1473 lines
216 KiB
XML
1473 lines
216 KiB
XML
<document id="33322B9A6D44A12A7283AD8C62D422E8" ID-DOI="10.11646/megataxa.11.1.1" ID-ISSN="2703-3090" ID-Zenodo-Dep="10993768" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713461317092" checkinUser="felipe" docAuthor="Kerr, Isaac A. R., Camens, Aaron B., Van Zoelen, Jacob D., Worthy, Trevor H. & Prideaux, Gavin J." docDate="2024" docId="03E587FDFF77D59DFF00770CFDD5FCA9" docLanguage="en" docName="Megataxa.11.1.001-261.pdf" docOrigin="Megataxa 11 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Protemnodon dawsonae Kerr & Camens & Van Zoelen & Worthy & Prideaux 2024, sp. nov." docType="treatment" docUuid="231A7C0D-FF39-458E-8703-1AF7B6FBCE77" docUuidSource="ZooBank" docVersion="4" lastPageNumber="169" masterDocId="FFDCFF85FFECD535FF887446FFD0FFEC" masterDocTitle="Systematics and palaeobiology of kangaroos of the late Cenozoic genus Protemnodon (Marsupialia, Macropodidae)" masterLastPageNumber="261" masterPageNumber="1" pageNumber="156" updateTime="1713463467402" updateUser="ExternalLinkService">
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<mods:title id="F68841485B19388BB8676E457F2C8D4B">Systematics and palaeobiology of kangaroos of the late Cenozoic genus Protemnodon (Marsupialia, Macropodidae)</mods:title>
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<mods:name id="C74D4F43CB289594E720D4C04D3CA3CF" type="personal">
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<mods:namePart id="F0DC853A990A88E7E051FCB337E03983">Kerr, Isaac A. R.</mods:namePart>
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<mods:affiliation id="59181C03C4367D7D3AF66FF78A188C64">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:roleTerm id="9FCEA6A2F70D40D2E22FE64E519D500D">Author</mods:roleTerm>
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<mods:namePart id="1A187225864A5B7206389ADDC5C94300">Camens, Aaron B.</mods:namePart>
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<mods:nameIdentifier id="1CE5BDCCD5F740B740FEE4D17F8E9A9E" type="ORCID">0000-0003-0464-0665</mods:nameIdentifier>
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<mods:affiliation id="DDDC013009413D58A7070F13B51CF6F6">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:roleTerm id="83DBBA3239649AE96D625542FFBCDAB4">Author</mods:roleTerm>
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<mods:namePart id="AC34C86D0A0C1C20A89525E6ECDF9483">Van Zoelen, Jacob D.</mods:namePart>
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<mods:nameIdentifier id="0CB292CA4D2F8E4B01470E0D4A5994E6" type="ORCID">0000-0003-2952-8549</mods:nameIdentifier>
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<mods:affiliation id="4AD54567DD77C9646357D4D8A0A8D462">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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</mods:role>
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<mods:namePart id="9AB76D434FE6CA441392C4A13AA3C83A">Worthy, Trevor H.</mods:namePart>
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<mods:nameIdentifier id="C6E4DCA3BFA83DD7778AE430E92922BB" type="ORCID">0000-0001-7047-4680</mods:nameIdentifier>
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<mods:affiliation id="71A16DB527FD47D01B40C3A722C41FB6">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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</mods:role>
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<mods:namePart id="FDCCEB864CEB6556055F45ED37F5088C">Prideaux, Gavin J.</mods:namePart>
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<mods:nameIdentifier id="C81301CD216D1732F7FB033162507BD5" type="ORCID">0000-0002-9958-0265</mods:nameIdentifier>
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<mods:affiliation id="4EFBD126061D877966312D0989FAC647">College of Science and Engineering, Flinders University, Bedford Park, South Australia, Australia, 5042</mods:affiliation>
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<mods:typeOfResource id="65190814F7552FEF0B3880DA085FF24D">text</mods:typeOfResource>
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<mods:title id="8E9EAD424680C574A804163BE219A6E5">Megataxa</mods:title>
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<mods:part id="D6627AC86AA357BDB893FC8BE5E0B317">
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<mods:date id="120300CD84F8C14D8F24C16529F15922">2024</mods:date>
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<mods:number id="4905BD1BDC742F1F06D458AA0FEC74FC">2024-04-15</mods:number>
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<mods:detail id="27042CE8A0EE2F381E545AA7E3008CCB" type="volume">
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<mods:number id="3D7345AA92A5BB79FD3158B90F645F65">11</mods:number>
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<mods:number id="36484535B58BAE3F5722187104A44850">1</mods:number>
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<mods:start id="26E2D9F9A13AB89E4788A391E510C557">1</mods:start>
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<mods:end id="0119678BB47C90CCF4C175584CFC20DA">261</mods:end>
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<mods:classification id="8C919A8DB4985FAF1713E33C128FA052">journal article</mods:classification>
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<mods:identifier id="F62A1B92CA1B63A7B5246CA491D6073C" type="DOI">10.11646/megataxa.11.1.1</mods:identifier>
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<mods:identifier id="1B97C04DFE2B98D0CDBE3152A33DBEE7" type="ISSN">2703-3090</mods:identifier>
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<mods:identifier id="71DD1328AEC387283AC98926C50B31B2" type="Zenodo-Dep">10993768</mods:identifier>
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<mods:identifier id="DC80045BE9E06CAE6EEDB39E3663480A" type="ZooBank">5F42E7FE-C154-4979-9691-E6F74BBBBC10</mods:identifier>
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<treatment id="03E587FDFF77D59DFF00770CFDD5FCA9" ID-DOI="http://doi.org/10.5281/zenodo.10993698" ID-Zenodo-Dep="10993698" LSID="urn:lsid:zoobank.org:act:231A7C0D-FF39-458E-8703-1AF7B6FBCE77" httpUri="http://treatment.plazi.org/id/03E587FDFF77D59DFF00770CFDD5FCA9" lastPageId="168" lastPageNumber="169" pageId="155" pageNumber="156">
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<subSubSection id="C3566560FF77D5AEFF00770CFDC0FC89" box="[136,528,842,869]" pageId="155" pageNumber="156" type="nomenclature">
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<paragraph id="8BF336EBFF77D5AEFF00770CFDC0FC89" blockId="155.[136,778,842,934]" box="[136,528,842,869]" pageId="155" pageNumber="156">
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<heading id="D0BB8187FF77D5AEFF00770CFDC0FC89" box="[136,528,842,869]" fontSize="11" level="2" pageId="155" pageNumber="156" reason="2">
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<emphasis id="B938EAF9FF77D5AEFF00770CFDC0FC89" bold="true" box="[136,528,842,869]" pageId="155" pageNumber="156">
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<taxonomicName id="4C4C4D68FF77D5AEFF00770CFE7BFC89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,427,842,869]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFF00770CFE7BFC89" bold="true" box="[136,427,842,869]" italics="true" pageId="155" pageNumber="156">Protemnodon dawsonae</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A20B5782FF77D5AEFE3A770DFDC0FC89" box="[434,528,843,869]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3566560FF77D5AEFF00772DFD17FC4A" pageId="155" pageNumber="156" type="description">
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<paragraph id="8BF336EBFF77D5AEFF00772DFD17FC4A" blockId="155.[136,778,842,934]" pageId="155" pageNumber="156">
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||
LSID of new species:
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urn:lsid:zoobank.org: act:
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<uuid id="FFEA0C3EFF77D5AEFF2677CDFD17FC4A" box="[174,711,907,934]" pageId="155" pageNumber="156">231A7C0D-FF39-458E-8703-1AF7B6FBCE77</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3566560FF77D5AEFF007788FF00FBA8" pageId="155" pageNumber="156" type="reference_group">
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||
<paragraph id="8BF336EBFF77D5AEFF007788FE2EFBE8" blockId="155.[136,779,974,1092]" pageId="155" pageNumber="156">
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||
<taxonomicName id="4C4C4D68FF77D5AEFF007788FE17FC08" authority="Bartholomai" authorityName=", Bartholomai" authorityYear="1973" box="[136,455,974,996]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF77D5AEFF007788FE97FC0F" box="[136,327,974,995]" italics="true" pageId="155" pageNumber="156">Protemnodon devisi</emphasis>
|
||
Bartholomai
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFDD4B1AFF77D5AEFE477788FD5CFC09" author="Bartholomai, A." box="[463,652,974,997]" pageId="155" pageNumber="156" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai (1973)</bibRefCitation>
|
||
pl. 22, fig. 3. Not
|
||
<taxonomicName id="4C4C4D68FF77D5AEFF6F77A9FE2AFBE8" authority=", Bartholomai, 1973" authorityName=", Bartholomai" authorityYear="1973" box="[231,506,1006,1028]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF77D5AEFF6F77A9FEE7FBEF" box="[231,311,1006,1028]" italics="true" pageId="155" pageNumber="156">P. devisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFDD4B1AFF77D5AEFECB77A8FE2AFBE8" author="Bartholomai, A." box="[323,506,1006,1028]" pageId="155" pageNumber="156" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
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||
<paragraph id="8BF336EBFF77D5AEFF007048FF00FBA8" blockId="155.[136,779,974,1092]" pageId="155" pageNumber="156">
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||
<taxonomicName id="4C4C4D68FF77D5AEFF007048FEFCFBC8" box="[136,300,1038,1060]" class="Mammalia" family="Macropodidae" genus="Protemnodon" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B938EAF9FF77D5AEFF007048FEDAFBCF" box="[136,266,1038,1059]" italics="true" pageId="155" pageNumber="156">Protemnodon</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
<taxonomicName id="4C4C4D68FF77D5AEFEB97048FE58FBCF" authorityName=", Bartholomai" authorityYear="1973" box="[305,392,1038,1060]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="devisi">
|
||
cf.
|
||
<emphasis id="B938EAF9FF77D5AEFED87048FE58FBCF" box="[336,392,1038,1059]" italics="true" pageId="155" pageNumber="156">devisi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFDD4B1AFF77D5AEFE1C7049FD8FFBC9" author="Dawson, L. & Muirhead, J. & Wroe, S." box="[404,607,1038,1061]" pageId="155" pageNumber="156" pagination="265 - 290" refId="ref145233" refString="Dawson, L., Muirhead, J. & Wroe, S. (1999) The Big Sink Local Fauna: a lower Pliocene mammalian fauna from the Wellington Caves complex, Wellington, New South Wales. Records of the Western Australian Museum, Supplement No. 57, 265 - 290." type="book chapter" year="1999">
|
||
Dawson
|
||
<emphasis id="B938EAF9FF77D5AEFE617049FDCAFBCF" box="[489,538,1038,1060]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
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||
(1999)
|
||
</bibRefCitation>
|
||
pp. 276–278, fig. 7.
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C3566560FF77D5AEFF00702DFB51FD29" pageId="155" pageNumber="156" type="materials_examined">
|
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<paragraph id="8BF336EBFF77D5AEFF00702DFEBDFAE9" blockId="155.[136,778,1131,1285]" pageId="155" pageNumber="156">
|
||
<materialsCitation id="3B243CB6FF77D5AEFF00702DFEBDFAE9" collectionCode="AM" latitude="-32.6" location="Big Sink doline, Bone Cave - Phosphate Mine system, Wellington Caves" longLatPrecision="1209" longitude="148.91667" pageId="155" pageNumber="156" specimenCode="F161915" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
|
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<emphasis id="B938EAF9FF77D5AEFF00702DFED4FB69" bold="true" box="[136,260,1131,1157]" pageId="155" pageNumber="156">
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<typeStatus id="54F78849FF77D5AEFF00702DFF2FFB69" box="[136,255,1131,1157]" pageId="155" pageNumber="156" type="holotype">Holotype</typeStatus>
|
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:
|
||
</emphasis>
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF0070CDFF65FB49" box="[136,181,1163,1189]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
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<specimenCode id="DBEA9E90FF77D5AEFF4070CDFEFCFB4A" box="[200,300,1163,1190]" pageId="155" pageNumber="156">F161915</specimenCode>
|
||
R partial cranium, preserving M1–4, maxilla, and partial lacrimal, frontal, palatine, and jugal. Collected on an excavation at Big Sink by Prideaux and Fusco
|
||
<emphasis id="B938EAF9FF77D5AEFF5A70AAFEDAFAE9" box="[210,266,1259,1285]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
|
||
in 2016.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFF00716DFEE2FAA8" blockId="155.[136,778,1322,1605]" box="[136,306,1322,1349]" pageId="155" pageNumber="156">
|
||
<emphasis id="B938EAF9FF77D5AEFF00716DFEE2FAA8" bold="true" box="[136,306,1322,1349]" pageId="155" pageNumber="156">Type locality:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFF00710DFD5CF9A9" blockId="155.[136,778,1322,1605]" pageId="155" pageNumber="156">
|
||
<location id="8E936030FF77D5AEFF00710DFE81FA69" LSID="urn:lsid:plazi:treatment:03E587FDFF77D59DFF00770CFDD5FCA9:8E936030FF77D5AEFF00710DFE81FA69" latitude="-32.6" longLatPrecision="1209" longitude="148.91667" name="Big Sink doline, Bone Cave - Phosphate Mine system, Wellington Caves" pageId="155" pageNumber="156" stateProvince="New South Wales">Big Sink doline, Bone Cave–Phosphate Mine system, Wellington Caves</location>
|
||
(
|
||
<geoCoordinate id="EE78502CFF77D5AEFEEC712DFE19FA69" box="[356,457,1387,1413]" degrees="32" direction="south" minutes="36" orientation="latitude" pageId="155" pageNumber="156" precision="925" value="-32.6">32°36’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE78502CFF77D5AEFE50712DFD9EFA69" box="[472,590,1387,1414]" degrees="148" direction="east" minutes="55" orientation="longitude" pageId="155" pageNumber="156" precision="925" value="148.91667">148°55’ E</geoCoordinate>
|
||
), central eastern
|
||
<collectingRegion id="4988F809FF77D5AEFF0071CDFE86FA49" box="[136,342,1419,1445]" country="Australia" name="New South Wales" pageId="155" pageNumber="156">New South Wales</collectingRegion>
|
||
. The Big Sink unit is made up of pale red-orange, consolidated sediments on the southern wall of the Big Sink doline (
|
||
<bibRefCitation id="EFDD4B1AFF77D5AEFE46718DFDA4FA09" author="Osborne, R. A. L." box="[462,628,1483,1509]" pageId="155" pageNumber="156" pagination="175 - 180" refId="ref152711" refString="Osborne, R. A. L. (1997) Rehabilitation of the Wellington Caves Phosphate Mine: implications for Cainozoic stratigraphy. Proceedings of the Linnean Society of New South Wales, 117, 175 - 180." type="journal article" year="1997">Osborne 1997</bibRefCitation>
|
||
). Though the deposit has not been dated directly, biochronology and biocorrelation with dated assemblages suggest a Pliocene age for the Big Sink LF (
|
||
<bibRefCitation id="EFDD4B1AFF77D5AEFE28726DFDAFF9A9" author="Dawson, L. & Muirhead, J. & Wroe, S." box="[416,639,1579,1605]" pageId="155" pageNumber="156" pagination="265 - 290" refId="ref145233" refString="Dawson, L., Muirhead, J. & Wroe, S. (1999) The Big Sink Local Fauna: a lower Pliocene mammalian fauna from the Wellington Caves complex, Wellington, New South Wales. Records of the Western Australian Museum, Supplement No. 57, 265 - 290." type="book chapter" year="1999">
|
||
Dawson
|
||
<emphasis id="B938EAF9FF77D5AEFD8C726AFDEDF9A9" box="[516,573,1579,1605]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFF00722DFEDFF969" blockId="155.[136,779,1643,2085]" box="[136,271,1643,1669]" pageId="155" pageNumber="156">
|
||
<emphasis id="B938EAF9FF77D5AEFF00722DFEDFF969" bold="true" box="[136,271,1643,1669]" pageId="155" pageNumber="156">
|
||
<typeStatus id="54F78849FF77D5AEFF00722DFEDAF969" box="[136,266,1643,1669]" pageId="155" pageNumber="156" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFF0072CDFAC9FF09" blockId="155.[136,779,1643,2085]" lastBlockId="155.[810,1451,171,229]" pageId="155" pageNumber="156">
|
||
Big Sink doline:
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEC672CDFEABF949" box="[334,379,1675,1701]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69835 partial L premaxilla, with I1–I3;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF5A72EDFF2FF929" box="[210,255,1707,1733]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69836 L premaxilla fragment;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFDE172EDFD46F929" box="[617,662,1707,1733]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69838 L I1;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF24728DFF09F909" box="[172,217,1739,1765]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69839
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEB4728DFE9FF909" box="[316,335,1739,1765]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
I1;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEF1728DFE76F909" box="[377,422,1739,1765]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69840b partial L maxilla;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFD55728DFCDAF909" box="[733,778,1739,1765]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69841 partial L maxilla;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFE3872ADFE0DF8E9" box="[432,477,1771,1797]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69845 partial
|
||
<collectionCode id="ED5DAE2EFF77D5AEFD1B72ADFD76F8E9" box="[659,678,1771,1797]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
maxilla;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF00734DFF65F8C9" box="[136,181,1803,1829]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69842 L P3;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEE5734DFE4AF8C9" box="[365,410,1803,1829]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69844 L P3;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFDDA734DFDAFF8C9" box="[594,639,1803,1829]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161923 L P3;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF32736DFF37F8A9" box="[186,231,1835,1861]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69858 &
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEF5736DFE7AF8A9" box="[381,426,1835,1861]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F65859 partial LR dentaries (reassociated);
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEB8730DFE8DF889" box="[304,349,1867,1893]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69860 partial L dentary;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFD09730DFD7EF889" box="[641,686,1867,1893]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69868 L i1;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF49732DFF3EF869" box="[193,238,1899,1925]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F69869
|
||
<collectionCode id="ED5DAE2EFF77D5AEFED8732DFEB3F869" box="[336,355,1899,1925]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
i1;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFE04732DFE69F869" box="[396,441,1899,1925]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161921
|
||
<collectionCode id="ED5DAE2EFF77D5AEFDA1732DFDECF869" box="[553,572,1899,1925]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
dentary fragment;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF0073CDFF65F849" box="[136,181,1931,1957]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161920 partial axis vertebra;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFD9773CDFD9CF849" box="[543,588,1931,1957]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161914 partial axis vertebra;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEA073EDFE85F829" box="[296,341,1963,1989]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161918 partial lumbar vertebra L2?;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF00738DFF65F809" box="[136,181,1995,2021]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161911 caudal vertebra Ca2?;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFDA9738DFD9EF809" box="[545,590,1995,2021]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161912 caudal vertebra Ca3?;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEB973ADFE8EF7E9" box="[305,350,2027,2053]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161919 ilial fragment;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFDFB73ADFD70F7E9" box="[627,672,2027,2053]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161913 partial
|
||
<collectionCode id="ED5DAE2EFF77D5AEFF5E7C4DFF39F7C9" box="[214,233,2059,2085]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
calcaneus;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFEE27C4DFE47F7C9" box="[362,407,2059,2085]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161917 partial L metatarsal
|
||
<collectionCode id="ED5DAE2EFF77D5AEFD6F7C4DFCD6F7C9" box="[743,774,2059,2085]" pageId="155" pageNumber="156">IV</collectionCode>
|
||
;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFCA274EDFC87FF29" box="[810,855,171,197]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F109039
|
||
<collectionCode id="ED5DAE2EFF77D5AEFC4074EDFC0BFF29" box="[968,987,171,197]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
metatarsal
|
||
<collectionCode id="ED5DAE2EFF77D5AEFBD374EDFBAAFF29" box="[1115,1146,171,197]" pageId="155" pageNumber="156">IV</collectionCode>
|
||
;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFB0B74EDFB60FF29" box="[1155,1200,171,197]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F106040 L metatarsal
|
||
<collectionCode id="ED5DAE2EFF77D5AEFCA2748DFC90FF09" box="[810,832,203,229]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="155" pageNumber="156" type="Museum">V</collectionCode>
|
||
;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFCC1748DFCA6FF09" box="[841,886,203,229]" country="Australia" name="Australian Museum" pageId="155" pageNumber="156" type="Museum">AM</collectionCode>
|
||
F161916 proximal pedal phalanx
|
||
<collectionCode id="ED5DAE2EFF77D5AEFB7F748DFAC5FF09" box="[1271,1301,203,229]" pageId="155" pageNumber="156">IV</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA2754CFBFAFEC8" blockId="155.[810,1066,266,292]" box="[810,1066,266,292]" pageId="155" pageNumber="156">
|
||
<emphasis id="B938EAF9FF77D5AEFCA2754CFBFAFEC8" bold="true" box="[810,1066,266,292]" pageId="155" pageNumber="156">Referred specimens:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA2750DFC30FE89" blockId="155.[810,1452,331,518]" box="[810,992,331,357]" pageId="155" pageNumber="156">
|
||
<emphasis id="B938EAF9FF77D5AEFCA2750DFC30FE89" box="[810,992,331,357]" italics="true" pageId="155" pageNumber="156">
|
||
<collectingRegion id="4988F809FF77D5AEFCA2750DFC30FE89" box="[810,992,331,357]" country="Australia" name="South Australia" pageId="155" pageNumber="156">South Australia</collectingRegion>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA2752DFC28FE29" blockId="155.[810,1452,331,518]" pageId="155" pageNumber="156">
|
||
Stirton Quarry, Lake Kanunka:
|
||
<collectionCode id="ED5DAE2EFF77D5AEFB2C752DFB25FE69" box="[1188,1269,363,389]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="155" pageNumber="156" type="Museum">SAMA</collectionCode>
|
||
P50399 partial juvenile
|
||
<collectionCode id="ED5DAE2EFF77D5AEFC4275CDFC0DFE49" box="[970,989,395,421]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
maxilla with P3 from crypt;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFAD575CDFA7CFE49" box="[1373,1452,395,421]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="155" pageNumber="156" type="University or college">UCMP</collectionCode>
|
||
322655
|
||
<collectionCode id="ED5DAE2EFF77D5AEFC3275EDFC1DFE29" box="[954,973,427,453]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
P3.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA2758DFA44FDE9" blockId="155.[810,1452,331,518]" pageId="155" pageNumber="156">
|
||
Site 4 (
|
||
<collectionCode id="ED5DAE2EFF77D5AEFC0F758DFC4CFE0A" box="[903,924,459,486]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="155" pageNumber="156" type="Museum">V</collectionCode>
|
||
77015), Lake Kanunka:
|
||
<collectionCode id="ED5DAE2EFF77D5AEFB4A758DFAC1FE09" box="[1218,1297,459,485]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="155" pageNumber="156" type="University or college">UCMP</collectionCode>
|
||
<quantity id="4CB49B0EFF77D5AEFA96758DFA48FE09" box="[1310,1432,459,485]" metricMagnitude="5" metricUnit="m" metricValue="1.7067799999999997" pageId="155" pageNumber="156" unit="m" value="170678.0">170678 m</quantity>
|
||
4;
|
||
<collectionCode id="ED5DAE2EFF77D5AEFCD775ADFC7EFDE9" box="[863,942,491,517]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="155" pageNumber="156" type="University or college">UCMP</collectionCode>
|
||
156881
|
||
<collectionCode id="ED5DAE2EFF77D5AEFB8775ADFBF2FDE9" box="[1039,1058,491,517]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="155" pageNumber="156">R</collectionCode>
|
||
ilium, partial pubis and ischium.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA2766DFC60FDA9" blockId="155.[810,1452,555,709]" box="[810,944,555,581]" pageId="155" pageNumber="156">
|
||
<emphasis id="B938EAF9FF77D5AEFCA2766DFC60FDA9" box="[810,944,555,581]" italics="true" pageId="155" pageNumber="156">
|
||
<collectingRegion id="4988F809FF77D5AEFCA2766DFC60FDA9" box="[810,944,555,581]" country="Australia" name="Queensland" pageId="155" pageNumber="156">Queensland</collectingRegion>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA2760DFC69FD69" blockId="155.[810,1452,555,709]" pageId="155" pageNumber="156">
|
||
Sand Scree Locality, Chinchilla:
|
||
<collectionCode id="ED5DAE2EFF77D5AEFB3E760DFB33FD89" box="[1206,1251,587,613]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="155" pageNumber="156" type="Museum">QM</collectionCode>
|
||
F7071 partial L dentary.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA276CDFB51FD29" blockId="155.[810,1452,555,709]" pageId="155" pageNumber="156">
|
||
Chinchilla, Darling Downs (site unknown):
|
||
<collectionCode id="ED5DAE2EFF77D5AEFAA376CDFA88FD49" box="[1323,1368,651,677]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="155" pageNumber="156" type="Museum">QM</collectionCode>
|
||
F4712 partial juvenile L maxilla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF77D5A9FCA276ACFE9DF9A9" lastPageId="156" lastPageNumber="157" pageId="155" pageNumber="156" type="diagnosis">
|
||
<paragraph id="8BF336EBFF77D5AEFCA276ACFBC0FCE8" blockId="155.[810,1452,746,2085]" box="[810,1040,746,772]" pageId="155" pageNumber="156">
|
||
<emphasis id="B938EAF9FF77D5AEFCA276ACFBC0FCE8" bold="true" box="[810,1040,746,772]" pageId="155" pageNumber="156">Specific diagnosis:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCA2774DFBBBFBA9" blockId="155.[810,1452,746,2085]" pageId="155" pageNumber="156">
|
||
<taxonomicName id="4C4C4D68FF77D5AEFCA2774DFBE7FCC9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[810,1079,779,805]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFCA2774DFBE7FCC9" box="[810,1079,779,805]" italics="true" pageId="155" pageNumber="156">Protemnodon dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFBB6774DFB47FCC9" bold="true" box="[1086,1175,779,805]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFBB6774DFB47FCC9" box="[1086,1175,779,805]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is differentiated from all other species of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFC50776DFBBFFCA9" authorityName="Owen" authorityYear="1874" box="[984,1135,811,837]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF77D5AEFC50776DFBBFFCA9" box="[984,1135,811,837]" italics="true" pageId="155" pageNumber="156">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
by one dental autapomorphy and a unique combination of dental and postcranial attributes. The upper dentition of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB3C772AFA93FC69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1204,1347,875,901]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFB3C772AFA93FC69" box="[1204,1347,875,901]" italics="true" pageId="155" pageNumber="156">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFAC6772DFA7BFC69" bold="true" box="[1358,1451,875,901]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFAC6772DFA7BFC69" box="[1358,1451,875,901]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of all species of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB2B77CDFAEAFC49" authorityName="Owen" authorityYear="1874" box="[1187,1338,907,933]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF77D5AEFB2B77CDFAEAFC49" box="[1187,1338,907,933]" italics="true" pageId="155" pageNumber="156">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
in having upper molars with the postparacrista distinctly inflected lingually then buccally (‘kinked’) in the interloph valley. The i1 of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFC1077AAFBF4FBE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[920,1060,1003,1029]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFC1077AAFBF4FBE9" box="[920,1060,1003,1029]" italics="true" pageId="155" pageNumber="156">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFBA377ADFB54FBE9" bold="true" box="[1067,1156,1003,1029]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFBA377ADFB54FBE9" box="[1067,1156,1003,1029]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all species of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFCA2704DFC11FBC9" authorityName="Owen" authorityYear="1874" box="[810,961,1035,1061]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF77D5AEFCA2704DFC11FBC9" box="[810,961,1035,1061]" italics="true" pageId="155" pageNumber="156">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB97704AFB70FBC9" box="[1055,1184,1035,1061]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF77D5AEFB97704AFB70FBC9" box="[1055,1184,1035,1061]" italics="true" pageId="155" pageNumber="156">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having a thick, low ventrolingual crest on the i1.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCD2700DFBDCF9C9" blockId="155.[810,1452,746,2085]" pageId="155" pageNumber="156">
|
||
The dentary and dentition of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB3B700AFA93FB89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1203,1347,1099,1125]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFB3B700AFA93FB89" box="[1203,1347,1099,1125]" italics="true" pageId="155" pageNumber="156">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFAC6700DFA7BFB89" bold="true" box="[1358,1451,1099,1125]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFAC6700DFA7BFB89" box="[1358,1451,1099,1125]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are most similar to those of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFBD6702AFB37FB69" authorityName="Plane" authorityYear="1967" box="[1118,1255,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF77D5AEFBD6702AFB37FB69" box="[1118,1255,1131,1157]" italics="true" pageId="155" pageNumber="156">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB7C702AFA54FB69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1268,1412,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFB7C702AFA54FB69" box="[1268,1412,1131,1157]" italics="true" pageId="155" pageNumber="156">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFA02702DFC8BFB49" bold="true" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFA02702DFC8BFB49" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF77D5AEFC0770CAFC3DFB49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[911,1005,1164,1189]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFC0770CAFC3DFB49" box="[911,1005,1164,1189]" italics="true" pageId="155" pageNumber="156">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFC7B70CDFB9BFB49" bold="true" box="[1011,1099,1163,1189]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFC7B70CDFB9BFB49" box="[1011,1099,1163,1189]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The dentary of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB7170CAFA54FB49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1273,1412,1163,1189]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFB7170CAFA54FB49" box="[1273,1412,1163,1189]" italics="true" pageId="155" pageNumber="156">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFA0270CDFC8BFB29" bold="true" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFA0270CDFC8BFB29" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
cannot be differentiated from that of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB7D70EAFA55FB29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1269,1413,1195,1221]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFB7D70EAFA55FB29" box="[1269,1413,1195,1221]" italics="true" pageId="155" pageNumber="156">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFA0270EDFC8BFB09" bold="true" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFA0270EDFC8BFB09" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
or
|
||
<taxonomicName id="4C4C4D68FF77D5AEFCF4708AFC09FB09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[892,985,1228,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFCF4708AFC09FB09" box="[892,985,1228,1253]" italics="true" pageId="155" pageNumber="156">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFC56708DFBE6FB09" bold="true" box="[990,1078,1227,1253]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFC56708DFBE6FB09" box="[990,1078,1227,1253]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but it differs from
|
||
<taxonomicName id="4C4C4D68FF77D5AEFA80708AFA40FB09" authorityName="Plane" authorityYear="1967" box="[1288,1424,1227,1253]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF77D5AEFA80708AFA40FB09" box="[1288,1424,1227,1253]" italics="true" pageId="155" pageNumber="156">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in having a taller, more robust anterior dentary, particularly around the i1. The dentition further differs from that of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFA1F714AFC48FAA9" authorityName="Plane" authorityYear="1967" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF77D5AEFA1F714AFC48FAA9" italics="true" pageId="155" pageNumber="156">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in having: a broader, anteroposteriorly shorter I1; relatively larger, broader and more robust i1; and a broader p3 relative to length. It further differs from that of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFCC371CAFC0EFA49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[843,990,1419,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFCC371CAFC0EFA49" box="[843,990,1419,1445]" italics="true" pageId="155" pageNumber="156">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFC6F71CDFB93FA49" bold="true" box="[999,1091,1419,1445]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFC6F71CDFB93FA49" box="[999,1091,1419,1445]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF77D5AEFBF671CAFB0FFA49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1150,1247,1420,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFBF671CAFB0FFA49" box="[1150,1247,1420,1445]" italics="true" pageId="155" pageNumber="156">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFB6071CDFA94FA49" bold="true" box="[1256,1348,1419,1445]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFB6071CDFA94FA49" box="[1256,1348,1419,1445]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being lower crowned, with: I1 absolutely narrower and narrower relative to I3; and upper molars with a narrower protoloph relative to the loph base when unworn, and a thinner, more raised preparacrista.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5AEFCD2726DFBCEF889" blockId="155.[810,1452,746,2085]" pageId="155" pageNumber="156">
|
||
The upper dentition of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFBEE726AFB24F9A9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1126,1268,1579,1605]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFBEE726AFB24F9A9" box="[1126,1268,1579,1605]" italics="true" pageId="155" pageNumber="156">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFB75726DFA89F9A9" bold="true" box="[1277,1369,1579,1605]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFB75726DFA89F9A9" box="[1277,1369,1579,1605]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFC35720AFBE0F989" authorityName="Bartholomai" authorityYear="1978" box="[957,1072,1612,1637]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF77D5AEFC35720AFBE0F989" box="[957,1072,1612,1637]" italics="true" pageId="155" pageNumber="156">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in having: relatively broader P3; and molars with a higher, more distinct preparacrista. The dentary differs from that of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB1A72CAFAD3F949" authorityName="Bartholomai" authorityYear="1978" box="[1170,1283,1676,1701]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF77D5AEFB1A72CAFAD3F949" box="[1170,1283,1676,1701]" italics="true" pageId="155" pageNumber="156">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being larger and more robust, with: a more robust and more dorsally deflected diastema, and a taller mandibular corpus. The lower dentition differs from that of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB4572AAFA92F8E9" authorityName="Bartholomai" authorityYear="1978" box="[1229,1346,1772,1797]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF77D5AEFB4572AAFA92F8E9" box="[1229,1346,1772,1797]" italics="true" pageId="155" pageNumber="156">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being larger and higher crowned, with: broader, more robust i1; relatively broader p3; and relatively broader molars with a smaller precingulid.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF77D5A9FCD2732DFE70FD29" blockId="155.[810,1452,746,2085]" lastBlockId="156.[136,778,171,1605]" lastPageId="156" lastPageNumber="157" pageId="155" pageNumber="156">
|
||
The postcranial skeleton of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB13732AFAF9F869" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1179,1321,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFB13732AFAF9F869" box="[1179,1321,1899,1925]" italics="true" pageId="155" pageNumber="156">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFABB732DFA5FF869" bold="true" box="[1331,1423,1899,1925]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFABB732DFA5FF869" box="[1331,1423,1899,1925]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to that of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFBB373CAFB1FF849" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1083,1231,1931,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="mamkurra">
|
||
<emphasis id="B938EAF9FF77D5AEFBB373CAFB1FF849" box="[1083,1231,1931,1957]" italics="true" pageId="155" pageNumber="156">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF77D5AEFA8473CAFABFF849" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1292,1391,1932,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="viator">
|
||
<emphasis id="B938EAF9FF77D5AEFA8473CAFABFF849" box="[1292,1391,1932,1957]" italics="true" pageId="155" pageNumber="156">P. viator</emphasis>
|
||
</taxonomicName>
|
||
. The proximal phalanx IV of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFBCC73EAFB03F829" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1092,1235,1963,1989]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="dawsonae">
|
||
<emphasis id="B938EAF9FF77D5AEFBCC73EAFB03F829" box="[1092,1235,1963,1989]" italics="true" pageId="155" pageNumber="156">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
differs from those of all species of
|
||
<taxonomicName id="4C4C4D68FF77D5AEFC66738DFB55F809" authorityName="Owen" authorityYear="1874" box="[1006,1157,1995,2021]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF77D5AEFC66738DFB55F809" box="[1006,1157,1995,2021]" italics="true" pageId="155" pageNumber="156">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C4C4D68FF77D5AEFB6B738AFA95F809" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1251,1349,1996,2021]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="155" pageNumber="156" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF77D5AEFB6B738AFA95F809" box="[1251,1349,1996,2021]" italics="true" pageId="155" pageNumber="156">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF77D5AEFAC7738DFA7BF809" bold="true" box="[1359,1451,1995,2021]" pageId="155" pageNumber="156">
|
||
<taxonomicNameLabel id="A20B5782FF77D5AEFAC7738DFA7BF809" box="[1359,1451,1995,2021]" pageId="155" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a more proximodorsally extensive distal articular surface. Its axial skeleton is differentiated from
|
||
<taxonomicName id="4C4C4D68FF70D5A9FF0074EAFECCFF29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,284,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FF0074EAFECCFF29" box="[136,284,171,197]" italics="true" pageId="156" pageNumber="157">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FEAE74EDFE54FF29" bold="true" box="[294,388,171,197]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FEAE74EDFE54FF29" box="[294,388,171,197]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having: a relatively narrower, deeper axis vertebra; lumbar vertebrae with deeper transverse processes and a centrum with relatively slightly broader cranial and caudal extremities, more deeply concave ventrolateral surfaces, a more deeply concave ventral margin in lateral view, and a less raised ventral ridge. The hindlimb differs from that of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FDD8752AFD31FE69" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[592,737,363,389]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FDD8752AFD31FE69" box="[592,737,363,389]" italics="true" pageId="156" pageNumber="157">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FD60752DFF69FE49" bold="true" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FD60752DFF69FE49" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a pelvis with a relatively smaller caudal iliac spine, less deeply concave gluteal fossa, and a less convex, less rugose caudal margin of the dorsal section of the ischium. The pes differs in having: a relatively taller, narrower calcaneus with a caudally displaced medial talar facet relative to the lateral talar facet and a sustentaculum tali with a rounded (rather than pointed) cranioplantar peak; metatarsal IV with a smaller plantar cuboid facet; and narrower metatarsal V with a more distally extensive lateral plantar tuberosity.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FF30768DFE23FAE9" blockId="156.[136,778,171,1605]" pageId="156" pageNumber="157">
|
||
The axial skeleton of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FE5F768AFDBEFD09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[471,622,715,741]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FE5F768AFDBEFD09" box="[471,622,715,741]" italics="true" pageId="156" pageNumber="157">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FD08768DFD35FD09" bold="true" box="[640,741,715,741]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FD08768DFD35FD09" box="[640,741,715,741]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from that of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FE4876AAFDF2FCE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[448,546,748,773]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FE4876AAFDF2FCE9" box="[448,546,748,773]" italics="true" pageId="156" pageNumber="157">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FDA576ADFD5AFCE9" bold="true" box="[557,650,747,773]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FDA576ADFD5AFCE9" box="[557,650,747,773]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having: a narrower axis vertebra with a thinner spinous process, more laterally tilted postzygopophyses with less concave articular surfaces, and a rounded (rather than bilobed) ventral margin of the caudal extremity of the centrum; and shorter, more robust caudal vertebrae. The hindlimb differs from
|
||
<taxonomicName id="4C4C4D68FF70D5A9FE9277EAFEABFC29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[282,379,940,965]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FE9277EAFEABFC29" box="[282,379,940,965]" italics="true" pageId="156" pageNumber="157">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FE0D77EDFE31FC29" bold="true" box="[389,481,939,965]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FE0D77EDFE31FC29" box="[389,481,939,965]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a pelvis with a less deeply concave gluteal fossa. The pes of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FDF7778AFCDAFC09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[639,778,971,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FDF7778AFCDAFC09" box="[639,778,971,997]" italics="true" pageId="156" pageNumber="157">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FF0077ADFF33FBE9" bold="true" box="[136,227,1003,1029]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FF0077ADFF33FBE9" box="[136,227,1003,1029]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FE4577AAFDFCFBE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[461,556,1004,1029]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FE4577AAFDFCFBE9" box="[461,556,1004,1029]" italics="true" pageId="156" pageNumber="157">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FDBD77ADFD5FFBE9" bold="true" box="[565,655,1003,1029]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FDBD77ADFD5FFBE9" box="[565,655,1003,1029]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having: a broader, dorsoplantarly shorter calcaneus with a more rounded calcaneal tuberosity in cross-section, less deep lateral talar facet, broader medial talar facet, and a more medially projected sustentaculum tali; relatively broader, dorsoplantarly shorter metatarsal V with a more elongate proximolateral process and a more proximally projected medial plantar tubercle; and a more robust proximal phalanx IV with a weaker waist.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FF30714DFE9DF9A9" blockId="156.[136,778,171,1605]" pageId="156" pageNumber="157">
|
||
The pes of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FEBF714AFE14FAC9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[311,452,1291,1317]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="dawsonae">
|
||
<emphasis id="B938EAF9FF70D5A9FEBF714AFE14FAC9" box="[311,452,1291,1317]" italics="true" pageId="156" pageNumber="157">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from that of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FF00716AFEC4FAA9" authorityName="Plane" authorityYear="1967" box="[136,276,1323,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF70D5A9FF00716AFEC4FAA9" box="[136,276,1323,1349]" italics="true" pageId="156" pageNumber="157">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in having: a calcaneus with a less medially displaced head and a narrower calcaneal tuberosity; metatarsal IV with continuous dorsal and plantar cuboid facets, a more concave, less laterally projected metatarsal V facet, larger plantar tubercle, more raised plantar ridge, and a relatively more plantarly projected keel; a more elongate metatarsal V, with a narrower, less dorsally projected cuboid facet; and a more elongate proximal pedal phalanx IV.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF70D5A9FF00722DFEC1F909" pageId="156" pageNumber="157" type="etymology">
|
||
<paragraph id="8BF336EBFF70D5A9FF00722DFEC9F969" blockId="156.[136,778,1643,1765]" box="[136,281,1643,1669]" pageId="156" pageNumber="157">
|
||
<emphasis id="B938EAF9FF70D5A9FF00722DFEC9F969" bold="true" box="[136,281,1643,1669]" pageId="156" pageNumber="157">Etymology:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FF0072CDFEC1F909" blockId="156.[136,778,1643,1765]" pageId="156" pageNumber="157">Named for Dr Lyndall Dawson, for her contributions to the study of macropodid systematics and the Big Sink assemblage.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF70D592FF00734CFC68F7E9" lastPageId="167" lastPageNumber="168" pageId="156" pageNumber="157" type="description">
|
||
<paragraph id="8BF336EBFF70D5A9FF00734CFE2FF8C8" blockId="156.[136,511,1802,1829]" box="[136,511,1802,1829]" pageId="156" pageNumber="157">
|
||
<emphasis id="B938EAF9FF70D5A9FF00734CFE2FF8C8" bold="true" box="[136,511,1802,1829]" pageId="156" pageNumber="157">Description and comparisons:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FF00730DFE4EF889" blockId="156.[136,779,1866,2085]" box="[136,414,1866,1893]" pageId="156" pageNumber="157">
|
||
<emphasis id="B938EAF9FF70D5A9FF00730DFE4EF889" bold="true" box="[136,414,1866,1893]" italics="true" pageId="156" pageNumber="157">Cranium and dentition</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FF00732DFBC2FF09" blockId="156.[136,779,1866,2085]" lastBlockId="156.[810,1452,171,2085]" pageId="156" pageNumber="157">
|
||
<emphasis id="B938EAF9FF70D5A9FF00732DFEA4F869" box="[136,372,1899,1925]" italics="true" pageId="156" pageNumber="157">
|
||
Cranium (
|
||
<figureCitation id="13772A6EFF70D5A9FE8C732DFEBCF869" box="[260,364,1899,1925]" captionStart="FIGURE 91" captionStartId="157.[136,235,1420,1444]" captionTargetBox="[162,1425,194,1388]" captionTargetId="figure-17@157.[151,1436,187,1399]" captionTargetPageId="157" captionText="FIGURE 91. (a–c) surface scan images of left I1–3 of P. dawsonae sp. nov. AM F69835 in (a) occlusal, (b) buccal/lateral, and (c) anterior views; and (d) partial right cranium of P. dawsonae sp. nov. holotype AM F161915 in right lateral view, showing M1–4, maxilla, and partial lacrimal, frontal, palatine, and jugal." figureDoi="http://doi.org/10.5281/zenodo.10993976" httpUri="https://zenodo.org/record/10993976/files/figure.png" pageId="156" pageNumber="157">Fig. 91d</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: jugal has a quite large, slightly posteriorly deflected and laterally projected masseteric process, with morphology very similar to that of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FD7D73EAFF2DF809" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FD7D73EAFF2DF809" italics="true" pageId="156" pageNumber="157">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FE8C738DFE8DF809" bold="true" box="[260,349,1995,2021]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FE8C738DFE8DF809" box="[260,349,1995,2021]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Ventral margin of the orbit is dorsally displaced relative to the maxillary shelf. Maxillary foramen small and round, opens posteriorly. Pterygoid is elongate. No vacuities are present on the preserved section of the palate.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FCD274ADFAA7FE69" blockId="156.[810,1452,171,2085]" pageId="156" pageNumber="157">
|
||
The known portion of the cranium of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FA9474AAFA7BFEE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1308,1451,235,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF70D5A9FA9474AAFA7BFEE9" box="[1308,1451,235,261]" italics="true" pageId="156" pageNumber="157">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF70D5A9FCA2754DFC54FEC9" bold="true" box="[810,900,267,293]" pageId="156" pageNumber="157">
|
||
<taxonomicNameLabel id="A20B5782FF70D5A9FCA2754DFC54FEC9" box="[810,900,267,293]" pageId="156" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF70D5A9FBE5754DFB2EFEC9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1133,1278,267,293]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF70D5A9FBE5754DFB2EFEC9" box="[1133,1278,267,293]" italics="true" pageId="156" pageNumber="157">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF70D5A9FABF754AFA7BFEC9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1335,1451,267,293]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="156" pageNumber="157" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF70D5A9FABF754AFA7BFEC9" box="[1335,1451,267,293]" italics="true" pageId="156" pageNumber="157">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in having a higher maxillary shelf relative to the position of the base of the masseteric process, and a higher ventral margin of the orbit with a less laterally projected lip.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FCD275CAFA90FC49" blockId="156.[810,1452,171,2085]" pageId="156" pageNumber="157">
|
||
<emphasis id="B938EAF9FF70D5A9FCD275CAFB03FE49" box="[858,1235,395,421]" italics="true" pageId="156" pageNumber="157">
|
||
Upper dentition (
|
||
<figureCitation id="13772A6EFF70D5A9FBAE75CDFB50FE49" box="[1062,1152,395,421]" captionStart="FIGURE 91" captionStartId="157.[136,235,1420,1444]" captionTargetBox="[162,1425,194,1388]" captionTargetId="figure-17@157.[151,1436,187,1399]" captionTargetPageId="157" captionText="FIGURE 91. (a–c) surface scan images of left I1–3 of P. dawsonae sp. nov. AM F69835 in (a) occlusal, (b) buccal/lateral, and (c) anterior views; and (d) partial right cranium of P. dawsonae sp. nov. holotype AM F161915 in right lateral view, showing M1–4, maxilla, and partial lacrimal, frontal, palatine, and jugal." figureDoi="http://doi.org/10.5281/zenodo.10993976" httpUri="https://zenodo.org/record/10993976/files/figure.png" pageId="156" pageNumber="157">Figs 91</figureCitation>
|
||
&
|
||
<figureCitation id="13772A6EFF70D5A9FB2675CDFB1BFE49" box="[1198,1227,395,421]" captionStart="FIGURE 92" captionStartId="158.[136,235,1580,1604]" captionTargetBox="[162,1425,192,1547]" captionTargetId="figure-27@158.[151,1436,185,1558]" captionTargetPageId="158" captionText="FIGURE 92. upper dentition of P. dawsonae sp. nov.: right DP2–M2 of SAMA P50399 in (a) buccal, (b) lingual, and (c) occlusal views; (d) right M1–4 of holotype AM F161915 in occlusal view; (e) left P3 of AM F161923 in occlusal view; and right P3 (removed from crypt) of SAMA P50399 in (f) lingual, (g) occlusal, and (h) buccal views." figureDoi="http://doi.org/10.5281/zenodo.10993978" httpUri="https://zenodo.org/record/10993978/files/figure.png" pageId="156" pageNumber="157">92</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: I1: robust, broad and arcuate. Enamel covers the anterior, buccal, and lingual surfaces, and is removed from the sides of the posterior surface by a small amount of wear. Occlusal surface broad and roughly rectangular in slightly worn specimens, becomes longer and rounder with increased wear. I2: arcuate, relatively small and narrow. Enamel covers the buccal and lingual surfaces, with the lingual enamel removed only by very extensive wear. Small posterobuccal crest is removed by a small amount of wear, leaving a slight concavity on buccal surface. I3: elongate, transversely compressed and trapezoidal to roughly triangular in buccal view; buccal enamel convex, buccal length greater than the width of I1 (
|
||
<figureCitation id="13772A6EFF70D5A9FB36776DFACFFCA9" box="[1214,1311,811,837]" captionStart="FIGURE 91" captionStartId="157.[136,235,1420,1444]" captionTargetBox="[162,1425,194,1388]" captionTargetId="figure-17@157.[151,1436,187,1399]" captionTargetPageId="157" captionText="FIGURE 91. (a–c) surface scan images of left I1–3 of P. dawsonae sp. nov. AM F69835 in (a) occlusal, (b) buccal/lateral, and (c) anterior views; and (d) partial right cranium of P. dawsonae sp. nov. holotype AM F161915 in right lateral view, showing M1–4, maxilla, and partial lacrimal, frontal, palatine, and jugal." figureDoi="http://doi.org/10.5281/zenodo.10993976" httpUri="https://zenodo.org/record/10993976/files/figure.png" pageId="156" pageNumber="157">Fig. 91a</figureCitation>
|
||
). Main crest curves anterolingually to sit lingual to the posterior margin of the similarly shaped anterobuccal crest. Anterobuccal crest is around half the length of the main crest.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A9FCD277EDFC4BF9C9" blockId="156.[810,1452,171,2085]" pageId="156" pageNumber="157">Cheek tooth row is straight to very slightly curved in adults. DP2: morphologically very similar to P3 but anteroposteriorly truncated; oblong, quite short and broad, broadens posteriorly, with thickened peaks over the anterior and posterior roots linked by the main crest. Main crest high, blade-like, anteroposteriorly to slightly posterobuccally orientated, jagged to gently undulating in buccal view, with a low, dorsoventrally aligned ridgelet on the midpoint of the buccal surface. Lingual crest low, extends from the lingual base of the anterior cusp to the small, secondary posterolingual peak; lingually borders a broad, anteriorly tapering lingual basin with a low transecting ridgelet. Posterior basin small, abuts the posterior margin, sits between the main and secondary posterior peaks; removed by a small amount of wear. DP3: molariform, fairly broad; anterior loph longer and narrower than the posterior loph; lingual bases of lophs swollen; interloph valley width subequal to that of lophs. Precingulum and cristae worn to absence in sole available specimen.</paragraph>
|
||
<paragraph id="8BF336EBFF70D5A8FCD2726DFC54F8C9" blockId="156.[810,1452,171,2085]" lastBlockId="157.[810,1452,1611,2086]" lastPageId="157" lastPageNumber="158" pageId="156" pageNumber="157">
|
||
P3: large, oblong and quite elongate, generally slightly broader posteriorly, with thickened peaks over the anterior and posterior roots linked by the main crest; straight to curving very slightly buccally toward the posterior, with posterior section very slightly rotated and swollen buccally, contributing to occasional slightly crescentic shape of the tooth. Main crest high, blade-like and roughly anteroposteriorly orientated with occasional very slight posterobuccal curvature in occlusal view; slightly jagged to gently undulating in buccal view; three dorsoventrally aligned ridgelets on the buccal surface, posterior ridgelets lower, with anteriormost ridgelet typically more raised and angular; in some specimens, small buccal ridgelets form enamel crenulations at the buccal base of the main crest (
|
||
<figureCitation id="13772A6EFF70D5A9FCE673ADFC01F7E9" box="[878,977,2027,2053]" captionStart="FIGURE 92" captionStartId="158.[136,235,1580,1604]" captionTargetBox="[162,1425,192,1547]" captionTargetId="figure-27@158.[151,1436,185,1558]" captionTargetPageId="158" captionText="FIGURE 92. upper dentition of P. dawsonae sp. nov.: right DP2–M2 of SAMA P50399 in (a) buccal, (b) lingual, and (c) occlusal views; (d) right M1–4 of holotype AM F161915 in occlusal view; (e) left P3 of AM F161923 in occlusal view; and right P3 (removed from crypt) of SAMA P50399 in (f) lingual, (g) occlusal, and (h) buccal views." figureDoi="http://doi.org/10.5281/zenodo.10993978" httpUri="https://zenodo.org/record/10993978/files/figure.png" pageId="156" pageNumber="157">
|
||
Fig.
|
||
<quantity id="4CB49B0EFF70D5A9FC2A73ADFC01F7E9" box="[930,977,2027,2053]" metricMagnitude="-2" metricUnit="kg" metricValue="9.2" pageId="156" pageNumber="157" unit="g" value="92.0">92g</quantity>
|
||
</figureCitation>
|
||
). Anterior peak quite tall, forms a rounded point; peak intersected by a very brief transverse crest, perpendicular to main crest, extends down buccal and lingual faces as a low ridgelet on posterior margins of anterior cusp, with lingual ridgelet merging into the anterior margin of the lingual crest; small, blunted cusp present at the anterior base, often lingually situated. Lingual crest low, undulating in lingual view, with a single low, smoothly rounded peak near the anterior margin; crest extends posteriorly, from or slightly posterior to the lingual base of the anterior cusp, to merge into the posterolingual peak; lingually borders a broad lingual basin, which is perpendicularly transected by three very low, indistinct ridgelets, roughly opposing the buccal ridgelets. Posterior cuspule broad and posteriorly rounded. Main posterior peak quite tall and rounded, continuous with the main crest, which extends and broadens posteriorly to merge with the low transverse posterior crest. Posterolingual peak rounded, lower than the main peak, linked to the main crest by a thin transverse crest that anteriorly borders the posterior basin. A small posterior basin sits between the main and posterolingual peaks, posteriorly bordered by the transverse posterior crest; removed by a small amount of wear.
|
||
</paragraph>
|
||
<caption id="DF336663FF71D5A8FF0071CAFD14FA09" ID-DOI="http://doi.org/10.5281/zenodo.10993976" ID-Zenodo-Dep="10993976" httpUri="https://zenodo.org/record/10993976/files/figure.png" pageId="157" pageNumber="158" startId="157.[136,235,1420,1444]" targetBox="[162,1425,194,1388]" targetPageId="157" targetType="figure">
|
||
<paragraph id="8BF336EBFF71D5A8FF0071CAFD14FA09" blockId="157.[136,1451,1420,1509]" pageId="157" pageNumber="158">
|
||
<emphasis id="B938EAF9FF71D5A8FF0071CAFEC0FA49" bold="true" box="[136,272,1420,1445]" pageId="157" pageNumber="158">FIGURE 91.</emphasis>
|
||
(a–c) surface scan images of left I1–3 of
|
||
<taxonomicName id="4C4C4D68FF71D5A8FD3C71CBFCE2FA49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[692,818,1421,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="157" pageNumber="158" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF71D5A8FD3C71CBFCE2FA49" box="[692,818,1421,1445]" italics="true" pageId="157" pageNumber="158">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF71D5A8FCB071CBFC59FA49" bold="true" box="[824,905,1421,1445]" pageId="157" pageNumber="158">
|
||
<taxonomicNameLabel id="A20B5782FF71D5A8FCB071CBFC59FA49" box="[824,905,1421,1445]" pageId="157" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
AM F69835 in (a) occlusal, (b) buccal/lateral, and (c) anterior views; and (d) partial right cranium of
|
||
<taxonomicName id="4C4C4D68FF71D5A8FDE371EBFD39FA29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[619,745,1453,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="157" pageNumber="158" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF71D5A8FDE371EBFD39FA29" box="[619,745,1453,1477]" italics="true" pageId="157" pageNumber="158">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF71D5A8FD7871EBFC91FA29" bold="true" box="[752,833,1453,1477]" pageId="157" pageNumber="158">
|
||
<taxonomicNameLabel id="A20B5782FF71D5A8FD7871EBFC91FA29" box="[752,833,1453,1477]" pageId="157" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype AM F161915 in right lateral view, showing M1–4, maxilla, and partial lacrimal, frontal, palatine, and jugal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF71D5ABFCD2736DFAF2F7C9" blockId="157.[810,1452,1611,2086]" lastBlockId="158.[810,1452,1771,2085]" lastPageId="158" lastPageNumber="159" pageId="157" pageNumber="158">
|
||
Molars: rounded-rectangular in occlusal view, with M1 and usually M2 more rounded than the posterior molars. Precingulum quite broad, slightly narrower than the anterior loph, gently medially tilted, extends from the anterior base of the preparacrista to adjacent to the anterolingual margin; generally becomes slightly larger, broader and more projected toward M4; flat, broad and shelf-like when worn; some specimens (see SAMA P50399) have a small preprotocrista extending very briefly up the anterior of the protoloph from around the midpoint of the precingulum. Preparacrista quite thin but distinct and straight, continuous with the buccal margin of the precingulum and extending to the paracone. Postparacrista thin, raised and distinctly kinked, extends straight posteriorly from the paracone before twisting strongly lingually near the base of the interloph valley and proceeding nearly to the end of the postprotocrista in the lingual valley (
|
||
<figureCitation id="13772A6EFF72D5ABFEE573ADFE35F7EA" box="[365,485,2027,2054]" captionStart="FIGURE 92" captionStartId="158.[136,235,1580,1604]" captionTargetBox="[162,1425,192,1547]" captionTargetId="figure-27@158.[151,1436,185,1558]" captionTargetPageId="158" captionText="FIGURE 92. upper dentition of P. dawsonae sp. nov.: right DP2–M2 of SAMA P50399 in (a) buccal, (b) lingual, and (c) occlusal views; (d) right M1–4 of holotype AM F161915 in occlusal view; (e) left P3 of AM F161923 in occlusal view; and right P3 (removed from crypt) of SAMA P50399 in (f) lingual, (g) occlusal, and (h) buccal views." figureDoi="http://doi.org/10.5281/zenodo.10993978" httpUri="https://zenodo.org/record/10993978/files/figure.png" pageId="158" pageNumber="159">Fig. 92c–d</figureCitation>
|
||
). Postprotocrista broad and quite straight, extends from protocone to around the midpoint of interloph valley; in most specimens, the postprotocrista extends onto the anterior face of the metaloph. Premetacrista slight, extends from metacone to meet base of postparacrista in interloph valley Postmetaconulecrista quite thick and raised, arises from metaconule and curves dorsobuccally to almost beneath metacone to form moderate oblique shelf beneath posterior basin. Postmetacrista lower, shorter and less distinct, extends straight from metacone to buccal margin of postmetaconulecrista or deflects slightly lingually to merge into buccal margin of posterior basin.
|
||
</paragraph>
|
||
<caption id="DF336663FF72D5ABFF00726AFBC1F969" ID-DOI="http://doi.org/10.5281/zenodo.10993978" ID-Zenodo-Dep="10993978" httpUri="https://zenodo.org/record/10993978/files/figure.png" pageId="158" pageNumber="159" startId="158.[136,235,1580,1604]" targetBox="[162,1425,192,1547]" targetPageId="158" targetType="figure">
|
||
<paragraph id="8BF336EBFF72D5ABFF00726AFBC1F969" blockId="158.[136,1451,1580,1669]" pageId="158" pageNumber="159">
|
||
<emphasis id="B938EAF9FF72D5ABFF00726AFEC0F9A9" bold="true" box="[136,272,1580,1605]" pageId="158" pageNumber="159">FIGURE 92.</emphasis>
|
||
upper dentition of
|
||
<taxonomicName id="4C4C4D68FF72D5ABFE5A726BFD80F9A9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[466,592,1581,1605]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="158" pageNumber="159" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF72D5ABFE5A726BFD80F9A9" box="[466,592,1581,1605]" italics="true" pageId="158" pageNumber="159">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF72D5ABFDDE726BFD76F9A9" bold="true" box="[598,678,1581,1605]" pageId="158" pageNumber="159">
|
||
<taxonomicNameLabel id="A20B5782FF72D5ABFDDE726BFD76F9A9" box="[598,678,1581,1605]" pageId="158" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: right DP2–M2 of SAMA P50399 in (a) buccal, (b) lingual, and (c) occlusal views; (d) right M1–4 of holotype AM F161915 in occlusal view; (e) left P3 of AM F161923 in occlusal view; and right P3 (removed from crypt) of SAMA P50399 in (f) lingual, (g) occlusal, and (h) buccal views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF336663FF73D5AAFF0072EAFB0CF8E9" ID-DOI="http://doi.org/10.5281/zenodo.10993980" ID-Zenodo-Dep="10993980" httpUri="https://zenodo.org/record/10993980/files/figure.png" pageId="159" pageNumber="160" startId="159.[136,235,1708,1732]" targetBox="[162,1426,202,1677]" targetPageId="159" targetType="figure">
|
||
<paragraph id="8BF336EBFF73D5AAFF0072EAFB0CF8E9" blockId="159.[136,1451,1708,1797]" pageId="159" pageNumber="160">
|
||
<emphasis id="B938EAF9FF73D5AAFF0072EAFEC2F929" bold="true" box="[136,274,1708,1733]" pageId="159" pageNumber="160">FIGURE 93.</emphasis>
|
||
dentaries and right lower dentition of
|
||
<taxonomicName id="4C4C4D68FF73D5AAFD2972EBFCF0F929" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[673,800,1709,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="159" pageNumber="160" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF73D5AAFD2972EBFCF0F929" box="[673,800,1709,1733]" italics="true" pageId="159" pageNumber="160">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF73D5AAFCA072EBFCAAF929" bold="true" box="[808,890,1709,1733]" pageId="159" pageNumber="160">
|
||
<taxonomicNameLabel id="A20B5782FF73D5AAFCA072EBFCAAF929" box="[808,890,1709,1733]" pageId="159" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: mandible (re-associated dentaries) AM F69858 (right dentary) and AM F69859 (left dentary) in (a) occlusal, and (b) ventral views; right dentary AM F69858 in (c) buccal, and (d) lingual views, and (e) lingual view of i1; and p3–m4 of AM F69858 in (f) occlusal, (g) buccal, and (h) lingual views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF73D595FF30732DFDC0FCCA" blockId="159.[136,779,1899,2085]" lastBlockId="160.[136,779,171,2085]" lastPageId="160" lastPageNumber="161" pageId="159" pageNumber="160">
|
||
The upper dentition of
|
||
<taxonomicName id="4C4C4D68FF73D5AAFE4C732AFD82F869" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[452,594,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="159" pageNumber="160" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF73D5AAFE4C732AFD82F869" box="[452,594,1899,1925]" italics="true" pageId="159" pageNumber="160">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF73D5AAFDD4732DFD68F869" bold="true" box="[604,696,1899,1925]" pageId="159" pageNumber="160">
|
||
<taxonomicNameLabel id="A20B5782FF73D5AAFDD4732DFD68F869" box="[604,696,1899,1925]" pageId="159" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of all other species of
|
||
<taxonomicName id="4C4C4D68FF73D5AAFE7673CDFD45F849" authorityName="Owen" authorityYear="1874" box="[510,661,1931,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="159" pageNumber="160" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF73D5AAFE7673CDFD45F849" box="[510,661,1931,1957]" italics="true" pageId="159" pageNumber="160">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
in having the postparacrista distinctly inflected lingually then buccally in the interloph valley. It further differs from
|
||
<taxonomicName id="4C4C4D68FF73D5AAFD7D738AFF6EF7E9" authorityName=", Owen" authorityYear="1874" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="159" pageNumber="160" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF73D5AAFD7D738AFF6EF7E9" italics="true" pageId="159" pageNumber="160">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in having larger P3 with lower, less distinct buccal and lingual ridgelets and a less jagged, more smoothly undulating and more anteriorly extensive lingual crest, and relatively broader molars with a narrower protoloph relative to the protoloph base when unworn; from
|
||
<taxonomicName id="4C4C4D68FF73D5AAFB7E73EAFA53F829" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1270,1411,1963,1989]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="159" pageNumber="160" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF73D5AAFB7E73EAFA53F829" box="[1270,1411,1963,1989]" italics="true" pageId="159" pageNumber="160">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF73D5AAFA0273EDFC8BF809" bold="true" pageId="159" pageNumber="160">
|
||
<taxonomicNameLabel id="A20B5782FF73D5AAFA0273EDFC8BF809" pageId="159" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF73D5AAFC1B738AFC22F809" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[915,1010,1996,2021]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="159" pageNumber="160" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF73D5AAFC1B738AFC22F809" box="[915,1010,1996,2021]" italics="true" pageId="159" pageNumber="160">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF73D5AAFC72738DFB85F809" bold="true" box="[1018,1109,1995,2021]" pageId="159" pageNumber="160">
|
||
<taxonomicNameLabel id="A20B5782FF73D5AAFC72738DFB85F809" box="[1018,1109,1995,2021]" pageId="159" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having I1 narrower relative to I3, P3 with slightly more raised and distinct buccal ridgelets, and molars with a thinner, more raised and more distinct preparacrista and a narrower protoloph relative to the loph base when unworn; additionally differs from
|
||
<taxonomicName id="4C4C4D68FF4CD595FF0074AAFEF2FEE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[136,290,235,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FF0074AAFEF2FEE9" box="[136,290,235,261]" italics="true" pageId="160" pageNumber="161">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FEBB74ADFE46FEE9" bold="true" box="[307,406,235,261]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FEBB74ADFE46FEE9" box="[307,406,235,261]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having relatively narrower posterior molars; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FE12754AFDCDFEC9" box="[410,541,267,293]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF4CD595FE12754AFDCDFEC9" box="[410,541,267,293]" italics="true" pageId="160" pageNumber="161">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in being larger and slightly higher crowned, with I1 lacking posterobuccal bulge, and generally less rounded molars in occlusal view with a more raised, more distinct preparacrista and lacking a urocrista; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FE2175CAFDE4FE49" authorityName="Plane" authorityYear="1967" box="[425,564,395,421]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF4CD595FE2175CAFDE4FE49" box="[425,564,395,421]" italics="true" pageId="160" pageNumber="161">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in being generally larger, with broader, anteroposteriorly shorter I1 lacking a posterobuccal bulge and molars lacking a urocrista; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FF0075AAFF2AFDE9" authorityName="Bartholomai" authorityYear="1978" box="[136,250,492,517]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF4CD595FF0075AAFF2AFDE9" box="[136,250,492,517]" italics="true" pageId="160" pageNumber="161">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with relatively broader P3 and molars with a higher, more distinct preparacrista; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FD78764DFF28FDA9" baseAuthorityName="Flannery" baseAuthorityYear="1989" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4CD595FD78764DFF28FDA9" italics="true" pageId="160" pageNumber="161">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger and higher crowned, with larger incisors relative to size of cranium and an absolutely and relatively more elongate P3; and from
|
||
<taxonomicName id="4C4C4D68FF4CD595FDBE762AFD79FD69" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[566,681,619,645]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4CD595FDBE762AFD79FD69" box="[566,681,619,645]" italics="true" pageId="160" pageNumber="161">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, and higher crowned, with P3 with a higher lingual crest and a larger posterior basin, and molars with a thicker postprotocrista distinctly extending to the protocone (rather than merging into the centre of the posterior surface of the protoloph).
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4CD595FF30776AFEDEFA29" blockId="160.[136,779,171,2085]" pageId="160" pageNumber="161">
|
||
<emphasis id="B938EAF9FF4CD595FF30776AFE63FCA9" box="[184,435,811,837]" italics="true" pageId="160" pageNumber="161">
|
||
Dentary (
|
||
<figureCitation id="13772A6EFF4CD595FEA1776DFE7BFCA9" box="[297,427,811,837]" captionStart="FIGURE 93" captionStartId="159.[136,235,1708,1732]" captionTargetBox="[162,1426,202,1677]" captionTargetId="figure-18@159.[151,1436,191,1687]" captionTargetPageId="159" captionText="FIGURE 93. dentaries and right lower dentition of P. dawsonae sp. nov.: mandible (re-associated dentaries) AM F69858 (right dentary) andAM F69859 (left dentary) in (a) occlusal, and (b) ventral views; right dentary AM F69858 in (c) buccal, and (d) lingual views, and (e) lingual view of i1; and p3–m4 of AM F69858 in (f) occlusal, (g) buccal, and (h) lingual views." figureDoi="http://doi.org/10.5281/zenodo.10993980" httpUri="https://zenodo.org/record/10993980/files/figure.png" pageId="160" pageNumber="161">Fig. 93a–d</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: robust, moderately tall, and gently transversely compressed, with a long, robust and gently tapering anterior dentary ventral to the diastema, projected straight anteriorly or slightly deflected dorsally (
|
||
<figureCitation id="13772A6EFF4CD595FF1877EDFF21FC29" box="[144,241,939,965]" captionStart="FIGURE 93" captionStartId="159.[136,235,1708,1732]" captionTargetBox="[162,1426,202,1677]" captionTargetId="figure-18@159.[151,1436,191,1687]" captionTargetPageId="159" captionText="FIGURE 93. dentaries and right lower dentition of P. dawsonae sp. nov.: mandible (re-associated dentaries) AM F69858 (right dentary) andAM F69859 (left dentary) in (a) occlusal, and (b) ventral views; right dentary AM F69858 in (c) buccal, and (d) lingual views, and (e) lingual view of i1; and p3–m4 of AM F69858 in (f) occlusal, (g) buccal, and (h) lingual views." figureDoi="http://doi.org/10.5281/zenodo.10993980" httpUri="https://zenodo.org/record/10993980/files/figure.png" pageId="160" pageNumber="161">Fig. 93a</figureCitation>
|
||
). Angle between the dentaries at the mandibular symphysis is ~35–40°. Mandibular symphyseal plate large and elongate, extends along the entirety of the anterior dentary to abut the posterior margin of i1. Mental foramen round to oval, opens dorsolaterally to anterodorsolaterally, located roughly one-fifth of distance from dp2/p3 to i1 and around one-quarter of distance from the diastema to the ventral margin. Mandibular corpus height generally subequal below p3 and m4, with the width at a minimum ventral to m1 and increasing gently posteriorly. Digastric sulcus shallow and broad, extends from beneath the posterior of the molar row to adjacent to the anteroventral margin of the medial pterygoid fossa. Buccinator sulcus distinct but quite shallow, extends along the buccal surface of the mandibular corpus with a slight ventral tilt posteriorly, slightly ventral to the three–four anteriormost cheek teeth.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4CD595FF30718DFD4DF849" blockId="160.[136,779,171,2085]" pageId="160" pageNumber="161">
|
||
The dentary of
|
||
<taxonomicName id="4C4C4D68FF4CD595FEF6718AFDC3FA09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[382,531,1483,1509]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FEF6718AFDC3FA09" box="[382,531,1483,1509]" italics="true" pageId="160" pageNumber="161">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FDAB718DFD55FA09" bold="true" box="[547,645,1483,1509]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FDAB718DFD55FA09" box="[547,645,1483,1509]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
cannot be differentiated from that of
|
||
<taxonomicName id="4C4C4D68FF4CD595FE3A71AAFD92F9E9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[434,578,1515,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FE3A71AAFD92F9E9" box="[434,578,1515,1541]" italics="true" pageId="160" pageNumber="161">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FDC171ADFD72F9E9" bold="true" box="[585,674,1515,1541]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FDC171ADFD72F9E9" box="[585,674,1515,1541]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
It differs from that of
|
||
<taxonomicName id="4C4C4D68FF4CD595FE94724AFEA0F9C9" authorityName=", Owen" authorityYear="1874" box="[284,368,1547,1573]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF4CD595FE94724AFEA0F9C9" box="[284,368,1547,1573]" italics="true" pageId="160" pageNumber="161">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being slightly broader and more robust, with a more dorsally deflected diastema; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FD7D726AFF1BF989" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FD7D726AFF1BF989" italics="true" pageId="160" pageNumber="161">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FF5C720DFEE0F989" bold="true" box="[212,304,1611,1637]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FF5C720DFEE0F989" box="[212,304,1611,1637]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being generally slightly smaller; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FF00722AFED7F969" box="[136,263,1643,1669]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF4CD595FF00722AFED7F969" box="[136,263,1643,1669]" italics="true" pageId="160" pageNumber="161">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a more dorsally situated mental foramen; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FE0C72CAFDDDF949" authorityName="Plane" authorityYear="1967" box="[388,525,1675,1701]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF4CD595FE0C72CAFDDDF949" box="[388,525,1675,1701]" italics="true" pageId="160" pageNumber="161">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in being slightly more robust, particularly in the diastema; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FDFB72EAFD39F929" authorityName="Bartholomai" authorityYear="1978" box="[627,745,1708,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF4CD595FDFB72EAFD39F929" box="[627,745,1708,1733]" italics="true" pageId="160" pageNumber="161">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being larger and more robust, with a more robust and more dorsally deflected diastema and a taller mandibular corpus; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FF4D734DFE85F8C9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[197,341,1803,1829]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4CD595FF4D734DFE85F8C9" box="[197,341,1803,1829]" italics="true" pageId="160" pageNumber="161">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger and more robust, with a relatively shorter, more dorsally deflected diastema, more posteroventrally situated mental foramen, and a taller mandibular corpus; and from
|
||
<taxonomicName id="4C4C4D68FF4CD595FE6A732AFD89F869" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[482,601,1899,1925]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4CD595FE6A732AFD89F869" box="[482,601,1899,1925]" italics="true" pageId="160" pageNumber="161">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, with a more dorsally deflected diastema.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4CD595FF3073EAFBF2FEC9" blockId="160.[136,779,171,2085]" lastBlockId="160.[809,1452,171,2085]" pageId="160" pageNumber="161">
|
||
<emphasis id="B938EAF9FF4CD595FF3073EAFDDFF829" box="[184,527,1963,1989]" italics="true" pageId="160" pageNumber="161">
|
||
Lower dentition (
|
||
<figureCitation id="13772A6EFF4CD595FE0E73EDFDD7F829" box="[390,519,1963,1989]" captionStart="FIGURE 93" captionStartId="159.[136,235,1708,1732]" captionTargetBox="[162,1426,202,1677]" captionTargetId="figure-18@159.[151,1436,191,1687]" captionTargetPageId="159" captionText="FIGURE 93. dentaries and right lower dentition of P. dawsonae sp. nov.: mandible (re-associated dentaries) AM F69858 (right dentary) andAM F69859 (left dentary) in (a) occlusal, and (b) ventral views; right dentary AM F69858 in (c) buccal, and (d) lingual views, and (e) lingual view of i1; and p3–m4 of AM F69858 in (f) occlusal, (g) buccal, and (h) lingual views." figureDoi="http://doi.org/10.5281/zenodo.10993980" httpUri="https://zenodo.org/record/10993980/files/figure.png" pageId="160" pageNumber="161">Fig. 93c–h</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: i1: procumbent and slightly dorsally deflected; broad, tilted dorsally and transversely compressed; acuminate when unworn, becomes shorter and rounder in cross-section with wear. Thick enamel completely covers the buccal surface. Dorsobuccal and ventrolingual crests low and thick (
|
||
<figureCitation id="13772A6EFF4CD595FAF6748DFC86FEE9" captionStart="FIGURE 93" captionStartId="159.[136,235,1708,1732]" captionTargetBox="[162,1426,202,1677]" captionTargetId="figure-18@159.[151,1436,191,1687]" captionTargetPageId="159" captionText="FIGURE 93. dentaries and right lower dentition of P. dawsonae sp. nov.: mandible (re-associated dentaries) AM F69858 (right dentary) andAM F69859 (left dentary) in (a) occlusal, and (b) ventral views; right dentary AM F69858 in (c) buccal, and (d) lingual views, and (e) lingual view of i1; and p3–m4 of AM F69858 in (f) occlusal, (g) buccal, and (h) lingual views." figureDoi="http://doi.org/10.5281/zenodo.10993980" httpUri="https://zenodo.org/record/10993980/files/figure.png" pageId="160" pageNumber="161">Fig. 93e</figureCitation>
|
||
). Enamel covers the ventral third of the lingual surface and tapers posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4CD595FCD2756DFB81FD29" blockId="160.[809,1452,171,2085]" pageId="160" pageNumber="161">
|
||
p3: very similar in morphology to that of
|
||
<taxonomicName id="4C4C4D68FF4CD595FA95756AFA7CFEA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1309,1452,299,325]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FA95756AFA7CFEA9" box="[1309,1452,299,325]" italics="true" pageId="160" pageNumber="161">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FCA2750DFC57FE89" bold="true" box="[810,903,331,357]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FCA2750DFC57FE89" box="[810,903,331,357]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4CD595FC4C750AFBF7FE89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[964,1063,332,357]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FC4C750AFBF7FE89" box="[964,1063,332,357]" italics="true" pageId="160" pageNumber="161">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FBB9750DFB5EFE89" bold="true" box="[1073,1166,331,357]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FBB9750DFB5EFE89" box="[1073,1166,331,357]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; oblong and elongate in occlusal view, typically with a slight waist; typically subequally broad across posterior and anterior moieties; tall and triangular in cross-section with bulging buccal and lingual bases. Anterior cuspid has a dorsoventrally aligned buccal and lingual ridgelet on posterior margin. Main crest linear, anteroposteriorly aligned and twists slightly lingually over the posterior root. Both surfaces of the main crest with two or three low, rounded, indistinct, roughly dorsoventrally aligned ridgelets extending to the peak of the crest, such that the crest appears slightly undulating in buccal view.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4CD595FCD2768DFB81F9A9" blockId="160.[809,1452,171,2085]" pageId="160" pageNumber="161">Molars: high-crowned, broad and rounded in occlusal view, with interlophid valley varying from slightly broader to slightly narrower than the lophid bases; when unworn or slightly worn, protolophid and hypolophid are posteriorly convex or have an oblique mesial kink toward the posterior in occlusal view; buccal margins of the lophids are slightly more convex than the lingual margins in posterior view. Precingulid narrow, anteriorly projected, usually forms a rounded corner; a slight raised lip is present around the anterior margin when unworn. Paracristid quite thick and raised; lingual component arises from the anterolingual edge of the trigonid basin, extends buccally then straight posteriorly before the buccal component ascends, curving gently buccally, to the protoconid; the degree of buccal curve and thickness increases with molar wear. Protoconid higher than the metaconid and slightly lingually displaced. Cristid obliqua thick and low, mostly contributed by the talonid; arises slightly buccal to the midpoint of the interlophid valley, curves very slightly buccally and extends to the hypoconid. Preentocristid absent to very low, broad and indistinct, arises midway between the base of the cristid obliqua and the lingual extremity of the interlophid valley and deflects slightly lingually to meet the entoconid. When unworn, the hypoconid is distinctly taller than the entoconid and sits slightly buccal of the midline of the tooth. Postcingulid narrow and slight on m1–m2, becomes broader and occasionally shelf-like in m3 and particularly in m4.</paragraph>
|
||
<paragraph id="8BF336EBFF4CD596FCD2720DFD20FDE9" blockId="160.[809,1452,171,2085]" lastBlockId="163.[136,778,171,517]" lastPageId="163" lastPageNumber="164" pageId="160" pageNumber="161">
|
||
The lower dentition of
|
||
<taxonomicName id="4C4C4D68FF4CD595FBEE720AFB24F989" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1126,1268,1611,1637]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FBEE720AFB24F989" box="[1126,1268,1611,1637]" italics="true" pageId="160" pageNumber="161">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FB76720DFA89F989" bold="true" box="[1278,1369,1611,1637]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FB76720DFA89F989" box="[1278,1369,1611,1637]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of all compared taxa except
|
||
<taxonomicName id="4C4C4D68FF4CD595FB64722AFAAFF969" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1260,1407,1643,1669]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FB64722AFAAFF969" box="[1260,1407,1643,1669]" italics="true" pageId="160" pageNumber="161">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FA02722DFC8AF949" bold="true" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FA02722DFC8AF949" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4CD595FC1472CAFC2FF949" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[924,1023,1676,1701]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FC1472CAFC2FF949" box="[924,1023,1676,1701]" italics="true" pageId="160" pageNumber="161">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FB8572CDFBBBF949" bold="true" box="[1037,1131,1675,1701]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FB8572CDFBBBF949" box="[1037,1131,1675,1701]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a lower, thicker ventrolingual crest on i1. It further differs from
|
||
<taxonomicName id="4C4C4D68FF4CD595FADF72EAFA7BF929" authorityName=", Owen" authorityYear="1874" box="[1367,1451,1707,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF4CD595FADF72EAFA7BF929" box="[1367,1451,1707,1733]" italics="true" pageId="160" pageNumber="161">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being slightly larger, with a more robust i1, and p3 with lower, less distinct ridgelets on the main crest and generally relatively broader molars; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FA9F734AFA7BF8C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1303,1451,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FA9F734AFA7BF8C9" box="[1303,1451,1803,1829]" italics="true" pageId="160" pageNumber="161">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FCA2736DFC54F8A9" bold="true" box="[810,900,1835,1861]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FCA2736DFC54F8A9" box="[810,900,1835,1861]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4CD595FC35736AFBCBF8A9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[957,1051,1836,1861]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FC35736AFBCBF8A9" box="[957,1051,1836,1861]" italics="true" pageId="160" pageNumber="161">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FBAC736DFBAEF8A9" bold="true" box="[1060,1150,1835,1861]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FBAC736DFBAEF8A9" box="[1060,1150,1835,1861]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being generally smaller and slightly lower crowned, with a ventrolingual crest present on i1; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FB88732AFBABF869" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1024,1147,1899,1925]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="tumbuna" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4CD595FB88732AFBABF869" box="[1024,1147,1899,1925]" italics="true" pageId="160" pageNumber="161">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4CD595FB09732DFB09F869" bold="true" box="[1153,1241,1899,1925]" pageId="160" pageNumber="161">
|
||
<taxonomicNameLabel id="A20B5782FF4CD595FB09732DFB09F869" box="[1153,1241,1899,1925]" pageId="160" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being larger and higher crowned, with relatively broader p3, and narrower molars with less convex buccal margins of lophids; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FCA2738AFC61F809" authorityName="Plane" authorityYear="1967" box="[810,945,1995,2021]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF4CD595FCA2738AFC61F809" box="[810,945,1995,2021]" italics="true" pageId="160" pageNumber="161">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in being generally larger, with larger, broader and more robust i1 and relatively broader p3; from
|
||
<taxonomicName id="4C4C4D68FF4CD595FA1F73AAFCAFF7C9" authorityName="Bartholomai" authorityYear="1978" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="160" pageNumber="161" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF4CD595FA1F73AAFCAFF7C9" italics="true" pageId="160" pageNumber="161">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being larger and higher crowned, with broader, more robust i1, relatively broader p3,and relatively broader molars with a smaller precingulid; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FDD0748DFD3BFF09" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[600,747,203,229]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4FD596FDD0748DFD3BFF09" box="[600,747,203,229]" italics="true" pageId="163" pageNumber="164">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger and higher crowned, with broader and more spatulate i1, relatively broader deciduous premolars, absolutely more elongate p3 that is longer relative to molar length, and molars with narrower lophid crests, higher cristid obliqua and a postcingulid present; and from
|
||
<taxonomicName id="4C4C4D68FF4FD596FF4D75CAFEE9FE49" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[197,313,395,421]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4FD596FF4D75CAFEE9FE49" box="[197,313,395,421]" italics="true" pageId="163" pageNumber="164">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, with broader, more spatulate i1, p3 with less raised and distinct ridgelets on the main crest, and molars with narrower lophid crests and a generally more anteriorly prominent precingulid.
|
||
</paragraph>
|
||
<caption id="DF336663FF4DD594FF0073CAFE00F809" ID-DOI="http://doi.org/10.5281/zenodo.10993982" ID-Zenodo-Dep="10993982" httpUri="https://zenodo.org/record/10993982/files/figure.png" pageId="161" pageNumber="162" startId="161.[136,235,1932,1956]" targetBox="[188,1399,183,1899]" targetPageId="161" targetType="figure">
|
||
<paragraph id="8BF336EBFF4DD594FF0073CAFE00F809" blockId="161.[136,1452,1932,2021]" pageId="161" pageNumber="162">
|
||
<emphasis id="B938EAF9FF4DD594FF0073CAFEC2F849" bold="true" box="[136,274,1932,1957]" pageId="161" pageNumber="162">FIGURE 94.</emphasis>
|
||
vertebrae of
|
||
<taxonomicName id="4C4C4D68FF4DD594FE1673CBFDCDF849" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[414,541,1933,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="161" pageNumber="162" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4DD594FE1673CBFDCDF849" box="[414,541,1933,1957]" italics="true" pageId="161" pageNumber="162">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4DD594FDAD73CBFDA7F849" bold="true" box="[549,631,1933,1957]" pageId="161" pageNumber="162">
|
||
<taxonomicNameLabel id="A20B5782FF4DD594FDAD73CBFDA7F849" box="[549,631,1933,1957]" pageId="161" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: partial axis vertebra AM F161920 in (a) left lateral, (b) cranial, and (c) dorsal views; partial axis vertebra AM F161914 in (d) right lateral, and (e) caudal views; and (f–g) lumbar vertebra L2? AM F161918 in (f) cranial, and (g) lateral views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF336663FF4ED597FF0073CAFEB0F809" ID-DOI="http://doi.org/10.5281/zenodo.10993986" ID-Zenodo-Dep="10993986" httpUri="https://zenodo.org/record/10993986/files/figure.png" pageId="162" pageNumber="163" startId="162.[136,235,1932,1956]" targetBox="[175,1424,185,1902]" targetPageId="162" targetType="figure">
|
||
<paragraph id="8BF336EBFF4ED597FF0073CAFEB0F809" blockId="162.[136,1452,1932,2021]" pageId="162" pageNumber="163">
|
||
<emphasis id="B938EAF9FF4ED597FF0073CAFEDFF849" bold="true" box="[136,271,1932,1957]" pageId="162" pageNumber="163">FIGURE 95.</emphasis>
|
||
caudal vertebra Ca2? of
|
||
<taxonomicName id="4C4C4D68FF4ED597FD8073CBFD54F849" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[520,644,1933,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="162" pageNumber="163" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4ED597FD8073CBFD54F849" box="[520,644,1933,1957]" italics="true" pageId="162" pageNumber="163">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4ED597FD0273CBFD09F849" bold="true" box="[650,729,1933,1957]" pageId="162" pageNumber="163">
|
||
<taxonomicNameLabel id="A20B5782FF4ED597FD0273CBFD09F849" box="[650,729,1933,1957]" pageId="162" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
AM F161911 in (a) lateral, (b) dorsal, and (c) cranial views.(d–f) partial pelvis, predominantly ilium and caudal component of ischium, of
|
||
<taxonomicName id="4C4C4D68FF4ED597FCA473EBFC7BF829" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[812,939,1965,1989]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="162" pageNumber="163" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4ED597FCA473EBFC7BF829" box="[812,939,1965,1989]" italics="true" pageId="162" pageNumber="163">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4ED597FC3A73EBFBD4F829" bold="true" box="[946,1028,1965,1989]" pageId="162" pageNumber="163">
|
||
<taxonomicNameLabel id="A20B5782FF4ED597FC3A73EBFBD4F829" box="[946,1028,1965,1989]" pageId="162" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
UCMP 156881 in: (d) caudal, (e) lateral, and (f) medial views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF4FD596FF00766CFEE4FDA9" blockId="163.[136,779,554,2085]" box="[136,308,554,581]" pageId="163" pageNumber="164">
|
||
<emphasis id="B938EAF9FF4FD596FF00766CFEE4FDA9" bold="true" box="[136,308,554,581]" italics="true" pageId="163" pageNumber="164">Axial skeleton</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4FD596FF00760DFE46FB09" blockId="163.[136,779,554,2085]" pageId="163" pageNumber="164">
|
||
<emphasis id="B938EAF9FF4FD596FF00760DFE58FD89" box="[136,392,587,613]" italics="true" pageId="163" pageNumber="164">
|
||
Axis (C2) (
|
||
<figureCitation id="13772A6EFF4FD596FE8D760DFE50FD89" box="[261,384,587,613]" captionStart="FIGURE 94" captionStartId="161.[136,235,1932,1956]" captionTargetBox="[188,1399,183,1899]" captionTargetId="figure-17@161.[177,1410,172,1910]" captionTargetPageId="161" captionText="FIGURE 94. vertebrae of P. dawsonae sp. nov.: partial axis vertebra AM F161920 in (a) left lateral, (b) cranial, and (c) dorsal views; partial axis vertebra AM F161914 in (d) right lateral, and (e) caudal views; and (f–g) lumbar vertebra L2? AM F161918 in (f) cranial, and (g) lateral views." figureDoi="http://doi.org/10.5281/zenodo.10993982" httpUri="https://zenodo.org/record/10993982/files/figure.png" pageId="163" pageNumber="164">Fig. 94a–e</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: moderately craniocaudally short, quite broad and robust. Dens short, blunt and gently dorsally deflected. Cranial articular surfaces broad and convex; angled laterally and slightly dorsally.Arch mostly crushed or absent in available specimens; appears thick and quite low. Vertebral canal roughly oval and gently domed. Spinous process crushed and caudally abraded, appears quite tall, thin and tilted slightly cranially, with gently convex dorsal margin and cranial extremity projecting past cranial margin of arch. Postzygopophyses fairly small and flared slightly laterally, caudally projected beyond caudal extremity of the centrum and subequally to the caudal extremity of the transverse processes; articular surfaces very slightly concave and facing caudoventrally, with a moderate lateral tilt. Transverse processes quite thick and long, with a strong caudal and slight ventral deflection. Ventral surface of the centrum broad, with a slightly raised medial ridge. Caudal extremity of the centrum incompletely preserved; gently caudally and ventrally projected and slightly dorsally tilted, with a rounded ventral margin.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4FD596FF3070ADFB95FF09" blockId="163.[136,779,554,2085]" lastBlockId="163.[810,1452,171,2085]" pageId="163" pageNumber="164">
|
||
The axis vertebra of
|
||
<taxonomicName id="4C4C4D68FF4FD596FE3F70AAFD98FAE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[439,584,1259,1285]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FE3F70AAFD98FAE9" box="[439,584,1259,1285]" italics="true" pageId="163" pageNumber="164">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FDDD70ADFD64FAE9" bold="true" box="[597,692,1259,1285]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FDDD70ADFD64FAE9" box="[597,692,1259,1285]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF4FD596FEAA714AFEA8FAC9" authorityName=", Owen" authorityYear="1874" box="[290,376,1291,1317]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF4FD596FEAA714AFEA8FAC9" box="[290,376,1291,1317]" italics="true" pageId="163" pageNumber="164">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being relatively and absolutely much less elongate, with a less elongate, slightly more dorsally deflected dens, dorsoventrally shorter, slightly more convex cranial articular surfaces, much less caudally projected caudal extremity of the centrum, and smaller, less elongate postzygopophyses; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FDC171EAFD0DFA29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[585,733,1451,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FDC171EAFD0DFA29" box="[585,733,1451,1477]" italics="true" pageId="163" pageNumber="164">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FD6071EDFF69FA09" bold="true" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FD6071EDFF69FA09" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being deeper relative to width, with the caudal extremity of the centrum more caudally projected; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FD7D71AAFF1BF9C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FD7D71AAFF1BF9C9" italics="true" pageId="163" pageNumber="164">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FF47724DFEF5F9C9" bold="true" box="[207,293,1547,1573]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FF47724DFEF5F9C9" box="[207,293,1547,1573]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller and narrower, with a thinner spinous process, more laterally tilted postzygopophyses with less concave articular surfaces and a rounded (rather than bilobed) ventral margin of the caudal extremity of the centrum; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FED472CDFE3DF949" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[348,493,1675,1701]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4FD596FED472CDFE3DF949" box="[348,493,1675,1701]" italics="true" pageId="163" pageNumber="164">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being broader relative to length, with a lower mesial ventral ridge, more laterally tilted caudal articular surfaces, and a relatively larger, broader caudal extremity of the centrum; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FD2372ADFCDAF8E9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[683,778,1771,1797]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF4FD596FD2372ADFCDAF8E9" box="[683,778,1771,1797]" italics="true" pageId="163" pageNumber="164">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being relatively slightly shorter and broader, with a less dorsally deflected dens and the dorsal margin of the spinous process rounded and convex (rather than level); from
|
||
<taxonomicName id="4C4C4D68FF4FD596FE9C732AFE64F869" authorityName="Desmarest" authorityYear="1817" box="[276,436,1899,1925]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF4FD596FE9C732AFE64F869" box="[276,436,1899,1925]" italics="true" pageId="163" pageNumber="164">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a slightly more convex dorsal margin of the spinous process, longer, more ventrally deflected transverse processes, less laterally flared, slightly more caudally projected postzygopophyses, and a more ventrally projected caudal extremity of the centrum; and from
|
||
<taxonomicName id="4C4C4D68FF4FD596FDA37C4AFD72F7C9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[555,674,2059,2085]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4FD596FDA37C4AFD72F7C9" box="[555,674,2059,2085]" italics="true" pageId="163" pageNumber="164">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a more rounded dens and more lateral facing cranial articular surfaces.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4FD596FCD274ADFAB6FCC9" blockId="163.[810,1452,171,2085]" pageId="163" pageNumber="164">
|
||
<emphasis id="B938EAF9FF4FD596FCD274ADFB20FEE9" box="[858,1264,235,261]" italics="true" pageId="163" pageNumber="164">
|
||
Lumbar vertebrae (L2?) (
|
||
<figureCitation id="13772A6EFF4FD596FBFC74ADFB39FEE9" box="[1140,1257,235,261]" captionStart="FIGURE 94" captionStartId="161.[136,235,1932,1956]" captionTargetBox="[188,1399,183,1899]" captionTargetId="figure-17@161.[177,1410,172,1910]" captionTargetPageId="161" captionText="FIGURE 94. vertebrae of P. dawsonae sp. nov.: partial axis vertebra AM F161920 in (a) left lateral, (b) cranial, and (c) dorsal views; partial axis vertebra AM F161914 in (d) right lateral, and (e) caudal views; and (f–g) lumbar vertebra L2? AM F161918 in (f) cranial, and (g) lateral views." figureDoi="http://doi.org/10.5281/zenodo.10993982" httpUri="https://zenodo.org/record/10993982/files/figure.png" pageId="163" pageNumber="164">Fig. 94f–g</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: robust; centrum short and broad, narrows to a waist; ventrolateral surfaces concave and laterally tilted, with the ventral margin concave in lateral view. Cranial extremity of the centrum gently convex and rounded with a flattened dorsal margin, slightly taller than the caudal extremity. Caudal extremity of the centrum gently concave and broadly rounded with a flattened dorsal margin. Vertebral canal quite low, broad and domed in cranial view. The prezygopophyses and spinous process are completely abraded. Transverse processes mostly abraded; bases highly dorsoventrally compressed, ventrally deflected and situated on the cranial part of the lateral surfaces. Postzygopophyses almost entirely abraded, bases appear thickened and robust. The base of a small anapophysis is present on the arch ventrolateral to the base of the postzygopophysis; transversely compressed and slightly caudally deflected, with a low ridge extending cranially from the base.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4FD596FCD2776DFC51FAC9" blockId="163.[810,1452,171,2085]" pageId="163" pageNumber="164">
|
||
The L2? of
|
||
<taxonomicName id="4C4C4D68FF4FD596FC61776AFBA9FCA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1001,1145,811,837]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FC61776AFBA9FCA9" box="[1001,1145,811,837]" italics="true" pageId="163" pageNumber="164">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FB0C776DFB31FCA9" bold="true" box="[1156,1249,811,837]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FB0C776DFB31FCA9" box="[1156,1249,811,837]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF4FD596FCC3770AFC0EFC89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[843,990,843,869]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FCC3770AFC0EFC89" box="[843,990,843,869]" italics="true" pageId="163" pageNumber="164">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FC6F770DFB93FC89" bold="true" box="[999,1091,843,869]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FC6F770DFB93FC89" box="[999,1091,843,869]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4FD596FBF6770AFB0FFC89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1150,1247,844,869]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FBF6770AFB0FFC89" box="[1150,1247,844,869]" italics="true" pageId="163" pageNumber="164">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FB60770DFA94FC89" bold="true" box="[1256,1348,843,869]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FB60770DFA94FC89" box="[1256,1348,843,869]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, with deeper transverse processes and a centrum with relatively lower, broader cranial and caudal extremities, more concave ventrolateral surfaces, more concave ventral margin in lateral view and a less raised ventral ridge; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FBA977ADFB6CFBE9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1057,1212,1003,1029]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4FD596FBA977ADFB6CFBE9" box="[1057,1212,1003,1029]" italics="true" pageId="163" pageNumber="164">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being relatively slightly longer; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FB9C704DFBBFFBC9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1044,1135,1035,1061]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF4FD596FB9C704DFBBFFBC9" box="[1044,1135,1035,1061]" italics="true" pageId="163" pageNumber="164">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being slightly larger, with a broader centrum relative to height and relatively deeper transverse processes; from
|
||
<taxonomicName id="4C4C4D68FF4FD596FBD2700AFB2BFB89" authorityName="Desmarest" authorityYear="1817" box="[1114,1275,1099,1125]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF4FD596FBD2700AFB2BFB89" box="[1114,1275,1099,1125]" italics="true" pageId="163" pageNumber="164">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with less dorsoventrally compressed cranial and caudal extremities of the centrum and a more concave ventral margin in lateral view; and from
|
||
<taxonomicName id="4C4C4D68FF4FD596FB4A70EAFAECFB29" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1218,1340,1195,1221]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4FD596FB4A70EAFAECFB29" box="[1218,1340,1195,1221]" italics="true" pageId="163" pageNumber="164">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, with a centrum having a less distinct waist in ventral view and a less ventrolaterally swollen caudal section.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4FD596FCD2716DFAA3F809" blockId="163.[810,1452,171,2085]" pageId="163" pageNumber="164">
|
||
<emphasis id="B938EAF9FF4FD596FCD2716DFAFEFAA9" box="[858,1326,1323,1349]" italics="true" pageId="163" pageNumber="164">
|
||
Caudal vertebrae (Ca2 & 3?) (
|
||
<figureCitation id="13772A6EFF4FD596FB25716DFAF7FAA9" box="[1197,1319,1323,1349]" captionStart="FIGURE 95" captionStartId="162.[136,235,1932,1956]" captionTargetBox="[175,1424,185,1902]" captionTargetId="figure-27@162.[164,1424,174,1913]" captionTargetPageId="162" captionText="FIGURE 95. caudal vertebra Ca2? of P. dawsonae sp. nov. AM F161911 in (a) lateral, (b) dorsal, and (c) cranial views.(d–f) partial pelvis, predominantly ilium and caudal component of ischium, of P. dawsonae sp. nov. UCMP 156881 in: (d) caudal, (e) lateral, and (f) medial views." figureDoi="http://doi.org/10.5281/zenodo.10993986" httpUri="https://zenodo.org/record/10993986/files/figure.png" pageId="163" pageNumber="164">Fig. 95a–c</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: numerical position of the vertebrae is not certain; vertebrae short and robust. Pre- and postzygopophyses heavily abraded in available specimens; prezygopophyses have a thick proximal section that is slightly dorsoventrally compressed and dorsally deflected; postzygopophyses short, robust and caudolaterally projected, with articular surfaces small, flat and round; bases of pre- and postzygopophyses merge at the centre of the arch around the base of the spinous process.
|
||
<collectingRegion id="4988F809FF4FD596FC72720DFB8CF989" box="[1018,1116,1611,1637]" country="France" name="Centre" pageId="163" pageNumber="164">Centrum</collectingRegion>
|
||
shallow, slightly arched in lateral view, with ventrally tilted cranial and caudal extremities; cranial extremity rounded; caudal extremity slightly dorsoventrally compressed and oval, with that of Ca2 slightly ventrally projected relative to the cranial extremity. Vertebral canal low and broad in Ca2; very low in Ca3. Transverse processes very deep in C2, extend from adjacent to the cranial margin of the centrum to adjacent to the caudal margin, dorsoventrally very thin, particularly cranially, caudally deflected, quite broad cranially and steadily broaden caudally; very abraded in Ca3, extend from slightly cranial of the midpoint of the centrum to abut the caudal margin with some caudal deflection.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4FD591FCD273ADFE82FE69" blockId="163.[810,1452,171,2085]" lastBlockId="164.[136,778,171,389]" lastPageId="164" lastPageNumber="165" pageId="163" pageNumber="164">
|
||
The Ca2? and 3? of
|
||
<taxonomicName id="4C4C4D68FF4FD596FBB573AAFB18F7E9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1085,1224,2027,2053]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FBB573AAFB18F7E9" box="[1085,1224,2027,2053]" italics="true" pageId="163" pageNumber="164">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FB4773ADFAF9F7E9" bold="true" box="[1231,1321,2027,2053]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FB4773ADFAF9F7E9" box="[1231,1321,2027,2053]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differ from those of
|
||
<taxonomicName id="4C4C4D68FF4FD596FC187C4AFBF3F7C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[912,1059,2059,2085]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FC187C4AFBF3F7C9" box="[912,1059,2059,2085]" italics="true" pageId="163" pageNumber="164">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FBA47C4DFB59F7C9" bold="true" box="[1068,1161,2059,2085]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FBA47C4DFB59F7C9" box="[1068,1161,2059,2085]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4FD596FB4D7C4AFAF6F7C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1221,1318,2060,2085]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4FD596FB4D7C4AFAF6F7C9" box="[1221,1318,2060,2085]" italics="true" pageId="163" pageNumber="164">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4FD596FAB87C4DFA5CF7C9" bold="true" box="[1328,1420,2059,2085]" pageId="163" pageNumber="164">
|
||
<taxonomicNameLabel id="A20B5782FF4FD596FAB87C4DFA5CF7C9" box="[1328,1420,2059,2085]" pageId="163" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller; from
|
||
<taxonomicName id="4C4C4D68FF48D591FEE374EDFE2BFF29" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[363,507,171,197]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF48D591FEE374EDFE2BFF29" box="[363,507,171,197]" italics="true" pageId="164" pageNumber="165">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being more elongate, with more laterally deflected prezygopophyses; from
|
||
<taxonomicName id="4C4C4D68FF48D591FD66748DFF12FEE9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF48D591FD66748DFF12FEE9" italics="true" pageId="164" pageNumber="165">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in having Ca2 broader and taller relative to length; from
|
||
<taxonomicName id="4C4C4D68FF48D591FF40754AFEBCFEC9" authorityName="Desmarest" authorityYear="1817" box="[200,364,267,293]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF48D591FF40754AFEBCFEC9" box="[200,364,267,293]" italics="true" pageId="164" pageNumber="165">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger, more robust and in having Ca3 slightly taller relative to length; and from
|
||
<taxonomicName id="4C4C4D68FF48D591FF00750AFF2EFE89" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[136,254,331,357]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF48D591FF00750AFF2EFE89" box="[136,254,331,357]" italics="true" pageId="164" pageNumber="165">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with more laterally deflected prezygopophyses.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FF0075ECFF2CFE29" blockId="164.[136,778,426,2085]" box="[136,252,426,453]" pageId="164" pageNumber="165">
|
||
<emphasis id="B938EAF9FF48D591FF0075ECFF2CFE29" bold="true" box="[136,252,426,453]" italics="true" pageId="164" pageNumber="165">Hindlimb</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FF00758DFEC8FB49" blockId="164.[136,778,426,2085]" pageId="164" pageNumber="165">
|
||
<emphasis id="B938EAF9FF48D591FF00758DFE86FE09" box="[136,342,459,485]" italics="true" pageId="164" pageNumber="165">
|
||
Pelvis (
|
||
<figureCitation id="13772A6EFF48D591FF52758DFE9FFE09" box="[218,335,459,485]" captionStart="FIGURE 95" captionStartId="162.[136,235,1932,1956]" captionTargetBox="[175,1424,185,1902]" captionTargetId="figure-27@162.[164,1424,174,1913]" captionTargetPageId="162" captionText="FIGURE 95. caudal vertebra Ca2? of P. dawsonae sp. nov. AM F161911 in (a) lateral, (b) dorsal, and (c) cranial views.(d–f) partial pelvis, predominantly ilium and caudal component of ischium, of P. dawsonae sp. nov. UCMP 156881 in: (d) caudal, (e) lateral, and (f) medial views." figureDoi="http://doi.org/10.5281/zenodo.10993986" httpUri="https://zenodo.org/record/10993986/files/figure.png" pageId="164" pageNumber="165">Fig. 95d–f</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: ilium robust, quite well-developed and roughly L-shaped in cross-section. Iliac crest epiphysis is not known; ilium gently increases to its maximum width at the iliac crest. Iliac fossa shallow craniomedial to the rectus tubercle, becomes shallower dorsally along the cranioventral surface of the ilium; extends around three-quarters of the length of the ilium. Gluteal fossa broad and moderately concave, particularly around the midpoint; extends to the iliac crest. Caudal iliac spine arises on the medial surface of the base of the ilium, opposite the rectus tubercule, thickens and rises rapidly, extends to the iliac crest; thick and deep ventrally for the origin of a large mm. gluteus, steadily becomes thinner and slightly shallower distally. Sacral surface large, very rugose and concave; articular surface for the wings of the sacrum deep and shelf-like with a strong cranial tilt; cranial margin of the sacral surface projects cranially in a broad, low ridge with a slight crest (cranial iliac spine) and contributes to the depth of the iliac fossa. Lateral iliac spine very broad, thin relative to the caudal spine; lateral margin slightly concave and curving laterally. Rectus tubercule quite rugose, roughly triangular, narrows strongly onto the lateral spine.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FF3070EDFDA4F989" blockId="164.[136,778,426,2085]" pageId="164" pageNumber="165">Acetabulum quite large, dorsoventrally tall and concave, taller than the craniocaudal depth, due chiefly to an enlarged caudoventral section; acetabular fossa deep, with the angle, width and curvature variable within individuals, partially covered cranially and caudally by lip projecting from the acetabular surface. Caudal surface of the dorsal section of the ischium (caudally adjacent to the acetabulum) broad and gently convex. Ischium badly crushed ventral to the acetabulum in the available specimen; craniocaudally deep and transversely compressed; caudal margin gently rugose and slightly convex from the origin of the mm. gemelli; deflected slightly caudally relative to the axis of the ilium; caudal section broadens toward the ventral margin.</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FF30722DFB8CFE49" blockId="164.[136,778,426,2085]" lastBlockId="164.[810,1451,171,421]" pageId="164" pageNumber="165">
|
||
The pelvis of
|
||
<taxonomicName id="4C4C4D68FF48D591FEE2722AFE2DF969" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[362,509,1643,1669]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF48D591FEE2722AFE2DF969" box="[362,509,1643,1669]" italics="true" pageId="164" pageNumber="165">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF48D591FD84722DFDBDF969" bold="true" box="[524,621,1643,1669]" pageId="164" pageNumber="165">
|
||
<taxonomicNameLabel id="A20B5782FF48D591FD84722DFDBDF969" box="[524,621,1643,1669]" pageId="164" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF48D591FF5372CAFEBEF949" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[219,366,1675,1701]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="mamkurra">
|
||
<emphasis id="B938EAF9FF48D591FF5372CAFEBEF949" box="[219,366,1675,1701]" italics="true" pageId="164" pageNumber="165">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF48D591FE2072CAFDD8F949" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[424,520,1676,1701]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="viator">
|
||
<emphasis id="B938EAF9FF48D591FE2072CAFDD8F949" box="[424,520,1676,1701]" italics="true" pageId="164" pageNumber="165">P. viator</emphasis>
|
||
</taxonomicName>
|
||
in being smaller, with a smaller caudal iliac spine, shallower gluteal fossa, narrower, less rugose and less convex caudal surface of the dorsal section of the ischium (
|
||
<figureCitation id="13772A6EFF48D591FD9172ADFDADF8EA" box="[537,637,1771,1798]" captionStart="FIGURE 95" captionStartId="162.[136,235,1932,1956]" captionTargetBox="[175,1424,185,1902]" captionTargetId="figure-27@162.[164,1424,174,1913]" captionTargetPageId="162" captionText="FIGURE 95. caudal vertebra Ca2? of P. dawsonae sp. nov. AM F161911 in (a) lateral, (b) dorsal, and (c) cranial views.(d–f) partial pelvis, predominantly ilium and caudal component of ischium, of P. dawsonae sp. nov. UCMP 156881 in: (d) caudal, (e) lateral, and (f) medial views." figureDoi="http://doi.org/10.5281/zenodo.10993986" httpUri="https://zenodo.org/record/10993986/files/figure.png" pageId="164" pageNumber="165">Fig. 95e</figureCitation>
|
||
), and a less deeply concave acetabulum; from
|
||
<taxonomicName id="4C4C4D68FF48D591FD81734AFD57F8C9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[521,647,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF48D591FD81734AFD57F8C9" box="[521,647,1803,1829]" italics="true" pageId="164" pageNumber="165">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having a more curved ilium in lateral view, less laterally projected rectus tubercle, acetabulum opening more laterally and less cranioventrally, and a more transversely compressed ischium; from
|
||
<taxonomicName id="4C4C4D68FF48D591FEB873CDFE12F849" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[304,450,1931,1957]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF48D591FEB873CDFE12F849" box="[304,450,1931,1957]" italics="true" pageId="164" pageNumber="165">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C4D68FF48D591FE5973CDFDFFF849" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[465,559,1931,1957]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF48D591FE5973CDFDFFF849" box="[465,559,1931,1957]" italics="true" pageId="164" pageNumber="165">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF48D591FDE073CAFCDAF849" authorityName="Desmarest" authorityYear="1817" box="[616,778,1931,1957]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF48D591FDE073CAFCDAF849" box="[616,778,1931,1957]" italics="true" pageId="164" pageNumber="165">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being more robust, with a more dorsally situated sacral surface relative to the position of the acetabulum, shallower, less distally extensive iliac fossa, broader, deeper gluteal fossa, and a smaller, less projected iliopubic eminence; and from
|
||
<taxonomicName id="4C4C4D68FF48D591FBAB74EAFB4DFF29" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[1059,1181,171,197]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="164" pageNumber="165" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF48D591FBAB74EAFB4DFF29" box="[1059,1181,171,197]" italics="true" pageId="164" pageNumber="165">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, with the iliac crest aligned more transversely and less craniocaudally, broader, deeper gluteal fossa, narrower, less distally extensive iliac fossa, broader lateral iliac spine, particularly dorsally, caudal iliac spine lacking a small, pointed eminence on the caudoventral shoulder, and the rectus tubercle having a much smaller, shallower fossa on the lateral surface.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FCA2758DFC84FE09" blockId="164.[810,1452,459,2085]" box="[810,852,459,485]" pageId="164" pageNumber="165">
|
||
<emphasis id="B938EAF9FF48D591FCA2758DFC84FE09" bold="true" box="[810,852,459,485]" italics="true" pageId="164" pageNumber="165">Pes</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FCA275ADFC1EFC29" blockId="164.[810,1452,459,2085]" pageId="164" pageNumber="165">
|
||
<emphasis id="B938EAF9FF48D591FCA275ADFBDFFDE9" box="[810,1039,491,517]" italics="true" pageId="164" pageNumber="165">
|
||
Calcaneus (
|
||
<figureCitation id="13772A6EFF48D591FC3B75ADFBD7FDE9" box="[947,1031,491,517]" captionStart="FIGURE 96" captionStartId="165.[136,235,1516,1540]" captionTargetBox="[162,1425,200,1485]" captionTargetId="figure-17@165.[151,1436,189,1495]" captionTargetPageId="165" captionText="FIGURE 96. partial calcanei of P. dawsonae sp. nov.: (a–e) surface scan images of partial left calcaneus AM F106042 in (a) dorsal, (b) plantar, (c) medial, (d) cranial, and (e) caudodorsal views; and (f) right calcaneus AM F161913 in dorsal view. AM F161913 is dorsoplantarly crushed, the tuberosity is warped medially, and it remains partially embedded in matrix and bone fragments, visible on the medial margin of the tuberosity." figureDoi="http://doi.org/10.5281/zenodo.10993988" httpUri="https://zenodo.org/record/10993988/files/figure.png" pageId="164" pageNumber="165">Fig. 96</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: moderately sized, robust and quite low. Calcaneal tuberosity short, broad and low, broadens plantarly and caudally, increases slightly in height caudally; domed in cross-section; lateral surface becomes deeply concave plantar to the fibular facets. Caudal epiphysis broad, thickened and rugose, oval to domed in posterior view; a shallow transverse groove extends across the caudal surface. Plantar surface thickened, rugose and quite broad, tapers gently cranially in plantar view and extends cranially to level with the cranial fibular facet, with the craniomedial margin deflected laterally around the cranial plantar tubercle. Cranial plantar tubercle generally small, rounded to oval, abuts or is caudally adjacent to the plantar margin of the plantomedial facet for the cuboid.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FCD2778DFBB3F969" blockId="164.[810,1452,459,2085]" pageId="164" pageNumber="165">The calcaneal head is large and broad, with a very slight medial rotation in dorsal view. Sustentaculum tali large, medially projected beyond the margin of the medial talar facet, not extensive caudally, caudal margin is rounded with a flat cranioplantar margin; flexor groove quite deep and broad. Medial talar facet slightly caudally displaced relative to lateral talar facet, broad and oval, craniocaudally compressed, slightly projected dorsally, tilted strongly cranially in medial view; caudal margin with a distinct rounded lip, particularly over the lateral part. A small, variable ridge extends caudally from the caudolateral margin of the medial talar facet, tapers and lowers caudally; occasionally present only as a slight swelling. Lateral talar facet broad, craniocaudally short, smoothly convex, roughly semicylindrical and tapering from the lateral margin to the midpoint.Fibular facets large relative to the lateral talar facet, bulbous and projected laterally; cranial component low, elongate, more laterally projected than the caudal component of the facet, and tilted cranially in lateral view; caudal component roughly rounded, gently convex, and facing caudally, with distinct plantar and medial margins.</paragraph>
|
||
<paragraph id="8BF336EBFF48D591FCD272CDFB8FF7C9" blockId="164.[810,1452,459,2085]" pageId="164" pageNumber="165">Facet for the talar head small; abuts the medial margin of the dorsomedial cuboid facet on the medial surface of the calcaneal head. Dorsomedial cuboid facet roughly oval in cranial view, gently convex, slightly broader than the dorsolateral facet, from which it is separated by a tall, quite bevelled step.Dorsolateral cuboid facet tall, cranially projected, very slightly concave and tilted plantarly and slightly medially, curves plantomedially to be continuous with the plantomedial facet. Plantomedial cuboid facet much smaller than the dorsal facets, dorsoplantarly compressed and broad, occasionally rounded. A shallow, oblong fossa is present between the dorsomedial and plantomedial cuboid facets.</paragraph>
|
||
<caption id="DF336663FF49D590FF0071AAFDC2F989" ID-DOI="http://doi.org/10.5281/zenodo.10993988" ID-Zenodo-Dep="10993988" httpUri="https://zenodo.org/record/10993988/files/figure.png" pageId="165" pageNumber="166" startId="165.[136,235,1516,1540]" targetBox="[162,1425,200,1485]" targetPageId="165" targetType="figure">
|
||
<paragraph id="8BF336EBFF49D590FF0071AAFDC2F989" blockId="165.[136,1452,1516,1637]" pageId="165" pageNumber="166">
|
||
<emphasis id="B938EAF9FF49D590FF0071AAFEDFF9E9" bold="true" box="[136,271,1516,1541]" pageId="165" pageNumber="166">FIGURE 96.</emphasis>
|
||
partial calcanei of
|
||
<taxonomicName id="4C4C4D68FF49D590FE4571ABFD9AF9E9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[461,586,1517,1541]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF49D590FE4571ABFD9AF9E9" box="[461,586,1517,1541]" italics="true" pageId="165" pageNumber="166">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF49D590FDC771ABFD4FF9E9" bold="true" box="[591,671,1517,1541]" pageId="165" pageNumber="166">
|
||
<taxonomicNameLabel id="A20B5782FF49D590FDC771ABFD4FF9E9" box="[591,671,1517,1541]" pageId="165" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: (a–e) surface scan images of partial left calcaneus AM F106042 in (a) dorsal, (b) plantar, (c) medial, (d) cranial, and (e) caudodorsal views; and (f) right calcaneus AM F161913 in dorsal view. AM F161913 is dorsoplantarly crushed, the tuberosity is warped medially, and it remains partially embedded in matrix and bone fragments, visible on the medial margin of the tuberosity.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF49D593FF30728DFE1AF809" blockId="165.[136,779,1739,2085]" lastBlockId="166.[136,779,1547,2085]" lastPageId="166" lastPageNumber="167" pageId="165" pageNumber="166">
|
||
The calcaneus of
|
||
<taxonomicName id="4C4C4D68FF49D590FE0C728AFDC2F909" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[388,530,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF49D590FE0C728AFDC2F909" box="[388,530,1739,1765]" italics="true" pageId="165" pageNumber="166">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF49D590FD94728DFDA8F909" bold="true" box="[540,632,1739,1765]" pageId="165" pageNumber="166">
|
||
<taxonomicNameLabel id="A20B5782FF49D590FD94728DFDA8F909" box="[540,632,1739,1765]" pageId="165" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF49D590FF6A72AAFEE8F8E9" authorityName=", Owen" authorityYear="1874" box="[226,312,1771,1797]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF49D590FF6A72AAFEE8F8E9" box="[226,312,1771,1797]" italics="true" pageId="165" pageNumber="166">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being dorsoplantarly shorter, with a narrower plantar surface, slightly shallower lateral talar facet, less rounded, less distinct caudal fibular facet, and a shallower flexor groove; from
|
||
<taxonomicName id="4C4C4D68FF49D590FE70730AFD5AF889" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[504,650,1867,1893]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF49D590FE70730AFD5AF889" box="[504,650,1867,1893]" italics="true" pageId="165" pageNumber="166">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF49D590FD19730DFD3CF889" bold="true" box="[657,748,1867,1893]" pageId="165" pageNumber="166">
|
||
<taxonomicNameLabel id="A20B5782FF49D590FD19730DFD3CF889" box="[657,748,1867,1893]" pageId="165" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, lower and narrower, with a more caudally displaced medial talar facet, less bulbous lateral talar and caudal fibular facets, fibular facets less plantarly extensive laterally, and a more rounded sustentaculum tali; from
|
||
<taxonomicName id="4C4C4D68FF49D590FD7D738AFF1BF7E9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF49D590FD7D738AFF1BF7E9" italics="true" pageId="165" pageNumber="166">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF49D590FF5973ADFEFAF7E9" bold="true" box="[209,298,2027,2053]" pageId="165" pageNumber="166">
|
||
<taxonomicNameLabel id="A20B5782FF49D590FF5973ADFEFAF7E9" box="[209,298,2027,2053]" pageId="165" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being smaller, lower and broader, with a more rounded, less dorsally pointed calcaneal tuberosity in cross-section, craniocaudally shorter lateral talar facet, broader, more transversely aligned medial talar facet, and more medially projected sustentaculum tali; from
|
||
<taxonomicName id="4C4C4D68FF49D590FA1F734AFC5CF8A9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF49D590FA1F734AFC5CF8A9" italics="true" pageId="165" pageNumber="166">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in having the head less medially displaced, a flatter, less convex plantar surface, and less medial flaring of the caudoplantar margin of the calcaneal tuberosity; from
|
||
<taxonomicName id="4C4C4D68FF49D590FCF973CAFBD3F849" authorityName="Plane" authorityYear="1967" box="[881,1027,1931,1957]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF49D590FCF973CAFBD3F849" box="[881,1027,1931,1957]" italics="true" pageId="165" pageNumber="166">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in having the head less medially displaced, with a narrower calcaneal tuberosity; from
|
||
<taxonomicName id="4C4C4D68FF49D590FCA2738DFC6DF809" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[810,957,1995,2021]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF49D590FCA2738DFC6DF809" box="[810,957,1995,2021]" italics="true" pageId="165" pageNumber="166">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger and relatively taller, with a less concave, less medially tilted plantar surface; from
|
||
<taxonomicName id="4C4C4D68FF49D590FA1873ADFCB6F7C9" baseAuthorityName="Desmarest" baseAuthorityYear="1822" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="165" pageNumber="166" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF49D590FA1873ADFCB6F7C9" italics="true" pageId="165" pageNumber="166">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being broader and relatively lower, with a less elongate tuberosity, no caudomedial facet for the posterior plantar process of the talus, less cranially tilted medial talar facet, shallower fossa cranial to the lateral talar facet, relatively narrower dorsolateral cuboid facet, more bevelled step between the dorsal cuboid facets, more medially projected sustentaculum tali, and a deeper flexor groove; from
|
||
<taxonomicName id="4C4C4D68FF4AD593FEAD728AFE17F909" authorityName="Desmarest" authorityYear="1817" box="[293,455,1739,1765]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="166" pageNumber="167" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF4AD593FEAD728AFE17F909" box="[293,455,1739,1765]" italics="true" pageId="166" pageNumber="167">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger, broader and relatively lower, with a less elongate tuberosity, shallower fossa cranial to the lateral talar facet, relatively narrower dorsolateral cuboid facet, more bevelled step between the dorsal cuboid facets, more medially projected sustentaculum tali, and a deeper flexor groove; and from
|
||
<taxonomicName id="4C4C4D68FF4AD593FF0073CAFED0F849" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[136,256,1931,1957]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="166" pageNumber="167" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4AD593FF0073CAFED0F849" box="[136,256,1931,1957]" italics="true" pageId="166" pageNumber="167">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, relatively broader and more robust, with broader medial talar facet and more medially projected sustentaculum tali.
|
||
</paragraph>
|
||
<caption id="DF336663FF4AD593FF00716AFB45FA49" ID-DOI="http://doi.org/10.5281/zenodo.10993990" ID-Zenodo-Dep="10993990" httpUri="https://zenodo.org/record/10993990/files/figure.png" pageId="166" pageNumber="167" startId="166.[136,235,1324,1348]" targetBox="[162,1425,195,1292]" targetPageId="166" targetType="figure">
|
||
<paragraph id="8BF336EBFF4AD593FF00716AFB45FA49" blockId="166.[136,1451,1324,1445]" pageId="166" pageNumber="167">
|
||
<emphasis id="B938EAF9FF4AD593FF00716AFEC0FAA9" bold="true" box="[136,272,1324,1349]" pageId="166" pageNumber="167">FIGURE 97.</emphasis>
|
||
pedal elements of
|
||
<taxonomicName id="4C4C4D68FF4AD593FE58716BFD9EFAA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[464,590,1325,1349]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="166" pageNumber="167" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4AD593FE58716BFD9EFAA9" box="[464,590,1325,1349]" italics="true" pageId="166" pageNumber="167">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4AD593FDDD716BFD75FAA9" bold="true" box="[597,677,1325,1349]" pageId="166" pageNumber="167">
|
||
<taxonomicNameLabel id="A20B5782FF4AD593FDDD716BFD75FAA9" box="[597,677,1325,1349]" pageId="166" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: (a) surface scan image of partial right metatarsal IV AM F106039 in dorsal view; (b–c) surface scan images of partial left metatarsal IV of AM F106041 in (b) proximal, and (c) lateral views; (d–h) surface scan images of left metatarsal V of AM F106040 in (d) dorsal, (e) plantar, (f) lateral, (g) medial, and (h) proximal views; and (i–k) partial right proximal pedal phalanx IV of AM F161916 in (i) dorsal, (j) medial, and (k) plantar views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336EBFF4AD592FF3073ADFE0BFE89" blockId="166.[136,779,1547,2085]" lastBlockId="167.[136,779,171,2085]" lastPageId="167" lastPageNumber="168" pageId="166" pageNumber="167">
|
||
<emphasis id="B938EAF9FF4AD593FF3073ADFE2AF7E9" box="[184,506,2027,2053]" italics="true" pageId="166" pageNumber="167">
|
||
Metatarsal IV (
|
||
<figureCitation id="13772A6EFF4AD593FEF973ADFE21F7E9" box="[369,497,2027,2053]" captionStart="FIGURE 97" captionStartId="166.[136,235,1324,1348]" captionTargetBox="[162,1425,195,1292]" captionTargetId="figure-27@166.[151,1436,184,1303]" captionTargetPageId="166" captionText="FIGURE 97. pedal elements of P. dawsonae sp. nov.: (a) surface scan image of partial right metatarsal IV AM F106039 in dorsal view; (b–c) surface scan images of partial left metatarsal IV of AM F106041 in (b) proximal, and (c) lateral views; (d–h) surface scan images of left metatarsal V of AM F106040 in (d) dorsal, (e) plantar, (f) lateral, (g) medial, and (h) proximal views; and (i–k) partial right proximal pedal phalanx IV of AM F161916 in (i) dorsal, (j) medial, and (k) plantar views." figureDoi="http://doi.org/10.5281/zenodo.10993990" httpUri="https://zenodo.org/record/10993990/files/figure.png" pageId="166" pageNumber="167">Fig. 97a–c</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: large, long and quite robust. Dorsal cuboid facet very similar to that of
|
||
<taxonomicName id="4C4C4D68FF4AD593FD7E7C4AFC4CF9C9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[758,924,1547,2085]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="166" pageNumber="167" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4AD593FD7E7C4AFC4CF9C9" box="[758,924,1547,2085]" italics="true" pageId="166" pageNumber="167">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4AD593FC2A724DFC2BF9C9" bold="true" box="[930,1019,1547,1573]" pageId="166" pageNumber="167">
|
||
<taxonomicNameLabel id="A20B5782FF4AD593FC2A724DFC2BF9C9" box="[930,1019,1547,1573]" pageId="166" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; proximal cuboid fossa small, shallow and plantolaterally situated; plantar cuboid facet quite small, round, very gently concave and tilted distinctly dorsally and slightly medially, extends plantarly partially onto proximal surface of deep, plantarly projected plantar tubercle. Proximal plantar sesamoid facet quite large, broad, flat, round to oval, faces distoplantarly and is tilted laterally. Articular surface for metatarsal III indistinct, situated in an elongate, shallow, rugose metatarsal III fossa on the medial surface of the proximal end, bordered dorsally by an indistinct, low ridge extending plantodistally from the dorsomedial corner of the dorsal cuboid facet. Dorsal facet for the ectocuneiform small and tapering plantarly, facing proximomedially, adjacent to the dorsal section of medial margin of dorsal cuboid facet; small facet for the plantolateral component of ectocuneiform situated on the proximomedial surface of the plantar tubercle, semicontinuous with the plantar cuboid facet. Facet for metatarsal V large, concave, slightly cranially tilted, moderately tall and proximodistally short; shape variable but generally oblong with rounded dorsal and plantar parts; extends plantarly onto the base of the lateral surface of the plantar tubercle.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4BD592FF30752DFE4FFD49" blockId="167.[136,779,171,2085]" pageId="167" pageNumber="168">
|
||
Plantar ridge quite broad, rugose, squarish to rounded in cross-section, extends distally with a very slight lateral deflection from the base of the plantar tubercle, merges into the plantar surface of the shaft before the midpoint; bordered medially by the fossa for metatarsal III and laterally by the fossa for metatarsal V. Shaft morphology very similar to that of
|
||
<taxonomicName id="4C4C4D68FF4BD592FE19766AFDF4FDA9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[401,548,555,581]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FE19766AFDF4FDA9" box="[401,548,555,581]" italics="true" pageId="167" pageNumber="168">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FDA6766DFD5AFDA9" bold="true" box="[558,650,555,581]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FDA6766DFD5AFDA9" box="[558,650,555,581]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Distal end broad; fossae for the collateral ligaments circular and quite deep; keel slightly more plantarly projected than the lateral and medial crests.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4BD592FF3076EDFE7BFA89" blockId="167.[136,779,171,2085]" pageId="167" pageNumber="168">
|
||
The metatarsal IV of
|
||
<taxonomicName id="4C4C4D68FF4BD592FE2776EAFDECFD29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[431,572,683,709]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FE2776EAFDECFD29" box="[431,572,683,709]" italics="true" pageId="167" pageNumber="168">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FDCD76EDFD71FD29" bold="true" box="[581,673,683,709]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FDCD76EDFD71FD29" box="[581,673,683,709]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
does not differ from those of
|
||
<taxonomicName id="4C4C4D68FF4BD592FEF2768AFE1FFD09" authorityName=", Owen" authorityYear="1874" box="[378,463,715,741]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF4BD592FEF2768AFE1FFD09" box="[378,463,715,741]" italics="true" pageId="167" pageNumber="168">P. anak</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4BD592FD86768AFD72FD09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[526,674,715,741]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FD86768AFD72FD09" box="[526,674,715,741]" italics="true" pageId="167" pageNumber="168">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FD24768DFCDAFD09" bold="true" box="[684,778,715,741]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FD24768DFCDAFD09" box="[684,778,715,741]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
It differs from that of
|
||
<taxonomicName id="4C4C4D68FF4BD592FE0276AAFE3CFCE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[394,492,748,773]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FE0276AAFE3CFCE9" box="[394,492,748,773]" italics="true" pageId="167" pageNumber="168">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FE7D76ADFD82FCE9" bold="true" box="[501,594,747,773]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FE7D76ADFD82FCE9" box="[501,594,747,773]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being larger, with a relatively broader proximal plantar sesamoid facet and a larger plantar facet for the ectocuneiform; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FD7D776AFF3AFC89" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF4BD592FD7D776AFF3AFC89" italics="true" pageId="167" pageNumber="168">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in being larger; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FE5F770AFD8FFC89" authorityName="Plane" authorityYear="1967" box="[471,607,843,869]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF4BD592FE5F770AFD8FFC89" box="[471,607,843,869]" italics="true" pageId="167" pageNumber="168">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with continuous dorsal and plantar cuboid facets, a more concave, less laterally projected metatarsal V facet, larger plantar tubercle, more raised plantar ridge, and a relatively more plantarly projected keel; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FDA5778AFD72FC09" authorityName="Bartholomai" authorityYear="1978" box="[557,674,972,997]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="snewini">
|
||
<emphasis id="B938EAF9FF4BD592FDA5778AFD72FC09" box="[557,674,972,997]" italics="true" pageId="167" pageNumber="168">P. snewini</emphasis>
|
||
</taxonomicName>
|
||
in being larger, with a smaller, more plantolaterally situated cuboid fossa; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FE92704DFE60FBC9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[282,432,1035,1061]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4BD592FE92704DFE60FBC9" box="[282,432,1035,1061]" italics="true" pageId="167" pageNumber="168">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger and broader, with a larger, more plantarly projected proximal plantar tubercle and a more raised plantar ridge; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FD23700DFCDAFB89" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[683,778,1099,1125]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF4BD592FD23700DFCDAFB89" box="[683,778,1099,1125]" italics="true" pageId="167" pageNumber="168">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4BD592FF34702AFEB1FB69" authorityName="Desmarest" authorityYear="1817" box="[188,353,1131,1157]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF4BD592FF34702AFEB1FB69" box="[188,353,1131,1157]" italics="true" pageId="167" pageNumber="168">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being shorter, much broader and more robust, with a larger facet for metatarsal V, the shaft more dorsoplantarly compressed and broadening more to the distal end, a less plantarly projected proximoplantar ridge, and a broader, more planar proximal dorsal surface; and from
|
||
<taxonomicName id="4C4C4D68FF4BD592FE8C714AFEAFFAC9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[260,383,1291,1317]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4BD592FE8C714AFEAFFAC9" box="[260,383,1291,1317]" italics="true" pageId="167" pageNumber="168">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being much larger, relatively broader and more robust, with a relatively slightly larger proximal plantar tubercle.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4BD592FF30712DFAB0FF09" blockId="167.[136,779,171,2085]" lastBlockId="167.[810,1452,171,2053]" pageId="167" pageNumber="168">
|
||
<emphasis id="B938EAF9FF4BD592FF30712DFE30FA69" box="[184,480,1387,1413]" italics="true" pageId="167" pageNumber="168">
|
||
Metatarsal V (
|
||
<figureCitation id="13772A6EFF4BD592FED3712DFE08FA69" box="[347,472,1387,1413]" captionStart="FIGURE 97" captionStartId="166.[136,235,1324,1348]" captionTargetBox="[162,1425,195,1292]" captionTargetId="figure-27@166.[151,1436,184,1303]" captionTargetPageId="166" captionText="FIGURE 97. pedal elements of P. dawsonae sp. nov.: (a) surface scan image of partial right metatarsal IV AM F106039 in dorsal view; (b–c) surface scan images of partial left metatarsal IV of AM F106041 in (b) proximal, and (c) lateral views; (d–h) surface scan images of left metatarsal V of AM F106040 in (d) dorsal, (e) plantar, (f) lateral, (g) medial, and (h) proximal views; and (i–k) partial right proximal pedal phalanx IV of AM F161916 in (i) dorsal, (j) medial, and (k) plantar views." figureDoi="http://doi.org/10.5281/zenodo.10993990" httpUri="https://zenodo.org/record/10993990/files/figure.png" pageId="167" pageNumber="168">Fig. 97d–h</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: quite elongate and gently transversely compressed proximally, with a distinct lateral curve to the distal component in dorsal view accentuated by the lateral expansion of the distal end. Proximolateral process blunt, thick and rugose. Cuboid facet broad, broader than the facet for metatarsal IV, concave, particularly medially, extends across the medial surface of the proximolateral process onto the lateral base of the medial plantar tubercle; dorsolateral margin raised to slight lip over dorsomedial surface of proximolateral process. Facet for metatarsal IV proximodistally deep but tapering laterally, and gently convex; extends from the dorsal surface of the shaft onto the medial plantar tubercle. Lateral plantar tuberosity large, broad, rugose and quite raised, merges with the shaft just past the midpoint; separated from the medial plantar tubercle by a broad, shallow plantar groove curving distally along the medial margin. Medial plantar tubercle small, distinct, rounded, and proximomedially projected. Shaft gently and smoothly arched in lateral view with the highest point around the midpoint. Distal end broad, laterally flared and slightly laterally deflected at base. Distal articular surface very similar to that of
|
||
<taxonomicName id="4C4C4D68FF4BD592FBBA74EAFB15FF29" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1074,1221,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FBBA74EAFB15FF29" box="[1074,1221,171,197]" italics="true" pageId="167" pageNumber="168">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FB4674EDFAFBFF29" bold="true" box="[1230,1323,171,197]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FB4674EDFAFBFF29" box="[1230,1323,171,197]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Fossae for collateral ligaments large, rounded and quite deep.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4BD592FCD274ADFBCEFBA9" blockId="167.[810,1452,171,2053]" pageId="167" pageNumber="168">
|
||
The metatarsal V of
|
||
<taxonomicName id="4C4C4D68FF4BD592FBDF74AAFB39FEE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1111,1257,235,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FBDF74AAFB39FEE9" box="[1111,1257,235,261]" italics="true" pageId="167" pageNumber="168">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FB7E74ADFA86FEE9" bold="true" box="[1270,1366,235,261]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FB7E74ADFA86FEE9" box="[1270,1366,235,261]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that of
|
||
<taxonomicName id="4C4C4D68FF4BD592FC35754AFBC1FEC9" authorityName=", Owen" authorityYear="1874" box="[957,1041,267,293]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF4BD592FC35754AFBC1FEC9" box="[957,1041,267,293]" italics="true" pageId="167" pageNumber="168">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in having less raised lateral plantar tuberosity, less tall proximolateral process and broader cuboid facet; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FB9A750AFB79FE89" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1042,1193,331,357]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FB9A750AFB79FE89" box="[1042,1193,331,357]" italics="true" pageId="167" pageNumber="168">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FB3E750DFAC5FE89" bold="true" box="[1206,1301,331,357]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FB3E750DFAC5FE89" box="[1206,1301,331,357]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a narrower cuboid facet and a more distally extensive lateral plantar tuberosity; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FB0175CAFB38FE49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1161,1256,396,421]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FB0175CAFB38FE49" box="[1161,1256,396,421]" italics="true" pageId="167" pageNumber="168">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FB6775CDFA99FE49" bold="true" box="[1263,1353,395,421]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FB6775CDFA99FE49" box="[1263,1353,395,421]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being longer and slightly less transversely compressed, with a lower, more elongate proximolateral process, narrower cuboid facet, more proximally projected medial plantar tubercle, and a less raised lateral plantar tuberosity; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FCA2766AFC7DFDA9" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[810,941,555,581]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="tumbuna">
|
||
<emphasis id="B938EAF9FF4BD592FCA2766AFC7DFDA9" box="[810,941,555,581]" italics="true" pageId="167" pageNumber="168">P. tumbuna</emphasis>
|
||
</taxonomicName>
|
||
in being larger and more elongate, with a longer, narrower proximolateral process, larger medial plantar tubercle, and a narrower cuboid facet; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FA1F762AFC48FD49" authorityName="Plane" authorityYear="1967" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF4BD592FA1F762AFC48FD49" italics="true" pageId="167" pageNumber="168">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in being more elongate, with a narrower, less dorsally projected cuboid facet and a slightly narrower distal end; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FC7B768DFB59FD09" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1011,1161,715,741]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4BD592FC7B768DFB59FD09" box="[1011,1161,715,741]" italics="true" pageId="167" pageNumber="168">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being larger, broader and more robust, lacking the slight kink of the arch of the shaft immediately proximal to the midpoint in lateral view, with a larger medial plantar tubercle, more medially situated facet for metatarsal IV, and a more raised lateral plantar tuberosity; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FBCF772DFB78FC69" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1095,1192,875,901]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF4BD592FBCF772DFB78FC69" box="[1095,1192,875,901]" italics="true" pageId="167" pageNumber="168">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D68FF4BD592FB6E772AFA5AFC69" authorityName="Desmarest" authorityYear="1817" box="[1254,1418,875,901]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF4BD592FB6E772AFA5AFC69" box="[1254,1418,875,901]" italics="true" pageId="167" pageNumber="168">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being far shorter and broader, less arched and much less transversely compressed, with a longer proximolateral process and a larger medial plantar tubercle; and from
|
||
<taxonomicName id="4C4C4D68FF4BD592FCA277AAFC72FBE9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[810,930,1003,1029]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4BD592FCA277AAFC72FBE9" box="[810,930,1003,1029]" italics="true" pageId="167" pageNumber="168">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, relatively broader and more robust, with a larger proximolateral process and a more concave cuboid facet.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4BD592FCD2700DFC6AFA69" blockId="167.[810,1452,171,2053]" pageId="167" pageNumber="168">
|
||
<emphasis id="B938EAF9FF4BD592FCD2700DFAC9FB89" box="[858,1305,1099,1125]" italics="true" pageId="167" pageNumber="168">
|
||
Proximal pedal phalanx IV (
|
||
<figureCitation id="13772A6EFF4BD592FB13700DFAC1FB89" box="[1179,1297,1099,1125]" captionStart="FIGURE 97" captionStartId="166.[136,235,1324,1348]" captionTargetBox="[162,1425,195,1292]" captionTargetId="figure-27@166.[151,1436,184,1303]" captionTargetPageId="166" captionText="FIGURE 97. pedal elements of P. dawsonae sp. nov.: (a) surface scan image of partial right metatarsal IV AM F106039 in dorsal view; (b–c) surface scan images of partial left metatarsal IV of AM F106041 in (b) proximal, and (c) lateral views; (d–h) surface scan images of left metatarsal V of AM F106040 in (d) dorsal, (e) plantar, (f) lateral, (g) medial, and (h) proximal views; and (i–k) partial right proximal pedal phalanx IV of AM F161916 in (i) dorsal, (j) medial, and (k) plantar views." figureDoi="http://doi.org/10.5281/zenodo.10993990" httpUri="https://zenodo.org/record/10993990/files/figure.png" pageId="167" pageNumber="168">Fig. 97i–k</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: fairly short, robust, and dorsoplantarly compressed. Proximal articular surface mostly abraded in available specimen, appears domed and gently concave. Proximal plantar tubercles partially abraded, appear low and rugose, separated by a shallow valley. Shaft narrows slightly to a waist, broadens very slightly to the distal end. Distal articular surface proximodorsally quite extensive; trochlea broad and quite shallow. Fossae for the collateral ligaments shallow and semicircular.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF4BD592FCD271CDFC68F7E9" blockId="167.[810,1452,171,2053]" pageId="167" pageNumber="168">
|
||
The proximal phalanx IV of
|
||
<taxonomicName id="4C4C4D68FF4BD592FB3971CAFA92FA49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1201,1346,1419,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FB3971CAFA92FA49" box="[1201,1346,1419,1445]" italics="true" pageId="167" pageNumber="168">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FAC571CDFA7BFA49" bold="true" box="[1357,1451,1419,1445]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FAC571CDFA7BFA49" box="[1357,1451,1419,1445]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from those of
|
||
<taxonomicName id="4C4C4D68FF4BD592FBAA71EDFB51FA29" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1058,1153,1451,1477]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF4BD592FBAA71EDFB51FA29" box="[1058,1153,1451,1477]" italics="true" pageId="167" pageNumber="168">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
and all compared species of
|
||
<taxonomicName id="4C4C4D68FF4BD592FCC6718DFC35FA09" authorityName="Owen" authorityYear="1874" box="[846,997,1483,1509]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF4BD592FCC6718DFC35FA09" box="[846,997,1483,1509]" italics="true" pageId="167" pageNumber="168">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C4C4D68FF4BD592FBCF718AFB7BFA09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[1095,1195,1484,1509]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FBCF718AFB7BFA09" box="[1095,1195,1484,1509]" italics="true" pageId="167" pageNumber="168">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FB30718DFAC7FA09" bold="true" box="[1208,1303,1483,1509]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FB30718DFAC7FA09" box="[1208,1303,1483,1509]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a more proximodorsally extensive distal articular surface. It further differs from
|
||
<taxonomicName id="4C4C4D68FF4BD592FBA1724AFBACF9C9" authorityName=", Owen" authorityYear="1874" box="[1065,1148,1547,1573]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF4BD592FBA1724AFBACF9C9" box="[1065,1148,1547,1573]" italics="true" pageId="167" pageNumber="168">P. anak</emphasis>
|
||
</taxonomicName>
|
||
in being shorter and more robust, with less projected plantar tubercles, a slightly broader waist, and a less dorsoplantarly compressed shaft; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FC3A722AFB95F969" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[946,1093,1643,1669]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="mamkurra" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FC3A722AFB95F969" box="[946,1093,1643,1669]" italics="true" pageId="167" pageNumber="168">P. mamkurra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FBC7722DFB7BF969" bold="true" box="[1103,1195,1643,1669]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FBC7722DFB7BF969" box="[1103,1195,1643,1669]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being shorter, with a slightly broader waist, and a slightly narrower distal end; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FC2872EAFBD0F929" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[928,1024,1708,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF4BD592FC2872EAFBD0F929" box="[928,1024,1708,1733]" italics="true" pageId="167" pageNumber="168">P. viator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF4BD592FB8072EDFBB2F929" bold="true" box="[1032,1122,1707,1733]" pageId="167" pageNumber="168">
|
||
<taxonomicNameLabel id="A20B5782FF4BD592FB8072EDFBB2F929" box="[1032,1122,1707,1733]" pageId="167" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in being more robust, with a broader waist and a narrower distal end; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FAAB728AFA7BF909" authorityName="Plane" authorityYear="1967" box="[1315,1451,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="otibandus">
|
||
<emphasis id="B938EAF9FF4BD592FAAB728AFA7BF909" box="[1315,1451,1739,1765]" italics="true" pageId="167" pageNumber="168">P. otibandus</emphasis>
|
||
</taxonomicName>
|
||
in being more elongate; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FB2872ADFAE9F8E9" baseAuthorityName="Flannery" baseAuthorityYear="1989" box="[1184,1337,1771,1797]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="kitcheneri">
|
||
<emphasis id="B938EAF9FF4BD592FB2872ADFAE9F8E9" box="[1184,1337,1771,1797]" italics="true" pageId="167" pageNumber="168">C. kitcheneri</emphasis>
|
||
</taxonomicName>
|
||
in being much broader and more robust, and lacking a small fossa immediately proximal to the dorsal margin of the distal articular surface; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FBB8730DFB5EF889" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[1072,1166,1867,1893]" class="Mammalia" family="Macropodidae" genus="Osphranter" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="B938EAF9FF4BD592FBB8730DFB5EF889" box="[1072,1166,1867,1893]" italics="true" pageId="167" pageNumber="168">O. rufus</emphasis>
|
||
</taxonomicName>
|
||
in being slightly shorter, much broader and more robust, with a broader waist and a less dorsoplantarly compressed distal shaft; from
|
||
<taxonomicName id="4C4C4D68FF4BD592FA0573CAFC75F829" authorityName="Desmarest" authorityYear="1817" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="fuliginosus">
|
||
<emphasis id="B938EAF9FF4BD592FA0573CAFC75F829" italics="true" pageId="167" pageNumber="168">M. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
in being larger, far broader and more robust; and from
|
||
<taxonomicName id="4C4C4D68FF4BD592FC1C738AFBD7F809" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[916,1031,1995,2021]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="167" pageNumber="168" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B938EAF9FF4BD592FC1C738AFBD7F809" box="[916,1031,1995,2021]" italics="true" pageId="167" pageNumber="168">W. bicolor</emphasis>
|
||
</taxonomicName>
|
||
in being larger, relatively broader and more robust.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566560FF44D59DFF0074EDFDD5FCA9" pageId="168" pageNumber="169" type="discussion">
|
||
<paragraph id="8BF336EBFF44D59DFF0074EDFED4FF29" blockId="168.[136,778,171,837]" box="[136,260,171,197]" pageId="168" pageNumber="169">
|
||
<emphasis id="B938EAF9FF44D59DFF0074EDFED4FF29" bold="true" box="[136,260,171,197]" pageId="168" pageNumber="169">Remarks:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFF00748DFD51FEA9" blockId="168.[136,778,171,837]" pageId="168" pageNumber="169">
|
||
For discussion of the taxon
|
||
<taxonomicName id="4C4C4D68FF44D59DFD90748DFE8FFEE9" authority="Bartholomai, 1973" authorityName=", Bartholomai" authorityYear="1973" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="devisi">
|
||
<emphasis id="B938EAF9FF44D59DFD90748DFCD9FF09" box="[536,777,203,229]" italics="true" pageId="168" pageNumber="169">Protemnodon devisi</emphasis>
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFF0074ADFE8FFEE9" author="Bartholomai, A." box="[136,351,235,261]" pageId="168" pageNumber="169" pagination="309 - 363" refId="ref143087" refString="Bartholomai, A. (1973) The genus Protemnodon Owen (Marsupialia: Macropodidae) in the Upper Cainozoic deposits of Queensland. Memoirs of the Queensland Museum, 16 (3), 309 - 363." type="journal article" year="1973">Bartholomai, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
and its relationship to
|
||
<taxonomicName id="4C4C4D68FF44D59DFDF174AAFCDAFEE9" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[633,778,235,261]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF44D59DFDF174AAFCDAFEE9" box="[633,778,235,261]" italics="true" pageId="168" pageNumber="169">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF44D59DFF00754DFF35FEC9" bold="true" box="[136,229,267,293]" pageId="168" pageNumber="169">
|
||
<taxonomicNameLabel id="A20B5782FF44D59DFF00754DFF35FEC9" box="[136,229,267,293]" pageId="168" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, see the segment below titled, ‘Status of other species previously referred to
|
||
<taxonomicName id="4C4C4D68FF44D59DFE51756DFDA0FEA9" authorityName="Owen" authorityYear="1874" box="[473,624,299,325]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF44D59DFE51756DFDA0FEA9" box="[473,624,299,325]" italics="true" pageId="168" pageNumber="169">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
’.
|
||
</paragraph>
|
||
<paragraph id="8BF336EBFF44D59DFF30750DFDD5FCA9" blockId="168.[136,778,171,837]" pageId="168" pageNumber="169">
|
||
The material from Big Sink LF provides evidence for only one species of
|
||
<taxonomicName id="4C4C4D68FF44D59DFEE7752DFCD5FE69" authority="(Dawson et al. 1999)" baseAuthorityName="Dawson" baseAuthorityYear="1999" box="[367,773,363,389]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF44D59DFEE7752DFDD6FE69" box="[367,518,363,389]" italics="true" pageId="168" pageNumber="169">Protemnodon</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDD4B1AFF44D59DFD9F752DFD2DFE69" author="Dawson, L. & Muirhead, J. & Wroe, S." box="[535,765,363,389]" pageId="168" pageNumber="169" pagination="265 - 290" refId="ref145233" refString="Dawson, L., Muirhead, J. & Wroe, S. (1999) The Big Sink Local Fauna: a lower Pliocene mammalian fauna from the Wellington Caves complex, Wellington, New South Wales. Records of the Western Australian Museum, Supplement No. 57, 265 - 290." type="book chapter" year="1999">
|
||
Dawson
|
||
<emphasis id="B938EAF9FF44D59DFDF6752AFD69FE69" box="[638,697,363,389]" italics="true" pageId="168" pageNumber="169">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, all of which is allocated here to
|
||
<taxonomicName id="4C4C4D68FF44D59DFD9875CAFD70FE49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[528,672,395,421]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF44D59DFD9875CAFD70FE49" box="[528,672,395,421]" italics="true" pageId="168" pageNumber="169">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF44D59DFD2375CDFCDAFE49" bold="true" box="[683,778,395,421]" pageId="168" pageNumber="169">
|
||
<taxonomicNameLabel id="A20B5782FF44D59DFD2375CDFCDAFE49" box="[683,778,395,421]" pageId="168" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
These fossils are of a large kangaroo, similar in size to
|
||
<taxonomicName id="4C4C4D68FF44D59DFF00758AFF0AFE09" authorityName=", Owen" authorityYear="1874" box="[136,218,459,485]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="anak">
|
||
<emphasis id="B938EAF9FF44D59DFF00758AFF0AFE09" box="[136,218,459,485]" italics="true" pageId="168" pageNumber="169">P. anak</emphasis>
|
||
</taxonomicName>
|
||
, separated from all other species of
|
||
<taxonomicName id="4C4C4D68FF44D59DFDFA758DFCD9FE09" box="[626,777,459,485]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAF9FF44D59DFDFA758DFCD9FE09" box="[626,777,459,485]" italics="true" pageId="168" pageNumber="169">Protemnodon</emphasis>
|
||
</taxonomicName>
|
||
by its upper molars with a kinked postparacrista and from other Pliocene species by its large, robust i1 with a low, thick ventrolingual crest. Craniodental fossils from Stirton Quarry, Lake Kanunka, and Sand Scree Locality, Chinchilla that match this morphotype are also allocated to
|
||
<taxonomicName id="4C4C4D68FF44D59DFF2276CAFEEAFD49" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[170,314,651,677]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF44D59DFF2276CAFEEAFD49" box="[170,314,651,677]" italics="true" pageId="168" pageNumber="169">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF44D59DFECE76CDFE74FD49" bold="true" box="[326,420,651,677]" pageId="168" pageNumber="169">
|
||
<taxonomicNameLabel id="A20B5782FF44D59DFECE76CDFE74FD49" box="[326,420,651,677]" pageId="168" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
As the m4 from Site 4, Lake Kanunka, falls within the dental dimensions of and is most similar in morphology to
|
||
<taxonomicName id="4C4C4D68FF44D59DFE6E768AFDA2FD09" authority="Kerr & Camens & Van Zoelen & Worthy & Prideaux, 2024" authorityName="Kerr & Camens & Van Zoelen & Worthy & Prideaux" authorityYear="2024" box="[486,626,715,741]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="168" pageNumber="169" phylum="Chordata" rank="species" species="dawsonae" status="sp. nov.">
|
||
<emphasis id="B938EAF9FF44D59DFE6E768AFDA2FD09" box="[486,626,715,741]" italics="true" pageId="168" pageNumber="169">P. dawsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EAF9FF44D59DFDF1768DFD03FD09" bold="true" box="[633,723,715,741]" pageId="168" pageNumber="169">
|
||
<taxonomicNameLabel id="A20B5782FF44D59DFDF1768DFD03FD09" box="[633,723,715,741]" pageId="168" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and as it was found in association with the partial pelvis, we allocate both to that species, albeit with less confidence than the other referred specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |