256 lines
31 KiB
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256 lines
31 KiB
XML
<document id="4C4CA968F316278675B00FB330D3AC0B" ID-CLB-Dataset="6402" ID-DOI="10.5252/g2013n4a5" ID-GBIF-Dataset="fbfe438c-1eae-40cb-b56f-ff8167b283c2" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538200" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613149818085" checkinUser="felipe" docAuthor="Boessenecker, Robert W." docDate="2013" docId="03E7DD69FFF6867A273178CBFCED38F2" docLanguage="en" docName="Geodiversitas.35.4.815-940.pdf" docOrigin="Geodiversitas 35 (4)" docSource="http://www.bioone.org/doi/abs/10.5252/g2013n4a5" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Balaenoptera" docType="treatment" docVersion="9" lastPageNumber="854" masterDocId="FFDEA511FFD0865D27477C65FF9B3B38" masterDocTitle="A new marine vertebrate assemblage from the Late Neogene Purisima Formation in Central California, part II: Pinnipeds and Cetaceans" masterLastPageNumber="940" masterPageNumber="815" pageNumber="853" updateTime="1698929391291" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="E5B973C8871273CCD7E7729A404A1F27">
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<mods:title id="525B0078B5B871415CFC07B46EC793AC">A new marine vertebrate assemblage from the Late Neogene Purisima Formation in Central California, part II: Pinnipeds and Cetaceans</mods:title>
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<mods:name id="8DC50789EC7CEC684B38C4F965A7C314" type="personal">
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<mods:namePart id="D4DAA9228ADE9AC0C4EDC856056203CC">Boessenecker, Robert W.</mods:namePart>
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<mods:typeOfResource id="0E1EE13256788072591AB7178B08AEEC">text</mods:typeOfResource>
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<mods:title id="FEA7B2F0FBF51AB3D9BFE90376A1FAD2">Geodiversitas</mods:title>
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<mods:part id="1CDEC78C1A88EE812146C2511AA3B468">
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<mods:date id="A60494F5AE054A45242D85074C459986">2013</mods:date>
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<mods:detail id="8CA114CAAE74E553C03D9AB622CF5328" type="pubDate">
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<mods:number id="489A190E57C9817CA56C1E32CEAE26EB">2013-12-31</mods:number>
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<mods:number id="7C66BB7977D6D3F201DB55AF8B373AFC">35</mods:number>
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<mods:number id="1203133F7A399E4D772D50FA78AB97C3">4</mods:number>
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<mods:start id="20E09134D9E2DDE693B52ECE164C8ACE">815</mods:start>
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<mods:end id="8BB3B4516F48351A1A2FF34C2D05506B">940</mods:end>
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<mods:classification id="BFEFCFF8FFCF5BF5C1C29E964C818EB4">journal article</mods:classification>
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<mods:identifier id="2C98F360F1F0C2B8C19E22479AA15D2C" type="CLB-Dataset">6402</mods:identifier>
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<mods:identifier id="7BE692620784E22C170A182C2B180A0F" type="DOI">10.5252/g2013n4a5</mods:identifier>
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<mods:identifier id="15801631AA799CCC58D06B2EB4F55DC8" type="ISSN">1638-9395</mods:identifier>
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<treatment id="03E7DD69FFF6867A273178CBFCED38F2" ID-DOI="http://doi.org/10.5281/zenodo.4818510" ID-GBIF-Taxon="182400953" ID-Zenodo-Dep="4818510" LSID="urn:lsid:plazi:treatment:03E7DD69FFF6867A273178CBFCED38F2" httpUri="http://treatment.plazi.org/id/03E7DD69FFF6867A273178CBFCED38F2" lastPageId="39" lastPageNumber="854" pageId="38" pageNumber="853">
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<subSubSection id="C3543FF4FFF6867B273178CBFF503E0C" pageId="38" pageNumber="853" type="nomenclature">
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<paragraph id="8BF16C7FFFF6867B273178CBFD193FF0" blockId="38.[118,642,1197,1224]" box="[118,642,1197,1224]" pageId="38" pageNumber="853">
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<heading id="D0B9DB13FFF6867B273178CBFD193FF0" box="[118,642,1197,1224]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="853" reason="6">
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“
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<taxonomicName id="4C4E17FCFFF6867B27C678C8FE8A3FF0" ID-CoL="37H3" box="[129,273,1197,1224]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="38" pageNumber="853" phylum="Chordata" rank="genus">
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<emphasis id="B93AB06DFFF6867B27C678C8FE8A3FF0" bold="true" box="[129,273,1197,1224]" italics="true" pageId="38" pageNumber="853">Balaenoptera</emphasis>
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</taxonomicName>
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”
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<emphasis id="B93AB06DFFF6867B266478CBFEFC3FF0" bold="true" box="[291,359,1198,1224]" italics="true" pageId="38" pageNumber="853">cortesi</emphasis>
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“var.”
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<emphasis id="B93AB06DFFF6867B26F478CBFE633FF0" bold="true" box="[435,504,1198,1224]" italics="true" pageId="38" pageNumber="853">portisi</emphasis>
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Sacco, 1890
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</heading>
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</paragraph>
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<paragraph id="8BF16C7FFFF6867B27367882FF503E0C" blockId="38.[112,646,1255,1332]" pageId="38" pageNumber="853">
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REFERRED MATERIAL. — UCMP 219135, a right petrosal collected by R.W. Boessenecker from UCMP locality
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<accessionNumber id="941DF19CFFF6867B27347979FF5C3E0C" box="[115,199,1308,1332]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V99840" pageId="38" pageNumber="853" type="EnaNcbi">V99840</accessionNumber>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3543FF4FFF6867A27367928FE943A49" lastPageId="39" lastPageNumber="854" pageId="38" pageNumber="853" type="description">
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<paragraph id="8BF16C7FFFF6867B27367928FD1C3EA0" blockId="38.[111,647,1355,1432]" pageId="38" pageNumber="853">
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STRATIGRAPHIC OCCURRENCE. — Middle part of the San Gregorio section of the Purisima Formation, Early Pliocene (
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<emphasis id="B93AB06DFFF6867B273079E4FF1C3EA0" box="[119,135,1409,1432]" italics="true" pageId="38" pageNumber="853">c.</emphasis>
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5-3.35 Ma; Zanclean-Piacenzian equivalent;
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<figureCitation id="137570FAFFF6867B250679E5FDE13EA0" box="[577,634,1408,1432]" captionStart="FIG" captionStartId="6.[113,124,1221,1239]" captionTargetBox="[761,1205,215,1066]" captionTargetId="figure-149@6.[759,1205,592,972]" captionTargetPageId="6" captionText="FIG. 2. — Stratigraphy of the Purisima Formation: A, Stratigraphic column of the San Gregorio section of the Purisima Formation (in meters), showing stratigraphic position of UCMP vertebrate localities, dated ash beds, diatom floras, and species of Carcharodon; thickness measurements start over above missing section in uppermost part due lack of overlapping strata; epoch and stage boundaries marked as "?" are estimated; modified from Boessenecker (2011a); Pliocene-Pleistocene boundary after Hilgen et al. (2012); B, photograph of 3-4 m thick ash within Purisima Formation at mouth of San Gregorio Creek correlated with 2.5 Ma Ishi Tuff (Powell et al.2007); C, trace fossils in pervasively bioturbated sandstone,Purisima Formation,mouth of San Gregorio Creek; D, swaley cross-stratified sandstone, mouth of San Gregorio Creek. Abbreviations: Qt, Pleistocene terrace deposits; Tp, Purisima Formation; C, claystone; St, siltstone; Ss, sandstone. Scale bars: A, 2 m; B, C, 10 cm." figureDoi="http://doi.org/10.5281/zenodo.4538204" httpUri="https://zenodo.org/record/4538204/files/figure.png" pageId="38" pageNumber="853">Fig. 2</figureCitation>
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).
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</paragraph>
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<paragraph id="8BF16C7FFFF6867B273679A9FE993EDC" blockId="38.[113,647,1484,1700]" box="[113,258,1484,1510]" pageId="38" pageNumber="853">DESCRIPTION</paragraph>
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<paragraph id="8BF16C7FFFF6867A2736798EFE943A49" blockId="38.[113,647,1484,1700]" lastBlockId="39.[113,649,216,369]" lastPageId="39" lastPageNumber="854" pageId="38" pageNumber="853">
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Ŋe pars cochlearis of this large petrosal (
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<tableCitation id="C6CC59C4FFF6867B256C798EFDE63D3D" box="[555,637,1515,1541]" captionStart="TABLE" captionStartId="38.[113,124,219,237]" captionTargetPageId="38" captionText="TABLE 7. — Measurements (in cm) of mysticete petrosals. Measurements given to nearest tenth of a millimeter. Total length refers to length of the petrosal body, and does not include the posterior process (preserved only in UCMP 219114 among these specimens)." httpUri="http://table.plazi.org/id/DF313CF7FFF6867B27367CBEFB2D3A3C" pageId="38" pageNumber="853" tableUuid="DF313CF7FFF6867B27367CBEFB2D3A3C">Table 7</tableCitation>
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) is dorsally elongate (
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<figureCitation id="137570FAFFF6867B26167A6FFE2F3D1C" box="[337,436,1546,1572]" captionStart="FIG" captionStartId="40.[113,124,1214,1232]" captionTargetBox="[120,1203,215,1176]" captionTargetId="figure-213@40.[634,1100,570,943]" captionTargetPageId="40" captionText="FIG. 19. — Right petrosal of “Balaenoptera” cortesi “var.” portisi (UCMP 219135); A, medial view; B, lateral view; C, ventral view; D, dorsal view. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4538249" httpUri="https://zenodo.org/record/4538249/files/figure.png" pageId="38" pageNumber="853">Fig. 19A</figureCitation>
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), with an elongate, posteroventrally directed posterior cochlear crest; adjacent to this crest is the small, circular fenestra cochleae. Ŋe groove for the tensor tympani muscle is situated anteroventral to the pars cochlearis, on the medial surface of the anterior process. Ŋe anterior process is triangular and bladelike, curves medially at its apex, and rugose along its medial margin (
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<figureCitation id="137570FAFFF6867B24437ED6FC1C39F5" box="[772,903,691,717]" captionStart="FIG" captionStartId="40.[113,124,1214,1232]" captionTargetBox="[120,1203,215,1176]" captionTargetId="figure-213@40.[634,1100,570,943]" captionTargetPageId="40" captionText="FIG. 19. — Right petrosal of “Balaenoptera” cortesi “var.” portisi (UCMP 219135); A, medial view; B, lateral view; C, ventral view; D, dorsal view. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4538249" httpUri="https://zenodo.org/record/4538249/files/figure.png" pageId="38" pageNumber="853">Fig. 19A, C</figureCitation>
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). In medial aspect, the ventral margin of the anterior process is straight and directed anterodorsally. Ŋe lateral tuberosity is oriented posteroventrally and positioned at the base of the anterior process. Ŋe anterior pedicle for the tympanic bulla is positioned just dorsal to the lateral tuberosity; dorsal to the pedicle is a shallow mallear fossa. Ŋe facial sulcus is oval and dorsal to the mallear fossa; an elongate trough is not developed as in some modern
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<taxonomicName id="4C4E17FCFFF6867B236C7FB5FB2F38D2" box="[1067,1204,976,1002]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="38" pageNumber="853" phylum="Chordata" rank="genus">
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<emphasis id="B93AB06DFFF6867B236C7FB5FB2F38D2" box="[1067,1204,976,1002]" italics="true" pageId="38" pageNumber="853">Balaenoptera</emphasis>
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</taxonomicName>
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. Ŋe oval-shaped fenestra vestibuli is medially adjacent to the facial sulcus and at the base of pars cochlearis. Ŋe neck for the posterior process is robust and quadrate in cross-section, and the small posterior pedicle is present anteromedially on the neck. Ŋe anteroposteriorly elongate, rectangular, and shallow stylomastoid fossa is positioned on the posterior surface of the pars cochlearis and base of the posterior process. Further anterior along the dorsal surface of the petrosal are the perilymphatic and endolymphatic foramina, which are situated in a common groove an separated from the stylomastoid fossa by a thin crest. Ŋe internal acoustic meatus is deep and cavernous (resulting from the extreme dorsal elongation of the pars cochlearis) with a circular canal for the vestibulocochlear nerve (= spiral cribriform tract of
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<bibRefCitation id="EFDF118EFFF6867B2485798EFB293D3D" author="MEAD J. G. & FORDYCE R. E." box="[962,1202,1515,1541]" pageId="38" pageNumber="853" pagination="1 - 248" refId="ref71975" refString="MEAD J. G. & FORDYCE R. E. 2009. - Ne therian skull: a lexicon with emphasis on the odontocetes. Smithsonian Contributions to Zoology 627: 1 - 248." type="journal article" year="2009">Mead & Fordyce 2009</bibRefCitation>
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) and an oval-shaped facial canal (
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<figureCitation id="137570FAFFF6867B23497A6FFBE13D1D" box="[1038,1146,1546,1573]" captionStart="FIG" captionStartId="40.[113,124,1214,1232]" captionTargetBox="[120,1203,215,1176]" captionTargetId="figure-213@40.[634,1100,570,943]" captionTargetPageId="40" captionText="FIG. 19. — Right petrosal of “Balaenoptera” cortesi “var.” portisi (UCMP 219135); A, medial view; B, lateral view; C, ventral view; D, dorsal view. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.4538249" httpUri="https://zenodo.org/record/4538249/files/figure.png" pageId="38" pageNumber="853">Fig. 19D</figureCitation>
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). Ŋe crista transversa is deeply recessed (
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<quantity id="4CB6C19AFFF6867B235F7A4FFBFE3D7D" box="[1048,1125,1578,1605]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="38" pageNumber="853" unit="mm" value="34.0">34mm</quantity>
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) within the internal acoustic meatus. Ŋe vestibulocochlear nerve canal is
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<quantity id="4CB6C19AFFF6867B247D7A0FFC103DBC" box="[826,907,1642,1668]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="38" pageNumber="853" unit="mm" value="43.0">43 mm</quantity>
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deep, and the facial canal is
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<quantity id="4CB6C19AFFF6867B25E27AECFD683D9C" box="[677,755,1673,1700]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="38" pageNumber="853" unit="mm" value="48.0">48 mm</quantity>
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deep. An additional unidentified foramen is present anterodorsally to the facial canal on the dorsal edge of the pars cochlearis; it is unclear whether or not this foramen is the hiatus fallopii. Ŋe lateral surface of the petrosal is rugose and gently convex.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3543FF4FFF7867A27367DF3FCED38F2" pageId="39" pageNumber="854" type="discussion">
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<paragraph id="8BF16C7FFFF7867A27367DF3FE3C3A96" blockId="39.[111,649,406,1700]" box="[113,423,406,432]" pageId="39" pageNumber="854">REMARKS AND COMPARISONS</paragraph>
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<paragraph id="8BF16C7FFFF7867A27367DD0FDE83E30" blockId="39.[111,649,406,1700]" pageId="39" pageNumber="854">
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Ŋis petrosal compares best with the
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<typeStatus id="54F5D2DDFFF7867A269D7DD0FDA83AF7" box="[474,563,437,463]" pageId="39" pageNumber="854" type="holotype">holotype</typeStatus>
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petrosal of “
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<taxonomicName id="4C4E17FCFFF7867A27D57DB0FE943AD7" box="[146,271,469,495]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
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<emphasis id="B93AB06DFFF7867A27D57DB0FE943AD7" box="[146,271,469,495]" italics="true" pageId="39" pageNumber="854">Balaenoptera</emphasis>
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</taxonomicName>
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”
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<emphasis id="B93AB06DFFF7867A26677DB3FEC03AD7" box="[288,347,470,495]" italics="true" pageId="39" pageNumber="854">cortesi</emphasis>
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“var.”
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<emphasis id="B93AB06DFFF7867A26DF7DB3FE4F3AD7" box="[408,468,470,495]" italics="true" pageId="39" pageNumber="854">portisi</emphasis>
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from the Pliocene of
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<collectingCountry id="F3592CEFFFF7867A27CC7D90FF223937" box="[139,185,501,527]" name="Italy" pageId="39" pageNumber="854">Italy</collectingCountry>
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(Sacco 1890), as well as SDNHM 68698, a petrosal preserved with a partial skull from the Upper Pliocene San Diego Formation of southern California referred to “
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<emphasis id="B93AB06DFFF7867A27AB7E30FE983956" box="[236,259,597,622]" italics="true" pageId="39" pageNumber="854">B.</emphasis>
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”
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<emphasis id="B93AB06DFFF7867A26507E30FEC83956" box="[279,339,597,622]" italics="true" pageId="39" pageNumber="854">cortesi</emphasis>
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“var.”
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<emphasis id="B93AB06DFFF7867A26D07E30FE4F3956" box="[407,468,597,622]" italics="true" pageId="39" pageNumber="854">portisi</emphasis>
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by Deméré
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<emphasis id="B93AB06DFFF7867A25127E30FD1C3956" box="[597,647,596,622]" italics="true" pageId="39" pageNumber="854">et al.</emphasis>
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(2005).
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<bibRefCitation id="EFDF118EFFF7867A278E7E11FEF539B7" author="DEMERE T. A." box="[201,366,628,655]" pageId="39" pageNumber="854" pagination="277 - 298" refId="ref67320" refString="DEMERE T. A. 1986. - Ne fossil whale, Balaenoptera davidsonii (Cope 1872), with a review of other Neogene species of Balaenoptera (Cetacea: Mysticeti). Marine Mammal Science 2: 277 - 298." type="journal article" year="1986">Deméré (1986)</bibRefCitation>
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and Deméré
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<emphasis id="B93AB06DFFF7867A25447E11FDAD39B6" box="[515,566,628,654]" italics="true" pageId="39" pageNumber="854">et al.</emphasis>
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(2005) further indicated that the
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<typeStatus id="54F5D2DDFFF7867A26327EF6FE4B3995" box="[373,464,659,685]" pageId="39" pageNumber="854" type="holotype">holotype</typeStatus>
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mandible of “
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<taxonomicName id="4C4E17FCFFF7867A251A7EF6FF5439F5" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
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<emphasis id="B93AB06DFFF7867A251A7EF6FF5439F5" italics="true" pageId="39" pageNumber="854">Balaenoptera</emphasis>
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</taxonomicName>
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”
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<taxonomicName id="4C4E17FCFFF7867A27987ED6FE4A39F5" authority="Kellogg, 1944" authorityName="Kellogg" authorityYear="1944" box="[223,465,691,717]" class="Mammalia" family="Physeteridae" genus="Kogiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="species" species="floridana">
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<emphasis id="B93AB06DFFF7867A27987ED6FEA139F5" box="[223,314,691,717]" italics="true" pageId="39" pageNumber="854">floridana</emphasis>
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<bibRefCitation id="EFDF118EFFF7867A26797ED6FE4A39F5" author="KELLOGG R." box="[318,465,691,717]" pageId="39" pageNumber="854" pagination="433 - 471" refId="ref70474" refString="KELLOGG R. 1944. - Fossil Cetaceans from the Florida Tertiary. Bulletin of the Museum of Comparative Zoology at Harvard College 44: 433 - 471." type="journal article" year="1944">Kellogg, 1944</bibRefCitation>
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</taxonomicName>
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, as well as the Italian Pliocene balaenopterid
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<taxonomicName id="4C4E17FCFFF7867A26C67EB6FD1339D5" baseAuthorityName="Brandt" baseAuthorityYear="1873" box="[385,648,723,749]" class="Mammalia" family="Balaenopteridae" genus="Cetotheriophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="species" species="capellinii">
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<emphasis id="B93AB06DFFF7867A26C67EB6FD1339D5" box="[385,648,723,749]" italics="true" pageId="39" pageNumber="854">Cetotheriophanes capellinii</emphasis>
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</taxonomicName>
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may also be congeneric or conspecific with this taxon. “
|
|
<emphasis id="B93AB06DFFF7867A273B7F76FF083814" box="[124,147,787,812]" italics="true" pageId="39" pageNumber="854">B.</emphasis>
|
|
”
|
|
<emphasis id="B93AB06DFFF7867A27E57F76FF453814" box="[162,222,787,812]" italics="true" pageId="39" pageNumber="854">cortesi</emphasis>
|
|
“var.”
|
|
<emphasis id="B93AB06DFFF7867A26587F76FEC73814" box="[287,348,787,812]" italics="true" pageId="39" pageNumber="854">portisi</emphasis>
|
|
does not belong in
|
|
<taxonomicName id="4C4E17FCFFF7867A25657F77FF0D3874" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
|
|
<emphasis id="B93AB06DFFF7867A25657F77FF0D3874" italics="true" pageId="39" pageNumber="854">Balaenoptera</emphasis>
|
|
</taxonomicName>
|
|
and requires a new genus name and reevaluation (Deméré
|
|
<emphasis id="B93AB06DFFF7867A27947F37FE9F3853" box="[211,260,849,875]" italics="true" pageId="39" pageNumber="854">et al.</emphasis>
|
|
2005). In particular, UCMP 219135 shares many distinctive features with of “
|
|
<emphasis id="B93AB06DFFF7867A25677F17FDAC38B3" box="[544,567,882,907]" italics="true" pageId="39" pageNumber="854">B.</emphasis>
|
|
”
|
|
<emphasis id="B93AB06DFFF7867A250D7F17FD1D38B3" box="[586,646,882,907]" italics="true" pageId="39" pageNumber="854">cortesi</emphasis>
|
|
“var.”
|
|
<emphasis id="B93AB06DFFF7867A27EC7FF7FF7C3893" box="[171,231,914,939]" italics="true" pageId="39" pageNumber="854">portisi</emphasis>
|
|
such as a dorsally elongate pars cochlearis, a straight anterior margin of the anterior process, an anterior process that is pointed anterodorsally rather than anteriorly,and a flattened,bladelike anterior process. Crania of “
|
|
<emphasis id="B93AB06DFFF7867A26567875FEB33F11" box="[273,296,1040,1065]" italics="true" pageId="39" pageNumber="854">B.</emphasis>
|
|
”
|
|
<emphasis id="B93AB06DFFF7867A26707875FEE83F11" box="[311,371,1040,1065]" italics="true" pageId="39" pageNumber="854">cortesi</emphasis>
|
|
“var.”
|
|
<emphasis id="B93AB06DFFF7867A26F57875FE743F11" box="[434,495,1040,1065]" italics="true" pageId="39" pageNumber="854">portisi</emphasis>
|
|
preserving this petrosal morphology differ from
|
|
<taxonomicName id="4C4E17FCFFF7867A2685784AFF1C3F51" authority="bertae n." authorityName="BERTAE N." class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="850" phylum="Chordata" rank="genus">
|
|
<emphasis id="B93AB06DFFF7867A2685784AFD1F3F71" box="[450,644,1071,1097]" italics="true" pageId="39" pageNumber="854">Balaenoptera bertae</emphasis>
|
|
<taxonomicNameLabel id="A2090D16FFF7867A2736782AFF1C3F51" box="[113,135,1103,1129]" pageId="39" pageNumber="854" rank="species">n.</taxonomicNameLabel>
|
|
</taxonomicName>
|
|
sp. in exhibiting more elongate and widely flaring zygomatic processes, lacking a squamosal crease, an anteroposteriorly more elongate supraorbital process of the frontal, and an occipital shield that is more anteriorly elongate and attenuate; it is thus unlikely that UCMP 219135 is referable to
|
|
<taxonomicName id="4C4E17FCFFF7867A269B788BFDAE3E3F" authority="Boessenecker, 2013" authorityName="Boessenecker" authorityYear="2013" box="[476,565,1261,1287]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="species" species="bertae" status="sp. nov.">
|
|
<emphasis id="B93AB06DFFF7867A269B788BFDAE3E3F" box="[476,565,1261,1287]" italics="true" pageId="39" pageNumber="854">B. bertae</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A2090D16FFF7867A257C788BFDE83E30" box="[571,627,1262,1288]" pageId="39" pageNumber="854" rank="species">n. sp.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="8BF16C7FFFF7867A27CF7968FB0C3A09" blockId="39.[111,649,406,1700]" lastBlockId="39.[674,1212,216,970]" pageId="39" pageNumber="854">
|
|
Some features of this specimen are shared with
|
|
<taxonomicName id="4C4E17FCFFF7867A27347948FEFB3E7F" authorityName="Brisson" authorityYear="1762" box="[115,352,1325,1351]" class="Mammalia" family="Balaenopteridae" genus="Megaptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="species" species="novaeangliae">
|
|
<emphasis id="B93AB06DFFF7867A27347948FEFB3E7F" box="[115,352,1325,1351]" italics="true" pageId="39" pageNumber="854">Megaptera novaeangliae</emphasis>
|
|
</taxonomicName>
|
|
, including the extreme elongation of the pars cochlearis, and having the perilymphatic and endolymphatic foramina situated in a common groove. However, UCMP 219135 differs from
|
|
<taxonomicName id="4C4E17FCFFF7867A27EB79C9FE8F3EFD" authorityName="Gray" authorityYear="1846" box="[172,276,1452,1477]" class="Mammalia" family="Balaenopteridae" genus="Megaptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
|
|
