria" authorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="mercedesae">
|
||
<emphasis id="B931EAAAFFE6FFA81F2036E729C3FAB5" italics="true" pageId="1" pageNumber="314">Chrysina mercedesae</emphasis>
|
||
Barria 2022
|
||
</taxonomicName>
|
||
”.
|
||
<typeStatus id="54FE881AFFE6FFA81ABF370728E3FAB5" box="[389,498,1309,1333]" pageId="1" pageNumber="314" type="paratype">Paratype</typeStatus>
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.
|
||
<specimenCount id="9D43FD31FFE6FFA8193B37072B42FAB5" box="[513,595,1309,1333]" count="1" pageId="1" pageNumber="314" type="male">1 male</specimenCount>
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||
(
|
||
<collectionCode id="ED54AE7DFFE6FFA8195D37072BACFAB5" box="[615,701,1309,1333]" pageId="1" pageNumber="314">MDBC</collectionCode>
|
||
), labeled: “
|
||
<collectingCountry id="F3527628FFE6FFA8187C37072AC4FAB5" box="[838,981,1309,1333]" name="Ecuador" pageId="1" pageNumber="314">ECUADOR</collectingCountry>
|
||
:
|
||
<collectingRegion id="4981F85AFFE6FFA818DE370729F7FAD5" country="Ecuador" name="Esmeraldas" pageId="1" pageNumber="314">Provincia de Esmeraldas</collectingRegion>
|
||
, Reserva Canandé,
|
||
<quantity id="4CBD9B5DFFE6FFA81AEA37272B0BFAD4" box="[464,538,1341,1365]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="314" unit="m" value="400.0">
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<elevation id="0068D18BFFE6FFA81AEA37272B0BFAD4" box="[464,538,1341,1365]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="314" unit="m" value="400.0">400 m</elevation>
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</quantity>
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,
|
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<geoCoordinate id="EE71507FFFE6FFA8198037272A49FAD5" box="[698,856,1341,1365]" degrees="79" direction="west" minutes="12" orientation="longitude" pageId="1" pageNumber="314" precision="1" seconds="51.0" value="-79.214165">79°12’51.0”W</geoCoordinate>
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,
|
||
<date id="FFFB1078FFE6FFA8185E37272AF4FAD5" box="[868,997,1341,1365]" pageId="1" pageNumber="314" value="2022-04-07">
|
||
<collectingDate id="EFBFE990FFE6FFA8185E37272AF4FAD5" box="[868,997,1341,1365]" pageId="1" pageNumber="314" value="2022-04-07">7/IV/2022</collectingDate>
|
||
</date>
|
||
, Barria M.D. &
|
||
<collectorName id="26B0536EFFE6FFA81B4D37472830FAF5" box="[119,289,1373,1397]" pageId="1" pageNumber="314">Robacker D.C.</collectorName>
|
||
coll., Metal-halide Hg lamp &
|
||
<collectionCode id="ED54AE7DFFE6FFA819B037472BA3FAF4" box="[650,690,1373,1396]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="1" pageNumber="314">UV</collectionCode>
|
||
light trap // (yellow label)
|
||
<typeStatus id="54FE881AFFE6FFA818C337472D6FFAF5" box="[1017,1150,1373,1397]" pageId="1" pageNumber="314" type="paratype">PARATIPO</typeStatus>
|
||
<specimenCount id="9D43FD31FFE6FFA81B60379A297CFA14" box="[90,109,1408,1428]" count="1" pageId="1" pageNumber="314" type="male">♂</specimenCount>
|
||
<taxonomicName id="4C454D3BFFE6FFA81B4E376728F7FA15" authority="Barria 2022" authorityName="Barria" authorityYear="2022" box="[116,486,1405,1429]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="mercedesae">
|
||
<emphasis id="B931EAAAFFE6FFA81B4E37672844FA15" box="[116,341,1405,1429]" italics="true" pageId="1" pageNumber="314">Chrysina mercedesae</emphasis>
|
||
Barria 2022
|
||
</taxonomicName>
|
||
”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6533FFE6FFAB1B6137A62D12FB95" lastPageId="2" lastPageNumber="315" pageId="1" pageNumber="314" type="description">
|
||
<paragraph id="8BFA36B8FFE6FFA81B6137A62D6CFA54" blockId="1.[89,1150,1471,1748]" box="[91,1149,1468,1493]" pageId="1" pageNumber="314">
|
||
<emphasis id="B931EAAAFFE6FFA81B6137A628A9FA55" bold="true" box="[91,440,1468,1493]" pageId="1" pageNumber="314">
|
||
Description. Male
|
||
<typeStatus id="54FE881AFFE6FFA81A7637A728A9FA55" box="[332,440,1469,1493]" pageId="1" pageNumber="314" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="137E2A3DFFE6FFA81AF537A72B0FFA55" box="[463,542,1469,1493]" captionStart="Figures 1" captionStartId="3.[91,172,1297,1319]" captionTargetBox="[104,1107,130,1279]" captionTargetId="figure-60@3.[91,1149,121,1286]" captionTargetPageId="3" captionText="Figures 1a-f. Chrysina mercedesae sp. nov. a-c) Male holotype, dorsal, ventral and lateral views. 1d-e) Female allotype, dorsal and ventral views. f) Female genital plates. / a-c) Holotipo macho, vistas dorsal, ventral y lateral. 1d-e) Alotipo hembra, vistas dorsal y ventral. f) Placas genitales de la hembra." figureDoi="http://doi.org/10.5281/zenodo.13205759" httpUri="https://zenodo.org/record/13205759/files/figure.png" pageId="1" pageNumber="314">Figs. 1</figureCitation>
|
||
a-c): Length:
|
||
<quantity id="4CBD9B5DFFE6FFA819F137A72A37FA54" box="[715,806,1469,1493]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="314" unit="mm" value="27.0">27 mm</quantity>
|
||
. Humeral width:
|
||
<quantity id="4CBD9B5DFFE6FFA81F3637A72D68FA54" box="[1036,1145,1469,1493]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="314" unit="mm" value="11.7">11.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFA36B8FFE6FFA81B6137C72D6FF935" blockId="1.[89,1150,1471,1748]" pageId="1" pageNumber="314">
|
||
Maximum elytra width:
|
||
<quantity id="4CBD9B5DFFE6FFA81ABF37C728FEFA74" box="[389,495,1501,1525]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="1" pageNumber="314" unit="mm" value="14.8">14.8 mm</quantity>
|
||
.
|
||
<emphasis id="B931EAAAFFE6FFA81AC737C72B29FA75" box="[509,568,1501,1525]" italics="true" pageId="1" pageNumber="314">Color</emphasis>
|
||
: Dorsally, bright pale yellowish-green; antennal scape, external border of clypeus, ocular margins, lateral pronotal margin beads pinkish-violet; dorsal and ventral side of the tibiae pinkish-violet; in dorsal and ventral view, meso- and metatibia mostly pinkish-brown with greenish tones; in lateral view, meso- and metatibiae pinkish-violaceous; elytra epipleura margins pinkish; elytral
|
||
<emphasis id="B931EAAAFFE6FFA8188F34472AF3F9F5" box="[949,994,1629,1653]" italics="true" pageId="1" pageNumber="314">calli</emphasis>
|
||
and external borders metallic golden; internal borders of the tibiae and lateral sides of the mesosternal process metallic greenish; femora ventral side greenish, internal borders pinkish-brownish;
|
||
</paragraph>
|
||
<paragraph id="8BFA36B8FFE6FFAB1B6334A72B5EFD34" blockId="1.[89,1150,1471,1748]" lastBlockId="2.[150,1211,125,693]" lastPageId="2" lastPageNumber="315" pageId="1" pageNumber="314">
|
||
protarsomeres and mesotarsomeres pinkish-violet, metatarsomeres greenish; venter and abdominal sternites matte metallic greenish with pinkish reflections.
|
||
<emphasis id="B931EAAAFFE5FFAB1F1632672D75FF15" box="[1068,1124,125,149]" italics="true" pageId="2" pageNumber="315">Head</emphasis>
|
||
: Frons depressed, surface densely and finely punctate; clypeus subtrapezoidal, apex slightly truncated, reflexed; labium with anterior margin strongly emarginated.
|
||
<emphasis id="B931EAAAFFE5FFAB18DD32A72D45FF54" box="[999,1108,189,212]" italics="true" pageId="2" pageNumber="315">Pronotum</emphasis>
|
||
: Surface like the frons; basal bead diffused centrally.
|
||
<emphasis id="B931EAAAFFE5FFAB19AC32C72BCBFF75" box="[662,730,221,245]" italics="true" pageId="2" pageNumber="315">Elytra</emphasis>
|
||
: Surface with about 8-9 striate; epipleura narrow, reaching level of second abdominal sternite.
|
||
<taxonomicName id="4C454D3BFFE5FFAB183C32E72A65FE95" box="[774,884,253,277]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="315" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EAAAFFE5FFAB183C32E72A65FE95" box="[774,884,253,277]" italics="true" pageId="2" pageNumber="315">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: Wider than long; posterior margin parabolic; surface sparsely punctate.
|
||
<taxonomicName id="4C454D3BFFE5FFAB198733072A39FEB5" box="[701,808,285,309]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="315" phylum="Chordata" rank="genus">
|
||
<emphasis id="B931EAAAFFE5FFAB198733072A39FEB5" box="[701,808,285,309]" italics="true" pageId="2" pageNumber="315">Pygidium</emphasis>
|
||
</taxonomicName>
|
||
: Plate (lateral view) weakly and uniformly convex; surface (frontal view) transversely and finely rugopunctate, with a few setae along rounded apex.
|
||
<emphasis id="B931EAAAFFE5FFAB1ADA33472B39FEF4" box="[480,552,349,372]" italics="true" pageId="2" pageNumber="315">Venter</emphasis>
|
||
: Surface glabrous; prosternal plate subtriangular with rounded apex, central region slightly concave; mesometasternal process moderately long, surpassing base of mesocoxae; in lateral view, process semiconical, poorly robust, process directed forward; fifth abdominal sternite with a broad emargination at its posterior end, with a slightly sclerosed subtriangular isosceliform membrane between fifth and sixth sternites; sixth abdominal sternite with an arcuate emargination at posterior margin, followed by a slightly sclerosed membrane; posterior margins of abdominal sternites with several transversely arranged setae.
|
||
<emphasis id="B931EAAAFFE5FFAB197030272B68FDD4" box="[586,633,573,596]" italics="true" pageId="2" pageNumber="315">Legs</emphasis>
|
||
: Protibia tridentate.
|
||
<emphasis id="B931EAAAFFE5FFAB184B30272AC5FDD5" box="[881,980,573,597]" italics="true" pageId="2" pageNumber="315">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="137E2A3DFFE5FFAB18D930272D23FDD5" box="[995,1074,573,597]" captionStart="Figure 2" captionStartId="4.[151,222,697,719]" captionTargetBox="[200,1161,123,682]" captionTargetId="figure-55@4.[200,1161,120,685]" captionTargetPageId="4" captionText="Figure 2. Chrysina mercedesae sp. nov. a-c) Aedeagus holotype, lateral, caudal and dorsal views. / a-c) Edeago del holotipo, vistas lateral, caudal y dorsal." figureDoi="http://doi.org/10.5281/zenodo.13205761" httpUri="https://zenodo.org/record/13205761/files/figure.png" pageId="2" pageNumber="315">Figs. 2</figureCitation>
|
||
a- c): Males parameres short and symmetrical, with two recurved and divergent dentiform processes becoming progressively more robust apically (appreciable in lateral view,
|
||
<figureCitation id="137E2A3DFFE5FFAB1F3930672D42FD15" box="[1027,1107,637,661]" captionStart="Figure 2" captionStartId="4.[151,222,697,719]" captionTargetBox="[200,1161,123,682]" captionTargetId="figure-55@4.[200,1161,120,685]" captionTargetPageId="4" captionText="Figure 2. Chrysina mercedesae sp. nov. a-c) Aedeagus holotype, lateral, caudal and dorsal views. / a-c) Edeago del holotipo, vistas lateral, caudal y dorsal." figureDoi="http://doi.org/10.5281/zenodo.13205761" httpUri="https://zenodo.org/record/13205761/files/figure.png" pageId="2" pageNumber="315">Fig. 2a</figureCitation>
|
||
), ventral plates protruding beyond parameres.
|
||
</paragraph>
|
||
<paragraph id="8BFA36B8FFE5FFAB1BAD30C72AC9FC34" blockId="2.[150,1211,733,949]" pageId="2" pageNumber="315">
|
||
<emphasis id="B931EAAAFFE5FFAB1BAD30C7284BFD75" bold="true" box="[151,346,733,757]" pageId="2" pageNumber="315">
|
||
Female
|
||
<typeStatus id="54FE881AFFE5FFAB1BCD30C7284BFD75" box="[247,346,733,757]" pageId="2" pageNumber="315" type="allotype">allotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="137E2A3DFFE5FFAB1A5630C728A4FD75" box="[364,437,733,757]" captionStart="Figures 1" captionStartId="3.[91,172,1297,1319]" captionTargetBox="[104,1107,130,1279]" captionTargetId="figure-60@3.[91,1149,121,1286]" captionTargetPageId="3" captionText="Figures 1a-f. Chrysina mercedesae sp. nov. a-c) Male holotype, dorsal, ventral and lateral views. 1d-e) Female allotype, dorsal and ventral views. f) Female genital plates. / a-c) Holotipo macho, vistas dorsal, ventral y lateral. 1d-e) Alotipo hembra, vistas dorsal y ventral. f) Placas genitales de la hembra." figureDoi="http://doi.org/10.5281/zenodo.13205759" httpUri="https://zenodo.org/record/13205759/files/figure.png" pageId="2" pageNumber="315">Figs. 1</figureCitation>
|
||
d-f): Length:
|
||
<quantity id="4CBD9B5DFFE5FFAB196E30C72BB8FD74" box="[596,681,733,757]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="315" unit="mm" value="31.0">31 mm</quantity>
|
||
. Humeral width:
|
||
<quantity id="4CBD9B5DFFE5FFAB184730C72AF7FD74" box="[893,998,733,757]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="2" pageNumber="315" unit="mm" value="14.2">14.2 mm</quantity>
|
||
. Maximum elytra width:
|
||
<quantity id="4CBD9B5DFFE5FFAB1BD730E72847FC94" box="[237,342,765,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="2" pageNumber="315" unit="mm" value="17.1">17.1 mm</quantity>
|
||
. Similar in coloration to the male; more robust and larger than males; the apex of the clypeus is mostly rounded (not truncated as in the male); without emargination between the fifth and sixth sternites, sixth abdominal sternite with a small arched emargination at apex; anterolateral surface of abdominal sternites coarsely punctate.
|
||
<emphasis id="B931EAAAFFE5FFAB1BAD316729EBFC15" box="[151,250,893,917]" italics="true" pageId="2" pageNumber="315">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="137E2A3DFFE5FFAB1A3031672847FC15" box="[266,342,893,917]" captionStart="Figures 1" captionStartId="3.[91,172,1297,1319]" captionTargetBox="[104,1107,130,1279]" captionTargetId="figure-60@3.[91,1149,121,1286]" captionTargetPageId="3" captionText="Figures 1a-f. Chrysina mercedesae sp. nov. a-c) Male holotype, dorsal, ventral and lateral views. 1d-e) Female allotype, dorsal and ventral views. f) Female genital plates. / a-c) Holotipo macho, vistas dorsal, ventral y lateral. 1d-e) Alotipo hembra, vistas dorsal y ventral. f) Placas genitales de la hembra." figureDoi="http://doi.org/10.5281/zenodo.13205759" httpUri="https://zenodo.org/record/13205759/files/figure.png" pageId="2" pageNumber="315">Fig. 1f</figureCitation>
|
||
): Female inferior plates pear-shaped, slightly narrower towards the apex, apical margins strongly serrate-ondulate with many erect rufous setae.
|
||
</paragraph>
|
||
<paragraph id="8BFA36B8FFE5FFAB1BAD31C62D12FB95" blockId="2.[151,1210,988,1045]" pageId="2" pageNumber="315">
|
||
<emphasis id="B931EAAAFFE5FFAB1BAD31C6281EFC74" bold="true" box="[151,271,988,1012]" pageId="2" pageNumber="315">Variation.</emphasis>
|
||
Male
|
||
<typeStatus id="54FE881AFFE5FFAB1A6731C728DBFC75" box="[349,458,989,1013]" pageId="2" pageNumber="315" type="paratype">paratype</typeStatus>
|
||
: Length:
|
||
<quantity id="4CBD9B5DFFE5FFAB190631C72BB4FC74" box="[572,677,989,1013]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="2" pageNumber="315" unit="mm" value="27.6">27.6 mm</quantity>
|
||
; humeral width:
|
||
<quantity id="4CBD9B5DFFE5FFAB184231C72AF2FC74" box="[888,995,989,1013]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="315" unit="mm" value="12.2">12.2 mm</quantity>
|
||
. Maximum elytra width:
|
||
<quantity id="4CBD9B5DFFE5FFAB1BD131E72843FB94" box="[235,338,1021,1045]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="2" pageNumber="315" unit="mm" value="14.9">14.9 mm</quantity>
|
||
. All tarsomeres of the male
|
||
<typeStatus id="54FE881AFFE5FFAB19A231E72A13FB95" box="[664,770,1021,1045]" pageId="2" pageNumber="315" type="paratype">paratype</typeStatus>
|
||
are metallic greenish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6533FFE5FFAB1BAD36262B95FA74" pageId="2" pageNumber="315" type="diagnosis">
|
||
<paragraph id="8BFA36B8FFE5FFAB1BAD36262B95FA74" blockId="2.[151,1211,1084,1525]" pageId="2" pageNumber="315">
|
||
<emphasis id="B931EAAAFFE5FFAB1BAD36262808FBD4" bold="true" box="[151,281,1084,1108]" pageId="2" pageNumber="315">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C454D3BFFE5FFAB1A1E36272B18FBD5" authorityName="Barria" authorityYear="2022" box="[292,521,1085,1109]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="mercedesae">
|
||
<emphasis id="B931EAAAFFE5FFAB1A1E36272B18FBD5" box="[292,521,1085,1109]" italics="true" pageId="2" pageNumber="315">Chrysina mercedesae</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from the other species of the genus by the combination of the following characters: striated elytra, pale yellowish-green dorsal coloration, venter matte metallic green with pinkish-violet reflections, tarsomeres pinkish-violet or greenish. The genitalia of the male have two symmetrical and recurved processes becoming progressively more robust apically, ventral processes of the phallus protrude beyond the parameres.
|
||
<taxonomicName id="4C454D3BFFE5FFAB1A8136C72BB0FB75" authorityName="Barria" authorityYear="2022" box="[443,673,1245,1269]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="mercedesae">
|
||
<emphasis id="B931EAAAFFE5FFAB1A8136C72BB0FB75" box="[443,673,1245,1269]" italics="true" pageId="2" pageNumber="315">Chrysina mercedesae</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C454D3BFFE5FFAB187336C72AACFB75" authorityName="Hawks" authorityYear="2017" box="[841,957,1245,1269]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="galbina">
|
||
<emphasis id="B931EAAAFFE5FFAB187336C72A4AFB74" box="[841,859,1245,1268]" italics="true" pageId="2" pageNumber="315">C</emphasis>
|
||
.
|
||
<emphasis id="B931EAAAFFE5FFAB185436C72AACFB75" box="[878,957,1245,1269]" italics="true" pageId="2" pageNumber="315">galbina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D3BFFE5FFAB1F3B36C72D86FB74" baseAuthorityName="Ohaus" baseAuthorityYear="1922" box="[1025,1175,1245,1268]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="veraguana">
|
||
<emphasis id="B931EAAAFFE5FFAB1F3B36C72D02FB74" box="[1025,1043,1245,1268]" italics="true" pageId="2" pageNumber="315">C</emphasis>
|
||
.
|
||
<emphasis id="B931EAAAFFE5FFAB1F1C36C72D86FB74" box="[1062,1175,1245,1268]" italics="true" pageId="2" pageNumber="315">veraguana</emphasis>
|
||
</taxonomicName>
|
||
in coloration and the genitalia
|
||
<taxonomicName id="4C454D3BFFE5FFAB1ADC36E72B45FA95" authorityName="Hawks" authorityYear="2017" box="[486,596,1277,1301]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="galbina">
|
||
<emphasis id="B931EAAAFFE5FFAB1ADC36E728E6FA94" box="[486,503,1277,1300]" italics="true" pageId="2" pageNumber="315">C</emphasis>
|
||
.
|
||
<emphasis id="B931EAAAFFE5FFAB193F36E72B45FA95" box="[517,596,1277,1301]" italics="true" pageId="2" pageNumber="315">galbina</emphasis>
|
||
</taxonomicName>
|
||
does not have a metallic venter, but it is yellowish-green, similar to dorsum, the male parameres are asymmetrical, with two dentiform projections oriented ventrally and to the right (third projection fused with central projection), left projection more robust and less acute than right.
|
||
<taxonomicName id="4C454D3BFFE5FFAB189B37472D20FAF4" baseAuthorityName="Ohaus" baseAuthorityYear="1922" box="[929,1073,1373,1396]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="veraguana">
|
||
<emphasis id="B931EAAAFFE5FFAB189B37472AA0FAF4" box="[929,945,1373,1396]" italics="true" pageId="2" pageNumber="315">C</emphasis>
|
||
.
|
||
<emphasis id="B931EAAAFFE5FFAB18FA37472D20FAF4" box="[960,1073,1373,1396]" italics="true" pageId="2" pageNumber="315">veraguana</emphasis>
|
||
</taxonomicName>
|
||
has silvery sternites and bluish tarsomeres, the male parameres present three projections, one short central dentiform process and two medians lateral dentiform processes, strongly curved towards ventral region, left process slightly more robust than right, both species are restricted to the central region of
|
||
<collectingCountry id="F3527628FFE5FFAB192437C72B91FA74" box="[542,640,1501,1524]" name="Panama" pageId="2" pageNumber="315">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6533FFE5FFAB1BAD34072DA9F9F4" pageId="2" pageNumber="315" type="etymology">
|
||
<paragraph id="8BFA36B8FFE5FFAB1BAD34072DA9F9F4" blockId="2.[149,1211,1565,1653]" pageId="2" pageNumber="315">
|
||
<emphasis id="B931EAAAFFE5FFAB1BAD3407280EF9B5" bold="true" box="[151,287,1565,1589]" pageId="2" pageNumber="315">Etymology.</emphasis>
|
||
The new species name is dedicated to my grandmother Mercedes A. Ortega, who during all my years of life and career as a student and researcher has always supported me and believed in me. The specific epithet is a singular feminine noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6533FFE5FFAC1BAD348629BCFB6D" lastPageId="5" lastPageNumber="318" pageId="2" pageNumber="315" type="distribution">
|
||
<paragraph id="8BFA36B8FFE5FFAB1BAD34862AFBF934" blockId="2.[151,1002,1692,1717]" box="[151,1002,1692,1717]" pageId="2" pageNumber="315">
|
||
<emphasis id="B931EAAAFFE5FFAB1BAD34862827F934" bold="true" box="[151,310,1692,1716]" pageId="2" pageNumber="315">Distribution.</emphasis>
|
||
<collectingCountry id="F3527628FFE5FFAB1A07348728B2F935" box="[317,419,1693,1717]" name="Ecuador" pageId="2" pageNumber="315">Ecuador</collectingCountry>
|
||
: Reserva Canandé (
|
||
<collectingRegion id="4981F85AFFE5FFAB19B034872A92F935" box="[650,899,1693,1717]" country="Ecuador" name="Esmeraldas" pageId="2" pageNumber="315">Esmeraldas Province</collectingRegion>
|
||
) (
|
||
<figureCitation id="137E2A3DFFE5FFAB18A334872ACBF935" box="[921,986,1693,1717]" captionStart="Figure 3" captionStartId="4.[151,222,1700,1722]" captionTargetBox="[252,1159,786,1686]" captionTargetId="figure-29@4.[201,1159,783,1688]" captionTargetPageId="4" captionText="Figure 3. Distribution map of the veraguana species-group. / Mapa de distribución del grupo de" figureDoi="http://doi.org/10.5281/zenodo.13205763" httpUri="https://zenodo.org/record/13205763/files/figure.png" pageId="2" pageNumber="315">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF3A6630FFE4FFAA1B61370B29DCFAFF" ID-DOI="http://doi.org/10.5281/zenodo.13205759" ID-Zenodo-Dep="13205759" httpUri="https://zenodo.org/record/13205759/files/figure.png" pageId="3" pageNumber="316" startId="3.[91,172,1297,1319]" targetBox="[104,1107,130,1279]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BFA36B8FFE4FFAA1B61370B29DCFAFF" blockId="3.[91,1149,1297,1407]" pageId="3" pageNumber="316">
|
||
<emphasis id="B931EAAAFFE4FFAA1B61370B29F7FAA7" bold="true" box="[91,230,1297,1319]" pageId="3" pageNumber="316">Figures 1a-f.</emphasis>
|
||
<taxonomicName id="4C454D3BFFE4FFAA1BD5370B28D1FAA6" authorityName="Barria" authorityYear="2022" box="[239,448,1297,1318]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="mercedesae" status="sp. nov.">
|
||
<emphasis id="B931EAAAFFE4FFAA1BD5370B28D1FAA6" box="[239,448,1297,1318]" italics="true" pageId="3" pageNumber="316">Chrysina mercedesae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EAAAFFE4FFAA1AF0370B2B44FAA7" bold="true" box="[458,597,1297,1319]" pageId="3" pageNumber="316">
|
||
<taxonomicNameLabel id="A20257D1FFE4FFAA1AF0370B2B32FAA6" box="[458,547,1297,1318]" pageId="3" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
|
||
a-c)
|
||
</emphasis>
|
||
Male holotype, dorsal, ventral and lateral views.
|
||
<emphasis id="B931EAAAFFE4FFAA1B6137342980FAC4" bold="true" box="[91,145,1326,1348]" pageId="3" pageNumber="316">1d-e)</emphasis>
|
||
Female allotype, dorsal and ventral views.
|
||
<emphasis id="B931EAAAFFE4FFAA194837342B95FAC4" bold="true" box="[626,644,1326,1348]" pageId="3" pageNumber="316">f)</emphasis>
|
||
Female genital plates. /
|
||
<emphasis id="B931EAAAFFE4FFAA18A237342AAEFAC4" bold="true" box="[920,959,1326,1348]" pageId="3" pageNumber="316">a-c)</emphasis>
|
||
Holotipo macho, vistas dorsal, ventral y lateral.
|
||
<emphasis id="B931EAAAFFE4FFAA1A93375628F1FAE2" bold="true" box="[425,480,1356,1378]" pageId="3" pageNumber="316">1d-e)</emphasis>
|
||
Alotipo hembra, vistas dorsal y ventral.
|
||
<emphasis id="B931EAAAFFE4FFAA18A137562ABDFAE2" bold="true" box="[923,940,1356,1378]" pageId="3" pageNumber="316">f)</emphasis>
|
||
Placas genitales de la hembra.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFA36B8FFE4FFAA1B6137A72B42F955" blockId="3.[91,1150,1469,1749]" pageId="3" pageNumber="316">
|
||
<emphasis id="B931EAAAFFE4FFAA1B6137A7287AFA55" bold="true" box="[91,363,1469,1493]" pageId="3" pageNumber="316">Distribution remarks.</emphasis>
|
||
I received a photo (
|
||
<date id="FFFB1078FFE4FFAA194F37A72A07FA55" box="[629,790,1469,1493]" pageId="3" pageNumber="316" value="2022-01">January 2022</date>
|
||
) in dorsal view of a female specimen from east of Riosucio (Chocó,
|
||
<collectingCountry id="F3527628FFE4FFAA196B37C72BD9FA75" box="[593,712,1501,1525]" name="Colombia" pageId="3" pageNumber="316">Colombia</collectingCountry>
|
||
), south of Pavarando Grande, very close to the limits between the department of Chocó and
|
||
<collectingRegion id="4981F85AFFE4FFAA19C237E72A7EF995" box="[760,879,1533,1557]" country="Colombia" name="Antioquia" pageId="3" pageNumber="316">Antioquia</collectingRegion>
|
||
(
|
||
<figureCitation id="137E2A3DFFE4FFAA184437E72AD0F995" box="[894,961,1533,1557]" captionStart="Figure 3" captionStartId="4.[151,222,1700,1722]" captionTargetBox="[252,1159,786,1686]" captionTargetId="figure-29@4.[201,1159,783,1688]" captionTargetPageId="4" captionText="Figure 3. Distribution map of the veraguana species-group. / Mapa de distribución del grupo de" figureDoi="http://doi.org/10.5281/zenodo.13205763" httpUri="https://zenodo.org/record/13205763/files/figure.png" pageId="3" pageNumber="316">Fig. 3</figureCitation>
|
||
, black asterisk). The female specimen appears identical to the one described in this work; however, I was unable to trace the collection where it was deposited to dissect the genitalia and confirm the identity. If this record is confirmed in the future, it would mean a great extension of the distribution of the species to the north of Chocó in
|
||
<collectingCountry id="F3527628FFE4FFAA19DD34672A4CF915" box="[743,861,1661,1685]" name="Colombia" pageId="3" pageNumber="316">Colombia</collectingCountry>
|
||
. Assuming this, it could be thought that the species is distributed throughout the biogeographical Chocó, from the north in
|
||
<collectingCountry id="F3527628FFE4FFAA1BFA34A72822F955" box="[192,307,1725,1749]" name="Colombia" pageId="3" pageNumber="316">Colombia</collectingCountry>
|
||
to the south in
|
||
<collectingCountry id="F3527628FFE4FFAA1AD034A72B5FF955" box="[490,590,1725,1749]" name="Ecuador" pageId="3" pageNumber="316">Ecuador</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF3A6630FFE3FFAD1BAD30A32BA5FD6D" ID-DOI="http://doi.org/10.5281/zenodo.13205761" ID-Zenodo-Dep="13205761" httpUri="https://zenodo.org/record/13205761/files/figure.png" pageId="4" pageNumber="317" startId="4.[151,222,697,719]" targetBox="[200,1161,123,682]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BFA36B8FFE3FFAD1BAD30A32BA5FD6D" blockId="4.[151,1209,697,749]" pageId="4" pageNumber="317">
|
||
<emphasis id="B931EAAAFFE3FFAD1BAD30A329E4FD4E" bold="true" box="[151,245,697,719]" pageId="4" pageNumber="317">Figure 2.</emphasis>
|
||
<taxonomicName id="4C454D3BFFE3FFAD1BC030A028D5FD4F" authorityName="Barria" authorityYear="2022" box="[250,452,698,719]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="mercedesae" status="sp. nov.">
|
||
<emphasis id="B931EAAAFFE3FFAD1BC030A028D5FD4F" box="[250,452,698,719]" italics="true" pageId="4" pageNumber="317">Chrysina mercedesae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EAAAFFE3FFAD1AF330A02B56FD4F" bold="true" box="[457,583,697,719]" pageId="4" pageNumber="317">
|
||
<taxonomicNameLabel id="A20257D1FFE3FFAD1AF330A02B0BFD4F" box="[457,538,698,719]" pageId="4" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
|
||
a-c)
|
||
</emphasis>
|
||
Aedeagus holotype, lateral, caudal and dorsal views. /
|
||
<emphasis id="B931EAAAFFE3FFAD1FA830A32DA8FD4F" bold="true" box="[1170,1209,697,719]" pageId="4" pageNumber="317">a-c)</emphasis>
|
||
Edeago del holotipo, vistas lateral, caudal y dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3A6630FFE3FFAD1BAD34BE2DA8F93A" ID-DOI="http://doi.org/10.5281/zenodo.13205763" ID-Zenodo-Dep="13205763" box="[151,1209,1700,1722]" httpUri="https://zenodo.org/record/13205763/files/figure.png" pageId="4" pageNumber="317" startId="4.[151,222,1700,1722]" targetBox="[252,1159,786,1686]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BFA36B8FFE3FFAD1BAD34BE2DA8F93A" blockId="4.[151,1209,1700,1748]" box="[151,1209,1700,1722]" pageId="4" pageNumber="317">
|
||
<emphasis id="B931EAAAFFE3FFAD1BAD34BE29E8F939" bold="true" box="[151,249,1700,1722]" pageId="4" pageNumber="317">Figure 3.</emphasis>
|
||
Distribution map of the
|
||
<taxonomicName id="4C454D3BFFE3FFAD193534BF2B65F93A" baseAuthorityName="Ohaus" baseAuthorityYear="1922" box="[527,628,1701,1722]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="veraguana">
|
||
<emphasis id="B931EAAAFFE3FFAD193534BF2B65F93A" box="[527,628,1701,1722]" italics="true" pageId="4" pageNumber="317">veraguana</emphasis>
|
||
</taxonomicName>
|
||
species-group. / Mapa de distribución del grupo de
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFA36B8FFE3FFAC1BAD34D8284CFF54" blockId="4.[151,1209,1700,1748]" lastBlockId="5.[91,1149,124,213]" lastPageId="5" lastPageNumber="318" pageId="4" pageNumber="317">
|
||
especies
|
||
<taxonomicName id="4C454D3BFFE3FFAD1BCE34D82848F957" baseAuthorityName="Ohaus" baseAuthorityYear="1922" box="[244,345,1730,1751]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="veraguana">
|
||
<emphasis id="B931EAAAFFE3FFAD1BCE34D82848F957" box="[244,345,1730,1751]" italics="true" pageId="4" pageNumber="317">veraguana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B931EAAAFFE2FFAC1B61326629AFFF14" bold="true" box="[91,190,124,148]" pageId="5" pageNumber="318">Biology.</emphasis>
|
||
Immatures are unknown. The species inhabit in tropical premontane forest about the
|
||
<quantity id="4CBD9B5DFFE2FFAC1BE83287285BFF34" box="[210,330,157,181]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="5" pageNumber="318" unit="m" value="450.0" valueMax="600.0" valueMin="300.0">300-600 m</quantity>
|
||
(
|
||
<figureCitation id="137E2A3DFFE2FFAC1A633287288CFF35" box="[345,413,157,181]" captionStart="Figure 4" captionStartId="5.[91,162,975,997]" captionTargetBox="[91,1149,257,963]" captionTargetId="figure-69@5.[91,1149,257,963]" captionTargetPageId="5" captionText="Figure 4. Habitat of Chrysina mercedesae sp. nov., Reserva Canandé, Prov. Esmeraldas, Ecuador. / Hábitat de Chrysina mercedesae sp. nov., Reserva Canandé, Prov. Esmeraldas, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.13205765" httpUri="https://zenodo.org/record/13205765/files/figure.png" pageId="5" pageNumber="318">Fig. 4</figureCitation>
|
||
). Adults were captured at night with mercury vapor light and ultraviolet light traps.
|
||
</paragraph>
|
||
<caption id="DF3A6630FFE2FFAC1B6131D52AF4FB83" ID-DOI="http://doi.org/10.5281/zenodo.13205765" ID-Zenodo-Dep="13205765" httpUri="https://zenodo.org/record/13205765/files/figure.png" pageId="5" pageNumber="318" startId="5.[91,162,975,997]" targetBox="[91,1149,257,963]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BFA36B8FFE2FFAC1B6131D52AF4FB83" blockId="5.[91,1149,975,1027]" pageId="5" pageNumber="318">
|
||
<emphasis id="B931EAAAFFE2FFAC1B6131D529ADFC64" bold="true" box="[91,188,975,997]" pageId="5" pageNumber="318">Figure 4.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="4C454D3BFFE2FFAC1A0631D52B18FC64" authorityName="Barria" authorityYear="2022" box="[316,521,975,996]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="mercedesae" status="sp. nov.">
|
||
<emphasis id="B931EAAAFFE2FFAC1A0631D52B18FC64" box="[316,521,975,996]" italics="true" pageId="5" pageNumber="318">Chrysina mercedesae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EAAAFFE2FFAC192831CA2B77FC65" bold="true" box="[530,614,976,997]" pageId="5" pageNumber="318">
|
||
<taxonomicNameLabel id="A20257D1FFE2FFAC192831CA2B77FC65" box="[530,614,976,997]" pageId="5" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Reserva Canandé, Prov. Esmeraldas, Ecuador. / Hábitat de
|
||
<taxonomicName id="4C454D3BFFE2FFAC1BEB31F7288AFB82" authorityName="Barria" authorityYear="2022" box="[209,411,1005,1026]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="mercedesae" status="sp. nov.">
|
||
<emphasis id="B931EAAAFFE2FFAC1BEB31F7288AFB82" box="[209,411,1005,1026]" italics="true" pageId="5" pageNumber="318">Chrysina mercedesae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EAAAFFE2FFAC1A9B31F728E3FB82" bold="true" box="[417,498,1005,1026]" pageId="5" pageNumber="318">
|
||
<taxonomicNameLabel id="A20257D1FFE2FFAC1A9B31F728E3FB82" box="[417,498,1005,1026]" pageId="5" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Reserva Canandé, Prov. Esmeraldas, Ecuador.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFA36B8FFE2FFAC1AEC362F2A13FBCD" blockId="5.[470,770,1077,1101]" box="[470,770,1077,1101]" pageId="5" pageNumber="318">
|
||
<emphasis id="B931EAAAFFE2FFAC1AEC362F2A13FBCD" bold="true" box="[470,770,1077,1101]" pageId="5" pageNumber="318">
|
||
<taxonomicName id="4C454D3BFFE2FFAC1AEC362F2B42FBCC" baseAuthorityName="Ohaus" baseAuthorityYear="1922" box="[470,595,1077,1100]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="veraguana">
|
||
<emphasis id="B931EAAAFFE2FFAC1AEC362F2B42FBCC" bold="true" box="[470,595,1077,1100]" italics="true" pageId="5" pageNumber="318">veraguana</emphasis>
|
||
</taxonomicName>
|
||
species-group
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFA36B8FFE2FFAC1B61366F29BCFB6D" blockId="5.[91,1150,1141,1261]" pageId="5" pageNumber="318">
|
||
<emphasis id="B931EAAAFFE2FFAC1B61366F2867FB0D" bold="true" box="[91,374,1141,1165]" pageId="5" pageNumber="318">Diagnostic description.</emphasis>
|
||
Green dorsal coloration; striated elytra; abdominal sternites with or without metallic sheen; prosternal process short and subtriangular; male parameres ending in three or apparently two dentiform processes, parameres mostly “short” and robust.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6533FFE2FFAC1B61370F28E8FA4E" pageId="5" pageNumber="318" type="discussion">
|
||
<paragraph id="8BFA36B8FFE2FFAC1B61370F28E8FA4E" blockId="5.[90,1150,1301,1486]" pageId="5" pageNumber="318">
|
||
<emphasis id="B931EAAAFFE2FFAC1B61370F29DFFAAD" bold="true" box="[91,206,1301,1325]" pageId="5" pageNumber="318">Remarks.</emphasis>
|
||
The
|
||
<taxonomicName id="4C454D3BFFE2FFAC1A3C370C2866FAAD" baseAuthorityName="Ohaus" baseAuthorityYear="1922" box="[262,375,1302,1325]" class="Insecta" family="Scarabaeidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="veraguana">
|
||
<emphasis id="B931EAAAFFE2FFAC1A3C370C2866FAAD" box="[262,375,1302,1325]" italics="true" pageId="5" pageNumber="318">veraguana</emphasis>
|
||
</taxonomicName>
|
||
species-group is very close to the
|
||
<taxonomicName id="4C454D3BFFE2FFAC183E370C2A62FAAD" authorityName="Waterhouse" authorityYear="1871" box="[772,883,1302,1325]" class="Insecta" family="Scarabaeidae" genus="Plusiotis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="marginata">
|
||
<emphasis id="B931EAAAFFE2FFAC183E370C2A62FAAD" box="[772,883,1302,1325]" italics="true" pageId="5" pageNumber="318">marginata</emphasis>
|
||
</taxonomicName>
|
||
group in that the male genitalia have pronounced dentiform processes in both groups. Also, abdominal sternites of some species in each group are metallic while other species have matte appearance. Differences between the groups include elytral striation in the veraguana group compared with smooth elytra in the
|
||
<taxonomicName id="4C454D3BFFE2FFAC1A91378C2B0BFA2D" authorityName="Waterhouse" authorityYear="1871" box="[427,538,1430,1453]" class="Insecta" family="Scarabaeidae" genus="Plusiotis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="marginata">
|
||
<emphasis id="B931EAAAFFE2FFAC1A91378C2B0BFA2D" box="[427,538,1430,1453]" italics="true" pageId="5" pageNumber="318">marginata</emphasis>
|
||
</taxonomicName>
|
||
group and much shorter and more robust male parameres in the veraguana group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |