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<document id="3BC785578BC1C0097EB79118E82A4EF4" ID-DOI="10.5281/zenodo.195855" ID-GBIF-Dataset="894cdac4-4f41-4d3d-a4a0-d606f3f1e3c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="195855" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460235104592" checkinUser="plazi" docAuthor="Sokolov, Igor M. &amp; Carlton, Christopher E." docDate="2010" docId="03E751295D55FF59FF55FC7CFA50FB1A" docLanguage="en" docName="zt02502p023.pdf" docOrigin="Zootaxa 2502" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anillinus loweae" docType="key" docVersion="10" lastPageNumber="15" masterDocId="FFDE29515D58FF57FFC2FFFEFFCCFFF5" masterDocTitle="New species of Anillinus Casey (Carabidae: Trechinae: Bembidiini) from the Southern Appalachians and phylogeography of the A. loweae species group" masterLastPageNumber="23" masterPageNumber="1" pageNumber="14" updateTime="1698239394963" updateUser="plazi">
<mods:mods id="EE512815651DA8113DCF2F9632BAE156" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A4CF9450E5ED7F9F01E6D8F2251701F0">New species of Anillinus Casey (Carabidae: Trechinae: Bembidiini) from the Southern Appalachians and phylogeography of the A. loweae species group</mods:title>
</mods:titleInfo>
<mods:name id="1E2CE6F4FF7967072B2C6C516AEA06F8" type="personal">
<mods:role id="D4901067741EDAA8A58D9F6BDF447D6A">
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<mods:namePart id="E6927065FAE375FEA4E627721B9C157F">Sokolov, Igor M.</mods:namePart>
</mods:name>
<mods:name id="294AAFAB35F64DE39CA406C3917D9DCF" type="personal">
<mods:role id="D9C432E1CBE2995B41220C33F8708537">
<mods:roleTerm id="5DAF95A6A2D76CCF3E1C56E6938F06FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F344BA9C5D1BEB0D43DC4A87AED00C18">Carlton, Christopher E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A8C1D7C2FF0E9B9E31268C5A1AB8AE77">text</mods:typeOfResource>
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<mods:part id="76C82CDCDE7C462B936204F0448ACCE9">
<mods:date id="07FCF2E03B823F9BD812F38BE950F67D">2010</mods:date>
<mods:detail id="8E8C6DEAB1CC63FE8EE2A2EDBBDD26BE" type="volume">
<mods:number id="98A66621A3D8EF284F238F25385BC572">2502</mods:number>
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<mods:classification id="C8A64ADB5CE6B08E9AC7E0B29417E8C9">journal article</mods:classification>
<mods:identifier id="2A4B65832046DA14EAE5D2CE7688B674" type="DOI">10.5281/zenodo.195855</mods:identifier>
<mods:identifier id="BCA25E94D1FD2A86C18301D77977DF30" type="GBIF-Dataset">894cdac4-4f41-4d3d-a4a0-d606f3f1e3c4</mods:identifier>
<mods:identifier id="C1AA1787E5A87071C36E85C43E33E5DC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4066A8B2279C701B1E7A094553E53356" type="Zenodo-Dep">195855</mods:identifier>
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<treatment id="03E751295D55FF59FF55FC7CFA50FB1A" ID-DOI="http://doi.org/10.5281/zenodo.5694155" ID-GBIF-Taxon="119387937" ID-Zenodo-Dep="5694155" LSID="urn:lsid:plazi:treatment:03E751295D55FF59FF55FC7CFA50FB1A" httpUri="http://treatment.plazi.org/id/03E751295D55FF59FF55FC7CFA50FB1A" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C354B3B45D55FF5AFF55FC7CFB1AFBC7" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BF1E03F5D55FF5AFF55FC7CFB89FC69" blockId="13.[151,1093,898,924]" box="[151,1093,898,924]" pageId="13" pageNumber="14">
<heading id="D0B957535D55FF5AFF55FC7CFB89FC69" bold="true" box="[151,1093,898,924]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B93A3C2D5D55FF5AFF55FC7CFB89FC69" bold="true" box="[151,1093,898,924]" pageId="13" pageNumber="14">
Key for identification of the litter species within the
<taxonomicName id="4C4E9BBC5D55FF5AFCE3FC7CFC5AFC69" ID-CoL="66Q8D" box="[801,918,898,924]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="B93A3C2D5D55FF5AFCE3FC7CFC5AFC69" bold="true" box="[801,918,898,924]" italics="true" pageId="13" pageNumber="14">A. loweae</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF1E03F5D55FF5AFF55FC33FB1AFBC7" blockId="13.[151,1437,973,1074]" pageId="13" pageNumber="14">
This key will allow determination of the
<taxonomicName id="4C4E9BBC5D55FF5AFD47FC33FD15FC13" box="[645,729,973,998]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="loweae">
<emphasis id="B93A3C2D5D55FF5AFD47FC33FD15FC13" box="[645,729,973,998]" italics="true" pageId="13" pageNumber="14">loweae</emphasis>
</taxonomicName>
-group species and their allies in the southern Appalachian Mountains. External features are of limited diagnostic value, and the key is based mostly on genitalia structures of both sexes, so correct identification of the specimen usually requires dissection.
</paragraph>
</subSubSection>
<subSubSection id="C354B3B45D55FF59FF55FB9EFA50FB1A" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="key">
<keyStep id="30BAF89A5D55FF5AFF55FB9EFA59FB62" pageId="13" pageNumber="14">
<paragraph id="8BF1E03F5D55FF5AFF55FB9EFA57FB82" blockId="13.[151,1436,1120,2007]" box="[151,1435,1120,1143]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FB9EFA57FB82" box="[151,1435,1120,1143]" pageId="13" pageNumber="14">1. Head with paramedian patches of effaced microsculpture on each side of the vertex ................................................3</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D55FF5AFF55FB7EFA59FB62" blockId="13.[151,1436,1120,2007]" box="[151,1429,1152,1175]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FB7EFA59FB62" box="[151,1429,1152,1175]" pageId="13" pageNumber="14">- Vertex totally covered with polygonal microsculpture (often all head covered with microsculpture)........................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D55FF5AFF55FB5EFA57FAE2" pageId="13" pageNumber="14">
<paragraph id="8BF1E03F5D55FF5AFF55FB5EFA50FB22" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FB5EFA50FB22" pageId="13" pageNumber="14">
2. Pronotum greatly constricted posteriad: ratio WPa/WPp≥1.00, and elytra subdepressed and parallel with 3 and more interneurs
<emphasis id="B93A3C2D5D55FF5AFEF0FB3EFB6EFB22" box="[306,1186,1216,1239]" italics="true" pageId="13" pageNumber="14">.................................................................................................................................. (</emphasis>
other
<taxonomicName id="4C4E9BBC5D55FF5AFB1DFB3EFA8CFB22" ID-CoL="8KV4T" box="[1247,1344,1216,1239]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3C2D5D55FF5AFB1DFB3EFA8CFB22" box="[1247,1344,1216,1239]" italics="true" pageId="13" pageNumber="14">Anillinus</emphasis>
</taxonomicName>
species)
</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D55FF5AFF55FB1EFA57FAE2" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FB1EFA57FAE2" pageId="13" pageNumber="14">
- Pronotum less constricted posteriad: ratio WPa/WPp ≤1.00, and elytra convex and ovoid with 1–2 interneurs only... ............................................................................................................................................................ (
<taxonomicName id="4C4E9BBC5D55FF5AFB2FFAFEFA80FAE2" ID-CoL="6728C" box="[1261,1356,1280,1303]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="langdoni">
<emphasis id="B93A3C2D5D55FF5AFB2FFAFEFA80FAE2" box="[1261,1356,1280,1303]" italics="true" pageId="13" pageNumber="14">langdoni</emphasis>
</taxonomicName>
-group)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D55FF5AFF55FADEFA50FA62" pageId="13" pageNumber="14">
<paragraph id="8BF1E03F5D55FF5AFF55FADEFA57FAA2" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FADEFA57FAA2" pageId="13" pageNumber="14">3. Frons and usually pronotum with distinct microsculpture, microsculpture on frons effaced on beetles from Alabama and Mississippi.............................................................................................................................................................. 4</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D55FF5AFF55FA9EFA50FA62" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FA9EFA50FA62" pageId="13" pageNumber="14">
- Frons, vertex (except small triangular area in the middle) and pronotum without microsculpture; beetles from eastern Tennessee, North Carolina,
<collectingCountry id="F359A0AF5D55FF5AFDDAFA7EFDA2FA62" box="[536,622,1408,1431]" name="Georgia" pageId="13" pageNumber="14">Georgia</collectingCountry>
and South Carolina .................................................. (other
<taxonomicName id="4C4E9BBC5D55FF5AFB1DFA7EFA8CFA62" ID-CoL="8KV4T" box="[1247,1344,1408,1431]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3C2D5D55FF5AFB1DFA7EFA8CFA62" box="[1247,1344,1408,1431]" italics="true" pageId="13" pageNumber="14">Anillinus</emphasis>
</taxonomicName>
species)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D55FF5AFF55FA5EFA57FA02" pageId="13" pageNumber="14">
<paragraph id="8BF1E03F5D55FF5AFF55FA5EFA50FA42" blockId="13.[151,1436,1120,2007]" box="[151,1436,1440,1463]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FA5EFA50FA42" box="[151,1436,1440,1463]" pageId="13" pageNumber="14">4. Beetles from Appalachian Mountains of Tennessee, North Carolina, and Virginia north of French Broad River ...... 5</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D55FF5AFF55FA3EFA57FA02" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55FA3EFA57FA02" pageId="13" pageNumber="14">
- Beetles from Appalachian Mountains of Tennessee, North Carolina south of French Broad River and South Carolina,
<collectingCountry id="F359A0AF5D55FF5AFF3EFA1EFE98FA02" box="[252,340,1504,1527]" name="Georgia" pageId="13" pageNumber="14">Georgia</collectingCountry>
, Alabama and Mississippi ....................................................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D55FF5AFF55F9FEFA57F902" pageId="13" pageNumber="14">
<paragraph id="8BF1E03F5D55FF5AFF55F9FEFA50F982" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55F9FEFA50F982" pageId="13" pageNumber="14">
5. Pronotum with very fine polygonal meshes on the disc visible only at certain angle. Dorsal sclerites in the form of filament-like structures (
<figureCitation id="1375FCBA5D55FF5AFE13F9DEFDEDF9C2" box="[465,545,1568,1591]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
a). Left paramere with long apical seta (
<figureCitation id="1375FCBA5D55FF5AFC16F9DEFBE8F9C2" box="[980,1060,1568,1591]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
b). Right paramere short with long setae (
<figureCitation id="1375FCBA5D55FF5AFECFF9BEFE92F9A2" box="[269,350,1600,1623]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
c). Spermatheca S-shaped with proximal curvature greatly reduced in length (
<figureCitation id="1375FCBA5D55FF5AFB69F9BEFB31F9A2" box="[1195,1277,1600,1623]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="13" pageNumber="14">Fig. 24</figureCitation>
). Stylomers as illustrated (
<figureCitation id="1375FCBA5D55FF5AFE82F99EFE5CF982" box="[320,400,1632,1655]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="13" pageNumber="14">Fig. 32</figureCitation>
). Beetles (
<figureCitation id="1375FCBA5D55FF5AFDC2F99EFD8CF982" box="[512,576,1632,1655]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
) from higher elevations (&gt;
<quantity id="4CB64DDA5D55FF5AFC9FF99EFC24F982" box="[861,1000,1632,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="13" pageNumber="14" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200-1300m</quantity>
)
<emphasis id="B93A3C2D5D55FF5AFC39F99EFB06F982" box="[1019,1226,1632,1655]" italics="true" pageId="13" pageNumber="14">
....................
<taxonomicName id="4C4E9BBC5D55FF5AFB44F99EFB06F982" ID-CoL="66PWB" authority="Sokolov &amp; Carlton" authorityName="Sokolov &amp; Carlton" box="[1158,1226,1632,1655]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="erwini">erwini</taxonomicName>
</emphasis>
Sokolov &amp; Carlton
</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D55FF5AFF55F97EFA57F902" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55F97EFA57F902" pageId="13" pageNumber="14">
- Pronotum with distinct polygonal meshes across entire surface. Dorsal sclerites in the form of blade-like structures (
<figureCitation id="1375FCBA5D55FF5AFF0DF95EFED0F942" box="[207,284,1696,1719]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="13" pageNumber="14">Fig. 17</figureCitation>
a). Left paramere with short apical seta (
<figureCitation id="1375FCBA5D55FF5AFD05F95EFCD8F942" box="[711,788,1696,1719]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="13" pageNumber="14">Fig. 17</figureCitation>
b). Right paramere elongate with short setae (
<figureCitation id="1375FCBA5D55FF5AFB37F95EFA8EF942" box="[1269,1346,1696,1719]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="13" pageNumber="14">Fig. 17</figureCitation>
c). Spermatheca S-shaped with both curvatures of equal width (
<figureCitation id="1375FCBA5D55FF5AFCE2F93EFCBDF922" box="[800,881,1728,1751]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="13" pageNumber="14">Fig. 25</figureCitation>
). Stylomers as illustrated (
<figureCitation id="1375FCBA5D55FF5AFB52F93EFB2DF922" box="[1168,1249,1728,1751]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="13" pageNumber="14">Fig. 33</figureCitation>
). Beetles (
<figureCitation id="1375FCBA5D55FF5AFA91F93EFA58F922" box="[1363,1428,1728,1751]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
) from low and middle altitudes (&lt;
<quantity id="4CB64DDA5D55FF5AFDE7F91EFD7CF902" box="[549,688,1760,1783]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="13" pageNumber="14" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200-1300m</quantity>
)
<emphasis id="B93A3C2D5D55FF5AFD79F91EFAC1F902" box="[699,1293,1760,1783]" italics="true" pageId="13" pageNumber="14">.........................................................................................</emphasis>
<taxonomicName id="4C4E9BBC5D55FF5AFAD6F91EFA57F902" ID-CoL="5V28F" authority="(Horn)" authorityName="Horn" baseAuthorityName="Horn" box="[1300,1435,1760,1783]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="B93A3C2D5D55FF5AFAD6F91EFA80F902" box="[1300,1356,1760,1783]" italics="true" pageId="13" pageNumber="14">fortis</emphasis>
(Horn)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D55FF5AFF55F8FEFA50F862" pageId="13" pageNumber="14">
<paragraph id="8BF1E03F5D55FF5AFF55F8FEFA57F8A2" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55F8FEFA57F8A2" pageId="13" pageNumber="14">
6. Males with right paramere short and wide (
<figureCitation id="1375FCBA5D55FF5AFD5FF8FEFD38F8E2" box="[669,756,1792,1815]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="13" pageNumber="14">Figs. 11</figureCitation>
c, 13c). Beetles from the southwestern Appalachian Mountains approximately from the eastern border of Blount County, Tennessee to north-western
<collectingCountry id="F359A0AF5D55FF5AFBA2F8DEFB74F8C2" box="[1120,1208,1824,1847]" name="Georgia" pageId="13" pageNumber="14">Georgia</collectingCountry>
, northern counties of Alabama, with outlying records from Mississippi ........................................................................................................ 7
</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D55FF5AFF55F89EFA50F862" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55F89EFA50F862" pageId="13" pageNumber="14">
- Males with right paramere elongate (
<figureCitation id="1375FCBA5D55FF5AFDA2F89EFD77F882" box="[608,699,1888,1911]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="13" pageNumber="14">Figs. 10</figureCitation>
c, 12c, 18c, 19c). Beetles mainly from eastern portions of southern Appalachian Mountains, including much of Great Smoky Mountains National Park.................................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D55FF59FF55F85EFA59FE9A" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<paragraph id="8BF1E03F5D55FF5AFF55F85EFA50F822" blockId="13.[151,1436,1120,2007]" pageId="13" pageNumber="14">
<keyLead id="30BF430A5D55FF5AFF55F85EFA50F822" pageId="13" pageNumber="14">
7. Median lobe without a patch of setae on ventral surface. Dorsal sclerites very long, in the form of filament-like structures (
<figureCitation id="1375FCBA5D55FF5AFEFFF83EFE46F822" box="[317,394,1984,2007]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
a). Left paramere of normal size and form (
<figureCitation id="1375FCBA5D55FF5AFC9AF83EFC68F822" box="[856,932,1984,2007]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
b). Spermatheca S-shaped with both curvatures
</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D56FF59FF05FF66FA59FE9A" blockId="14.[151,1436,152,1263]" pageId="14" pageNumber="15">
of equal width (
<figureCitation id="1375FCBA5D56FF59FEB1FF66FE0FFF5A" box="[371,451,152,175]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 21</figureCitation>
). Stylomers as illustrated (
<figureCitation id="1375FCBA5D56FF59FD21FF66FCF8FF5A" box="[739,820,152,175]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="14" pageNumber="15">Fig. 29</figureCitation>
). Beetles (
<figureCitation id="1375FCBA5D56FF59FC64FF66FC2BFF5A" box="[934,999,152,175]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="14" pageNumber="15">Fig. 2</figureCitation>
) from Starr Mountain, McMinn County, Tennessee
<emphasis id="B93A3C2D5D56FF59FEFDFF46FA8BFF3A" box="[319,1351,184,207]" italics="true" pageId="14" pageNumber="15">
.................................................................................................................................................
<taxonomicName id="4C4E9BBC5D56FF59FAC8FF46FA8BFF3A" ID-CoL="5V28C" box="[1290,1351,184,207]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="juliae" status="sp. nov.">juliae</taxonomicName>
</emphasis>
<emphasis id="B93A3C2D5D56FF59FA8FFF49FA50FF3A" bold="true" box="[1357,1436,183,207]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A20981565D56FF59FA8FFF49FA50FF3A" box="[1357,1436,183,207]" pageId="14" pageNumber="15" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
<keyLead id="30BF430A5D56FF59FF55FF26FA59FE9A" pageId="14" pageNumber="15">
- Median lobe with a patch of setae on ventral surface. Dorsal sclerites of moderate size in form of blade-like structures (
<figureCitation id="1375FCBA5D56FF59FECFFF06FEACFEFA" box="[269,352,248,271]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="14" pageNumber="15">Fig. 13</figureCitation>
a). Left paramere noticeably enlarged (
<figureCitation id="1375FCBA5D56FF59FCD0FF06FCAFFEFA" box="[786,867,248,271]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="14" pageNumber="15">Fig. 13</figureCitation>
b). Spermatheca S-shaped, with proximal curvature reduced, noticeably narrower in comparison to the distal curvature (
<figureCitation id="1375FCBA5D56FF59FC59FEE6FC27FEDA" box="[923,1003,280,303]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 23</figureCitation>
). Stylomers as illustrated (
<figureCitation id="1375FCBA5D56FF59FAC5FEE6FA9AFEDA" box="[1287,1366,280,303]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="14" pageNumber="15">Fig. 31</figureCitation>
). Beetles (
<figureCitation id="1375FCBA5D56FF59FF3EFEC6FEF2FEBA" box="[252,318,312,335]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="14" pageNumber="15">Fig. 4</figureCitation>
) from north-eastern corner of
<collectingCountry id="F359A0AF5D56FF59FD4BFEC6FD2DFEBA" box="[649,737,312,335]" name="Georgia" pageId="14" pageNumber="15">Georgia</collectingCountry>
, the northern counties of Alabama and Mississippi and from the western half of Great Smoky Mountains National Park (
<figureCitation id="1375FCBA5D56FF59FCDEFEA6FC90FE9A" box="[796,860,344,367]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
)
<emphasis id="B93A3C2D5D56FF59FCADFEA6FA93FE9A" box="[879,1375,344,367]" italics="true" pageId="14" pageNumber="15">
..............................................................
<taxonomicName id="4C4E9BBC5D56FF59FAD3FEA6FA93FE9A" ID-CoL="5V27L" authority="Barr" authorityName="Barr" box="[1297,1375,344,367]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="steevesi">steevesi</taxonomicName>
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Barr
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</paragraph>
</keyStep>
<keyStep id="30BAF89A5D56FF59FF55FE86FA57FD7A" pageId="14" pageNumber="15">
<paragraph id="8BF1E03F5D56FF59FF55FE86FA50FDFA" blockId="14.[151,1436,152,1263]" pageId="14" pageNumber="15">
<keyLead id="30BF430A5D56FF59FF55FE86FA50FDFA" pageId="14" pageNumber="15">
8. Dorsal sclerites very long, in form of filament-like structures with greatly elongated basal processes (
<figureCitation id="1375FCBA5D56FF59FAF3FE86FA4EFE7A" box="[1329,1410,376,399]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
a). Setae of right paramere long, much longer than the length of paramere itself (
<figureCitation id="1375FCBA5D56FF59FBC6FE66FB9CFE5A" box="[1028,1104,408,431]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
c). Spermatheca S-shaped with both curvatures of equal width, and distal curvature with angulated sinuation below (
<figureCitation id="1375FCBA5D56FF59FBA8FE46FB70FE3A" box="[1130,1212,440,463]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 20</figureCitation>
). Stylomers as illustrated (
<figureCitation id="1375FCBA5D56FF59FED1FE26FEAFFE1A" box="[275,355,472,495]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
). Beetles (
<figureCitation id="1375FCBA5D56FF59FE16FE26FDDEFE1A" box="[468,530,472,495]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
) from the limestone area at the eastern part of Rich Mountain ridge in Great Smoky Mountains National Park
<emphasis id="B93A3C2D5D56FF59FE1BFE06FA8AFDFA" box="[473,1350,504,527]" italics="true" pageId="14" pageNumber="15">
......................................................................................................................
<taxonomicName id="4C4E9BBC5D56FF59FB32FE06FA8AFDFA" ID-CoL="66PWF" box="[1264,1350,504,527]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gimmeli" status="sp. nov.">gimmeli</taxonomicName>
</emphasis>
<emphasis id="B93A3C2D5D56FF59FA8FFE09FA50FDFA" bold="true" box="[1357,1436,503,527]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A20981565D56FF59FA8FFE09FA50FDFA" box="[1357,1436,503,527]" pageId="14" pageNumber="15" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D56FF59FF55FDE6FA57FD7A" blockId="14.[151,1436,152,1263]" pageId="14" pageNumber="15">
<keyLead id="30BF430A5D56FF59FF55FDE6FA57FD7A" pageId="14" pageNumber="15">
- Dorsal sclerites shorter, in form of blade-like structures (
<figureCitation id="1375FCBA5D56FF59FCF2FDE6FC45FDDA" box="[816,905,536,559]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="14" pageNumber="15">Figs. 12</figureCitation>
a, 18a, 19a). Setae of right paramere shorter than the length of paramere itself (
<figureCitation id="1375FCBA5D56FF59FDC1FDC6FD97FDBA" box="[515,603,568,591]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="14" pageNumber="15">Figs. 12</figureCitation>
c, 18c, 19c). Spermatheca S-shaped, EITHER with proximal curvature noticeably narrower than distal curvature (
<figureCitation id="1375FCBA5D56FF59FD8EFDA6FD0EFD9A" box="[588,706,600,623]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 22, 27</figureCitation>
) OR with distal curvature widely rounded and without angulate sinuation below (
<figureCitation id="1375FCBA5D56FF59FEA2FD86FE62FD7A" box="[352,430,632,655]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 26</figureCitation>
)................................................................................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D56FF59FF55FD66FA57FC5A" pageId="14" pageNumber="15">
<paragraph id="8BF1E03F5D56FF59FF55FD66FA50FCDA" blockId="14.[151,1436,152,1263]" pageId="14" pageNumber="15">
<keyLead id="30BF430A5D56FF59FF55FD66FA50FCDA" pageId="14" pageNumber="15">
9. Median lobe with a patch of spines near apical orifice (
<figureCitation id="1375FCBA5D56FF59FCE6FD66FCB8FD5A" box="[804,884,664,687]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="14" pageNumber="15">Fig. 18</figureCitation>
a). Apex of median lobe with angulate sinuation at dorsal side. Spermatheca S-shaped with both curvatures of equal size and distal curvature widely rounded below (
<figureCitation id="1375FCBA5D56FF59FF0DFD26FEEDFD1A" box="[207,289,728,751]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 26</figureCitation>
). Parameres (
<figureCitation id="1375FCBA5D56FF59FE75FD26FDCAFD1A" box="[439,518,728,751]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="14" pageNumber="15">Fig. 18</figureCitation>
b, c) and stylomers as illustrated (
<figureCitation id="1375FCBA5D56FF59FCBEFD26FC03FD1A" box="[892,975,728,751]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="14" pageNumber="15">Fig. 34</figureCitation>
). Beetles (
<figureCitation id="1375FCBA5D56FF59FB86FD26FB4AFD1A" box="[1092,1158,728,751]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
) from the eastern half of Great Smoky Mountains National Park and from ridges extending from Cataloochee Divide (Haywood County, North Carolina) to the south-eastern corner of Macon County, North Carolina
<emphasis id="B93A3C2D5D56FF59FBCAFCE6FB06FCDA" box="[1032,1226,792,815]" italics="true" pageId="14" pageNumber="15">
.................
<taxonomicName id="4C4E9BBC5D56FF59FBBDFCE6FB06FCDA" ID-CoL="66Q8D" authority="Sokolov &amp; Carlton" authorityName="Sokolov &amp; Carlton" box="[1151,1226,792,815]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="loweae">loweae</taxonomicName>
</emphasis>
Sokolov &amp; Carlton
</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D56FF59FF55FCC6FA57FC5A" blockId="14.[151,1436,152,1263]" pageId="14" pageNumber="15">
<keyLead id="30BF430A5D56FF59FF55FCC6FA57FC5A" pageId="14" pageNumber="15">- Median lobe without a patch of spines near apical orifice. Apex of median lobe at most with very obtuse sinuation in dorsal part or simply rounded. Spermatheca S-shaped with proximal curvature narrower than the distal. Beetles from the western half of Great Smoky Mountains National Park and ridges extending from Fontana Village (Swain County, North Carolina) to the north-western counties of South Carolina................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAF89A5D56FF59FF55FC46FA50FB1A" pageId="14" pageNumber="15">
<paragraph id="8BF1E03F5D56FF59FF55FC46FA50FBBA" blockId="14.[151,1436,152,1263]" pageId="14" pageNumber="15">
<keyLead id="30BF430A5D56FF59FF55FC46FA50FBBA" pageId="14" pageNumber="15">
10. Dorsal sclerites of median lobe larger, with short processes radiating from base (
<figureCitation id="1375FCBA5D56FF59FB82FC46FB5EFC3A" box="[1088,1170,952,975]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,271,1397,1421]" captionTargetBox="[175,1409,184,1396]" captionTargetId="figure@6.[172,1414,177,1396]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Male aedeagus of (10) Anillinus gimmeli, new species, (TN, Blount Co., White Oak Sink); (11) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (12) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (13) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195857/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
a). Apex of median lobe symmetrically rounded. Spermatheca comparatively wide, only 1.5 times longer than its maximal width (
<figureCitation id="1375FCBA5D56FF59FAFCFC26FA5CFC1A" box="[1342,1424,984,1007]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 22</figureCitation>
). Stylomers as illustrated (
<figureCitation id="1375FCBA5D56FF59FE15FC06FDE5FBFA" box="[471,553,1016,1039]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="14" pageNumber="15">Fig. 30</figureCitation>
). Beetles from low altitudes (
<quantity id="4CB64DDA5D56FF59FCADFC06FC2CFBFA" box="[879,992,1016,1039]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="14" pageNumber="15" unit="m" value="550.0" valueMax="700.0" valueMin="400.0">400-700m</quantity>
) from the Twentymile Creek valley (the western corner of Swain County, North Carolina to Gregory Bald) and Macon County, North Carolina .....................
<emphasis id="B93A3C2D5D56FF59FF0CFBC6FA8BFBBA" box="[206,1351,1080,1103]" italics="true" pageId="14" pageNumber="15">
...............................................................................................................................................................
<taxonomicName id="4C4E9BBC5D56FF59FB37FBC6FA8BFBBA" ID-CoL="66PVH" box="[1269,1351,1080,1103]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="merritti" status="sp. nov.">merritti</taxonomicName>
</emphasis>
<emphasis id="B93A3C2D5D56FF59FA8FFBC9FA50FBBA" bold="true" box="[1357,1436,1079,1103]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A20981565D56FF59FA8FFBC9FA50FBBA" box="[1357,1436,1079,1103]" pageId="14" pageNumber="15" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF1E03F5D56FF59FF55FBA6FA50FB1A" blockId="14.[151,1436,152,1263]" pageId="14" pageNumber="15">
<keyLead id="30BF430A5D56FF59FF55FBA6FA50FB1A" pageId="14" pageNumber="15">
- Dorsal sclerites of median lobe smaller, base prolonged and acute (
<figureCitation id="1375FCBA5D56FF59FC68FBA6FC36FB9A" box="[938,1018,1112,1135]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="14" pageNumber="15">Fig. 19</figureCitation>
a). Apex of median lobe less rounded, noticeably asymmetrical, its upper margin slightly elevated. Spermatheca elongate, more than 2 times longer than its maximal width (
<figureCitation id="1375FCBA5D56FF59FEB4FB66FE0AFB5A" box="[374,454,1176,1199]" captionStart="FIGURES 20 – 27" captionStartId="9.[151,274,1602,1626]" captionTargetBox="[366,1232,380,1576]" captionTargetId="figure@9.[360,1235,374,1586]" captionTargetPageId="9" captionText="FIGURES 20 – 27. Spermathecae of (20) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (21) Anillinus juliae, new species, (TN, McMinn Co., Starr Mtn.); (22) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (23) A. steevesi Barr (GA, Dade Co., Cloudland SP); (24) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.); (25) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (26) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (27) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 50 μm." httpUri="https://zenodo.org/record/195860/files/figure.png" pageId="14" pageNumber="15">Fig. 27</figureCitation>
). Parameres (
<figureCitation id="1375FCBA5D56FF59FD94FB66FD6FFB5A" box="[598,675,1176,1199]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,269,1554,1578]" captionTargetBox="[161,1426,363,1525]" captionTargetId="figure@8.[151,1436,343,1533]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Male aedeagus of (16) Anillinus erwini Sokolov &amp; Carlton, (NC, Avery Co., Grandfather Mtn.); (17) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (18) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (19) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest); left lateral aspect. a – median lobe, b – left paramere, c – right paramere. Scale = 100 μm." httpUri="https://zenodo.org/record/195859/files/figure.png" pageId="14" pageNumber="15">Fig. 19</figureCitation>
b, c) and stylomers as illustrated (
<figureCitation id="1375FCBA5D56FF59FBC8FB66FB95FB5A" box="[1034,1113,1176,1199]" captionStart="FIGURES 28 – 35" captionStartId="12.[151,269,1446,1470]" captionTargetBox="[175,1407,412,1421]" captionTargetId="figure@12.[172,1415,401,1429]" captionTargetPageId="12" captionText="FIGURES 28 – 35. Right stylomers and sternum IX of (28) Anillinus gimmeli, new species (TN, Blount Co., White Oak Sink); (29) A. juliae, new species, (TN, McMinn Co., Starr Mtn.); (30) A. merritti, new species, (NC, Swain Co., Twentymile Creek Trail); (31) A. steevesi Barr (GA, Dade Co., Cloudland SP); (32) A. erwini Sokolov &amp; Carlton (VA, Smyth Co., Mt. Rogers); (33) A. fortis (Horn), (NC, McDowell Co., Deerlick Gap Overlook); (34) A. loweae Sokolov &amp; Carlton, (NC, Haywood Co., Cataloochee Divide Trail); (35) A. cherokee Sokolov &amp; Carlton (NC, Graham Co., Nantahala National Forest). Scale = 100 μm." httpUri="https://zenodo.org/record/195861/files/figure.png" pageId="14" pageNumber="15">Fig. 35</figureCitation>
). Beetles (
<figureCitation id="1375FCBA5D56FF59FB0BFB66FAC5FB5A" box="[1225,1289,1176,1199]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,273,1846,1870]" captionTargetBox="[280,1310,185,1810]" captionTargetId="figure@4.[266,1325,178,1839]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Habitus of (1) A. gimmeli, new species (TN, Blount Co., GSMNP), (2) A. juliae, new species (TN, McMinn Co., Starr Mtn.), (3) A. merritti, new species (NC, Swain Co., GSMNP), (4) A. steevesi Barr (GA, Dade Co., Cloudland Canyon State Park), (5) A. steevesi Barr (TN, Blount Co., Chilhowee Mtn.), (6) A. erwini Sokolov &amp; Carlton (NC, Avery Co., Grandfather Mtn.), (7) A. fortis (Horn) (NC, McDowell Co., Deerlick Gap Overlook), (8) A. loweae Sokolov &amp; Carlton (NC, Haywood Co., GSMNP), (9) A. cherokee Sokolov &amp; Carlton (TN, Blount Co., GSMNP), dorsal aspect." httpUri="https://zenodo.org/record/195856/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
) from middle and high altitudes (
<quantity id="4CB64DDA5D56FF59FE58FB46FDD4FB3A" box="[410,536,1208,1231]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="14" pageNumber="15" unit="m" value="950.0" valueMax="1300.0" valueMin="600.0">600-1300m</quantity>
) from Laurel Falls (eastern Sevier County, Tennessee) and Gregory Bald areas of GSMNP south-east to the north-western corner of South Carolina.
<emphasis id="B93A3C2D5D56FF59FC48FB26FB06FB1A" box="[906,1226,1240,1263]" italics="true" pageId="14" pageNumber="15">
.................................
<taxonomicName id="4C4E9BBC5D56FF59FBA9FB26FB06FB1A" ID-CoL="5V288" authority="Sokolov &amp; Carlton" authorityName="Sokolov &amp; Carlton" box="[1131,1226,1240,1263]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cherokee">cherokee</taxonomicName>
</emphasis>
Sokolov &amp; Carlton
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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