<emphasis id="B93AB06DFFF7867A27EB79C9FE8F3EFD" box="[172,276,1452,1477]" italics="true" pageId="39" pageNumber="854">Megaptera</emphasis>
|
|
</taxonomicName>
|
|
in having deep canals for the facial and vestibulocochlear nerves, in having the internal acoustic meatus positioned in the middle of the pars cochlearis (instead of the posterior aspect as in extant
|
|
<taxonomicName id="4C4E17FCFFF7867A27367A4EFF433D7C" authorityName="Gray" authorityYear="1846" box="[113,216,1579,1604]" class="Mammalia" family="Balaenopteridae" genus="Megaptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
|
|
<emphasis id="B93AB06DFFF7867A27367A4EFF433D7C" box="[113,216,1579,1604]" italics="true" pageId="39" pageNumber="854">Megaptera</emphasis>
|
|
</taxonomicName>
|
|
), and in having a smaller and less acutely pointed lateral tuberosity.UCMP 219135 differs from extant
|
|
<taxonomicName id="4C4E17FCFFF7867A27FD7A0CFEA73DBB" box="[186,316,1641,1667]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
|
|
<emphasis id="B93AB06DFFF7867A27FD7A0CFEA73DBB" box="[186,316,1641,1667]" italics="true" pageId="39" pageNumber="854">Balaenoptera</emphasis>
|
|
</taxonomicName>
|
|
spp. in the extreme elongation of the pars cochlearis, and an anterior process that is directed anterodorsally (relative to the long axis of the pars cochlearis), rather than anteriorly in extant
|
|
<taxonomicName id="4C4E17FCFFF7867A25E27D72FCBE3A09" box="[677,805,279,305]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
|
|
<emphasis id="B93AB06DFFF7867A25E27D72FCBE3A09" box="[677,805,279,305]" italics="true" pageId="39" pageNumber="854">Balaenoptera</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C4E17FCFFF7867A241F7D7DFB083A09" authority="(Ekdale et al. 2011)" baseAuthorityName="Ekdale" baseAuthorityYear="2011" box="[856,1171,279,305]" class="Mammalia" family="Balaenopteridae" genus="Megaptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
|
|
<emphasis id="B93AB06DFFF7867A241F7D7DFC5B3A09" box="[856,960,280,305]" italics="true" pageId="39" pageNumber="854">Megaptera</emphasis>
|
|
(
|
|
<bibRefCitation id="EFDF118EFFF7867A248A7D72FB103A09" author="EKDALE E. G. & BERTA A. & DEMERE T. A." box="[973,1163,279,305]" pageId="39" pageNumber="854" pagination="1 - 42" refId="ref68110" refString="EKDALE E. G., BERTA A. & DEMERE T. A. 2011. - Ne comparative osteology of the petrotympanic complex (ear region) of extant baleen whales (Cetacea: Mysticeti). Plos One 6 (6): 1 - 42." type="journal article" year="2011">
|
|
Ekdale
|
|
<emphasis id="B93AB06DFFF7867A235C7D7DFBD73A09" box="[1051,1100,279,305]" italics="true" pageId="39" pageNumber="854">et al.</emphasis>
|
|
2011
|
|
</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BF16C7FFFF7867A25FC7D52FCEB3957" blockId="39.[674,1212,216,970]" pageId="39" pageNumber="854">
|
|
<materialsCitation id="3B266622FFF7867A25FC7D52FB2D3AF7" ID-GBIF-Occurrence="3120744303" collectionCode="UCMP" pageId="39" pageNumber="854" specimenCode="FP 192" specimenCount="1">
|
|
Two additional specimens from the Purisima Formation undoubtedly represent this taxon.A large cranial fragment (
|
|
<collectionCode id="ED5FF4BAFFF7867A24547D13FCFE3AA8" box="[787,869,374,400]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="39" pageNumber="854" type="University or college">UCMP</collectionCode>
|
|
uncataloged, field no.
|
|
<specimenCode id="DBE8C404FFF7867A23197D12FB2B3AA8" box="[1118,1200,374,401]" collectionCode="FP" pageId="39" pageNumber="854">FP 192</specimenCode>
|
|
) from the Santa Cruz section of the Purisima Formation exhibits a well-preserved and identical petrosal
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="3B266622FFF7867A25E37DB0FCF63957" ID-GBIF-Occurrence="3120744304" collectionCode="SCMNH, SDNHM" pageId="39" pageNumber="854" specimenCode="SCMNH 21292, SDNHM 65769" specimenCount="1" typeStatus="holotype">
|
|
furthermore, a partial cranium (
|
|
<specimenCode id="DBE8C404FFF7867A24BB7DB0FB293AD7" box="[1020,1202,469,495]" collectionCode="SCMNH" pageId="39" pageNumber="854">SCMNH 21292</specimenCode>
|
|
) exhibits squamosal morphology that is identical to the
|
|
<typeStatus id="54F5D2DDFFF7867A258A7E70FCB33917" box="[717,808,533,559]" pageId="39" pageNumber="854" type="holotype">holotype</typeStatus>
|
|
of “
|
|
<emphasis id="B93AB06DFFF7867A24137E70FCF03916" box="[852,875,533,558]" italics="true" pageId="39" pageNumber="854">B.</emphasis>
|
|
”
|
|
<emphasis id="B93AB06DFFF7867A243D7E70FC2D3916" box="[890,950,533,558]" italics="true" pageId="39" pageNumber="854">cortesi</emphasis>
|
|
“var.”
|
|
<emphasis id="B93AB06DFFF7867A24BD7E70FBAC3916" box="[1018,1079,533,558]" italics="true" pageId="39" pageNumber="854">portisi</emphasis>
|
|
and referred skulls from the San Diego Formation (
|
|
<specimenCode id="DBE8C404FFF7867A23097E51FD713957" collectionCode="SDNHM" country="USA" httpUri="http://grbio.org/cool/4ksk-xsw5" name="San Diego Natural History Museum" pageId="39" pageNumber="854">SDNHM 65769</specimenCode>
|
|
and 68698)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BF16C7FFFF7867A25FC7E11FCED38F2" blockId="39.[674,1212,216,970]" pageId="39" pageNumber="854">
|
|
Ŋis fossil represents an additional occurrence of “
|
|
<emphasis id="B93AB06DFFF7867A25E87EF1FD5D3995" box="[687,710,660,685]" italics="true" pageId="39" pageNumber="854">B.</emphasis>
|
|
”
|
|
<emphasis id="B93AB06DFFF7867A259F7EF1FC8F3995" box="[728,788,660,685]" italics="true" pageId="39" pageNumber="854">cortesi</emphasis>
|
|
“var.”
|
|
<emphasis id="B93AB06DFFF7867A241D7EF1FC0C3995" box="[858,919,660,685]" italics="true" pageId="39" pageNumber="854">portisi</emphasis>
|
|
from the eastern North Pacific along with that from the San Diego Formation reported by Deméré
|
|
<emphasis id="B93AB06DFFF7867A243B7EB6FC3539D5" box="[892,942,723,749]" italics="true" pageId="39" pageNumber="854">et al.</emphasis>
|
|
(2005). Along with other records of this taxon from the Pliocene of Florida and
|
|
<collectingCountry id="F3592CEFFFF7867A25E57F77FD543814" box="[674,719,786,812]" name="Italy" pageId="39" pageNumber="854">Italy</collectingCountry>
|
|
(
|
|
<bibRefCitation id="EFDF118EFFF7867A259D7F77FCCA3815" author="PORTIS A." box="[730,849,786,813]" pageId="39" pageNumber="854" pagination="247 - 365" refId="ref73764" refString="PORTIS A. 1885. - Catalogo descrittivo dei Talassoterii rinvenuti nei terreni terziarii del Piemonte e della Liguria. Memoire della R. Accademia delle scienze di Torino, Turin 37: 247 - 365." type="journal article" year="1885">Portis1885</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFDF118EFFF7867A241E7F77FC783814" author="KELLOGG R." box="[857,995,786,812]" pageId="39" pageNumber="854" pagination="433 - 471" refId="ref70474" refString="KELLOGG R. 1944. - Fossil Cetaceans from the Florida Tertiary. Bulletin of the Museum of Comparative Zoology at Harvard College 44: 433 - 471." type="journal article" year="1944">Kellogg 1944</bibRefCitation>
|
|
; Deméré
|
|
<emphasis id="B93AB06DFFF7867A23067F76FBE93814" box="[1089,1138,786,812]" italics="true" pageId="39" pageNumber="854">et al.</emphasis>
|
|
2005), this indicates that this taxon (whether several species within a genus, or a single species) was cosmopolitan and formerly inhabited the eastern North Pacific,the western North Atlantic, and the Mediterranean Sea during the Pliocene.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |