256 lines
33 KiB
XML
256 lines
33 KiB
XML
<document id="C4CD0E2546C143153E77B810F1EB39A7" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFDE502FFF23FF5BFE16434E" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus sikorai Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="48" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="47" updateTime="1726053143081" updateUser="ExternalLinkService">
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<mods:title id="57F2E8916C7528A45425674DA62EC71C">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="974C21F38B52DF85F3C9520492EEA5BA">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="1A237B2F0746CAD85F16ED6B5C6B8E76">text</mods:typeOfResource>
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<mods:title id="575839BF4F78635B1EBF45F783D5EECA">Zootaxa</mods:title>
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<mods:part id="0C2659320B709A7D83577FFF41B9FD08">
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<mods:date id="C6EA97E897D8E39FFC0507FF9F4F62C3">2024</mods:date>
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<mods:number id="B0788D624F9355CB4A17CB8C1AFAFBC7">2024-09-04</mods:number>
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<mods:number id="223602DD0DD90F11A1EC7227DB6F9616">5505</mods:number>
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<mods:url id="322E8482B50250CF0DF7BD7FC08CE8E0">http://dx.doi.org/10.11646/zootaxa.5505.1.1</mods:url>
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<mods:classification id="2B4FE64FBE03384493BEF486E936B80F">journal article</mods:classification>
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<mods:identifier id="B4025E6A6E226B14699468CBD7B4BBCF" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
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<mods:identifier id="51F1666713C1DCA4344D3B96567B62B3" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03E7FF50FFDE502FFF23FF5BFE16434E" ID-DOI="http://doi.org/10.5281/zenodo.13746077" ID-Zenodo-Dep="13746077" LSID="urn:lsid:plazi:treatment:03E7FF50FFDE502FFF23FF5BFE16434E" httpUri="http://treatment.plazi.org/id/03E7FF50FFDE502FFF23FF5BFE16434E" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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<subSubSection id="C3541DCDFFDE502EFF23FF5BFD7044C2" box="[151,693,151,178]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="8BF14E46FFDE502EFF23FF5BFD7044C2" blockId="46.[151,693,151,213]" box="[151,693,151,178]" pageId="46" pageNumber="47">
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<heading id="D0B9F92AFFDE502EFF23FF5BFD7044C2" bold="true" box="[151,693,151,178]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
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<emphasis id="B93A9254FFDE502EFF23FF5BFD7044C2" bold="true" box="[151,693,151,178]" pageId="46" pageNumber="47">
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<taxonomicName id="4C4E35C5FFDE502EFF23FF5BFDF844C2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,573,151,178]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sikorai" status="comb. nov.">
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<emphasis id="B93A9254FFDE502EFF23FF5BFE1344C1" bold="true" box="[151,470,151,178]" italics="true" pageId="46" pageNumber="47">Anhoraeomorphus sikorai</emphasis>
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(Franz)
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2092F2FFFDE502EFDFEFF5BFD7044C2" box="[586,693,151,177]" pageId="46" pageNumber="47" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3541DCDFFDE502EFF23FF77FE8644A6" box="[151,323,187,213]" pageId="46" pageNumber="47" type="description">
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<paragraph id="8BF14E46FFDE502EFF23FF77FE8644A6" blockId="46.[151,693,151,213]" box="[151,323,187,213]" pageId="46" pageNumber="47">
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(
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<figureCitation id="137552C3FFDE502EFF2BFF77FEFF44A6" box="[159,314,187,213]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 171–178</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3541DCDFFDE502EFF23FECCFD954564" box="[151,592,255,280]" pageId="46" pageNumber="47" type="reference_group">
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<paragraph id="8BF14E46FFDE502EFF23FECCFD954564" blockId="46.[151,592,255,280]" box="[151,592,255,280]" pageId="46" pageNumber="47">
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<treatmentCitationGroup id="AB5E6968FFDE502EFF23FECCFD954564" box="[151,592,255,280]" pageId="46" pageNumber="47">
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<treatmentCitation id="0AEF6857FFDE502EFF23FECCFD894564" author="Franz, H." box="[151,588,255,280]" page="162" pageId="46" pageNumber="47" year="1986">
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<taxonomicName id="4C4E35C5FFDE502EFF23FECCFD894564" ID-CoL="3MNLK" authority="Franz, 1986 b: 162" authorityName="Franz" authorityPageNumber="162" authorityYear="1986" box="[151,588,255,280]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sikorai">
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<emphasis id="B93A9254FFDE502EFF23FECCFE4F456B" box="[151,394,256,280]" italics="true" pageId="46" pageNumber="47">Horaeomorphus sikorai</emphasis>
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<bibRefCitation id="EFDF33B7FFDE502EFE24FECCFD894564" author="Franz, H." box="[400,588,255,279]" pageId="46" pageNumber="47" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 162</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3541DCDFFDE502EFF23FE8AFD05469F" pageId="46" pageNumber="47" type="materials_examined">
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<paragraph id="8BF14E46FFDE502EFF23FE8AFA8245DB" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
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<emphasis id="B93A9254FFDE502EFF23FE8AFE77452C" bold="true" box="[151,434,325,352]" pageId="46" pageNumber="47">Type material studied.</emphasis>
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<materialsCitation id="3B26441BFFDE502EFE75FE8AFECC45DB" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B93A9254FFDE502EFE75FE8AFD26452C" bold="true" box="[449,739,325,352]" pageId="46" pageNumber="47">
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<typeStatus id="54F5F0E4FFDE502EFE75FE8AFDEB4513" box="[449,558,326,352]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
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(
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<collectingCountry id="F3590ED6FFDE502EFDF0FE89FD1E452C" box="[580,731,325,351]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
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)
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</emphasis>
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:
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<specimenCount id="9D4885CFFFDE502EFD4DFE8AFCD54513" box="[761,784,326,352]" count="1" pageId="46" pageNumber="47" type="male">♂</specimenCount>
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, three labels: “Madagaskar / Sikora” [white, printed], “
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<taxonomicName id="4C4E35C5FFDE502EFF15FEA6FE9D45F7" box="[161,344,362,388]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
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/ sikorai m. / det.
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<determinerName id="7E2DA282FFDE502EFD81FEA6FD5645F7" box="[565,659,362,388]" pageId="46" pageNumber="47">H.Franz</determinerName>
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” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
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<collectionCode id="ED5FD683FFDE502EFF2BFE42FEC745DB" box="[159,258,398,424]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47" type="Museum">NHMW</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B26441BFFDE502EFEA0FE42FA8145DB" box="[276,1348,398,424]" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
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<emphasis id="B93A9254FFDE502EFEA0FE42FE5045DB" bold="true" box="[276,405,398,424]" pageId="46" pageNumber="47">
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<typeStatus id="54F5F0E4FFDE502EFEA0FE42FE5545DB" box="[276,400,398,424]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D4885CFFFDE502EFE28FE42FE1945DB" box="[412,476,398,424]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
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, same data as for holotype, but with yellow “
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<typeStatus id="54F5F0E4FFDE502EFC61FE42FBBD45DB" box="[981,1144,398,424]" pageId="46" pageNumber="47" type="paratype">PARATYPUS</typeStatus>
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” labels (
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<collectionCode id="ED5FD683FFDE502EFB6DFE42FAF945DB" box="[1241,1340,398,424]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="46" pageNumber="47" type="Museum">NHMW</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BF14E46FFDE502EFF73FE7DFD05469F" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
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<emphasis id="B93A9254FFDE502EFF73FE7DFE6345B8" bold="true" box="[199,422,433,459]" pageId="46" pageNumber="47">Revised diagnosis.</emphasis>
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Body (
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<figureCitation id="137552C3FFDE502EFDB7FE7EFDA245BF" box="[515,615,434,460]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 171</figureCitation>
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) slender; head (
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<figureCitation id="137552C3FFDE502EFC9DFE7EFC0C45BF" box="[809,969,434,460]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 172–173</figureCitation>
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) round, with vertex posteriorly broadly rounded; pronotum (
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<figureCitation id="137552C3FFDE502EFE37FE1AFE204583" box="[387,485,470,496]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 171</figureCitation>
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) bell-shaped, broadest in anterior third, with one pair of large transverse antebasal pits; antennomere 3 (
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<figureCitation id="137552C3FFDE502EFECBFE36FE184667" box="[383,477,506,532]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
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) as long as broad, 4–5 each slightly elongate, 6 as long as broad, 7–10 each indistinctly transverse; profemora similarly slender as meso- and metafemora; metatibiae in male modified, but only slightly strongly broadening to distal third and then narrowing to apex; aedeagus (
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<figureCitation id="137552C3FFDE502EFC6AFD8EFBBE462E" box="[990,1147,578,605]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 175–178</figureCitation>
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) in ventral view broadest near basal third, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites separated except their apices touching at middle, each strongly elongate and with distal margin concave with outer and inner subtriangular apex, parameres with several subapical setae distributed along more than half of length and increasing in length from proximal to distal ones.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3541DCDFFDE502EFF73FD39FAB24373" pageId="46" pageNumber="47" type="description">
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<paragraph id="8BF14E46FFDE502EFF73FD39FCAF4747" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
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<emphasis id="B93A9254FFDE502EFF73FD39FEB0477C" bold="true" box="[199,373,757,783]" pageId="46" pageNumber="47">Redescription.</emphasis>
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Body of male (
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<figureCitation id="137552C3FFDE502EFDABFD3AFDB84763" box="[543,637,758,784]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 171</figureCitation>
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) moderately dark brown, elytra slightly lighter than head and pronotum, tarsi and palps distinctly lighter, setae light brown; BL
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<quantity id="4CB6E3A3FFDE502EFD4AFCD6FCA04747" box="[766,869,794,821]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="46" pageNumber="47" unit="mm" value="2.15">2.15 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BF14E46FFDE502EFF73FCF2FDF44027" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
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Head (
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<figureCitation id="137552C3FFDE502EFEA0FCF2FEB0472B" box="[276,373,830,856]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 172</figureCitation>
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) in anterodorsal view round, broadest at eyes, HL
|
||
<quantity id="4CB6E3A3FFDE502EFC0BFCF2FBE2472B" box="[959,1063,830,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="46" pageNumber="47" unit="mm" value="0.38">0.38 mm</quantity>
|
||
, HW
|
||
<quantity id="4CB6E3A3FFDE502EFBD8FCF2FB10472B" box="[1132,1237,830,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="46" pageNumber="47" unit="mm" value="0.38">0.38 mm</quantity>
|
||
; temple in lateral view (
|
||
<figureCitation id="137552C3FFDE502EFF69FCAEFEF9470F" box="[221,316,866,892]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
|
||
) subequal to eye; vertex and frons confluent, together weakly convex and weakly transverse; vertex posteriorly broadly rounded; each eye large and weakly convex, shallowly emarginate posteriorly, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
|
||
<figureCitation id="137552C3FFDE502EFCCAFC02FC1B479B" box="[894,990,974,1000]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
|
||
) slender but short, slightly shorter than half BL, AnL
|
||
<quantity id="4CB6E3A3FFDE502EFE81FC3EFE59407F" box="[309,412,1010,1036]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="47" unit="mm" value="0.9">0.90 mm</quantity>
|
||
, antennomeres 1–2 each elongate, 3 as long as broad, 4–5 each slightly elongate, 6 as long as broad, 7–10 each indistinctly transverse, 11 distinctly shorter than 9 and 10 combined, indistinctly narrower than 10, about 1.7 times as long as broad.
|
||
</paragraph>
|
||
<paragraph id="8BF14E46FFDE502EFF73FB92FF35417B" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
|
||
Pronotum bell-shaped, broadest in anterior third; PL
|
||
<quantity id="4CB6E3A3FFDE502EFCA4FB92FCB0400B" box="[784,885,1118,1145]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="mm" value="0.5">0.50 mm</quantity>
|
||
, PW
|
||
<quantity id="4CB6E3A3FFDE502EFC04FB92FBD3400B" box="[944,1046,1118,1145]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="mm" value="0.5">0.50 mm</quantity>
|
||
. Anterior margin strongly rounded, lateral margins rounded in anterior half, sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large transversely oval pits, and with short but distinct sublateral carinae. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
|
||
</paragraph>
|
||
<paragraph id="8BF14E46FFDE502EFF73FADEFEAE4107" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
|
||
Elytra together oval, broadest distinctly in front of middle; EL
|
||
<quantity id="4CB6E3A3FFDE502EFC39FADEFC31415F" box="[909,1012,1298,1324]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="46" pageNumber="47" unit="mm" value="1.28">1.28 mm</quantity>
|
||
, EW
|
||
<quantity id="4CB6E3A3FFDE502EFB87FADEFB5F415F" box="[1075,1178,1298,1324]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="46" pageNumber="47" unit="mm" value="0.8">0.80 mm</quantity>
|
||
, EI 1.59; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
|
||
</paragraph>
|
||
<paragraph id="8BF14E46FFDE502EFF73FAB2FE8C41CF" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">Legs moderately long, slender, mesotibiae distinctly curved, metatibiae slightly broadening to distal third and then narrowing.</paragraph>
|
||
<paragraph id="8BF14E46FFDE502EFF73FA0AFAFE42CB" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
|
||
Aedeagus (
|
||
<figureCitation id="137552C3FFDE502EFEF0FA0AFE1A4192" box="[324,479,1478,1505]" captionStart="FIGURES 171–178" captionStartId="47.[151,264,1540,1564]" captionTargetBox="[151,1407,181,1508]" captionTargetId="figure-16@47.[151,1435,181,1516]" captionTargetPageId="47" captionText="FIGURES 171–178. Anhoraeomorphus sikorai (Franz), holotype. Habitus in dorsal view (171); head in anterodorsal view (172); head and prothorax in lateral view (only bristles on head are shown) (173); antenna (174); aedeagus in ventral (175, 177) and lateral (176, 178) views." figureDoi="http://doi.org/10.5281/zenodo.13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="46" pageNumber="47">Figs 175–178</figureCitation>
|
||
) stout, AeL
|
||
<quantity id="4CB6E3A3FFDE502EFDD2FA0AFD084193" box="[614,717,1478,1505]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="46" pageNumber="47" unit="mm" value="0.35">0.35 mm</quantity>
|
||
; in ventral view drop-shaped, distally rapidly narrowing towards broadly subtrapezoidal apex with rounded distal margin; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with asymmetrical and long distolateral sclerites and with strongly asymmetrical assemblage of median proximal sclerotized structures, distolateral sclerites separated except for their tips touching at middle, each with concave distal margin and outer and inner subtriangular apices; parameres strongly broadening distally, their apices slightly projecting beyond apex of median lobe, each paramere with subapical group of setae distributed along more than distal half of outer lateral margin, setae increasing in length from proximal to distal ones.
|
||
</paragraph>
|
||
<paragraph id="8BF14E46FFDE502EFF73F90EFAB24373" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
|
||
Female. Externally similar to male, but with slenderer hind tibiae. BL
|
||
<quantity id="4CB6E3A3FFDE502EFC6CF90EFBB842AF" box="[984,1149,1730,1757]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" metricValueMax="2.23" metricValueMin="2.15" pageId="46" pageNumber="47" unit="mm" value="2.19" valueMax="2.23" valueMin="2.15">2.15–2.23 mm</quantity>
|
||
; HL 0.40–0.43–
|
||
<quantity id="4CB6E3A3FFDE502EFA87F90EFA5D42AF" box="[1331,1432,1730,1756]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="46" pageNumber="47" unit="mm" value="0.43">0.43 mm</quantity>
|
||
, HW
|
||
<quantity id="4CB6E3A3FFDE502EFF78F92AFEF74373" box="[204,306,1766,1792]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="46" pageNumber="47" unit="mm" value="0.43">0.43 mm</quantity>
|
||
, AnL
|
||
<quantity id="4CB6E3A3FFDE502EFEC2F92AFDD94373" box="[374,540,1766,1792]" metricMagnitude="-4" metricUnit="m" metricValue="9.55" metricValueMax="10.3" metricValueMin="8.8" pageId="46" pageNumber="47" unit="mm" value="0.955" valueMax="1.03" valueMin="0.88">0.88–1.03 mm</quantity>
|
||
; PL
|
||
<quantity id="4CB6E3A3FFDE502EFDFAF92AFD364373" box="[590,755,1766,1793]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="46" pageNumber="47" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
|
||
, PW
|
||
<quantity id="4CB6E3A3FFDE502EFC9BF92AFC104373" box="[815,981,1766,1793]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="46" pageNumber="47" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
|
||
; EL
|
||
<quantity id="4CB6E3A3FFDE502EFBBCF92AFBAB4373" box="[1032,1134,1766,1793]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="46" pageNumber="47" unit="mm" value="1.25">1.25 mm</quantity>
|
||
, EW
|
||
<quantity id="4CB6E3A3FFDE502EFB18F92AFAD74373" box="[1196,1298,1766,1793]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="mm" value="0.85">0.85 mm</quantity>
|
||
, EI 1.47.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3541DCDFFDE502EFF73F8C5FD5B4357" box="[199,670,1801,1828]" pageId="46" pageNumber="47" type="distribution">
|
||
<paragraph id="8BF14E46FFDE502EFF73F8C5FD5B4357" blockId="46.[151,1437,325,1936]" box="[199,670,1801,1828]" pageId="46" pageNumber="47">
|
||
<emphasis id="B93A9254FFDE502EFF73F8C5FEA44350" bold="true" box="[199,353,1801,1827]" pageId="46" pageNumber="47">Distribution.</emphasis>
|
||
<collectingCountry id="F3590ED6FFDE502EFED3F8C6FE2B4357" box="[359,494,1802,1828]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
|
||
(see Remarks).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3541DCDFFDE502FFF73F8E2FE16434E" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="discussion">
|
||
<paragraph id="8BF14E46FFDE502EFF73F8E2FB2043E3" blockId="46.[151,1437,325,1936]" pageId="46" pageNumber="47">
|
||
<emphasis id="B93A9254FFDE502EFF73F8E2FEFE433B" bold="true" box="[199,315,1838,1864]" pageId="46" pageNumber="47">Remarks.</emphasis>
|
||
<taxonomicName id="4C4E35C5FFDE502EFEF2F8E2FDB5433B" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[326,624,1838,1864]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="sikorai">
|
||
<emphasis id="B93A9254FFDE502EFEF2F8E2FDB5433B" box="[326,624,1838,1864]" italics="true" pageId="46" pageNumber="47">Anhoraeomorphus sikorai</emphasis>
|
||
</taxonomicName>
|
||
is unremarkable and the aedeagus must be examined to identify this species. The large asymmetrical endophallic distolateral sclerites touching distally at middle, each with concave distal margin with inner and outer subtriangular process are unique among
|
||
<taxonomicName id="4C4E35C5FFDE502EFC6BF8BAFB2043E3" box="[991,1253,1910,1936]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B93A9254FFDE502EFC6BF8BAFB6A43E3" box="[991,1199,1910,1936]" italics="true" pageId="46" pageNumber="47">Anhoraeomorphus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<caption id="DF311ECEFFDF502FFF23F9C8FE7F4217" ID-DOI="http://doi.org/10.5281/zenodo.13745981" ID-Zenodo-Dep="13745981" httpUri="https://zenodo.org/record/13745981/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,264,1540,1564]" targetBox="[151,1407,181,1508]" targetPageId="47" targetType="figure">
|
||
<paragraph id="8BF14E46FFDF502FFF23F9C8FE7F4217" blockId="47.[151,1436,1540,1637]" pageId="47" pageNumber="48">
|
||
<emphasis id="B93A9254FFDF502FFF23F9C8FEB5426F" bold="true" box="[151,368,1540,1564]" pageId="47" pageNumber="48">FIGURES 171–178.</emphasis>
|
||
<taxonomicName id="4C4E35C5FFDF502FFECDF9C8FD1E426F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[377,731,1540,1564]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="sikorai">
|
||
<emphasis id="B93A9254FFDF502FFECDF9C8FD40426F" box="[377,645,1540,1564]" italics="true" pageId="47" pageNumber="48">Anhoraeomorphus sikorai</emphasis>
|
||
(Franz)
|
||
</taxonomicName>
|
||
, holotype. Habitus in dorsal view (
|
||
<emphasis id="B93A9254FFDF502FFBE4F9C8FBB3426F" bold="true" box="[1104,1142,1540,1564]" pageId="47" pageNumber="48">171</emphasis>
|
||
); head in anterodorsal view (
|
||
<emphasis id="B93A9254FFDF502FFF14F9E4FF034233" bold="true" box="[160,198,1576,1600]" pageId="47" pageNumber="48">172</emphasis>
|
||
); head and prothorax in lateral view (only bristles on head are shown) (
|
||
<emphasis id="B93A9254FFDF502FFC23F9E4FC784233" bold="true" box="[919,957,1576,1600]" pageId="47" pageNumber="48">173</emphasis>
|
||
); antenna (
|
||
<emphasis id="B93A9254FFDF502FFB9BF9E4FB904233" bold="true" box="[1071,1109,1576,1600]" pageId="47" pageNumber="48">174</emphasis>
|
||
); aedeagus in ventral (
|
||
<emphasis id="B93A9254FFDF502FFA88F9E4FA514233" bold="true" box="[1340,1428,1576,1600]" pageId="47" pageNumber="48">175, 177</emphasis>
|
||
) and lateral (
|
||
<emphasis id="B93A9254FFDF502FFEA4F980FEAC4217" bold="true" box="[272,361,1612,1636]" pageId="47" pageNumber="48">176, 178</emphasis>
|
||
) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF14E46FFDF502FFF73F97AFE16434E" blockId="47.[151,1437,1718,1853]" pageId="47" pageNumber="48">
|
||
The specimens used by Franz to describe
|
||
<taxonomicName id="4C4E35C5FFDF502FFD1DF97BFCDD42A3" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[681,792,1718,1745]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="sikorai">
|
||
<emphasis id="B93A9254FFDF502FFD1DF97BFCDD42A3" box="[681,792,1718,1745]" italics="true" pageId="47" pageNumber="48">A. sikorai</emphasis>
|
||
</taxonomicName>
|
||
were deposited at NHMW before he worked in Vienna. The exact collecting site remains unknown. These specimens were collected by Franz Sikora (1863–1902), an Austrian traveler and collector of various biological specimens active in
|
||
<collectingCountry id="F3590ED6FFDF502FFC68F932FBA6436B" box="[988,1123,1790,1816]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3590ED6FFDF502FFB2BF932FB3B436B" box="[1183,1278,1790,1816]" name="Réunion" pageId="47" pageNumber="48">Réunion</collectingCountry>
|
||
during 1890s (
|
||
<emphasis id="B93A9254FFDF502FFF15F8EFFF0C434F" box="[161,201,1827,1852]" italics="true" pageId="47" pageNumber="48">e.g.</emphasis>
|
||
,
|
||
<bibRefCitation id="EFDF33B7FFDF502FFF63F8EEFE0A434E" author="Bolotov, I. N. & Eliseeva, T. A. & Tsiplenkina, I. G. & Gofarov, M. Y. & Kondakov, A. V." box="[215,463,1826,1853]" pageId="47" pageNumber="48" pagination="56 - 67" refId="ref52681" refString="Bolotov, I. N., Eliseeva, T. A., Tsiplenkina, I. G., Gofarov, M. Y. & Kondakov, A. V. (2023) Long forgotten record of a freshwater leech on Madagascar indicates a host-associated dispersal event from continental Africa. Ecologica Montenegrina, 61, 56 - 67. https: // doi. org / 10.37828 / em. 2023.61.7" type="journal article" year="2023">
|
||
Bolotov
|
||
<emphasis id="B93A9254FFDF502FFE8CF8EFFEB5434F" box="[312,368,1826,1852]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
|
||
(2023))
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |