plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/E5/87/03E587FAFFABFFAEFF2A5B052E7F7733.xml

554 lines
84 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="04594E23BA7935957186C7C6AFB9046C" ID-CLB-Dataset="8302" ID-DOI="10.11646/zootaxa.4926.2.1" ID-GBIF-Dataset="bc32a3b9-3917-47ef-8b02-b914c660daac" ID-ISSN="1175-5326" ID-Zenodo-Dep="4505942" ID-ZooBank="740D55C2-35EA-4D83-98D0-AC924C38CDD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612506571182" checkinUser="plazi" docAuthor="Fresneda, Javier, Rizzo, Valeria, Comas, Jordi &amp; Ribera, Ignacio" docDate="2021" docId="03E587FAFFABFFAEFF2A5B052E7F7733" docLanguage="en" docName="zootaxa.4926.2.1.pdf" docOrigin="Zootaxa 4926 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Troglocharinus Reitter 1908" docType="treatment" docVersion="9" lastPageNumber="164" masterDocId="FFDCFF82FFA7FFA0FFBD5F132D13762E" masterDocTitle="A redefinition of the genus Troglocharinus Reitter, 1908, with description of new species (Coleoptera, Leiodidae, Cholevinae, Leptodirini)" masterLastPageNumber="188" masterPageNumber="151" pageNumber="163" updateTime="1732703630073" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F3714FDA4813A1332D52956299F1F96F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AF6C764B37FA27DC9D4F53A1019F28D7">
<mods:title id="03B4F60DBFF1755C5960BE75EB760577">A redefinition of the genus Troglocharinus Reitter, 1908, with description of new species (Coleoptera, Leiodidae, Cholevinae, Leptodirini)</mods:title>
</mods:titleInfo>
<mods:name id="524D5289159A6D9E67FA52E7C80FE02D" type="personal">
<mods:role id="C1FF5726B3DDADB07E6C070B5BB999D7">
<mods:roleTerm id="7EE2BE89FF11C0649DA37F4410176AD0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9180BD437498C5DC609FC0DFE230DB14">Fresneda, Javier</mods:namePart>
<mods:affiliation id="64F84479E0801EA5EA56B51711D7AB25">Ca de Massa, 25526 Llesp-El Pont de Suert, Lleida, Spain. &amp; Museu de CieÌncies Naturals (Zoologia), Passeig Picasso s / n, 08003 Barcelona, Spain. comasj 2 @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="F0DE1840EA57642CC0AF4AF6B18A7B3E" type="personal">
<mods:role id="9FC7C4D1A85099B90ABFA9BD382B7384">
<mods:roleTerm id="77A714BECE540A3E9897B0BB40320D3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD8AE9D67CB4B894D324F4BEB2426230">Rizzo, Valeria</mods:namePart>
<mods:affiliation id="4A41415FA880A10C3311868AB33514BF">Myosotis M. M. Coop, Via Ulisse Aldrovandi, 18, 00197, Roma RM, Italy. valeria. rizzo 80 @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="0A0CBBCD95CD4C5D259F10FAD013F86B" type="personal">
<mods:role id="AD5F3731E25C5D5156D748BA0C871FB3">
<mods:roleTerm id="6544EAE52D77D3D630A33AC7AAB9D99D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1FDFA557FF5B933FEA1C211BDB57653">Comas, Jordi</mods:namePart>
<mods:affiliation id="2EECA3D135B4CA9A8EBD5001D9B35F8E">Museu de CieÌncies Naturals (Zoologia), Passeig Picasso s / n, 08003 Barcelona, Spain. comasj 2 @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="B2DA07D15003805356EE3EAE7FF22ED1" type="personal">
<mods:role id="90250418C6C1812F209D170E3602F0AF">
<mods:roleTerm id="5C7CC59CA4312B1EFA8F906A9290B187">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC19831E75CB5AE7BA6491DCEC37C583">Ribera, Ignacio</mods:namePart>
<mods:nameIdentifier id="B8B649DDE2800685A6F629ACC0D4FAC7" type="ORCID">0000-0003-0701-6016</mods:nameIdentifier>
<mods:affiliation id="80A2E4D505478E9CFD3441C75E5B0A10">Institut de Biologia Evolutiva (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37 - 49, 08003 Barcelona, Spain. Corresponding author. ffresned @ gmail. com; https: // orcid. org / 0000 - 0003 - 0701 - 6016</mods:affiliation>
<mods:nameIdentifier id="56B0AE5D80D6E4FBE146D3ADD039273A" type="email">ffresned@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7FF48B311FF3C73AF605818396BC8869">text</mods:typeOfResource>
<mods:relatedItem id="6875652B3BAE55CB452BC539380C17D1" type="host">
<mods:titleInfo id="2C513B410F7741235ABA59E94CA4494F">
<mods:title id="AF04437B98EA9FC776771ED843723A53">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3EB9AB0A9FAB41637E8576C6BBEDD7DE">
<mods:date id="CDE837AF07CDFC42EE7C1175FED6C373">2021</mods:date>
<mods:detail id="21A36D0B9F8003B7567F2096D07E1557" type="pubDate">
<mods:number id="EF4FDF9E2F17CF651E32347791AFE781">2021-02-05</mods:number>
</mods:detail>
<mods:detail id="C8B60E7BB0917C6B2794603666A7C9FA" type="volume">
<mods:number id="E1A774053A44DAD99E1CD046E5B5DEB1">4926</mods:number>
</mods:detail>
<mods:detail id="2E8E7D3756518B0BB6A4B188A1695FEF" type="issue">
<mods:number id="17C3BE3936F2A7BB64F32251BC75C1EE">2</mods:number>
</mods:detail>
<mods:extent id="01790A6A75E57AE913F8EA10979A754C" unit="page">
<mods:start id="347E0CFED367A4F13E14DF7D944C8D4B">151</mods:start>
<mods:end id="48449EE3B79AEE715AEC387BD8EF322B">188</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0AE45D9B0F3DB6C8F8B47A0FFDC1D4FF">journal article</mods:classification>
<mods:identifier id="0804F6227B09925CFEE762278BED8FD0" type="CLB-Dataset">8302</mods:identifier>
<mods:identifier id="3BE22E25AF3334F008D6F0064248F9F8" type="DOI">10.11646/zootaxa.4926.2.1</mods:identifier>
<mods:identifier id="33E6F5184ED135AE7E91A39325BCFDED" type="GBIF-Dataset">bc32a3b9-3917-47ef-8b02-b914c660daac</mods:identifier>
<mods:identifier id="DA76ADF2852393C1930426E97D65EB08" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7000719A1D05238A9652D8CDFCDD12ED" type="Zenodo-Dep">4505942</mods:identifier>
<mods:identifier id="BF63414EDD1435DE1BCF1A0D893ABF9C" type="ZooBank">740D55C2-35EA-4D83-98D0-AC924C38CDD5</mods:identifier>
</mods:mods>
<treatment id="03E587FAFFABFFAEFF2A5B052E7F7733" ID-DOI="http://doi.org/10.5281/zenodo.4534246" ID-GBIF-Taxon="177318788" ID-Zenodo-Dep="4534246" LSID="urn:lsid:plazi:treatment:03E587FAFFABFFAEFF2A5B052E7F7733" httpUri="http://treatment.plazi.org/id/03E587FAFFABFFAEFF2A5B052E7F7733" lastPageId="14" lastPageNumber="164" pageId="12" pageNumber="163" scope_family="Leiodidae" scope_order="Coleoptera" scope_subFamily="Cholevinae" scope_tribe="Leptodirini">
<subSubSection id="C3566567FFABFFACFF2A5B052CEA721E" box="[151,505,1046,1073]" pageId="12" pageNumber="163" type="nomenclature">
<paragraph id="8BF336ECFFABFFACFF2A5B052CEA721E" blockId="12.[151,505,1046,1073]" box="[151,505,1046,1073]" pageId="12" pageNumber="163">
<heading id="D0BB8180FFABFFACFF2A5B052CEA721E" bold="true" box="[151,505,1046,1073]" fontSize="11" level="1" pageId="12" pageNumber="163" reason="1">
<taxonomicName id="4C4C4D6FFFABFFACFF2A5B052CEA721E" ID-CoL="8X7XM" authority="Reitter, 1908" authorityName="Reitter" authorityYear="1908" box="[151,505,1046,1073]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFABFFACFF2A5B052CEA721E" bold="true" box="[151,505,1046,1073]" pageId="12" pageNumber="163">
<emphasis id="B938EAFEFFABFFACFF2A5B052C40721F" bold="true" box="[151,339,1046,1073]" italics="true" pageId="12" pageNumber="163">Troglocharinus</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFEE65B052CEA721E" author="Reitter, E." box="[347,505,1046,1072]" pageId="12" pageNumber="163" pagination="103 - 118" refId="ref18553" refString="Reitter, E. (1908) Dichotomische Ubersicht der Blinden Silphiden-Gattungen. Wiener entomologische Zeitung, 27, 103 - 118." type="journal article" year="1908">Reitter, 1908</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3566567FFABFFACFF2A5B4F2CE672C0" pageId="12" pageNumber="163" type="reference_group">
<paragraph id="8BF336ECFFABFFACFF2A5B4F2E60725D" blockId="12.[151,883,1115,1262]" box="[151,883,1115,1139]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB5C11C2FFABFFACFF2A5B4F2E60725D" box="[151,883,1115,1139]" pageId="12" pageNumber="163">
<treatmentCitation id="0AED10FDFFABFFACFF2A5B4F2C8E725D" author="Reitter, E." box="[151,413,1115,1139]" page="16" pageId="12" pageNumber="163" year="1885">
<taxonomicName id="4C4C4D6FFFABFFACFF2A5B4F2C8E725D" authority="Reitter, 1885: 16" authorityName="Reitter" authorityPageNumber="16" authorityYear="1885" box="[151,413,1115,1139]" class="Gastropoda" family="Chilodontidae" genus="Perrinia" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="163" phylum="Mollusca" rank="genus">
<emphasis id="B938EAFEFFABFFACFF2A5B4F2DFE725D" box="[151,237,1116,1139]" italics="true" pageId="12" pageNumber="163">Perrinia</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFF4E5B482C8E725D" author="Reitter, E." box="[243,413,1115,1139]" pageId="12" pageNumber="163" pagination="3 - 122" refId="ref18522" refString="Reitter, E. (1885) Bestimmungs-Tabellen der europaischen Coleopteren, 12 Necrophaga. Verhandlungen des naturforschenden Vereines in Brunn, 23, 3 - 122. [1884]" type="journal article" year="1885">Reitter, 1885: 16</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<taxonomicName id="4C4C4D6FFFABFFACFE115B4F2F83725D" authority="Adams, 1853" authorityName="Adams" authorityYear="1853" box="[428,656,1115,1139]" class="Gastropoda" family="Chilodontidae" genus="Perrinia" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="163" phylum="Mollusca" rank="genus">
<emphasis id="B938EAFEFFABFFACFE115B4F2F11725D" box="[428,514,1116,1139]" italics="true" pageId="12" pageNumber="163">Perrinia</emphasis>
Adams, 1853
</taxonomicName>
, Mollusca,
