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<document id="2669E701EC1682FFEA260B1C18C9003A" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFF8500BFF23FDEEFCA846DA" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus ankaratranus Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="9" updateTime="1726053143081" updateUser="ExternalLinkService">
<mods:mods id="7CFCAAD5EDC22F2D96AD2A6B35EA21DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7F902C672156B2422FF3F0EF622E036B">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="A04DA0DDFBAD9447460D73E9A80E59A3">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="4B1EF3E71EE5924EA7C49C1167ACF36D">Zootaxa</mods:title>
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<mods:date id="92F327E8F036B9071F1296E94B745992">2024</mods:date>
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<mods:classification id="2E36C74E1633F1495EEA91045AEC9198">journal article</mods:classification>
<mods:identifier id="631D2DC0B4045B17FC9DAA2197DA4026" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
<mods:identifier id="32F6010BEDBCF907AF4F82DF4E572231" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03E7FF50FFF8500BFF23FDEEFCA846DA" ID-DOI="http://doi.org/10.5281/zenodo.13745989" ID-Zenodo-Dep="13745989" LSID="urn:lsid:plazi:treatment:03E7FF50FFF8500BFF23FDEEFCA846DA" httpUri="http://treatment.plazi.org/id/03E7FF50FFF8500BFF23FDEEFCA846DA" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
<subSubSection id="C3541DCDFFF85008FF23FDEEFCC2464E" box="[151,775,546,573]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BF14E46FFF85008FF23FDEEFCC2464E" blockId="8.[151,775,546,609]" box="[151,775,546,573]" pageId="8" pageNumber="9">
<heading id="D0B9F92AFFF85008FF23FDEEFCC2464E" bold="true" box="[151,775,546,573]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
<emphasis id="B93A9254FFF85008FF23FDEEFCC2464E" bold="true" box="[151,775,546,573]" pageId="8" pageNumber="9">
<taxonomicName id="4C4E35C5FFF85008FF23FDEEFD4B464F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,654,546,573]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankaratranus" status="comb. nov.">
<emphasis id="B93A9254FFF85008FF23FDEEFDED464E" bold="true" box="[151,552,546,573]" italics="true" pageId="8" pageNumber="9">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFF85008FD28FDEFFCC2464E" box="[668,775,547,573]" pageId="8" pageNumber="9" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85008FF23FD8BFEDC4612" box="[151,281,583,609]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF14E46FFF85008FF23FD8BFEDC4612" blockId="8.[151,775,546,609]" box="[151,281,583,609]" pageId="8" pageNumber="9">
(
<figureCitation id="137552C3FFF85008FF2BFD8BFED54612" box="[159,272,583,609]" captionStart-0="FIGURES 1–8" captionStart-1="FIGURES 9–11" captionStart-2="FIGURES 12–19" captionStartId-0="9.[151,264,1405,1429]" captionStartId-1="10.[151,264,705,729]" captionStartId-2="10.[151,264,1945,1969]" captionTargetBox-0="[151,1429,181,1369]" captionTargetBox-1="[183,1404,181,681]" captionTargetBox-2="[199,1390,902,1907]" captionTargetId-0="figure-16@9.[151,1435,181,1381]" captionTargetId-1="figure-28@10.[183,1404,181,681]" captionTargetId-2="figure-107@10.[189,1397,898,1922]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="10" captionText-0="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." captionText-1="FIGURES 9–11. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." captionText-2="FIGURES 12–19. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi-0="http://doi.org/10.5281/zenodo.13745916" figureDoi-1="http://doi.org/10.5281/zenodo.13745921" figureDoi-2="http://doi.org/10.5281/zenodo.13745923" httpUri-0="https://zenodo.org/record/13745916/files/figure.png" httpUri-1="https://zenodo.org/record/13745921/files/figure.png" httpUri-2="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">Figs 1–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85008FF23FD47FD3246B1" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8BF14E46FFF85008FF23FD47FD5646D0" blockId="8.[151,759,651,706]" box="[151,659,651,675]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB5E6968FFF85008FF23FD47FD5646D0" box="[151,659,651,675]" pageId="8" pageNumber="9">
<treatmentCitation id="0AEF6857FFF85008FF23FD47FD4A46D0" author="Franz, H." box="[151,655,651,675]" page="164" pageId="8" pageNumber="9" year="1986">
<taxonomicName id="4C4E35C5FFF85008FF23FD47FD4A46D0" ID-CoL="3MNJ7" authority="Franz, 1986 b: 164" authorityName="Franz" authorityPageNumber="164" authorityYear="1986" box="[151,655,651,675]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFF85008FF23FD47FE0846D0" box="[151,461,651,675]" italics="true" pageId="8" pageNumber="9">Horaeomorphus ankaratranus</emphasis>
<bibRefCitation id="EFDF33B7FFF85008FE66FD40FD4A46D0" author="Franz, H." box="[466,655,651,675]" pageId="8" pageNumber="9" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 164</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF14E46FFF85008FF23FD66FD3246B1" blockId="8.[151,759,651,706]" box="[151,759,681,706]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB5E6968FFF85008FF23FD66FD3246B1" box="[151,759,681,706]" pageId="8" pageNumber="9">
<treatmentCitation id="0AEF6857FFF85008FF23FD66FD6146B1" author="Franz, H." box="[151,676,682,706]" page="173" pageId="8" pageNumber="9" year="1986">
<taxonomicName id="4C4E35C5FFF85008FF23FD66FD6146B1" ID-CoL="3MNKM" authority="Franz, 1986 b: 173" authorityName="Franz" authorityPageNumber="173" authorityYear="1986" box="[151,676,682,706]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFF85008FF23FD66FE2746B1" box="[151,482,682,706]" italics="true" pageId="8" pageNumber="9">Horaeomorphus manjakotompoi</emphasis>
<bibRefCitation id="EFDF33B7FFF85008FE5CFD66FD6146B1" author="Franz, H." box="[488,676,682,706]" pageId="8" pageNumber="9" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 173</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B93A9254FFF85008FD19FD65FD3246B1" bold="true" box="[685,759,681,706]" pageId="8" pageNumber="9">Syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85008FF23FD3CFD2440C9" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BF14E46FFF85008FF23FD3CFC154791" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
<emphasis id="B93A9254FFF85008FF23FD3CFE744779" bold="true" box="[151,433,752,778]" pageId="8" pageNumber="9">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFF85008FE0BFD3CFC364721" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFF85008FE0BFD3CFC204779" bold="true" box="[447,997,751,778]" pageId="8" pageNumber="9">
<typeStatus id="54F5F0E4FFF85008FE0BFD3CFDE94779" box="[447,556,752,778]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFF85008FDEBFD3CFCE24779" authorityName="Franz" authorityYear="1986" box="[607,807,751,778]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFF85008FDEBFD3CFCE24779" bold="true" box="[607,807,751,778]" italics="true" pageId="8" pageNumber="9">H. ankaratranus</emphasis>
</taxonomicName>
(
<collectingCountry id="F3590ED6FFF85008FC89FD3CFC1D4779" box="[829,984,752,778]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFF85008FC47FD3DFBCF4778" box="[1011,1034,753,779]" count="1" pageId="8" pageNumber="9" type="male">♂</specimenCount>
, three labels: “Massif de l’An- / karatra,Madagaskar / lg.
<collectorName id="26BB2B90FFF85008FE01FCD8FDD7475D" box="[437,530,788,814]" pageId="8" pageNumber="9">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFF85008FCF8FCD8FBC6475D" box="[844,1027,788,814]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ ankaratranus m. / det.
<determinerName id="7E2DA282FFF85008FA80FCD8FA57475D" box="[1332,1426,788,814]" pageId="8" pageNumber="9">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFF85008FC3CFCF4FC2E4721" box="[904,1003,824,850]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFF85008FC4BFCF4FC914705" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="8" pageNumber="9" specimenCount="2" typeStatus="paralectotype">
<emphasis id="B93A9254FFF85008FC4BFCF4FB6A4721" bold="true" box="[1023,1199,824,850]" pageId="8" pageNumber="9">
<typeStatus id="54F5F0E4FFF85008FC4BFCF4FB6A4721" box="[1023,1199,824,850]" pageId="8" pageNumber="9" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
: 26 exx., data as for holotype, except each with yellow “
<typeStatus id="54F5F0E4FFF85008FD96FC90FD504705" box="[546,661,860,886]" pageId="8" pageNumber="9" type="paratype">Paratypus</typeStatus>
” label (
<collectionCode id="ED5FD683FFF85008FD5EFC90FC884705" box="[746,845,860,886]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26441BFFF85008FCE9FC90FC094791" collectionCode="R, VII, NHMW" country="Madagascar" determinerName="H. Franz" location="Manjakotompo" pageId="8" pageNumber="9" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFF85008FCE9FC90FABF4705" bold="true" box="[861,1402,859,886]" pageId="8" pageNumber="9">
<typeStatus id="54F5F0E4FFF85008FCE9FC90FC0F4705" box="[861,970,860,886]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFF85008FC59FC90FB014705" box="[1005,1220,859,886]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFF85008FC59FC90FB014705" bold="true" box="[1005,1220,859,886]" italics="true" pageId="8" pageNumber="9">H. manjakotompoi</emphasis>
</taxonomicName>
(
<collectingCountry id="F3590ED6FFF85008FB66FC90FAA84705" box="[1234,1389,860,886]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFF85008FA34FC91FA524704" box="[1408,1431,861,887]" count="1" pageId="8" pageNumber="9" type="male">♂</specimenCount>
, five labels: “
<location id="8E91189DFFF85008FE94FC4CFE0B47E9" LSID="urn:lsid:plazi:treatment:03E7FF50FFF8500BFF23FDEEFCA846DA:8E91189DFFF85008FE94FC4CFE0B47E9" box="[288,462,896,922]" country="Madagascar" name="Manjakotompo" pageId="8" pageNumber="9">Manjakotompo</location>
/ 6.vii.56 (E.
<collectionCode id="ED5FD683FFF85008FDEEFC4CFDAA47E9" box="[602,623,896,922]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="9">R</collectionCode>
{or K})” [brownish, handwritten], “
<specimenCount id="9D4885CFFFF85008FBB5FC4CFBDD47E9" box="[1025,1048,896,922]" count="1" pageId="8" pageNumber="9" type="male">♂</specimenCount>
,” [white, printed],
<taxonomicName id="4C4E35C5FFF85008FB5CFC4CFA5947E9" box="[1256,1436,896,922]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ manjakotompoi m. / det.
<determinerName id="7E2DA282FFF85008FE61FC68FDF747CD" box="[469,562,932,958]" pageId="8" pageNumber="9">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Manjakotompo /
<collectionCode id="ED5FD683FFF85008FB5EFC68FAD647CC" box="[1258,1299,932,959]" pageId="8" pageNumber="9">VII</collectionCode>
.56 (E.
<collectionCode id="ED5FD683FFF85008FAD7FC69FAB347CC" box="[1379,1398,933,959]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="9">R</collectionCode>
.) /
<collectingCountry id="F3590ED6FFF85008FF23FC04FEE44791" box="[151,289,968,994]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
” [white, handwritten], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFF85008FCD5FC04FC014791" box="[865,964,968,994]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73FC20FD2440C9" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
<emphasis id="B93A9254FFF85008FF73FC20FE634075" bold="true" box="[199,422,1004,1030]" pageId="8" pageNumber="9">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFF85008FDB0FC20FD8C4075" box="[516,585,1004,1030]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
) stout; head (
<figureCitation id="137552C3FFF85008FD41FC20FCF84075" box="[757,829,1004,1030]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
) about as long as wide, with vertex subconical, in anterodorsal view its posterior margin subtriangular; pronotum (
<figureCitation id="137552C3FFF85008FCDAFBDCFC774059" box="[878,946,1040,1066]" captionStart="FIGURES 9–11" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 9–11. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
) subtrapezoidal, broadest at base, with two pairs of distinct antebasal pits; antennomeres 3–10 not elongate; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFF85008FE28FB94FE3A4000" box="[412,511,1112,1139]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 5–8</figureCitation>
) in ventral view broadest near base, with apical region of dorsal wall abruptly bent dorsad, distolateral endophallic sclerites widely separated and with rounded apices, each paramere with a row of subapical setae distributed on more than distal third.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF85009FF73FB08FD894319" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF14E46FFF85008FF73FB08FB434170" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
<emphasis id="B93A9254FFF85008FF73FB08FEB040AD" bold="true" box="[199,373,1220,1246]" pageId="8" pageNumber="9">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFF85008FDA9FB08FDA740AD" box="[541,610,1220,1246]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 1</figureCitation>
,
<figureCitation id="137552C3FFF85008FDDBFB08FD4940AD" box="[623,652,1220,1246]" captionStart="FIGURES 12–19" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 12–19. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">12</figureCitation>
) moderately dark to strongly dark brown with reddish hue, appendages indistinctly lighter, setae lighter than cuticle, yellowish to light brown; BL
<quantity id="4CB6E3A3FFF85008FC68FB24FB474170" box="[988,1154,1256,1283]" metricMagnitude="-3" metricUnit="m" metricValue="1.635" metricValueMax="1.68" metricValueMin="1.59" pageId="8" pageNumber="9" unit="mm" value="1.635" valueMax="1.68" valueMin="1.59">1.59–1.68 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73FAC0FB784251" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
Head (
<figureCitation id="137552C3FFF85008FEA5FAC0FEB34155" box="[273,374,1292,1318]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 2–3</figureCitation>
,
<figureCitation id="137552C3FFF85008FE35FAC0FE7E4155" box="[385,443,1292,1318]" captionStart="FIGURES 9–11" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 9–11. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">9–10</figureCitation>
,
<figureCitation id="137552C3FFF85008FE73FAC0FDCB4155" box="[455,526,1292,1318]" captionStart="FIGURES 12–19" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 12–19. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">13–14</figureCitation>
) in anterodorsal view almost triangular, broadest at eyes, HL
<quantity id="4CB6E3A3FFF85008FB0AFAC0FAA64154" box="[1214,1379,1292,1319]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="8" pageNumber="9" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.30–0.33 mm</quantity>
, HW
<quantity id="4CB6E3A3FFF85008FF23FAFCFEF94138" box="[151,316,1328,1355]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="8" pageNumber="9" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.28–0.30 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFF85008FDF1FAFCFD434139" box="[581,646,1328,1354]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
) subequal to length of eye; vertex and frons confluent, together weakly convex and slightly elongate; vertex subconical, with its posterior margin nearly triangular; each eye moderately large and weakly convex, with small posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; posterior region of frons and entire vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFF85008FCA1FA0CFC9841A9" box="[789,861,1472,1498]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
,
<figureCitation id="137552C3FFF85008FCDCFA0CFC4041A8" box="[872,901,1472,1499]" captionStart="FIGURES 12–19" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 12–19. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="8" pageNumber="9">15</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFF85008FF65FA28FEB3418C" box="[209,374,1508,1535]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="8" pageNumber="9" unit="mm" value="0.665" valueMax="0.68" valueMin="0.65">0.65–0.68 mm</quantity>
, antennomeres 1 and 2 each weakly elongate, 3–10 each about as long as broad or indistinctly transverse; 11 shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F9E1FCE842A5" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
Pronotum (
<figureCitation id="137552C3FFF85008FEFCF9E0FE4B4235" box="[328,398,1580,1606]" captionStart="FIGURES 9–11" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 9–11. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
) subtrapezoidal, broadest at base; PL
<quantity id="4CB6E3A3FFF85008FCF3F9E0FC2A4234" box="[839,1007,1580,1607]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.3" metricValueMin="4.1" pageId="8" pageNumber="9" unit="mm" value="0.42" valueMax="0.43" valueMin="0.41">0.41–0.43 mm</quantity>
, PW
<quantity id="4CB6E3A3FFF85008FB84F9E0FB1C4234" box="[1072,1241,1580,1607]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="8" pageNumber="9" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.38–0.43 mm</quantity>
. Anterior margin weakly arcuate, lateral margins weakly convex and strongly diverging posteriorly; posterior corners blunt; posterior margin indistinctly convex. Base with two pairs of round and deep pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F92CFC414331" blockId="8.[151,1437,751,1931]" pageId="8" pageNumber="9">
Elytra (
<figureCitation id="137552C3FFF85008FEA9F92CFE9A4289" box="[285,351,1760,1786]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
) together oval, broadest slightly in front of middle; EL
<quantity id="4CB6E3A3FFF85008FC51F92CFB494288" box="[997,1164,1760,1787]" metricMagnitude="-4" metricUnit="m" metricValue="9.05" metricValueMax="9.3" metricValueMin="8.8" pageId="8" pageNumber="9" unit="mm" value="0.905" valueMax="0.93" valueMin="0.88">0.88–0.93 mm</quantity>
, EW
<quantity id="4CB6E3A3FFF85008FB79F92CFAB14288" box="[1229,1396,1760,1787]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.0" metricValueMin="6.8" pageId="8" pageNumber="9" unit="mm" value="0.69" valueMax="0.7" valueMin="0.68">0.68–0.70 mm</quantity>
, EI 1.25–1.32; humeral calli weakly elevated, basal elytral foveae (
<figureCitation id="137552C3FFF85008FCE7F8C8FC50436D" box="[851,917,1796,1822]" captionStart="FIGURES 9–11" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 9–11. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
) vestigial, barely discernible even under SEM. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F880FB934315" blockId="8.[151,1437,751,1931]" box="[199,1110,1868,1895]" pageId="8" pageNumber="9">
Metaventrite (
<figureCitation id="137552C3FFF85008FED2F880FE704315" box="[358,437,1868,1894]" captionStart="FIGURES 9–11" captionStartId="10.[151,264,705,729]" captionTargetBox="[183,1404,181,681]" captionTargetId="figure-28@10.[183,1404,181,681]" captionTargetPageId="10" captionText="FIGURES 9–11. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
) with setae similar to those on elytra but nearly impunctate.
</paragraph>
<paragraph id="8BF14E46FFF85008FF73F8BDFAAD43F9" blockId="8.[151,1437,751,1931]" box="[199,1384,1904,1931]" pageId="8" pageNumber="9">Legs moderately long, slender, unmodified except slightly recurved tibiae (mesotibiae most distinctly so).</paragraph>
<caption id="DF311ECEFFF95009FF23FAB1FF3041AE" ID-DOI="http://doi.org/10.5281/zenodo.13745916" ID-Zenodo-Dep="13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,264,1405,1429]" targetBox="[151,1429,181,1369]" targetPageId="9" targetType="figure">
<paragraph id="8BF14E46FFF95009FF23FAB1FF3041AE" blockId="9.[151,1437,1405,1502]" pageId="9" pageNumber="10">
<emphasis id="B93A9254FFF95009FF23FAB1FEFE41E6" bold="true" box="[151,315,1405,1429]" pageId="9" pageNumber="10">FIGURES 1–8.</emphasis>
<taxonomicName id="4C4E35C5FFF95009FEF6FAB1FD2441E6" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[322,737,1405,1429]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFF95009FEF6FAB1FD4B41E6" box="[322,654,1405,1429]" italics="true" pageId="9" pageNumber="10">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFF95009FBF3FAB1FB9141E6" bold="true" box="[1095,1108,1405,1429]" pageId="9" pageNumber="10">1</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFF95009FA34FAB1FA4841E6" bold="true" box="[1408,1421,1405,1429]" pageId="9" pageNumber="10">2</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFF95009FCD4FA6DFCA841CA" bold="true" box="[864,877,1441,1465]" pageId="9" pageNumber="10">3</emphasis>
); antenna (
<emphasis id="B93A9254FFF95009FC55FA6DFC2B41CA" bold="true" box="[993,1006,1441,1465]" pageId="9" pageNumber="10">4</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFF95009FB6DFA6DFAC541CA" bold="true" box="[1241,1280,1441,1465]" pageId="9" pageNumber="10">5, 7</emphasis>
) and lateral (
<emphasis id="B93A9254FFF95009FA3DFA6DFF6141AE" bold="true" pageId="9" pageNumber="10">6, 8</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFF95009FF73F9E3FE2F428E" blockId="9.[151,1437,1583,1933]" pageId="9" pageNumber="10">
Aedeagus (
<figureCitation id="137552C3FFF95009FEF2F9E3FE684239" box="[326,429,1583,1610]" captionStart="FIGURES 1–8" captionStartId="9.[151,264,1405,1429]" captionTargetBox="[151,1429,181,1369]" captionTargetId="figure-16@9.[151,1435,181,1381]" captionTargetPageId="9" captionText="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.13745916" httpUri="https://zenodo.org/record/13745916/files/figure.png" pageId="9" pageNumber="10">Figs 5–8</figureCitation>
,
<figureCitation id="137552C3FFF95009FE0FF9E3FDC4423A" box="[443,513,1583,1609]" captionStart="FIGURES 12–19" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 12–19. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="9" pageNumber="10">16–19</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFF95009FD25F9E3FD3F4239" box="[657,762,1583,1610]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="mm" value="0.25">0.25 mm</quantity>
; in ventral view broadest near base and narrowing towards truncate apex; in lateral view apical region of dorsal wall strongly and abruptly bent dorsally; endophallus with symmetrical and elongate distolateral sclerites and basilateral sclerites and with asymmetrical weakly sclerotized median structures, distolateral sclerites broadly separated and each with rounded apex; parameres slender and their apices only slightly projecting beyond aedeagal apex, each paramere with a row of sparse and short setae occupying slightly more than distal third.
</paragraph>
<paragraph id="8BF14E46FFF95009FF73F8CBFD894319" blockId="9.[151,1437,1583,1933]" pageId="9" pageNumber="10">
Female. Externally similar to male, with indistinctly shorter antennae in relation to body length. BL
<quantity id="4CB6E3A3FFF95009FA98F8CBFF034335" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.7" metricValueMin="1.58" pageId="9" pageNumber="10" unit="mm" value="1.64" valueMax="1.7" valueMin="1.58">1.58–1.70 mm</quantity>
; HL
<quantity id="4CB6E3A3FFF95009FF4AF8E0FE614335" box="[254,420,1836,1862]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="9" pageNumber="10" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.30–0.33 mm</quantity>
, HW
<quantity id="4CB6E3A3FFF95009FE53F8E0FD884335" box="[487,589,1836,1862]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="mm" value="0.3">0.30 mm</quantity>
, AnL
<quantity id="4CB6E3A3FFF95009FD26F8E7FCFC4335" box="[658,825,1835,1862]" metricMagnitude="-4" metricUnit="m" metricValue="5.9" metricValueMax="6.3" metricValueMin="5.5" pageId="9" pageNumber="10" unit="mm" value="0.59" valueMax="0.63" valueMin="0.55">0.55–0.63 mm</quantity>
; PL
<quantity id="4CB6E3A3FFF95009FCD9F8E7FBD64335" box="[877,1043,1835,1862]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="9" pageNumber="10" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
, PW
<quantity id="4CB6E3A3FFF95009FBE4F8E0FB324335" box="[1104,1271,1836,1862]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="9" pageNumber="10" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.38–0.43 mm</quantity>
; EL
<quantity id="4CB6E3A3FFF95009FA98F8E7FF004319" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="9" pageNumber="10" unit="mm" value="0.8999999999999999" valueMax="0.95" valueMin="0.85">0.85–0.95 mm</quantity>
, EW
<quantity id="4CB6E3A3FFF95009FEB7F883FE6D4319" box="[259,424,1871,1898]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.3" metricValueMin="6.8" pageId="9" pageNumber="10" unit="mm" value="0.705" valueMax="0.73" valueMin="0.68">0.68–0.73 mm</quantity>
, EI 1.25–1.31.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFF95009FF73F8BFFCAE43FE" box="[199,875,1907,1933]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BF14E46FFF95009FF73F8BFFCAE43FE" blockId="9.[151,1437,1583,1933]" box="[199,875,1907,1933]" pageId="9" pageNumber="10">
<emphasis id="B93A9254FFF95009FF73F8BFFEA443FE" bold="true" box="[199,353,1907,1933]" pageId="9" pageNumber="10">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFF95009FE75F8BFFD8D43FE" box="[449,584,1907,1933]" name="Madagascar" pageId="9" pageNumber="10">Madagascar</collectingCountry>
(Vakinankaratra Region).
</paragraph>
</subSubSection>
<caption id="DF311ECEFFFA500AFF23FD0DFED04719" ID-DOI="http://doi.org/10.5281/zenodo.13745921" ID-Zenodo-Dep="13745921" httpUri="https://zenodo.org/record/13745921/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,705,729]" targetBox="[183,1404,181,681]" targetPageId="10" targetType="figure">
<paragraph id="8BF14E46FFFA500AFF23FD0DFED04719" blockId="10.[151,1436,705,874]" pageId="10" pageNumber="11">
<emphasis id="B93A9254FFFA500AFF23FD0DFE8346AA" bold="true" box="[151,326,705,729]" pageId="10" pageNumber="11">FIGURES 9–11.</emphasis>
<taxonomicName id="4C4E35C5FFFA500AFEFFFD0EFD2C46A9" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[331,745,706,730]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFA500AFEFFFD0EFD5246A9" box="[331,663,706,730]" italics="true" pageId="10" pageNumber="11">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
, holotype. Head, prothorax and elytral base in dorsal view (
<emphasis id="B93A9254FFFA500AFAF0FD0DFA9446AA" bold="true" box="[1348,1361,705,729]" pageId="10" pageNumber="11">9</emphasis>
); head, prothorax and anterior region of mesothorax in ventral view (
<emphasis id="B93A9254FFFA500AFD48FD29FCD0468E" bold="true" box="[764,789,741,765]" pageId="10" pageNumber="11">10</emphasis>
); pterothorax in ventral view (
<emphasis id="B93A9254FFFA500AFBF3FD29FB9A468E" bold="true" box="[1095,1119,741,765]" pageId="10" pageNumber="11">11</emphasis>
). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum.
</paragraph>
</caption>
<caption id="DF311ECEFFFA500AFF23F855FC8B438A" ID-DOI="http://doi.org/10.5281/zenodo.13745923" ID-Zenodo-Dep="13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1945,1969]" targetBox="[199,1390,902,1907]" targetPageId="10" targetType="figure">
<paragraph id="8BF14E46FFFA500AFF23F855FC8B438A" blockId="10.[151,1436,1945,2042]" pageId="10" pageNumber="11">
<emphasis id="B93A9254FFFA500AFF23F855FE9243C2" bold="true" box="[151,343,1945,1969]" pageId="10" pageNumber="11">FIGURES 12–19.</emphasis>
<taxonomicName id="4C4E35C5FFFA500AFED4F855FCC143C1" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[352,772,1945,1970]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFA500AFED4F855FD6A43C2" box="[352,687,1945,1969]" italics="true" pageId="10" pageNumber="11">Anhoraeomorphus ankaratranus</emphasis>
(Franz)
</taxonomicName>
, holotype of
<taxonomicName id="4C4E35C5FFFA500AFC25F855FA4543C2" authority="Franz. Habitus" authorityName="Franz. Habitus" box="[913,1408,1945,1969]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFFA500AFC25F855FB1A43C2" box="[913,1247,1945,1969]" italics="true" pageId="10" pageNumber="11">Horaeomorphus manjakotompoi</emphasis>
Franz. Habitus
</taxonomicName>
in dorsal view (
<emphasis id="B93A9254FFFA500AFEABF871FEFD43A6" bold="true" box="[287,312,1981,2005]" pageId="10" pageNumber="11">12</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFFA500AFDDFF871FD4143A6" bold="true" box="[619,644,1981,2005]" pageId="10" pageNumber="11">13</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFFA500AFAC7F871FA4943A6" bold="true" box="[1395,1420,1981,2005]" pageId="10" pageNumber="11">14</emphasis>
); antenna (
<emphasis id="B93A9254FFFA500AFF41F82DFECB438A" bold="true" box="[245,270,2017,2041]" pageId="10" pageNumber="11">15</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFFA500AFE42F82DFDF0438A" bold="true" box="[502,565,2017,2041]" pageId="10" pageNumber="11">16, 18</emphasis>
) and lateral (
<emphasis id="B93A9254FFFA500AFD09F82DFD39438A" bold="true" box="[701,764,2017,2041]" pageId="10" pageNumber="11">17, 19</emphasis>
) views.
</paragraph>
</caption>
<subSubSection id="C3541DCDFFFB500BFF73FF5BFCA846DA" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BF14E46FFFB500BFF73FF5BFC2C4532" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="12">
<emphasis id="B93A9254FFFB500BFF73FF5BFEFE44C2" bold="true" box="[199,315,151,177]" pageId="11" pageNumber="12">Remarks.</emphasis>
The
<typeStatus id="54F5F0E4FFFB500BFEC7FF5BFE6644C2" box="[371,419,151,177]" pageId="11" pageNumber="12">type</typeStatus>
specimens of
<taxonomicName id="4C4E35C5FFFB500BFDF6FF5BFD3E44C2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[578,763,151,177]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFDF6FF5BFD3E44C2" box="[578,763,151,177]" italics="true" pageId="11" pageNumber="12">H. ankaratranus</emphasis>
</taxonomicName>
(
<figureCitation id="137552C3FFFB500BFCBDFF5BFCBC44C2" box="[777,889,151,177]" captionStart-0="FIGURES 1–8" captionStart-1="FIGURES 9–11" captionStartId-0="9.[151,264,1405,1429]" captionStartId-1="10.[151,264,705,729]" captionTargetBox-0="[151,1429,181,1369]" captionTargetBox-1="[183,1404,181,681]" captionTargetId-0="figure-16@9.[151,1435,181,1381]" captionTargetId-1="figure-28@10.[183,1404,181,681]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 1–8. Anhoraeomorphus ankaratranus (Franz), holotype. Habitus in dorsal view (1); head in anterodorsal view (2); head and prothorax in lateral view (only bristles on head are shown) (3); antenna (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." captionText-1="FIGURES 9–11. Anhoraeomorphus ankaratranus (Franz), holotype. Head, prothorax and elytral base in dorsal view (9); head, prothorax and anterior region of mesothorax in ventral view (10); pterothorax in ventral view (11). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; gp, gular plate; gs, gular suture; hr, hypomeral ridge; hyr, hypomeral ridge; msvp, mesoventral intermesocoxal process; mtvp, metaventral process; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum." figureDoi-0="http://doi.org/10.5281/zenodo.13745916" figureDoi-1="http://doi.org/10.5281/zenodo.13745921" httpUri-0="https://zenodo.org/record/13745916/files/figure.png" httpUri-1="https://zenodo.org/record/13745921/files/figure.png" pageId="11" pageNumber="12">Figs 1–11</figureCitation>
) and
<taxonomicName id="4C4E35C5FFFB500BFC03FF5BFB4344C2" box="[951,1158,151,177]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFFB500BFC03FF5BFB4344C2" box="[951,1158,151,177]" italics="true" pageId="11" pageNumber="12">H. manjakotompoi</emphasis>
</taxonomicName>
(
<figureCitation id="137552C3FFFB500BFB21FF5BFAD144C2" box="[1173,1300,151,177]" captionStart="FIGURES 12–19" captionStartId="10.[151,264,1945,1969]" captionTargetBox="[199,1390,902,1907]" captionTargetId="figure-107@10.[189,1397,898,1922]" captionTargetPageId="10" captionText="FIGURES 12–19. Anhoraeomorphus ankaratranus (Franz), holotype of Horaeomorphus manjakotompoi Franz. Habitus in dorsal view (12); head in anterodorsal view (13); head and prothorax in lateral view (only bristles on head are shown) (14); antenna (15); aedeagus in ventral (16, 18) and lateral (17, 19) views." figureDoi="http://doi.org/10.5281/zenodo.13745923" httpUri="https://zenodo.org/record/13745923/files/figure.png" pageId="11" pageNumber="12">Figs 12–19</figureCitation>
) come from the same area of the Ankaratra volcano range (including specimens of
<taxonomicName id="4C4E35C5FFFB500BFC29FF77FB9044A6" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[925,1109,187,213]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFC29FF77FB9044A6" box="[925,1109,187,213]" italics="true" pageId="11" pageNumber="12">H. ankaratranus</emphasis>
</taxonomicName>
collected in Manjakotompo). They differ only in the pigmentation (
<taxonomicName id="4C4E35C5FFFB500BFDF8FF13FCCD448A" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[588,776,223,249]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFDF8FF13FCCD448A" box="[588,776,223,249]" italics="true" pageId="11" pageNumber="12">H. ankaratranus</emphasis>
</taxonomicName>
slightly darker) and the preservation of setae and bristles on the head (in
<taxonomicName id="4C4E35C5FFFB500BFEF9FECFFDDA456E" box="[333,543,259,285]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="manjakotompoi">
<emphasis id="B93A9254FFFB500BFEF9FECFFDDA456E" box="[333,543,259,285]" italics="true" pageId="11" pageNumber="12">H. manjakotompoi</emphasis>
</taxonomicName>
most of them are broken off). The aedeagi have the same shapes and internal structures, and there is no doubt that these specimens belong to one species.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73FE87FDC945DE" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="12">
<taxonomicName id="4C4E35C5FFFB500BFF73FE87FDFC4516" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,569,331,357]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFF73FE87FDFC4516" box="[199,569,331,357]" italics="true" pageId="11" pageNumber="12">Anhoraeomorphus ankaratranus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4E35C5FFFB500BFCADFE87FC0F4516" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[793,970,331,357]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFFB500BFCADFE87FC0F4516" box="[793,970,331,357]" italics="true" pageId="11" pageNumber="12">A. sakarahanus</emphasis>
</taxonomicName>
, including external morphology and the aedeagus. These species clearly differ in the shape of the pronotum, in
<taxonomicName id="4C4E35C5FFFB500BFC63FEA3FB5545FA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[983,1168,367,393]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFC63FEA3FB5545FA" box="[983,1168,367,393]" italics="true" pageId="11" pageNumber="12">A. ankaratranus</emphasis>
</taxonomicName>
broadest at base, in
<taxonomicName id="4C4E35C5FFFB500BFA30FEA3FEE345DE" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFFB500BFA30FEA3FEE345DE" italics="true" pageId="11" pageNumber="12">A. sakarahanus</emphasis>
</taxonomicName>
near posterior third.
</paragraph>
<paragraph id="8BF14E46FFFB500BFF73FE7BFCA846DA" blockId="11.[151,1437,151,681]" pageId="11" pageNumber="12">
<specimenCount id="9D4885CFFFFB500BFF73FE7BFEA345A2" box="[199,358,439,465]" count="1" pageId="11" pageNumber="12" type="generic">One specimen</specimenCount>
of
<taxonomicName id="4C4E35C5FFFB500BFE3DFE7BFDFB45A2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[393,574,439,465]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFFB500BFE3DFE7BFDFB45A2" box="[393,574,439,465]" italics="true" pageId="11" pageNumber="12">A. ankaratranus</emphasis>
</taxonomicName>
in the type series is labeled as “Typus”, while all others as “
<typeStatus id="54F5F0E4FFFB500BFB66FE7BFA8D45A2" box="[1234,1352,439,465]" pageId="11" pageNumber="12" type="paratype">Paratypus</typeStatus>
”.
<bibRefCitation id="EFDF33B7FFFB500BFAE8FE7BFF354586" author="Franz, H." pageId="11" pageNumber="12" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
uses the terms “
<typeStatus id="54F5F0E4FFFB500BFE17FE17FDD94586" box="[419,540,475,501]" pageId="11" pageNumber="12" type="holotype">Holotypus</typeStatus>
” and “
<typeStatus id="54F5F0E4FFFB500BFDD1FE17FD124586" box="[613,727,475,501]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
” in his description, so the “Typus” is a correctly fixed
<typeStatus id="54F5F0E4FFFB500BFA87FE17FA5C4586" box="[1331,1433,475,501]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
. All 27 studied specimens (from Franz Coll. in NHMW) were collected in Ankaratra volcanic range situated ~
<quantity id="4CB6E3A3FFFB500BFACBFE32FF7E464E" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="km" value="50.0">50 km</quantity>
SW of
<collectingRegion id="498A80A4FFFB500BFEA4FDEFFE69464E" box="[272,428,547,573]" country="Madagascar" name="Antananarivo" pageId="11" pageNumber="12">Antananarivo</collectingRegion>
.
<bibRefCitation id="EFDF33B7FFFB500BFE02FDEFFD93464E" author="Franz, H." box="[438,598,547,573]" pageId="11" pageNumber="12" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
states that he examined
<specimenCount id="9D4885CFFFFB500BFCDDFDEFFBC4464E" box="[873,1025,547,573]" count="39" pageId="11" pageNumber="12" type="generic">39 specimens</specimenCount>
coming from Ankaratra, some taken in Manjakotompo, and some in Col de Manontongana. Only “Massif de l’Ankaratra” is given on all labels, without exact localities. According to
<bibRefCitation id="EFDF33B7FFFB500BFE51FDA7FD4246F6" author="Franz, H." box="[485,647,619,645]" pageId="11" pageNumber="12" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
, only a part of the
<typeStatus id="54F5F0E4FFFB500BFCEEFDA7FC0046F6" box="[858,965,619,645]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
is deposited in his collection (currently at NHMW); the depository of the remaining
<typeStatus id="54F5F0E4FFFB500BFDC6FD43FD1846DA" box="[626,733,655,681]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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