<taxonomicName id="4C4C4D6FFFABFFACFCBB5B482E7F725D" ID-CoL="1ef1ab08-077b-44dc-beee-be487cd82ff7" authorityName="Rafinesque" authorityYear="1815" box="[774,876,1115,1139]" class="Gastropoda" family="Trochidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="12" pageNumber="163" phylum="Mollusca" rank="family">
<emphasis id="B938EAFEFFABFFACFCBB5B482E7F725D" box="[774,876,1115,1139]" italics="true" pageId="12" pageNumber="163">Trochidae</emphasis>
</taxonomicName>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336ECFFABFFACFF2A5B692F9172BC" blockId="12.[151,883,1115,1262]" box="[151,642,1146,1170]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB5C11C2FFABFFACFF2A5B692F9172BC" box="[151,642,1146,1170]" pageId="12" pageNumber="163">
<treatmentCitation id="0AED10FDFFABFFACFF2A5B692F9172BC" author="Reitter, E." box="[151,642,1146,1170]" page="116" pageId="12" pageNumber="163" year="1908">
<taxonomicName id="4C4C4D6FFFABFFACFF2A5B692F9172BC" authority="Reitter, 1908: 116" authorityName="Reitter" authorityPageNumber="116" authorityYear="1908" box="[151,642,1146,1170]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="subGenus" subGenus="Troglocharinus">
<emphasis id="B938EAFEFFABFFACFF2A5B692C0072BC" box="[151,275,1146,1170]" italics="true" pageId="12" pageNumber="163">Troglophyes</emphasis>
(
<emphasis id="B938EAFEFFABFFACFE9F5B692CAE72BC" box="[290,445,1146,1170]" italics="true" pageId="12" pageNumber="163">Troglocharinus</emphasis>
)
<bibRefCitation id="EFDD4B1DFFABFFACFE715B692F9172BC" author="Reitter, E." box="[460,642,1146,1170]" pageId="12" pageNumber="163" pagination="103 - 118" refId="ref18553" refString="Reitter, E. (1908) Dichotomische Ubersicht der Blinden Silphiden-Gattungen. Wiener entomologische Zeitung, 27, 103 - 118." type="journal article" year="1908">Reitter, 1908: 116</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336ECFFABFFACFF2A5B8B2F2E729F" blockId="12.[151,883,1115,1262]" box="[151,573,1176,1201]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB5C11C2FFABFFACFF2A5B8B2F2E729F" box="[151,573,1176,1201]" pageId="12" pageNumber="163">
<taxonomicName id="4C4C4D6FFFABFFACFF2A5B8B2C91729E" ID-CoL="8X7XM" authority="Reitter" authorityName="Reitter" authorityYear="1908" box="[151,386,1176,1200]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFABFFACFF2A5B8B2C21729E" box="[151,306,1176,1200]" italics="true" pageId="12" pageNumber="163">Troglocharinus</emphasis>
Reitter
</taxonomicName>
:
<treatmentCitation id="0AED10FDFFABFFACFE365B8B2F2E729F" author="Jeannel, R." box="[395,573,1176,1201]" page="34" pageId="12" pageNumber="163" year="1910">
<bibRefCitation id="EFDD4B1DFFABFFACFE365B8B2F2E729F" author="Jeannel, R." box="[395,573,1176,1201]" pageId="12" pageNumber="163" pagination="149 - 163" refId="ref18237" refString="Jeannel, R. (1910) Nouveaux Silphides cavernicoles des Pyrenees catalanes. Archives de Zoologie experimentale et generale, 45, 149 - 163." type="journal article" year="1910">Jeannel, 1910: 34</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336ECFFABFFACFF2A5BA42CAE72E1" blockId="12.[151,883,1115,1262]" box="[151,445,1207,1231]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB5C11C2FFABFFACFF2A5BA42CAE72E1" box="[151,445,1207,1231]" pageId="12" pageNumber="163">
<treatmentCitation id="0AED10FDFFABFFACFF2A5BA42CAE72E1" author="Jeannel, R." box="[151,445,1207,1231]" page="93" pageId="12" pageNumber="163" year="1911">
<taxonomicName id="4C4C4D6FFFABFFACFF2A5BA42CAE72E1" ID-CoL="8XPY2" authority="Jeannel, 1911: 93" authorityName="Jeannel" authorityPageNumber="93" authorityYear="1911" box="[151,445,1207,1231]" class="Insecta" family="Leiodidae" genus="Speophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFABFFACFF2A5BA42C1572E1" box="[151,262,1207,1231]" italics="true" pageId="12" pageNumber="163">Speophilus</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFEB05BA42CAE72E1" author="Jeannel, R." box="[269,445,1207,1231]" pageId="12" pageNumber="163" pagination="1 - 641" refId="ref18264" refString="Jeannel, R. (1911) Revision des Bathysciinae (Coleopteres, Silphides). Morphologie, distribution geographique, Systematique. Archives de Zoologie experimentale et generale, 47, 1 - 641." type="journal article" year="1911">Jeannel, 1911: 93</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336ECFFABFFACFF2A5BC52CE672C0" blockId="12.[151,883,1115,1262]" box="[151,501,1238,1262]" pageId="12" pageNumber="163">
<treatmentCitationGroup id="AB5C11C2FFABFFACFF2A5BC52CE672C0" box="[151,501,1238,1262]" pageId="12" pageNumber="163">
<treatmentCitation id="0AED10FDFFABFFACFF2A5BC52CE672C0" author="Jeannel, R." box="[151,501,1238,1262]" page="283" pageId="12" pageNumber="163" year="1910">
<taxonomicName id="4C4C4D6FFFABFFACFF2A5BC52CE672C0" ID-CoL="8XJMM" authority="Jeannel, 1910: 283" authorityName="Jeannel" authorityPageNumber="283" authorityYear="1910" box="[151,501,1238,1262]" class="Insecta" family="Leiodidae" genus="Antrocharidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFABFFACFF2A5BC52C3C72C0" box="[151,303,1238,1262]" italics="true" pageId="12" pageNumber="163">Antrocharidius</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFE8B5BC52CE672C0" author="Jeannel, R." box="[310,501,1238,1262]" pageId="12" pageNumber="163" pagination="149 - 163" refId="ref18237" refString="Jeannel, R. (1910) Nouveaux Silphides cavernicoles des Pyrenees catalanes. Archives de Zoologie experimentale et generale, 45, 149 - 163." type="journal article" year="1910">Jeannel, 1910: 283</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3566567FFABFFAEFF2A5A072E7F7733" lastPageId="14" lastPageNumber="165" pageId="12" pageNumber="163" type="materials_examined">
<paragraph id="8BF336ECFFABFFACFF2A5A072E347305" blockId="12.[151,807,1299,1324]" box="[151,807,1299,1324]" pageId="12" pageNumber="163">
<typeStatus id="54F7884EFFABFFACFF2A5A072DDA7302" box="[151,201,1300,1324]" pageId="12" pageNumber="163">Type</typeStatus>
species:
<taxonomicName id="4C4C4D6FFFABFFACFE9B5A002E347305" authority="Reitter, 1908" authorityName="Reitter" authorityYear="1908" box="[294,807,1299,1323]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ferreri" subGenus="Troglocharinus">
<emphasis id="B938EAFEFFABFFACFE9B5A002CB17305" box="[294,418,1299,1323]" italics="true" pageId="12" pageNumber="163">Troglophyes</emphasis>
(
<emphasis id="B938EAFEFFABFFACFE0C5A002F5F7305" box="[433,588,1299,1323]" italics="true" pageId="12" pageNumber="163">Troglocharinus</emphasis>
)
<emphasis id="B938EAFEFFABFFACFDE15A002F8E7305" box="[604,669,1299,1323]" italics="true" pageId="12" pageNumber="163">ferreri</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFD1E5A002E347305" author="Reitter, E." box="[675,807,1299,1323]" pageId="12" pageNumber="163" pagination="103 - 118" refId="ref18553" refString="Reitter, E. (1908) Dichotomische Ubersicht der Blinden Silphiden-Gattungen. Wiener entomologische Zeitung, 27, 103 - 118." type="journal article" year="1908">Reitter, 1908</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BF336ECFFABFFACFF2A5A40287A73D3" blockId="12.[151,1437,1363,2037]" pageId="12" pageNumber="163">
Species of
<taxonomicName id="4C4C4D6FFFABFFACFEAE5A402CAC7343" authorityName="Reitter" authorityYear="1908" box="[275,447,1363,1389]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFABFFACFEAE5A402CAC7343" box="[275,447,1363,1389]" italics="true" pageId="12" pageNumber="163">Troglocharinus</emphasis>
</taxonomicName>
are hypogean, anophthalmous, apterous and depigmented, with a pholeuonoid (i.e. pronotum bell-shaped) or bathyscioid (i.e. pronotum trapezoidal) habitus (
<figureCitation id="13772A69FFABFFACFC755A64292373BF" box="[968,1072,1398,1425]" captionStart-0="FIGURES 3–4" captionStart-1="FIGURES 5–6" captionStartId-0="16.[151,264,1518,1542]" captionStartId-1="17.[151,264,1938,1962]" captionTargetBox-0="[151,1436,181,1495]" captionTargetBox-1="[157,1429,535,1903]" captionTargetId-0="figure-24@16.[151,1436,181,1495]" captionTargetId-1="figure-180@17.[151,1436,525,1915]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 3–4. Habitus of the holotypus of Troglocharinus sendrai sp. nov. (3) and T. fadriquei sp. nov. (4)." captionText-1="FIGURES 5–6. morphometric measurements. LA: Length of antenna; LE: Length of elytra; LP: Length of pronotum; LT: Total length; LL: Length of metatibiae; WE: Maximum width of elytra; WP: Maximum width of pronotum; WPA: Width of pronotal anteriad; WPB: Width of pronotal base. Troglocharinus ferreri (5) and Speonomites velox (6)." figureDoi-0="http://doi.org/10.5281/zenodo.4505948" figureDoi-1="http://doi.org/10.5281/zenodo.4505950" httpUri-0="https://zenodo.org/record/4505948/files/figure.png" httpUri-1="https://zenodo.org/record/4505950/files/figure.png" pageId="12" pageNumber="163">Figs 3–5</figureCitation>
). Size ranging between 2.5–4.5 mm, with a yellowish-reddish general colouration; pubescence yellowish, fine, dense and recumbent, although in some cases with longer erect setae, mostly in the apical area of the elytra (as e.g. in
<taxonomicName id="4C4C4D6FFFABFFACFBF85AAC288B73F7" authority="(Zariquiey, 1922))" authorityName="" baseAuthorityName="Zariquiey" baseAuthorityYear="1922" box="[1093,1432,1471,1497]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="patracoi">
<emphasis id="B938EAFEFFABFFACFBF85AAC29D373F6" box="[1093,1216,1471,1496]" italics="true" pageId="12" pageNumber="163">T. patracoi</emphasis>
(Zariquiey, 1922))
</taxonomicName>
; with irregular punctation on the pronotum and arranged transversally in the elytra, forming transverse striolas.
</paragraph>
<paragraph id="8BF336ECFFABFFACFF7A591429727068" blockId="12.[151,1437,1363,2037]" pageId="12" pageNumber="163">
Antennae longer than the body, particularly in the species with more developed pholeuonoid habitus. Antennomeres i–vi slender, vii with engrossed apex, viii slender and ix-xi enlarged (
<tableCitation id="C6CE0357FFABFFACFBBE593829477068" box="[1027,1108,1579,1606]" captionStart="TABLE 3" captionStartId="31.[151,244,1410,1436]" captionTargetPageId="31" captionText="TABLE 3. Length of the antennomeres. All values in mm." httpUri="http://table.plazi.org/id/DF336664FFB8FFBFFF2A5A912E3573B3" pageId="12" pageNumber="163" tableUuid="DF336664FFB8FFBFFF2A5A912E3573B3">Table3</tableCitation>
).
</paragraph>
<paragraph id="8BF336ECFFABFFACFF7A595C2CD57133" blockId="12.[151,1437,1363,2037]" pageId="12" pageNumber="163">
Pronotum narrowing posteriad in some species (
<taxonomicName id="4C4C4D6FFFABFFACFD54595C2E5E7047" authorityName="Reitter" authorityYear="1908" box="[745,845,1615,1641]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ferreri">
<emphasis id="B938EAFEFFABFFACFD54595C2E5E7047" box="[745,845,1615,1641]" italics="true" pageId="12" pageNumber="163">T. ferreri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFCE6595C29957047" authority="Jeannel, 1911" authorityName="Jeannel" authorityYear="1911" box="[859,1158,1615,1641]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="hustachei">
<emphasis id="B938EAFEFFABFFACFCE6595C2EF07047" box="[859,995,1615,1641]" italics="true" pageId="12" pageNumber="163">T. hustachei</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFC56595C29957047" author="Jeannel, R." box="[1003,1158,1615,1641]" pageId="12" pageNumber="163" pagination="1 - 641" refId="ref18264" refString="Jeannel, R. (1911) Revision des Bathysciinae (Coleopteres, Silphides). Morphologie, distribution geographique, Systematique. Archives de Zoologie experimentale et generale, 47, 1 - 641." type="journal article" year="1911">Jeannel, 1911</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFB2E595C2DDC70A3" authority="Espanol, 1955" authorityName="Espanol" authorityYear="1955" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="impellitierii">
<emphasis id="B938EAFEFFABFFACFB2E595C28207047" box="[1171,1331,1615,1641]" italics="true" pageId="12" pageNumber="163">T. impellitierii</emphasis>
Español, 1955
</taxonomicName>
and
<taxonomicName id="4C4C4D6FFFABFFACFEB959602F1A70A0" authority="Escola, 1967" authorityName="Escola" authorityYear="1967" box="[260,521,1651,1678]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="senenti">
<emphasis id="B938EAFEFFABFFACFEB959602C7D70A2" box="[260,366,1651,1676]" italics="true" pageId="12" pageNumber="163">T. senenti</emphasis>
Escolà, 1967
</taxonomicName>
), with sinusoid lateral margins (
<taxonomicName id="4C4C4D6FFFABFFACFCD1596029BD70A3" authority="Zariquiey, 1950" authorityName="Zariquiey" authorityYear="1950" box="[876,1198,1650,1677]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B938EAFEFFABFFACFCD159602EE770A3" box="[876,1012,1651,1677]" italics="true" pageId="12" pageNumber="163">T. elongatus</emphasis>
Zariquiey, 1950
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFB0559602DC8709F" authority="(Jeannel, 1930)" baseAuthorityName="Jeannel" baseAuthorityYear="1930" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="espanoli">
<emphasis id="B938EAFEFFABFFACFB055960282170A3" box="[1208,1330,1651,1677]" italics="true" pageId="12" pageNumber="163">T. espanoli</emphasis>
(Jeannel, 1930)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFF5A59842CFE709F" authority="(Jeannel, 1910)" baseAuthorityName="Jeannel" baseAuthorityYear="1910" box="[231,493,1687,1713]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fonti">
<emphasis id="B938EAFEFFABFFACFF5A59842C25709F" box="[231,310,1687,1713]" italics="true" pageId="12" pageNumber="163">T. fonti</emphasis>
(
<bibRefCitation id="EFDD4B1DFFABFFACFEF859842CF6709F" author="Jeannel, R." box="[325,485,1687,1713]" pageId="12" pageNumber="163" pagination="149 - 163" refId="ref18237" refString="Jeannel, R. (1910) Nouveaux Silphides cavernicoles des Pyrenees catalanes. Archives de Zoologie experimentale et generale, 45, 149 - 163." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFE4459842FE9709C" authority="(Lagar, 1966)" baseAuthorityName="Lagar" baseAuthorityYear="1966" box="[505,762,1687,1714]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="jacasi">
<emphasis id="B938EAFEFFABFFACFE4459842F4B709E" box="[505,600,1687,1712]" italics="true" pageId="12" pageNumber="163">T. jacasi</emphasis>
(Lagar, 1966)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFCBB59842948709C" authority="(Dieck, 1869)" baseAuthorityName="Dieck" baseAuthorityYear="1869" box="[774,1115,1687,1714]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="kiesenwetteri">
<emphasis id="B938EAFEFFABFFACFCBB59842EA5709F" box="[774,950,1687,1713]" italics="true" pageId="12" pageNumber="163">T. kiesenwetteri</emphasis>
(Dieck, 1869)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFBDA59842DC170FB" authority="Belles &amp; Deliot, 1983" authorityName="Belles &amp; Deliot" authorityYear="1983" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ludovici">
<emphasis id="B938EAFEFFABFFACFBDA598429CE709F" box="[1127,1245,1687,1713]" italics="true" pageId="12" pageNumber="163">T. ludovici</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFB5859842DC170FB" author="Belles, X. &amp; Deliot, Ph." pageId="12" pageNumber="163" pagination="237 - 243" refId="ref17124" refString="Belles, X. &amp; Deliot, Ph. (1983) Nouveaux Bathysciinae (Coleopteres Catopidae) des Pyrenees francaises et espagnoles. Memoires de Biospeologie, 10, 237 - 243." type="journal article" year="1983">Bellés &amp; Déliot, 1983</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFF6059A82CFE70FB" authority="Lagar, 1952" authorityName="Lagar" authorityYear="1952" box="[221,493,1722,1749]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="olerdolai">
<emphasis id="B938EAFEFFABFFACFF6059A82C4D70FB" box="[221,350,1723,1749]" italics="true" pageId="12" pageNumber="163">T. olerdolai</emphasis>
Lagar, 1952
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFE4559A82E0870FB" authority="(Jeannel, 1910)" baseAuthorityName="Jeannel" baseAuthorityYear="1910" box="[504,795,1723,1749]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B938EAFEFFABFFACFE4559A82F7570FA" box="[504,614,1723,1748]" italics="true" pageId="12" pageNumber="163">T. orcinus</emphasis>
(
<bibRefCitation id="EFDD4B1DFFABFFACFDC959A82E0070FB" author="Jeannel, R." box="[628,787,1723,1749]" pageId="12" pageNumber="163" pagination="149 - 163" refId="ref18237" refString="Jeannel, R. (1910) Nouveaux Silphides cavernicoles des Pyrenees catalanes. Archives de Zoologie experimentale et generale, 45, 149 - 163." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFC9B59A8298570FB" authority="Rizzo &amp; Comas, 2015" authorityName="Rizzo &amp; Comas" authorityYear="2015" box="[806,1174,1722,1749]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pallisei">
<emphasis id="B938EAFEFFABFFACFC9B59A82E8770FB" box="[806,916,1723,1749]" italics="true" pageId="12" pageNumber="163">T. pallisei</emphasis>
<bibRefCitation id="EFDD4B1DFFABFFACFC2659A8298570FB" author="Rizzo, V. &amp; Comas, J." box="[923,1174,1722,1749]" pageId="12" pageNumber="163" pagination="104 - 112" refId="ref18637" refString="Rizzo, V. &amp; Comas, J. (2015) A new species of Troglocharinus Reitter, 1908 (Coleoptera, Leiodidae, Cholevinae, Leptodirini) from southern Catalonia, with a molecular phylogeny of the related species group. Zootaxa, 3946, 104 - 112." type="journal article" year="2015">Rizzo &amp; Comas, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFB1D59A82DC870D7" authority="(Zariquiey, 1922)" authorityName="" baseAuthorityName="Zariquiey" baseAuthorityYear="1922" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="patracoi">
<emphasis id="B938EAFEFFABFFACFB1D59A8280970FA" box="[1184,1306,1723,1748]" italics="true" pageId="12" pageNumber="163">T. patracoi</emphasis>
(Zariquiey, 1922)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFF5B59CC2F2F70D7" authority="(Jeannel, 1911)" baseAuthorityName="Jeannel" baseAuthorityYear="1911" box="[230,572,1759,1785]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="quadricollis">
<emphasis id="B938EAFEFFABFFACFF5B59CC2C9B70D7" box="[230,392,1759,1785]" italics="true" pageId="12" pageNumber="163">T. quadricollis</emphasis>
(
<bibRefCitation id="EFDD4B1DFFABFFACFE2A59CC2F2770D7" author="Jeannel, R." box="[407,564,1759,1785]" pageId="12" pageNumber="163" pagination="1 - 641" refId="ref18264" refString="Jeannel, R. (1911) Revision des Bathysciinae (Coleopteres, Silphides). Morphologie, distribution geographique, Systematique. Archives de Zoologie experimentale et generale, 47, 1 - 641." type="journal article" year="1911">Jeannel, 1911</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFDFA59CC2E7570D7" authority="(Espanol, 1972)" baseAuthorityName="Espanol" baseAuthorityYear="1972" box="[583,870,1759,1785]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="schibii">
<emphasis id="B938EAFEFFABFFACFDFA59CC2FBE70D7" box="[583,685,1759,1785]" italics="true" pageId="12" pageNumber="163">T. schibii</emphasis>
(Español, 1972)
</taxonomicName>
and
<taxonomicName id="4C4C4D6FFFABFFACFC2659CC29D170D7" authority="(Espanol, 1965))" authorityName="" baseAuthorityName="Espanol" baseAuthorityYear="1965" box="[923,1218,1758,1785]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="subilsi">
<emphasis id="B938EAFEFFABFFACFC2659CC2EEC70D7" box="[923,1023,1759,1785]" italics="true" pageId="12" pageNumber="163">T. subilsi</emphasis>
(
<bibRefCitation id="EFDD4B1DFFABFFACFBB359CC29BC70D7" author="Espanol, F." box="[1038,1199,1758,1785]" pageId="12" pageNumber="163" pagination="1 - 8" refId="ref17634" refString="Espanol, F. (1965) Sobre Bathysciinae catalanes (Col. Catopidae). International Journal of Speleology, 2, 1 - 8." type="journal article" year="1965">Español, 1965</bibRefCitation>
))
</taxonomicName>
or regularly arched (
<taxonomicName id="4C4C4D6FFFABFFACFF1C58102CD17133" authority="(Escola, 1971))" authorityName="Escola" authorityYear="1971" baseAuthorityName="Escola" baseAuthorityYear="1971" box="[161,450,1795,1821]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFABFFACFF1C58102C187132" box="[161,267,1795,1820]" italics="true" pageId="12" pageNumber="163">T. vinyasi</emphasis>
(
<bibRefCitation id="EFDD4B1DFFABFFACFEA758102CBC7133" author="Escola, O." box="[282,431,1795,1821]" pageId="12" pageNumber="163" pagination="31 - 34" refId="ref17428" refString="Escola, O. (1971) Un nuevo Speonomus de la provincia de Lleida. Speleon, 18, 31 - 34." type="journal article" year="1971">Escolà, 1971</bibRefCitation>
))
</taxonomicName>
.
</paragraph>
<paragraph id="8BF336ECFFABFFACFF7A583429BB716F" blockId="12.[151,1437,1363,2037]" box="[199,1192,1831,1857]" pageId="12" pageNumber="163">Elytra long and narrow in pholeuonoid species, shorter and wider in bathyscioid species.</paragraph>
<paragraph id="8BF336ECFFABFFACFF7A585828427183" blockId="12.[151,1437,1363,2037]" pageId="12" pageNumber="163">
Mesoventral keel well developed (
<figureCitation id="13772A69FFABFFACFDF258582FB7714B" box="[591,676,1867,1893]" captionStart="FIGURES 12–14" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[262,1325,182,1883]" captionTargetId="figure-17@19.[262,1325,181,1922]" captionTargetPageId="19" captionText="FIGURES 12–14. Mesoventral keel of Troglocharinus fadriquei sp. nov. (12), T. hustachei (13)—Cova del Gel, Llimiana—and T. ferreri pallaresi (14)—Avenc de l’Esquerda, La Palma de Cervelló—." figureDoi="http://doi.org/10.5281/zenodo.4505954" httpUri="https://zenodo.org/record/4505954/files/figure.png" pageId="12" pageNumber="163">Fig. 12</figureCitation>
;
<taxonomicName id="4C4C4D6FFFABFFACFD0C58582E12714B" baseAuthorityName="Jeannel" baseAuthorityYear="1910" box="[689,769,1867,1893]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fonti">
<emphasis id="B938EAFEFFABFFACFD0C58582E12714B" box="[689,769,1867,1893]" italics="true" pageId="12" pageNumber="163">T. fonti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFCAC58582ED1714B" baseAuthorityName="Dieck" baseAuthorityYear="1869" box="[785,962,1867,1893]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="kiesenwetteri">
<emphasis id="B938EAFEFFABFFACFCAC58582ED1714B" box="[785,962,1867,1893]" italics="true" pageId="12" pageNumber="163">T. kiesenwetteri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFC6C5858295B714B" authorityName="Belles &amp; Deliot" authorityYear="1983" box="[977,1096,1867,1893]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ludovici">
<emphasis id="B938EAFEFFABFFACFC6C5858295B714B" box="[977,1096,1867,1893]" italics="true" pageId="12" pageNumber="163">T. ludovici</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFBE5585829EF714B" baseAuthorityName="Jeannel" baseAuthorityYear="1911" box="[1112,1276,1867,1893]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="quadricollis">
<emphasis id="B938EAFEFFABFFACFBE5585829EF714B" box="[1112,1276,1867,1893]" italics="true" pageId="12" pageNumber="163">T. quadricollis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFAB15858286B714A" authorityName="Escola" authorityYear="1967" box="[1292,1400,1867,1892]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="senenti">
<emphasis id="B938EAFEFFABFFACFAB15858286B714A" box="[1292,1400,1867,1892]" italics="true" pageId="12" pageNumber="163">T. senenti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFA3558582DF371A7" authorityName="" baseAuthorityName="Espanol" baseAuthorityYear="1965" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="subilsi">
<emphasis id="B938EAFEFFABFFACFA3558582DF371A7" italics="true" pageId="12" pageNumber="163">T. subilsi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D6FFFABFFACFEA5587C2C9771A6" authorityName="Escola" authorityYear="1971" baseAuthorityName="Escola" baseAuthorityYear="1971" box="[280,388,1903,1928]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="B938EAFEFFABFFACFEA5587C2C9771A6" box="[280,388,1903,1928]" italics="true" pageId="12" pageNumber="163">T. vinyasi</emphasis>
</taxonomicName>
) or reduced (
<figureCitation id="13772A69FFABFFACFDA0587C2FB371A7" box="[541,672,1903,1929]" captionStart="FIGURES 12–14" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[262,1325,182,1883]" captionTargetId="figure-17@19.[262,1325,181,1922]" captionTargetPageId="19" captionText="FIGURES 12–14. Mesoventral keel of Troglocharinus fadriquei sp. nov. (12), T. hustachei (13)—Cova del Gel, Llimiana—and T. ferreri pallaresi (14)—Avenc de l’Esquerda, La Palma de Cervelló—." figureDoi="http://doi.org/10.5281/zenodo.4505954" httpUri="https://zenodo.org/record/4505954/files/figure.png" pageId="12" pageNumber="163">Figs 13–14</figureCitation>
;
<taxonomicName id="4C4C4D6FFFABFFACFD11587C2E2571A7" authorityName="Zariquiey" authorityYear="1950" box="[684,822,1903,1929]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B938EAFEFFABFFACFD11587C2E2571A7" box="[684,822,1903,1929]" italics="true" pageId="12" pageNumber="163">T. elongatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFCF9587C2ED371A7" baseAuthorityName="Jeannel" baseAuthorityYear="1930" box="[836,960,1903,1929]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="espanoli">
<emphasis id="B938EAFEFFABFFACFCF9587C2ED371A7" box="[836,960,1903,1929]" italics="true" pageId="12" pageNumber="163">T. espanoli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFC73587C292071A7" authorityName="Reitter" authorityYear="1908" box="[974,1075,1903,1929]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ferreri">
<emphasis id="B938EAFEFFABFFACFC73587C292071A7" box="[974,1075,1903,1929]" italics="true" pageId="12" pageNumber="163">T. ferreri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFBFF587C29DA71A7" authorityName="Jeannel" authorityYear="1911" box="[1090,1225,1903,1929]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="hustachei">
<emphasis id="B938EAFEFFABFFACFBFF587C29DA71A7" box="[1090,1225,1903,1929]" italics="true" pageId="12" pageNumber="163">T. hustachei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFB65587C286A71A7" authorityName="Espanol" authorityYear="1955" box="[1240,1401,1903,1929]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="impellitierii">
<emphasis id="B938EAFEFFABFFACFB65587C286A71A7" box="[1240,1401,1903,1929]" italics="true" pageId="12" pageNumber="163">T. impellitierii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFA35587C2DC97182" baseAuthorityName="Lagar" baseAuthorityYear="1966" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="jacasi">
<emphasis id="B938EAFEFFABFFACFA35587C2DC97182" italics="true" pageId="12" pageNumber="163">T. jacasi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFF5558802F847180" authority="(Escola 1966)" baseAuthorityName="Escola" baseAuthorityYear="1966" box="[232,663,1939,1966]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="subSpecies" species="kiesenwetteri" subSpecies="andresi">
<emphasis id="B938EAFEFFABFFACFF5558802CE27183" box="[232,497,1939,1965]" italics="true" pageId="12" pageNumber="163">T. kiesenwetteri andresi</emphasis>
(Escolà 1966)
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFD1E58802E377183" authorityName="Lagar" authorityYear="1952" box="[675,804,1939,1965]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="olerdolai">
<emphasis id="B938EAFEFFABFFACFD1E58802E377183" box="[675,804,1939,1965]" italics="true" pageId="12" pageNumber="163">T. olerdolai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFC8F58802EB37182" baseAuthorityName="Jeannel" baseAuthorityYear="1910" box="[818,928,1939,1964]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B938EAFEFFABFFACFC8F58802EB37182" box="[818,928,1939,1964]" italics="true" pageId="12" pageNumber="163">T. orcinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFC135880290F7183" authorityName="Rizzo &amp; Comas" authorityYear="2015" box="[942,1052,1939,1965]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pallisei">
<emphasis id="B938EAFEFFABFFACFC135880290F7183" box="[942,1052,1939,1965]" italics="true" pageId="12" pageNumber="163">T. pallisei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D6FFFABFFACFB97588029B67182" authorityName="" baseAuthorityName="Zariquiey" baseAuthorityYear="1922" box="[1066,1189,1939,1964]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="patracoi">
<emphasis id="B938EAFEFFABFFACFB97588029B67182" box="[1066,1189,1939,1964]" italics="true" pageId="12" pageNumber="163">T. patracoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D6FFFABFFACFB66588028527183" baseAuthorityName="Espanol" baseAuthorityYear="1972" box="[1243,1345,1939,1965]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="schibii">
<emphasis id="B938EAFEFFABFFACFB66588028527183" box="[1243,1345,1939,1965]" italics="true" pageId="12" pageNumber="163">T. schibii</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF336ECFFABFFACFF7A58A42E5671DB" blockId="12.[151,1437,1363,2037]" pageId="12" pageNumber="163">Legs exceptionally long, in particular the metafemur of the more pholeuonoid species. Males with dilated first three protarsomeres; females with non dilated protarsomeres.</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5F842CCC76D4" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Female urite viii with short spiculum, with apex narrowly rounded (
<figureCitation id="13772A69FFAAFFADFC7D5F84297A769C" box="[960,1129,150,178]" captionStart="FIGURES 58–63" captionStartId="29.[151,264,1784,1808]" captionTargetBox="[151,1436,489,1757]" captionTargetId="figure-128@29.[151,1436,489,1760]" captionTargetPageId="29" captionText="FIGURES 58–63. viii female urite of Troglocharinus sendrai sp. nov. (58), T. fadriquei sp. nov. (59), T. mengeli (60)—Coves Encantades de Toloriu—, T. mercedesi (61)—Bòfia de Sant Jaume—, T. vinyasi (62)—Avenc de l’Home Mort—and Speonomites velox (63)—Querant del riu Merlé, Vilanova de Meià—. Spermathecal complex of T. mercedesi: CE, spermaduct; E, spermatheca; LA, apical lobe; LBS, basal lobe; RM middle region; VIII, eight urite." figureDoi="http://doi.org/10.5281/zenodo.4505970" httpUri="https://zenodo.org/record/4505970/files/figure.png" pageId="13" pageNumber="164">Figs 58, 61, 62</figureCitation>
) or, more frequently, bifurcated (
<figureCitation id="13772A69FFAAFFADFF5E5FA82C7B76FB" box="[227,360,186,213]" captionStart="FIGURES 58–63" captionStartId="29.[151,264,1784,1808]" captionTargetBox="[151,1436,489,1757]" captionTargetId="figure-128@29.[151,1436,489,1760]" captionTargetPageId="29" captionText="FIGURES 58–63. viii female urite of Troglocharinus sendrai sp. nov. (58), T. fadriquei sp. nov. (59), T. mengeli (60)—Coves Encantades de Toloriu—, T. mercedesi (61)—Bòfia de Sant Jaume—, T. vinyasi (62)—Avenc de l’Home Mort—and Speonomites velox (63)—Querant del riu Merlé, Vilanova de Meià—. Spermathecal complex of T. mercedesi: CE, spermaduct; E, spermatheca; LA, apical lobe; LBS, basal lobe; RM middle region; VIII, eight urite." figureDoi="http://doi.org/10.5281/zenodo.4505970" httpUri="https://zenodo.org/record/4505970/files/figure.png" pageId="13" pageNumber="164">Figs 59–60</figureCitation>
). The related genera from the Catalonian Pyrenees
<taxonomicName id="4C4C4D6FFFAAFFADFC0C5FA8295276FB" authorityName="Fresneda" authorityYear="1998" box="[945,1089,187,213]" class="Insecta" family="Leiodidae" genus="Pallaresiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFC0C5FA8295276FB" box="[945,1089,187,213]" italics="true" pageId="13" pageNumber="164">Pallaresiella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D6FFFAAFFADFBC65FA8281A76FB" box="[1147,1289,187,213]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFBC65FA8281A76FB" box="[1147,1289,187,213]" italics="true" pageId="13" pageNumber="164">Speonomites</emphasis>
</taxonomicName>
have similar bifurcated urite viii (
<figureCitation id="13772A69FFAAFFADFEC25FCC2CC176D4" box="[383,466,223,250]" captionStart="FIGURES 58–63" captionStartId="29.[151,264,1784,1808]" captionTargetBox="[151,1436,489,1757]" captionTargetId="figure-128@29.[151,1436,489,1760]" captionTargetPageId="29" captionText="FIGURES 58–63. viii female urite of Troglocharinus sendrai sp. nov. (58), T. fadriquei sp. nov. (59), T. mengeli (60)—Coves Encantades de Toloriu—, T. mercedesi (61)—Bòfia de Sant Jaume—, T. vinyasi (62)—Avenc de l’Home Mort—and Speonomites velox (63)—Querant del riu Merlé, Vilanova de Meià—. Spermathecal complex of T. mercedesi: CE, spermaduct; E, spermatheca; LA, apical lobe; LBS, basal lobe; RM middle region; VIII, eight urite." figureDoi="http://doi.org/10.5281/zenodo.4505970" httpUri="https://zenodo.org/record/4505970/files/figure.png" pageId="13" pageNumber="164">Fig. 63</figureCitation>
).
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5E10283D776F" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Metatergal apparatus very reduced (
<figureCitation id="13772A69FFAAFFADFDE65E102FCD7730" box="[603,734,259,286]" captionStart="FIGURES 64–69" captionStartId="30.[151,264,1151,1175]" captionTargetBox="[158,1429,186,1123]" captionTargetId="figure-24@30.[152,1436,181,1127]" captionTargetPageId="30" captionText="FIGURES 64–69. Metendosternite of Troglocharinus impellitierii (64)—Cova Palomera de Taús—, T. quadricollis (65)— Graller de Castellet de Tremp—, T. mengeli (66)—Coves Encantades de Toloriu—, and Speonomites velox (67)—Querant del riu Merlé, Vilanova de Meià—. Metatergal apparatus of Troglocharinus senenti (68)—Querant Gran de Paús, Vilanova de Meià—and T. mengeli (69)—Coves Encantades de Toloriu." figureDoi="http://doi.org/10.5281/zenodo.4505972" httpUri="https://zenodo.org/record/4505972/files/figure.png" pageId="13" pageNumber="164">Figs 68, 69</figureCitation>
), with narrow and straight metascutum (SC3) (sensu
<bibRefCitation id="EFDD4B1DFFAAFFADFA8B5E102C53776C" author="Friedrich, F. &amp; Beutel, R. G." pageId="13" pageNumber="164" pagination="290 - 315" refId="ref18081" refString="Friedrich, F. &amp; Beutel, R. G. (2006) The pterothoracic skeletomuscular system of Scirtoidea (Coleoptera: Polyphaga) and its implications for the high-level phylogeny of beetles. Journal of Zoological Systematics and Evolutionary Research, 44, 290 - 315." type="journal article" year="2006">Friedrich &amp; Beutel 2006</bibRefCitation>
), without sutures; alacrista (
<collectionCode id="ED5DAE29FFAAFFADFDC75E342FA6776F" box="[634,693,295,321]" pageId="13" pageNumber="164">ALC</collectionCode>
) without apical node and very short basal apophisis (A).
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5E582C9977FF" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Metendosternite with two long symmetric lateral arms, very narrow and diverging; stalk also narrow and as long as the arms (
<figureCitation id="13772A69FFAAFFADFEDF5E7C2CF577A4" box="[354,486,367,394]" captionStart="FIGURES 64–69" captionStartId="30.[151,264,1151,1175]" captionTargetBox="[158,1429,186,1123]" captionTargetId="figure-24@30.[152,1436,181,1127]" captionTargetPageId="30" captionText="FIGURES 64–69. Metendosternite of Troglocharinus impellitierii (64)—Cova Palomera de Taús—, T. quadricollis (65)— Graller de Castellet de Tremp—, T. mengeli (66)—Coves Encantades de Toloriu—, and Speonomites velox (67)—Querant del riu Merlé, Vilanova de Meià—. Metatergal apparatus of Troglocharinus senenti (68)—Querant Gran de Paús, Vilanova de Meià—and T. mengeli (69)—Coves Encantades de Toloriu." figureDoi="http://doi.org/10.5281/zenodo.4505972" httpUri="https://zenodo.org/record/4505972/files/figure.png" pageId="13" pageNumber="164">Figs 64–66</figureCitation>
). In
<taxonomicName id="4C4C4D6FFFAAFFADFDA75E7C2FBB77A7" box="[538,680,367,393]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFDA75E7C2FBB77A7" box="[538,680,367,393]" italics="true" pageId="13" pageNumber="164">Speonomites</emphasis>
</taxonomicName>
(
<figureCitation id="13772A69FFAAFFADFD055E7C2E1977A4" box="[696,778,367,394]" captionStart="FIGURES 64–69" captionStartId="30.[151,264,1151,1175]" captionTargetBox="[158,1429,186,1123]" captionTargetId="figure-24@30.[152,1436,181,1127]" captionTargetPageId="30" captionText="FIGURES 64–69. Metendosternite of Troglocharinus impellitierii (64)—Cova Palomera de Taús—, T. quadricollis (65)— Graller de Castellet de Tremp—, T. mengeli (66)—Coves Encantades de Toloriu—, and Speonomites velox (67)—Querant del riu Merlé, Vilanova de Meià—. Metatergal apparatus of Troglocharinus senenti (68)—Querant Gran de Paús, Vilanova de Meià—and T. mengeli (69)—Coves Encantades de Toloriu." figureDoi="http://doi.org/10.5281/zenodo.4505972" httpUri="https://zenodo.org/record/4505972/files/figure.png" pageId="13" pageNumber="164">Fig. 67</figureCitation>
) the stalk is ca. half the length of the arms. In the species transferred to
<taxonomicName id="4C4C4D6FFFAAFFADFE895E802CF37783" authorityName="Reitter" authorityYear="1908" box="[308,480,403,429]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFE895E802CF37783" box="[308,480,403,429]" italics="true" pageId="13" pageNumber="164">Troglocharinus</emphasis>
</taxonomicName>
and the newly described ones the stalk is longer than in
<taxonomicName id="4C4C4D6FFFAAFFADFBE45E8029F47783" box="[1113,1255,403,429]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFBE45E8029F47783" box="[1113,1255,403,429]" italics="true" pageId="13" pageNumber="164">Speonomites</emphasis>
</taxonomicName>
, but still clearly shorter than the arms.
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5EC82EF9744F" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Median lobe of the aedeagus in lateral view strongly flexed in an obtuse or right angle between the middle part and the apical third (
<figureCitation id="13772A69FFAAFFADFEC75EEC2CEF7437" box="[378,508,510,537]" captionStart="FIGURES 15–19" captionStartId="20.[151,264,1699,1723]" captionTargetBox="[151,1424,382,1676]" captionTargetId="figure-113@20.[151,1436,381,1676]" captionTargetPageId="20" captionText="FIGURES 15–19. Aedeagus in lateral view of Troglocharinus sendrai sp. nov. (15), T. fadriquei sp. nov. (16), T. mengeli (17)—Coves Encantades de Toloriu—, T. mercedesi (18)—Bofia de Sant Jaume—and T. vinyasi (19)—Avenc de la Roca del Corb." figureDoi="http://doi.org/10.5281/zenodo.4505956" httpUri="https://zenodo.org/record/4505956/files/figure.png" pageId="13" pageNumber="164">Figs 15–19</figureCitation>
); narrowing progressively from base to apex, which is pointed with perhaps a slight preapical dilatation; the dorsal part of the aedeagus may have a depression in the apical third with a very narrow extended apex; the sternal part of the apical third may be somewhat sinuate.
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5D782DC7758B" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Lateral stylus of the tegmen (i.e. parameres) shorter than the median lobe (
<figureCitation id="13772A69FFAAFFADFBBB5D78299A74AB" box="[1030,1161,618,645]" captionStart-0="FIGURES 15–19" captionStart-1="FIGURES 20–24" captionStartId-0="20.[151,264,1699,1723]" captionStartId-1="22.[151,264,1951,1975]" captionTargetBox-0="[151,1424,382,1676]" captionTargetBox-1="[252,1333,182,1914]" captionTargetId-0="figure-113@20.[151,1436,381,1676]" captionTargetId-1="figure-24@22.[251,1336,181,1927]" captionTargetPageId-0="20" captionTargetPageId-1="22" captionText-0="FIGURES 15–19. Aedeagus in lateral view of Troglocharinus sendrai sp. nov. (15), T. fadriquei sp. nov. (16), T. mengeli (17)—Coves Encantades de Toloriu—, T. mercedesi (18)—Bofia de Sant Jaume—and T. vinyasi (19)—Avenc de la Roca del Corb." captionText-1="FIGURES 20–24. Aedeagus in dorsal view of Troglocharinus sendrai sp. nov. (20), T. fadriquei sp. nov. (21), T. mengeli (22)—Cabana d’en Garraba—, T. mercedesi (23)—Bòfia de Sant Jaume—and T. vinyasi (24)—Avenc de la Roca del Corb." figureDoi-0="http://doi.org/10.5281/zenodo.4505956" figureDoi-1="http://doi.org/10.5281/zenodo.4505958" httpUri-0="https://zenodo.org/record/4505956/files/figure.png" httpUri-1="https://zenodo.org/record/4505958/files/figure.png" pageId="13" pageNumber="164">Figs 15–24</figureCitation>
); with two setae inserted apically and one pre-apically, the first two long, robust and either straight, arched, slightly sinuous or abruptly curved in different sections; the third finer and as long as the others or much shorter and slightly sinuous or almost straight. With a well developed membranous lamina (
<figureCitation id="13772A69FFAAFFADFD445DC42E6874DF" box="[761,891,726,753]" captionStart="FIGURES 35–47" captionStartId="25.[151,264,1494,1518]" captionTargetBox="[151,1430,193,1464]" captionTargetId="figure-17@25.[151,1436,181,1470]" captionTargetPageId="25" captionText="FIGURES 35–47. Apex of the lateral stylus of the aedeagus of Troglocharinus ferreri (35)—Avenc Gran del Sant Roc, Begues—, T. fonti (36)—Cova d’Ormini, Montanissell—, T. hustachei (37)—Cova del Gel, Llimiana—, T. impellitierii (38)—Cova Palomera, Taús—, T. senenti (39)—Querant Gran de Paús, Vilanova de Meià—, T. subilsi (40)—Grallera de Cambrils—, T. mengeli (41)—Coves Encantades de Toloriu—, T. sendrai sp. nov. (42), T. fadriquei sp. nov. (43), T. orcinus (44)—Cova Gran de la Febró, Montblanc—, Speonomites velox (45)—Querant del riu Merlé, Vilanova de Meià—, S. antemi (46)—Cova dels Porredons, Baén—and S. latrunculus (47)—Cova del Lladre, Santa Linya—. Membranous lamina in yellow." figureDoi="http://doi.org/10.5281/zenodo.4505964" httpUri="https://zenodo.org/record/4505964/files/figure.png" pageId="13" pageNumber="164">Figs 35–43</figureCitation>
) or short and trapezoidal (e.g. in
<taxonomicName id="4C4C4D6FFFAAFFADFB465DC4287874DE" baseAuthorityName="Jeannel" baseAuthorityYear="1910" box="[1275,1387,727,752]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B938EAFEFFAAFFADFB465DC4287874DE" box="[1275,1387,727,752]" italics="true" pageId="13" pageNumber="164">T. orcinus</emphasis>
</taxonomicName>
and subspecies and in
<taxonomicName id="4C4C4D6FFFAAFFADFEDB5DE82FC5753B" authority="Rizzo &amp; Comas 2015" authorityName="Rizzo &amp; Comas" authorityYear="2015" box="[358,726,762,789]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pallisei">
<emphasis id="B938EAFEFFAAFFADFEDB5DE82CC7753B" box="[358,468,763,789]" italics="true" pageId="13" pageNumber="164">T. pallisei</emphasis>
<bibRefCitation id="EFDD4B1DFFAAFFADFE605DE82FC5753B" author="Rizzo, V. &amp; Comas, J." box="[477,726,762,789]" pageId="13" pageNumber="164" pagination="104 - 112" refId="ref18637" refString="Rizzo, V. &amp; Comas, J. (2015) A new species of Troglocharinus Reitter, 1908 (Coleoptera, Leiodidae, Cholevinae, Leptodirini) from southern Catalonia, with a molecular phylogeny of the related species group. Zootaxa, 3946, 104 - 112." type="journal article" year="2015">Rizzo &amp; Comas 2015</bibRefCitation>
</taxonomicName>
;
<figureCitation id="13772A69FFAAFFADFD5E5DE82E25753B" box="[739,822,763,789]" captionStart="FIGURES 35–47" captionStartId="25.[151,264,1494,1518]" captionTargetBox="[151,1430,193,1464]" captionTargetId="figure-17@25.[151,1436,181,1470]" captionTargetPageId="25" captionText="FIGURES 35–47. Apex of the lateral stylus of the aedeagus of Troglocharinus ferreri (35)—Avenc Gran del Sant Roc, Begues—, T. fonti (36)—Cova d’Ormini, Montanissell—, T. hustachei (37)—Cova del Gel, Llimiana—, T. impellitierii (38)—Cova Palomera, Taús—, T. senenti (39)—Querant Gran de Paús, Vilanova de Meià—, T. subilsi (40)—Grallera de Cambrils—, T. mengeli (41)—Coves Encantades de Toloriu—, T. sendrai sp. nov. (42), T. fadriquei sp. nov. (43), T. orcinus (44)—Cova Gran de la Febró, Montblanc—, Speonomites velox (45)—Querant del riu Merlé, Vilanova de Meià—, S. antemi (46)—Cova dels Porredons, Baén—and S. latrunculus (47)—Cova del Lladre, Santa Linya—. Membranous lamina in yellow." figureDoi="http://doi.org/10.5281/zenodo.4505964" httpUri="https://zenodo.org/record/4505964/files/figure.png" pageId="13" pageNumber="164">Fig. 44</figureCitation>
), a derived state according to our phylogenetic results (
<figureCitation id="13772A69FFAAFFADFF1D5C0C2DF17517" box="[160,226,799,825]" captionStart="FIGURE 1" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[420,1160,182,1917]" captionTargetId="figure-17@11.[414,1174,181,1919]" captionTargetPageId="11" captionText="FIGURE 1. Majority rule consensus tree of the analysis with Bayesian probabilities in BEAST. Numbers in nodes, posterior probability (BEAST) (if&gt;0.5) / Maximum likelihood boostrap (RAxML) (if&gt;50). Scale in million years B.P. See Table 1 for details of the locality of the specimens." figureDoi="http://doi.org/10.5281/zenodo.4505944" httpUri="https://zenodo.org/record/4505944/files/figure.png" pageId="13" pageNumber="164">Fig. 1</figureCitation>
); with a tuf of long hairs (“penicillum”) in the surface of the lamina, especially along the margins, absent only in
<taxonomicName id="4C4C4D6FFFAAFFADFF565C502C4A7572" baseAuthorityName="Jeannel" baseAuthorityYear="1910" box="[235,345,835,860]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B938EAFEFFAAFFADFF565C502C4A7572" box="[235,345,835,860]" italics="true" pageId="13" pageNumber="164">T. orcinus</emphasis>
</taxonomicName>
and subspecies and in
<taxonomicName id="4C4C4D6FFFAAFFADFDEB5C502FD07573" authorityName="Rizzo &amp; Comas" authorityYear="2015" box="[598,707,835,861]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pallisei">
<emphasis id="B938EAFEFFAAFFADFDEB5C502FD07573" box="[598,707,835,861]" italics="true" pageId="13" pageNumber="164">T. pallisei</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDD4B1DFFAAFFADFD6C5C502ED67573" author="Rizzo, V. &amp; Comas, J." box="[721,965,834,861]" pageId="13" pageNumber="164" pagination="104 - 112" refId="ref18637" refString="Rizzo, V. &amp; Comas, J. (2015) A new species of Troglocharinus Reitter, 1908 (Coleoptera, Leiodidae, Cholevinae, Leptodirini) from southern Catalonia, with a molecular phylogeny of the related species group. Zootaxa, 3946, 104 - 112." type="journal article" year="2015">Rizzo &amp; Comas 2015</bibRefCitation>
;
<figureCitation id="13772A69FFAAFFADFC6D5C5029307573" box="[976,1059,835,861]" captionStart="FIGURES 35–47" captionStartId="25.[151,264,1494,1518]" captionTargetBox="[151,1430,193,1464]" captionTargetId="figure-17@25.[151,1436,181,1470]" captionTargetPageId="25" captionText="FIGURES 35–47. Apex of the lateral stylus of the aedeagus of Troglocharinus ferreri (35)—Avenc Gran del Sant Roc, Begues—, T. fonti (36)—Cova d’Ormini, Montanissell—, T. hustachei (37)—Cova del Gel, Llimiana—, T. impellitierii (38)—Cova Palomera, Taús—, T. senenti (39)—Querant Gran de Paús, Vilanova de Meià—, T. subilsi (40)—Grallera de Cambrils—, T. mengeli (41)—Coves Encantades de Toloriu—, T. sendrai sp. nov. (42), T. fadriquei sp. nov. (43), T. orcinus (44)—Cova Gran de la Febró, Montblanc—, Speonomites velox (45)—Querant del riu Merlé, Vilanova de Meià—, S. antemi (46)—Cova dels Porredons, Baén—and S. latrunculus (47)—Cova del Lladre, Santa Linya—. Membranous lamina in yellow." figureDoi="http://doi.org/10.5281/zenodo.4505964" httpUri="https://zenodo.org/record/4505964/files/figure.png" pageId="13" pageNumber="164">Fig. 44</figureCitation>
). The length of the lamina and the tuff of setae seem to be related, as both characters always occur together. With some sensilla placoidea close to the apex.
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5CBC2F7E737B" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Endophallus with three sections as defined by
<bibRefCitation id="EFDD4B1DFFAAFFADFD595CBC2E9F75E7" author="Jeannel, R." box="[740,908,943,969]" pageId="13" pageNumber="164" pagination="1 - 436" refId="ref18333" refString="Jeannel, R. (1924) Monographie des Bathysciinae (Col. Catopidae). Archives de Zoologie experimentale et generale, 63, 1 - 436." type="journal article" year="1924">Jeannel (1924)</bibRefCitation>
(
<figureCitation id="13772A69FFAAFFADFC235CBC293675E7" box="[926,1061,943,969]" captionStart="FIGURES 27–34" captionStartId="24.[151,264,1944,1968]" captionTargetBox="[180,1409,181,1908]" captionTargetId="figure-24@24.[170,1418,181,1920]" captionTargetPageId="24" captionText="FIGURES 27–34. Endophallus and schematic ampliation of the middle region of Troglocharinus sendrai sp. nov. (27–28), T. fadriquei sp. nov. (29–30), T. mengeli (31–32)—Cabana d’en Garraba—and T. mercedesi—Bòfia de Sant Jaume—(33–34)." figureDoi="http://doi.org/10.5281/zenodo.4505962" httpUri="https://zenodo.org/record/4505962/files/figure.png" pageId="13" pageNumber="164">Figs 27–34</figureCitation>
). Basal region (RB) with an “Ypiece” (PY) with divergent and sinuous arms, articulated with the apical membranous extension through a nodule; these structures form a funnel with the floating pieces and the appendices of the dorsal feather-like structures of the medial region (
<collectionCode id="ED5DAE29FFAAFFADFED35B082CBD721B" box="[366,430,1051,1077]" pageId="13" pageNumber="164">FDM</collectionCode>
) in the upper part. Medial region with two enlarged feather-like structures, bifurcated at the base; one of the bifurcations extented until the funnel-like structure of the basal region; with a floating piece in between the two arms of the bifurcated feather-like structures also in contact with the funnel-like structure. With two ventral feather like structures (
<collectionCode id="ED5DAE29FFAAFFADFD985B942F76728F" box="[549,613,1159,1185]" pageId="13" pageNumber="164">FVM</collectionCode>
) enlarged and divergent at their base, with a nodule of variable shape in the apex; with two upper dorsal plates, each of them with a long and thin external, and another more robust internal, feather-like structures, the later with the apex bifurcated and the enlarged area folded in a different orientation, very sclerotised. With two large bags at both sides of the axial piece, with very sclerotised internal nodules (
<collectionCode id="ED5DAE29FFAAFFADFA9B5BE028437323" box="[1318,1360,1267,1293]" pageId="13" pageNumber="164">NC</collectionCode>
) and a long appendix at the base. Apical region with reinforcement bands (
<collectionCode id="ED5DAE29FFAAFFADFC2D5A042EDE731F" box="[912,973,1303,1329]" country="Slovakia" lsid="urn:lsid:biocol.org:col:14946" name="Slovak National Museum" pageId="13" pageNumber="164" type="Herbarium">BRA</collectionCode>
) with a bag in between them, with longitudinal stripes and the margins engrossed.
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5A4C2E5973CB" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Female genitalia (
<figureCitation id="13772A69FFAAFFADFE305A4C2F1E7357" box="[397,525,1374,1401]" captionStart="FIGURES 52–57" captionStartId="28.[151,264,1907,1931]" captionTargetBox="[151,1436,527,1884]" captionTargetId="figure-144@28.[151,1436,525,1884]" captionTargetPageId="28" captionText="FIGURES 52–57. Spermatheca of Troglocharinus sendrai sp. nov. (52), T. fadriquei sp. nov. (53), T. mengeli (54)—Coves Encantades de Toloriu—, T. mercedesi (55)—Bòfia de Sant Jaume—, T. mercedesi (56)—Avenc del Xato—and T. vinyasi (57)—Avenc de l’Home Mort." figureDoi="http://doi.org/10.5281/zenodo.4505968" httpUri="https://zenodo.org/record/4505968/files/figure.png" pageId="13" pageNumber="164">Figs 52–57</figureCitation>
) with an elongated spermatheca, with two well sclerotised lobes and a hyaline medial region, narrower than the terminal lobes; apical lobe hemispherical. Spermaduct and spermatic gland inserted in the basal region; spermatic duct between 40–70 times the length of the spermatheca, with two sections of different diameter and a basal process in the bursa copulatrix (
<figureCitation id="13772A69FFAAFFADFD545AD82E2F73CB" box="[745,828,1482,1509]" captionStart="FIGURES 48–51" captionStartId="27.[151,264,1954,1978]" captionTargetBox="[183,1405,181,1927]" captionTargetId="figure-17@27.[183,1405,181,1930]" captionTargetPageId="27" captionText="FIGURES 48–51. Spermathecal complex of Troglocharinus fadriquei sp. nov. (48), T. mengeli (49)—Coves Encantades de Toloriu—, T. mercedesi (50)—Avenc del Xato—and T. ferreri (51)—Avenc del Sant Roc de Begues." figureDoi="http://doi.org/10.5281/zenodo.4505966" httpUri="https://zenodo.org/record/4505966/files/figure.png" pageId="13" pageNumber="164">Fig. 51</figureCitation>
).
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5AFC292C7107" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
The presence of a large membranous lamina (in comparison to other genera with a smaller lamina, as e.g.
<taxonomicName id="4C4C4D6FFFAAFFADFAD75AFC2DEF7002" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFAD75AFC2DEF7002" italics="true" pageId="13" pageNumber="164">Speonomites</emphasis>
</taxonomicName>
,
<figureCitation id="13772A69FFAAFFADFEB759002C947003" box="[266,391,1554,1581]" captionStart="FIGURES 35–47" captionStartId="25.[151,264,1494,1518]" captionTargetBox="[151,1430,193,1464]" captionTargetId="figure-17@25.[151,1436,181,1470]" captionTargetPageId="25" captionText="FIGURES 35–47. Apex of the lateral stylus of the aedeagus of Troglocharinus ferreri (35)—Avenc Gran del Sant Roc, Begues—, T. fonti (36)—Cova d’Ormini, Montanissell—, T. hustachei (37)—Cova del Gel, Llimiana—, T. impellitierii (38)—Cova Palomera, Taús—, T. senenti (39)—Querant Gran de Paús, Vilanova de Meià—, T. subilsi (40)—Grallera de Cambrils—, T. mengeli (41)—Coves Encantades de Toloriu—, T. sendrai sp. nov. (42), T. fadriquei sp. nov. (43), T. orcinus (44)—Cova Gran de la Febró, Montblanc—, Speonomites velox (45)—Querant del riu Merlé, Vilanova de Meià—, S. antemi (46)—Cova dels Porredons, Baén—and S. latrunculus (47)—Cova del Lladre, Santa Linya—. Membranous lamina in yellow." figureDoi="http://doi.org/10.5281/zenodo.4505964" httpUri="https://zenodo.org/record/4505964/files/figure.png" pageId="13" pageNumber="164">Figs 45–47</figureCitation>
) with a tuff of setae in the apex of the lateral stylus of the median lobe (the “penicillum”), and the very long spermatic duct with two sections with a different diameter and a basal process in the bursa copulatrix are two potential synapomorphies of the genus
<taxonomicName id="4C4C4D6FFFAAFFADFD1659482E44705B" authorityName="Reitter" authorityYear="1908" box="[683,855,1627,1653]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFD1659482E44705B" box="[683,855,1627,1653]" italics="true" pageId="13" pageNumber="164">Troglocharinus</emphasis>
</taxonomicName>
. The first, presence of a large lamina with “penicillum” in the apex of the parameres, is secondarily lost in the species
<taxonomicName id="4C4C4D6FFFAAFFADFC30596C2EEF70B6" baseAuthorityName="Jeannel" baseAuthorityYear="1910" box="[909,1020,1663,1688]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B938EAFEFFAAFFADFC30596C2EEF70B6" box="[909,1020,1663,1688]" italics="true" pageId="13" pageNumber="164">T. orcinus</emphasis>
</taxonomicName>
and its subspecies (
<figureCitation id="13772A69FFAAFFADFB67596C283570B4" box="[1242,1318,1663,1690]" captionStart="FIGURES 35–47" captionStartId="25.[151,264,1494,1518]" captionTargetBox="[151,1430,193,1464]" captionTargetId="figure-17@25.[151,1436,181,1470]" captionTargetPageId="25" captionText="FIGURES 35–47. Apex of the lateral stylus of the aedeagus of Troglocharinus ferreri (35)—Avenc Gran del Sant Roc, Begues—, T. fonti (36)—Cova d’Ormini, Montanissell—, T. hustachei (37)—Cova del Gel, Llimiana—, T. impellitierii (38)—Cova Palomera, Taús—, T. senenti (39)—Querant Gran de Paús, Vilanova de Meià—, T. subilsi (40)—Grallera de Cambrils—, T. mengeli (41)—Coves Encantades de Toloriu—, T. sendrai sp. nov. (42), T. fadriquei sp. nov. (43), T. orcinus (44)—Cova Gran de la Febró, Montblanc—, Speonomites velox (45)—Querant del riu Merlé, Vilanova de Meià—, S. antemi (46)—Cova dels Porredons, Baén—and S. latrunculus (47)—Cova del Lladre, Santa Linya—. Membranous lamina in yellow." figureDoi="http://doi.org/10.5281/zenodo.4505964" httpUri="https://zenodo.org/record/4505964/files/figure.png" pageId="13" pageNumber="164">Fig.44</figureCitation>
), although some populations may present intermediate degrees of development of this structure (unpublished observations). There are two further apomorphies of the genus shared by related genera: antennae longer than the body, present also in
<taxonomicName id="4C4C4D6FFFAAFFADFF5559F82F3B712B" authority="Jeannel, 1919" authorityName="Jeannel" authorityYear="1919" box="[232,552,1771,1797]" class="Insecta" family="Leiodidae" genus="Speocharidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFF5559F82C9B712B" box="[232,392,1771,1797]" italics="true" pageId="13" pageNumber="164">Speocharidius</emphasis>
Jeannel, 1919
</taxonomicName>
from the Vasc Country and other Leptodirini of different faunas, and very long legs, specially the metafemora, also present in
<taxonomicName id="4C4C4D6FFFAAFFADFD1F581C2E517107" authorityName="Jeannel" authorityYear="1919" box="[674,834,1807,1833]" class="Insecta" family="Leiodidae" genus="Speocharidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFD1F581C2E517107" box="[674,834,1807,1833]" italics="true" pageId="13" pageNumber="164">Speocharidius</emphasis>
</taxonomicName>
and other Leptodirini.
</paragraph>
<paragraph id="8BF336ECFFAAFFADFF7A5820295D7197" blockId="13.[151,1437,150,2014]" pageId="13" pageNumber="164">
Both synapomorphies are present in the species formerly in
<taxonomicName id="4C4C4D6FFFAAFFADFCD858202EE07163" box="[869,1011,1843,1869]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFCD858202EE07163" box="[869,1011,1843,1869]" italics="true" pageId="13" pageNumber="164">Speonomites</emphasis>
</taxonomicName>
transferred here to
<taxonomicName id="4C4C4D6FFFAAFFADFB715820286B7163" authorityName="Reitter" authorityYear="1908" box="[1228,1400,1843,1869]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFAAFFADFB715820286B7163" box="[1228,1400,1843,1869]" italics="true" pageId="13" pageNumber="164">Troglocharinus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C4C4D6FFFAAFFADFA3558202DE3715F" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" baseAuthorityName="Jeannel" baseAuthorityYear="1910" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="mengeli">
<emphasis id="B938EAFEFFAAFFADFA3558202DE3715F" italics="true" pageId="13" pageNumber="164">T. mengeli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D6FFFAAFFADFE9A58442CA1715F" authorityName="Fresneda &amp; Rizzo &amp; Comas &amp; Ribera" authorityYear="2021" baseAuthorityName="Zariquiey" baseAuthorityYear="1922" box="[295,434,1879,1905]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="mercedesi">
<emphasis id="B938EAFEFFAAFFADFE9A58442CA1715F" box="[295,434,1879,1905]" italics="true" pageId="13" pageNumber="164">T. mercedesi</emphasis>
</taxonomicName>
) and the two newly described species (see below). These four species have a bathyscioid habitus, characterised by a rather rounded body form and shorter appendages and antennae (
<figureCitation id="13772A69FFAAFFADFB0C5868280B71BB" box="[1201,1304,1915,1941]" captionStart="FIGURES 3–4" captionStartId="16.[151,264,1518,1542]" captionTargetBox="[151,1436,181,1495]" captionTargetId="figure-24@16.[151,1436,181,1495]" captionTargetPageId="16" captionText="FIGURES 3–4. Habitus of the holotypus of Troglocharinus sendrai sp. nov. (3) and T. fadriquei sp. nov. (4)." figureDoi="http://doi.org/10.5281/zenodo.4505948" httpUri="https://zenodo.org/record/4505948/files/figure.png" pageId="13" pageNumber="164">Figs 3–4</figureCitation>
), compared with the pholeuonid habitus of other species, and a well developed mesoventral keel.
</paragraph>
<paragraph id="8BF336ECFFAAFFAEFF2A58D02E7F7733" blockId="13.[151,1437,150,2014]" lastBlockId="14.[151,1436,150,285]" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164">
The phylogenetic reconstruction (
<figureCitation id="13772A69FFAAFFADFD9F58D02F7171F3" box="[546,610,1987,2013]" captionStart="FIGURE 1" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[420,1160,182,1917]" captionTargetId="figure-17@11.[414,1174,181,1919]" captionTargetPageId="11" captionText="FIGURE 1. Majority rule consensus tree of the analysis with Bayesian probabilities in BEAST. Numbers in nodes, posterior probability (BEAST) (if&gt;0.5) / Maximum likelihood boostrap (RAxML) (if&gt;50). Scale in million years B.P. See Table 1 for details of the locality of the specimens." figureDoi="http://doi.org/10.5281/zenodo.4505944" httpUri="https://zenodo.org/record/4505944/files/figure.png" pageId="13" pageNumber="164">Fig 1</figureCitation>
) shows that
<taxonomicName id="4C4C4D6FFFAAFFADFCB958D029F271F0" authority="(Jeannel, 1911)" baseAuthorityName="Jeannel" baseAuthorityYear="1911" box="[772,1249,1987,2014]" class="Insecta" family="Leiodidae" genus="Pallaresiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pallaresana">
<emphasis id="B938EAFEFFAAFFADFCB958D0293571F3" box="[772,1062,1987,2013]" italics="true" pageId="13" pageNumber="164">Pallaresiella pallaresana</emphasis>
(
<bibRefCitation id="EFDD4B1DFFAAFFADFB8758D029CA71F0" author="Jeannel, R." box="[1082,1241,1987,2014]" pageId="13" pageNumber="164" pagination="1 - 641" refId="ref18264" refString="Jeannel, R. (1911) Revision des Bathysciinae (Coleopteres, Silphides). Morphologie, distribution geographique, Systematique. Archives de Zoologie experimentale et generale, 47, 1 - 641." type="journal article" year="1911">Jeannel, 1911</bibRefCitation>
)
</taxonomicName>
—code voucher AF123—is the sister species of
<taxonomicName id="4C4C4D6FFFA9FFAEFE4B5F842999769F" authority="(Escola, Belles &amp; Comas, 1985)" baseAuthorityName="Escola, Belles &amp; Comas" baseAuthorityYear="1985" box="[502,1162,150,177]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="tincatincensis">
<emphasis id="B938EAFEFFA9FFAEFE4B5F842E30769E" box="[502,803,151,177]" italics="true" pageId="14" pageNumber="165">Speonomites tincatincensis</emphasis>
(Escolà, Bellés &amp; Comas, 1985)
</taxonomicName>
—code voucher IRC6— with a support value (
<collectionCode id="ED5DAE29FFA9FFAEFE3A5FA82CA776FB" box="[391,436,187,213]" country="France" name="Musee de Lectoure" pageId="14" pageNumber="165">ML</collectionCode>
) of 100, and both are sister of the genus
<taxonomicName id="4C4C4D6FFFA9FFAEFCC85FA829B676FB" authority="Jeannel, 1910" authorityName="Jeannel" authorityYear="1910" box="[885,1189,187,213]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFA9FFAEFCC85FA8291076FB" box="[885,1027,187,213]" italics="true" pageId="14" pageNumber="165">Speonomites</emphasis>
<bibRefCitation id="EFDD4B1DFFA9FFAEFBB45FA829B676FB" author="Jeannel, R." box="[1033,1189,187,213]" pageId="14" pageNumber="165" pagination="149 - 163" refId="ref18237" refString="Jeannel, R. (1910) Nouveaux Silphides cavernicoles des Pyrenees catalanes. Archives de Zoologie experimentale et generale, 45, 149 - 163." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
</taxonomicName>
. Since the structure of the endophallus is similar and the two genera present the same distribution area, we establish the synonymy between the genus
<taxonomicName id="4C4C4D6FFFA9FFAEFEB45E102C847733" authorityName="Jeannel" authorityYear="1910" box="[265,407,259,285]" class="Insecta" family="Leiodidae" genus="Speonomites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFA9FFAEFEB45E102C847733" box="[265,407,259,285]" italics="true" pageId="14" pageNumber="165">Speonomites</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D6FFFA9FFAEFE705E102E1D7733" authority="Fresneda, 1998" authorityName="Fresneda" authorityYear="1998" box="[461,782,259,285]" class="Insecta" family="Leiodidae" genus="Pallaresiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAFEFFA9FFAEFE705E102F4E7733" box="[461,605,259,285]" italics="true" pageId="14" pageNumber="165">Pallaresiella</emphasis>
<bibRefCitation id="EFDD4B1DFFA9FFAEFDDE5E102E1D7733" author="Fresneda, J." box="[611,782,259,285]" pageId="14" pageNumber="165" pagination="53 - 86" refId="ref17908" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del sur de Pirineos: generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille 1894. (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 86." type="journal article" year="1998">Fresneda, 1998</bibRefCitation>
</taxonomicName>
syn. nv.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink