treatments-xml/data/03/E5/87/03E587F0624EFFB3FF0A3484BCB82E1E.xml

<document id="09A7CA7DDD5A23A2E9F6B5153BD5D1FC" ID-CLB-Dataset="8216" ID-DOI="10.11646/zootaxa.4858.4.2" ID-GBIF-Dataset="2b7f92c6-3055-451d-bdd1-11fa5283d226" ID-ISSN="1175-5326" ID-Zenodo-Dep="4412668" ID-ZooBank="820D3461-81A6-4AED-B717-FDCBFEADE7DA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609691026015" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Anker, Arthur" docDate="2020" docId="03E587F0624EFFB3FF0A3484BCB82E1E" docLanguage="en" docName="zootaxa.4858.4.2.pdf" docOrigin="Zootaxa 4858 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Manaxius paullus Komai &amp; Anker 2020, n. sp." docType="treatment" docUuid="D29CA70A-F389-4D75-B6F0-BB6EC629A0AF" docUuidSource="ZooBank" docVersion="10" lastPageNumber="519" masterDocId="FFDCFF88624FFFBFFF9D3653BD282B2F" masterDocTitle="A new species of the axiid shrimp genus Manaxius Kensley, 2003 (Decapoda Axiidea) from shallow coral reefs of the western Pacific" masterLastPageNumber="520" masterPageNumber="507" pageNumber="508" updateTime="1702002806933" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="07F7015BCD6D07C68BEF3AB10FEE232A">A new species of the axiid shrimp genus Manaxius Kensley, 2003 (Decapoda Axiidea) from shallow coral reefs of the western Pacific</mods:title>
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<mods:namePart id="400627FC33069A6960F5FE993DBF5CF9">Komai, Tomoyuki</mods:namePart>
<mods:affiliation id="9EE152BCC7597B23E081F7E8AA93324E">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba, 260 - 8682 Japan.</mods:affiliation>
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<mods:namePart id="E0C506BAA1E51AB9FC1C1066FAF99E54">Anker, Arthur</mods:namePart>
<mods:affiliation id="F1D387466AE51E352865663B40C4AFFC">Universidade Federal de Goiás (UFG), Campus Samambaia, Instituto de Ciências Biológicas, ICB- 5, Avenida Esperança s / n, Goiânia, GO, 74690 - 900, Brazil. arthuranker 7 @ gmail. com; http: // orcid. org / 0000 - 0002 - 5350 - 4267</mods:affiliation>
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<mods:title id="2F42D9E91AD4121C312147BA9BD218AC">Zootaxa</mods:title>
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<mods:date id="EC8359122E08E6FD3A76DFB43C7119A7">2020</mods:date>
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<treatment id="03E587F0624EFFB3FF0A3484BCB82E1E" ID-DOI="http://doi.org/10.5281/zenodo.4412654" ID-GBIF-Taxon="177326249" ID-Zenodo-Dep="4412654" LSID="urn:lsid:zoobank.org:act:D29CA70A-F389-4D75-B6F0-BB6EC629A0AF" httpUri="http://treatment.plazi.org/id/03E587F0624EFFB3FF0A3484BCB82E1E" lastPageId="12" lastPageNumber="519" pageId="1" pageNumber="508">
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<heading id="D0BB818A624EFFBEFF0A3484BC9429DE" bold="true" box="[151,444,726,753]" fontSize="11" level="1" pageId="1" pageNumber="508" reason="1">
<emphasis id="B938EAF4624EFFBEFF0A3484BC9429DE" bold="true" box="[151,444,726,753]" pageId="1" pageNumber="508">
<taxonomicName id="4C4C4D65624EFFBEFF0A3484BC5B29DE" ID-CoL="BPJDB" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[151,371,726,753]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF4624EFFBEFF0A3484BC5B29DE" bold="true" box="[151,371,726,753]" italics="true" pageId="1" pageNumber="508">Manaxius paullus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B578F624EFFBEFEE73484BC9429DE" box="[378,444,727,753]" pageId="1" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BF336E6624EFFBEFF0A34A8BF15283A" blockId="1.[151,573,726,825]" box="[151,573,762,789]" pageId="1" pageNumber="508">[New Japanese name: Sango-ana-ebi]</paragraph>
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<paragraph id="8BF336E6624EFFBEFF0A354CBC3A2816" blockId="1.[151,573,726,825]" box="[151,274,798,825]" pageId="1" pageNumber="508">
(
<figureCitation id="13772A63624EFFBEFF02354CBC212816" box="[159,265,798,825]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1764,1788]" captionStartId-1="3.[151,250,1831,1855]" captionStartId-2="6.[151,250,1915,1939]" captionStartId-3="7.[151,250,1828,1852]" captionStartId-4="8.[151,250,1812,1836]" captionTargetBox-0="[158,1427,571,1730]" captionTargetBox-1="[163,1424,539,1797]" captionTargetBox-2="[162,1426,199,1883]" captionTargetBox-3="[163,1424,190,1794]" captionTargetBox-4="[167,1401,216,1758]" captionTargetId-0="figure-179@2.[151,1436,561,1741]" captionTargetId-1="figure-172@3.[151,1436,525,1808]" captionTargetId-2="figure-19@6.[151,1436,181,1892]" captionTargetId-3="figure-17@7.[151,1436,181,1804]" captionTargetId-4="figure-19@8.[164,1423,181,1788]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344.A, cephalothorax and cephalic appendages, lateral view (antennal flagellum partially omitted); B, pleon, telson and left uropod, lateral view." captionText-1="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." captionText-2="FIGURE 3. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344, showing setation of chelae. A, right major chela, lateral view; B, same, mesial view; C, left minor chela, lateral view; D, same, mesial view." captionText-3="FIGURE 4. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344, chelipeds (setae omitted except for A, F). A, right (major) cheliped, ischium and merus, lateral view; B, same, carpus and chela, lateral view; C, same, ischium and merus, mesial view; D, same, carpus and chela, mesial view; E, left (minor) cheliped, ischium and merus, lateral view; F, same, carpus and chela, lateral view; G, same, ischium and merus, mesial view; H, same, carpus and chela, mesial view." captionText-4="FIGURE 5. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.4412672" figureDoi-1="http://doi.org/10.5281/zenodo.4412674" figureDoi-2="http://doi.org/10.5281/zenodo.4412676" figureDoi-3="http://doi.org/10.5281/zenodo.4412678" figureDoi-4="http://doi.org/10.5281/zenodo.4412680" httpUri-0="https://zenodo.org/record/4412672/files/figure.png" httpUri-1="https://zenodo.org/record/4412674/files/figure.png" httpUri-2="https://zenodo.org/record/4412676/files/figure.png" httpUri-3="https://zenodo.org/record/4412678/files/figure.png" httpUri-4="https://zenodo.org/record/4412680/files/figure.png" pageId="1" pageNumber="508">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C356656D624EFFBEFF0A3534B81B2F3E" pageId="1" pageNumber="508" type="materials_examined">
<paragraph id="8BF336E6624EFFBEFF0A3534B84228E5" blockId="1.[151,1437,870,2013]" pageId="1" pageNumber="508">
<emphasis id="B938EAF4624EFFBEFF0A3534BC1628AF" bold="true" box="[151,318,870,897]" pageId="1" pageNumber="508">Type material</emphasis>
. 
<materialsCitation id="3B243CBB624EFFBEFED03534B84228E5" ID-GBIF-Occurrence="3034657302" collectingDate="2008-11-10" collectionCode="FLMNH-UF" collectorName="S. McKeon &amp; J. Moore " country="French Polynesia" elevation="6" latitude="-17.4764" location="Society Islands" longLatPrecision="7" longitude="-149.8327" municipality="Moorea" pageId="1" pageNumber="508" specimenCode="FLMNH-UF 16344" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F78844624EFFBEFED03534BC9228AE" box="[333,442,871,897]" pageId="1" pageNumber="508" type="holotype">Holotype</typeStatus>
: 
<specimenCode id="DBEA9E9D624EFFBEFE583534BF8D28AE" box="[453,677,871,897]" collectionCode="FLMNH-UF" pageId="1" pageNumber="508">FLMNH-UF 16344</specimenCode>
, female (pocl 
<quantity id="4CB49B03624EFFBEFCD83535BE8828AE" box="[837,928,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="508" unit="mm" value="3.5">3.5 mm</quantity>
), 
<collectingCountry id="F35B7676624EFFBEFC293534B95128AE" box="[948,1145,871,897]" name="French Polynesia" pageId="1" pageNumber="508">French Polynesia</collectingCountry>
, 
<location id="8E93603D624EFFBEFB193534B81A28AE" LSID="urn:lsid:plazi:treatment:03E587F0624EFFB3FF0A3484BCB82E1E:8E93603D624EFFBEFB193534B81A28AE" box="[1156,1330,871,897]" country="French Polynesia" latitude="-17.4764" longLatPrecision="7" longitude="-149.8327" municipality="Moorea" name="Society Islands" pageId="1" pageNumber="508">Society Islands</location>
, 
<collectingMunicipality id="6B97AC9C624EFFBEFAA33534B8B028AE" box="[1342,1432,871,897]" pageId="1" pageNumber="508">Moorea</collectingMunicipality>
, 
<locationDeviation id="9A412FC4624EFFBEFF0A35D8BF48288A" box="[151,608,907,933]" firstLocation="Cook's Bay" pageId="1" pageNumber="508" secondLocation="Sheraton Hotel">between Cook’s Bay and Sheraton Hotel</locationDeviation>
, 
<geoCoordinate id="EE785021624EFFBEFDF635D8BFCC288A" box="[619,740,907,933]" degrees="17.4764" direction="south" orientation="latitude" pageId="1" pageNumber="508" precision="5" value="-17.4764">17.4764°S</geoCoordinate>
, 
<geoCoordinate id="EE785021624EFFBEFD7235D8BE56288A" box="[751,894,907,933]" degrees="149.8327" direction="west" orientation="longitude" pageId="1" pageNumber="508" precision="5" value="-149.8327">149.8327°W</geoCoordinate>
, depth 
<quantity id="4CB49B03624EFFBEFC5335D8B93F288A" box="[974,1047,907,933]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="508" unit="m" value="5.5" valueMax="7.0" valueMin="4.0">
<elevation id="0061D1D5624EFFBEFC5335D8B93F288A" box="[974,1047,907,933]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="508" unit="m" value="5.5" valueMax="7.0" valueMin="4.0">4–7 m</elevation>
</quantity>
, outer reef slope, inside dead coral head (
<taxonomicName id="4C4C4D65624EFFBEFF4135FCBC6E28E6" box="[220,326,943,969]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="1" pageNumber="508" phylum="Cnidaria" rank="genus">
<emphasis id="B938EAF4624EFFBEFF4135FCBC6E28E6" box="[220,326,943,969]" italics="true" pageId="1" pageNumber="508">Acropora</emphasis>
</taxonomicName>
sp.), 
<date id="FFF21026624EFFBEFE1835FCBF4828E6" box="[389,608,943,969]" pageId="1" pageNumber="508" value="2008-11-10">
<collectingDate id="EFB6E9CE624EFFBEFE1835FCBF4828E6" box="[389,608,943,969]" pageId="1" pageNumber="508" value="2008-11-10">10 November 2008</collectingDate>
</date>
, coll. 
<collectorName id="26B95330624EFFBEFD3E35FCBE0B28E6" box="[675,803,943,969]" pageId="1" pageNumber="508">S. McKeon</collectorName>
&amp; 
<collectorName id="26B95330624EFFBEFCDB35FCBE8228E6" box="[838,938,943,969]" pageId="1" pageNumber="508">J. Moore</collectorName>
(extracted by 
<collectorName id="26B95330624EFFBEFBD435FCB99828E6" box="[1097,1200,943,969]" pageId="1" pageNumber="508">A. Anker</collectorName>
&amp; 
<collectorName id="26B95330624EFFBEFB4935FCB87628E5" box="[1236,1374,943,970]" pageId="1" pageNumber="508">S. Ivanenko</collectorName>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF336E6624EFFBEFF0A3580B81B2F3E" blockId="1.[151,1437,870,2013]" pageId="1" pageNumber="508">
<materialsCitation id="3B243CBB624EFFBEFF0A3580B81B2F3E" ID-GBIF-Occurrence="3034657301" collectingDate="1998-09-03" collectorName="K. Nomura." country="Japan" county="Fuki Beach" elevation="2" latitude="24.2542" location="Ryukyu Islands" longLatPrecision="7" longitude="123.9936" municipality="Kuro-shima Island" pageId="1" pageNumber="508" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54F78844624EFFBEFF0A3580BDD528C2" box="[151,253,979,1005]" pageId="1" pageNumber="508" type="paratype">Paratype</typeStatus>
: CBM-ZC 16165, 
<specimenCount id="9D4AFD6F624EFFBEFE4B3580BF0B28C2" box="[470,547,979,1005]" pageId="1" pageNumber="508" type="male">1 male</specimenCount>
(pocl 
<quantity id="4CB49B03624EFFBEFDED3580BFE428C2" box="[624,716,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="508" unit="mm" value="4.2">4.2 mm</quantity>
), 
<collectingCountry id="F35B7676624EFFBEFD7F3580BE0D28C2" box="[738,805,979,1005]" name="Japan" pageId="1" pageNumber="508">Japan</collectingCountry>
, 
<location id="8E93603D624EFFBEFCAF3580BECD28C2" LSID="urn:lsid:plazi:treatment:03E587F0624EFFB3FF0A3484BCB82E1E:8E93603D624EFFBEFCAF3580BECD28C2" box="[818,997,979,1005]" country="Japan" county="Fuki Beach" latitude="24.2542" longLatPrecision="7" longitude="123.9936" municipality="Kuro-shima Island" name="Ryukyu Islands" pageId="1" pageNumber="508">Ryukyu Islands</location>
, 
<location id="8E93603D624EFFBEFC6C3580B99C28C2" LSID="urn:lsid:plazi:treatment:03E587F0624EFFB3FF0A3484BCB82E1E:8E93603D624EFFBEFC6C3580B99C28C2" box="[1009,1204,979,1005]" country="Japan" county="Fuki Beach" latitude="24.2542" longLatPrecision="7" longitude="123.9936" municipality="Kuro-shima Island" name="Yaeyama Islands" pageId="1" pageNumber="508">Yaeyama Islands</location>
, 
<collectingMunicipality id="6B97AC9C624EFFBEFB5F3580B8B028C2" box="[1218,1432,979,1005]" pageId="1" pageNumber="508">Kuro-shima Island</collectingMunicipality>
, 
<collectingCounty id="62924E6A624EFFBEFF0A35A4BC332F3E" box="[151,283,1015,1041]" pageId="1" pageNumber="508">Fuki Beach</collectingCounty>
, 
<geoCoordinate id="EE785021624EFFBEFEBB35A5BC8B2F3E" box="[294,419,1014,1041]" degrees="24.2542" direction="north" orientation="latitude" pageId="1" pageNumber="508" precision="5" value="24.2542">24.2542°N</geoCoordinate>
, 
<geoCoordinate id="EE785021624EFFBEFE3335A4BF1E2F3E" box="[430,566,1015,1041]" degrees="123.9936" direction="east" orientation="longitude" pageId="1" pageNumber="508" precision="5" value="123.9936">123.9936°E</geoCoordinate>
, lagoon on reef flat, depth 
<quantity id="4CB49B03624EFFBEFCFF35A4BEA72F3E" box="[866,911,1015,1041]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="508" unit="m" value="2.0">
<elevation id="0061D1D5624EFFBEFCFF35A4BEA72F3E" box="[866,911,1015,1041]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="508" unit="m" value="2.0">2 m</elevation>
</quantity>
, 
<date id="FFF21026624EFFBEFC0635A4B9422F3E" box="[923,1130,1015,1041]" pageId="1" pageNumber="508" value="1998-09-03">
<collectingDate id="EFB6E9CE624EFFBEFC0635A4B9422F3E" box="[923,1130,1015,1041]" pageId="1" pageNumber="508" value="1998-09-03">3 September 1998</collectingDate>
</date>
, coll. 
<collectorName id="26B95330624EFFBEFB3035A4B81B2F3E" box="[1197,1331,1015,1041]" pageId="1" pageNumber="508">K. Nomura.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C356656D624EFFBEFF5A3249B9792D5A" pageId="1" pageNumber="508" type="diagnosis">
<paragraph id="8BF336E6624EFFBEFF5A3249B9792D5A" blockId="1.[151,1437,870,2013]" pageId="1" pageNumber="508">
<emphasis id="B938EAF4624EFFBEFF5A3249BC122F1B" bold="true" box="[199,314,1050,1076]" pageId="1" pageNumber="508">Diagnosis</emphasis>
. Rostrum triangular in dorsal view, reaching distal margin of article 2 of antennular peduncle, each lateral margin bearing 1 small spine at its mid-length. Carapace surface glabrous on lateral sides; gastric region with median carina bearing 1 small anterior spine followed by 1 small denticle, 1 obsolete (or trace of) tubercle and 1 minute denticle; submedian carinae each consisting of 3 spinules; lateral carinae each divided in 2 parts, anterior part with 1 conspicuous spine (supraocular spine), posterior part ending in 1 small spine and sometimes followed by 1 minute denticle; 1 pair of additional small spines present between first spine of submedian carinae and median carina; no trace of postcervical median carina. Pleura 2–4 rounded, posteroventral margin of pleuron 5 slightly angular. Telson with 2 pairs of small spines on dorsal surface; lateral margins each with 1 conspicuous proximal lobe, latter with 1 minute spine, 3 or 4 pairs of small spines and 1 pair of strong spiniform setae; posterolateral angles each with 2 unequal submarginal spiniform setae. Eyestalks reaching mid-length of rostrum, cornea pigmented. Chelipeds subequal in length, slightly dissimilar in armature and stoutness; each ischium with 1 small subdistal spine on dorsal margin and 1 subdistal spine followed by some small tubercles on ventral margin; each merus with 2 small subdistal spines on dorsal margin, ventromesial margin armed with 1 strong distal spine followed by serration consisting of 5 or 6 spinules; each carpus with small distodorsal spine; dorsal and lateral surfaces of chelae densely covered with tufts of short and long setae; dorsal margin of each palm armed with 4 or 5 spines, each lateral surface with 3 or 4 small spines adjacent to dorsal margin, but without spine near base of fingers; dactylus slightly shorter than palm on major cheliped side, subequal in length to palm on minor cheliped side.
</paragraph>
</subSubSection>
<subSubSection id="C356656D624EFFBAFF5A302DBFE52D01" lastPageId="5" lastPageNumber="512" pageId="1" pageNumber="508" type="description">
<paragraph id="8BF336E6624EFFBDFF5A302DBF102A32" blockId="1.[151,1437,870,2013]" lastBlockId="2.[151,1437,151,537]" lastPageId="2" lastPageNumber="509" pageId="1" pageNumber="508">
<emphasis id="B938EAF4624EFFBEFF5A302DBC792DB7" bold="true" box="[199,337,1662,1688]" pageId="1" pageNumber="508">Description</emphasis>
. 
<emphasis id="B938EAF4624EFFBEFEFD302CBF082DB6" box="[352,544,1663,1689]" italics="true" pageId="1" pageNumber="508">
<typeStatus id="54F78844624EFFBEFEFD302CBCEF2DB6" box="[352,455,1663,1689]" pageId="1" pageNumber="508" type="holotype">Holotype</typeStatus>
female.
</emphasis>
Rostrum (
<figureCitation id="13772A63624EFFBEFD08302CBFD12DB6" box="[661,761,1663,1689]" captionStart="FIGURE 1" captionStartId="2.[151,250,1764,1788]" captionTargetBox="[158,1427,571,1730]" captionTargetId="figure-179@2.[151,1436,561,1741]" captionTargetPageId="2" captionText="FIGURE 1. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344.A, cephalothorax and cephalic appendages, lateral view (antennal flagellum partially omitted); B, pleon, telson and left uropod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4412672" httpUri="https://zenodo.org/record/4412672/files/figure.png" pageId="1" pageNumber="508">Figs. 1A</figureCitation>
, 
<figureCitation id="13772A63624EFFBEFC98302CBE602DB6" box="[773,840,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="1" pageNumber="508">2A, B</figureCitation>
) moderately narrow, triangular in dorsal view, reaching distal end of article 2 of antennular peduncle, 0.3 times as long as distance between rostral base and cervical groove or 0.17 times as long as carapace, tapering into acute tip; lateral margins elevated, each with 1 spine at midlength and supraocular spine, continuous with lateral gastric carina on carapace; dorsal surface shallowly concave. Carapace (
<figureCitation id="13772A63624EFFBEFE92315CBC452C06" box="[271,365,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1764,1788]" captionTargetBox="[158,1427,571,1730]" captionTargetId="figure-179@2.[151,1436,561,1741]" captionTargetPageId="2" captionText="FIGURE 1. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344.A, cephalothorax and cephalic appendages, lateral view (antennal flagellum partially omitted); B, pleon, telson and left uropod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4412672" httpUri="https://zenodo.org/record/4412672/files/figure.png" pageId="1" pageNumber="508">Figs 1A</figureCitation>
, 
<figureCitation id="13772A63624EFFBEFEE4315CBCB52C06" box="[377,413,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="1" pageNumber="508">2A</figureCitation>
) with smooth surface, with either tufts of or individual short to long setae arranged in longitudinal rows on gastric region and scattered short setae on postcervical dorsum; dorsal surface nearly straight in lateral view, gastric region slightly sloping down to rostrum; median gastric carina distinct, extending anteriorly to level of supraocular spines, not reaching cervical groove, with 1 small anterior spine followed by 1 small denticle, 1 trace of minute tubercle and denticle; submedian carinae consisting of 3 spinules; lateral gastric carinae falling far short of midway between rostral base and cervical groove, divided into 2 widely separated parts, anterior part without armature other than supraocular spine, posterior part represented by 1 small spine; area between median and submedian carinae at level of anterior end of submedian carina with 1 small spine; no trace of postcervical carina; cervical groove distinct, but restricted to dorsum; suborbital lobe obtuse; pterygostomial margin slightly produced, broadly rounded; cardiac notch deep.
</paragraph>
<paragraph id="8BF336E6624DFFBDFF5A3774BE5B2A4A" blockId="2.[151,1437,151,537]" pageId="2" pageNumber="509">
Thoracic sternite 7 with shield (
<figureCitation id="13772A63624DFFBDFDAB3774BFA62A6E" box="[566,654,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="2" pageNumber="509">Fig. 2C</figureCitation>
) deeply depressed medially, divided by deep median groove, lateral margins strongly elevated, subtriangular with sharply pointed tip.
</paragraph>
<paragraph id="8BF336E6624DFFBDFF5A373CB84B2936" blockId="2.[151,1437,151,537]" pageId="2" pageNumber="509">
Pleon (
<figureCitation id="13772A63624DFFBDFE8A373CBC452AA6" box="[279,365,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1764,1788]" captionTargetBox="[158,1427,571,1730]" captionTargetId="figure-179@2.[151,1436,561,1741]" captionTargetPageId="2" captionText="FIGURE 1. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344.A, cephalothorax and cephalic appendages, lateral view (antennal flagellum partially omitted); B, pleon, telson and left uropod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4412672" httpUri="https://zenodo.org/record/4412672/files/figure.png" pageId="2" pageNumber="509">Fig. 1B</figureCitation>
) with scattered tufts of short to long setae on each somite. Pleomere 1 widening posteriorly in dorsal view, anterodorsal margin excavate; pleuron triangular with subacute tip, produced slightly beyond ventral margin of pleuron 2. Pleomere 2 with pleuron slightly asymmetrical; ventral margin unarmed, rounded at either angle. Pleomeres 3–5 with all pleura unarmed; pleura 3 and 4 rounded; pleuron 5 slightly angular on posterolateral margin. Pleomere 6 with pleuron bluntly pointed ventrally; posterolateral projection triangular, with blunt tip.
</paragraph>
<caption id="DF33666E624DFFBDFF0A30B7B9892C0E" ID-DOI="http://doi.org/10.5281/zenodo.4412672" ID-Zenodo-Dep="4412672" httpUri="https://zenodo.org/record/4412672/files/figure.png" pageId="2" pageNumber="509" startId="2.[151,250,1764,1788]" targetBox="[158,1427,571,1730]" targetPageId="2">
<paragraph id="8BF336E6624DFFBDFF0A30B7B9892C0E" blockId="2.[151,1437,1764,1825]" pageId="2" pageNumber="509">
<emphasis id="B938EAF4624DFFBDFF0A30B7BC3A2DD3" bold="true" box="[151,274,1764,1788]" pageId="2" pageNumber="509">FIGURE 1.</emphasis>
<taxonomicName id="4C4C4D65624DFFBDFE8A30B6BCE42DD2" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[279,460,1765,1789]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="509" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF4624DFFBDFE8A30B6BCE42DD2" box="[279,460,1765,1789]" italics="true" pageId="2" pageNumber="509">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF4624DFFBDFE4C30B6BF212DD2" bold="true" box="[465,521,1765,1789]" pageId="2" pageNumber="509">
<taxonomicNameLabel id="A20B578F624DFFBDFE4C30B6BF212DD2" box="[465,521,1765,1789]" pageId="2" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, cephalothorax and cephalic appendages, lateral view (antennal flagellum partially omitted); B, pleon, telson and left uropod, lateral view.
</paragraph>
</caption>
<paragraph id="8BF336E6624DFFBCFF5A311CB81E2B9E" blockId="2.[151,1436,1871,2042]" lastBlockId="3.[151,1437,151,502]" lastPageId="3" lastPageNumber="510" pageId="2" pageNumber="509">
Telson (
<figureCitation id="13772A63624DFFBDFEBC311CBC512C46" box="[289,377,1871,1897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="2" pageNumber="509">Fig. 2D</figureCitation>
) subrectangular, 1.2 times as long as maximal width at anterolateral lobes; lateral margin with conspicuous blunt anterolateral lobe latter bearing 1 submarginal spinule, 4 (left) or 3 (right) spines in posterior 0.4 of telson length and 1 strong spiniform seta adjacent to first spine; posterior margin convex, without posteromedian spine, fringed with numerous long setae, each posterolateral area with 2 unequal spiniform setae (lateral seta much smaller than mesial); dorsal face shallowly sulcate medially, armed with 2 pairs of small spines, anterior spine lo- cated at mid-length and posterior spine at 0.6 of telson length; long setae arising at bases of dorsal spines.
</paragraph>
<paragraph id="8BF336E6624CFFBCFF5A36E8BC8F2BD6" blockId="3.[151,1437,151,502]" pageId="3" pageNumber="510">
Eyestalk (
<figureCitation id="13772A63624CFFBCFEAA36E8BCA52BFA" box="[311,397,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="3" pageNumber="510">Fig. 2B</figureCitation>
) reaching mid-length of rostrum; cornea hemispherical, subequal in length and width to eyestalk, darkly pigmented.
</paragraph>
<paragraph id="8BF336E6624CFFBCFF5A3750BFCC2AD9" blockId="3.[151,1437,151,502]" pageId="3" pageNumber="510">
Antennular peduncle (
<figureCitation id="13772A63624CFFBCFE223750BF3C2A32" box="[447,532,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="3" pageNumber="510">Fig. 2B</figureCitation>
) reaching distal margin of article 4 of antennal peduncle; statocyst lobe on article 1 unarmed; flagella subequal in length to carapace. Antennal peduncle (
<figureCitation id="13772A63624CFFBCFC273774B93F2A6E" box="[954,1047,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1764,1788]" captionTargetBox="[158,1427,571,1730]" captionTargetId="figure-179@2.[151,1436,561,1741]" captionTargetPageId="2" captionText="FIGURE 1. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344.A, cephalothorax and cephalic appendages, lateral view (antennal flagellum partially omitted); B, pleon, telson and left uropod, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4412672" httpUri="https://zenodo.org/record/4412672/files/figure.png" pageId="3" pageNumber="510">Figs 1A</figureCitation>
, 
<figureCitation id="13772A63624CFFBCFBBE3774B96D2A6E" box="[1059,1093,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="3" pageNumber="510">2B</figureCitation>
) with article 1 bearing 1 small spine at ventromesial distal angle; article 2 with straight dorsolateral distal spine reaching beyond mid-length of scaphocerite; scaphocerite simple, acuminate, reaching almost to mid-length of article 4; article 3 with 1 small spine at ventromesial distal angle; article 4 slightly longer than article 2 (except for dorsolateral distal spine); article 5 about 0.6 length of article 4; flagellum twice as long as carapace, each article distally with some setae of various length (longest setae about twice length of 1 article).
</paragraph>
<caption id="DF33666E624CFFBCFF0A3174B9442CA8" ID-DOI="http://doi.org/10.5281/zenodo.4412674" ID-Zenodo-Dep="4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="3" pageNumber="510" startId="3.[151,250,1831,1855]" targetBox="[163,1424,539,1797]" targetPageId="3">
<paragraph id="8BF336E6624CFFBCFF0A3174B9442CA8" blockId="3.[151,1436,1831,1927]" pageId="3" pageNumber="510">
<emphasis id="B938EAF4624CFFBCFF0A3174BC3B2C10" bold="true" box="[151,275,1831,1855]" pageId="3" pageNumber="510">FIGURE 2.</emphasis>
<taxonomicName id="4C4C4D65624CFFBCFE87317BBCE72C10" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[282,463,1831,1855]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="510" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF4624CFFBCFE87317BBCE72C10" box="[282,463,1831,1855]" italics="true" pageId="3" pageNumber="510">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF4624CFFBCFE483174BF272C10" bold="true" box="[469,527,1831,1855]" pageId="3" pageNumber="510">
<taxonomicNameLabel id="A20B578F624CFFBCFE483174BF272C10" box="[469,527,1831,1855]" pageId="3" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted).
</paragraph>
</caption>
<paragraph id="8BF336E6624BFFBBFF5A36C4BFD32A71" blockId="4.[151,1437,150,2011]" pageId="4" pageNumber="511">
Mouthparts not dissected. Maxilliped 3 (
<figureCitation id="13772A63624BFFBBFD0636C4BFDD2B9E" box="[667,757,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1812,1836]" captionTargetBox="[167,1401,216,1758]" captionTargetId="figure-19@8.[164,1423,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4412680" httpUri="https://zenodo.org/record/4412680/files/figure.png" pageId="4" pageNumber="511">Fig. 5A</figureCitation>
) moderately slender. Coxa with small spine at distoventral angle. Basis with 1 small distoventral spine. Ischium with distomesial angle produced into strong projection; crista dentata consisting of row of sharp teeth diminishing in size proximally (
<figureCitation id="13772A63624BFFBBFC29368FB9232BD8" box="[948,1035,220,247]" captionStart="FIGURE 5" captionStartId="8.[151,250,1812,1836]" captionTargetBox="[167,1401,216,1758]" captionTargetId="figure-19@8.[164,1423,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4412680" httpUri="https://zenodo.org/record/4412680/files/figure.png" pageId="4" pageNumber="511">Fig. 5B</figureCitation>
); ventral margin with 2 small spines in proximal half. Merus bearing 2 unequal prominent spines on ventral margin (distal spine much stronger). Carpus with 1 small spine at ventrolateral distal angle. Dactylus shorter than propodus. Exopod almost reaching distal margin of merus, its distal half with multiple subdivisions.
</paragraph>
<paragraph id="8BF336E6624BFFBBFF5A3734BC922AE9" blockId="4.[151,1437,150,2011]" pageId="4" pageNumber="511">
Chelipeds (
<figureCitation id="13772A63624BFFBBFEDB3734BCA52AAE" box="[326,397,359,385]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1939]" captionTargetBox="[162,1426,199,1883]" captionTargetId="figure-19@6.[151,1436,181,1892]" captionTargetPageId="6" captionText="FIGURE 3. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344, showing setation of chelae. A, right major chela, lateral view; B, same, mesial view; C, left minor chela, lateral view; D, same, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4412676" httpUri="https://zenodo.org/record/4412676/files/figure.png" pageId="4" pageNumber="511">Figs 3</figureCitation>
, 
<figureCitation id="13772A63624BFFBBFE043734BC802AAE" box="[409,424,359,385]" captionStart="FIGURE 4" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[163,1424,190,1794]" captionTargetId="figure-17@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 4. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344, chelipeds (setae omitted except for A, F). A, right (major) cheliped, ischium and merus, lateral view; B, same, carpus and chela, lateral view; C, same, ischium and merus, mesial view; D, same, carpus and chela, mesial view; E, left (minor) cheliped, ischium and merus, lateral view; F, same, carpus and chela, lateral view; G, same, ischium and merus, mesial view; H, same, carpus and chela, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4412678" httpUri="https://zenodo.org/record/4412678/files/figure.png" pageId="4" pageNumber="511">4</figureCitation>
) subequal in length, right slightly stouter than left; mero-carpal articulation with about 30° of torsion (left with clockwise torsion, right with counter-clockwise torsion); chelae with dense setation particularly on lateral surface (
<figureCitation id="13772A63624BFFBBFEFA37FFBC842AE9" box="[359,428,428,454]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1939]" captionTargetBox="[162,1426,199,1883]" captionTargetId="figure-19@6.[151,1436,181,1892]" captionTargetPageId="6" captionText="FIGURE 3. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344, showing setation of chelae. A, right major chela, lateral view; B, same, mesial view; C, left minor chela, lateral view; D, same, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4412676" httpUri="https://zenodo.org/record/4412676/files/figure.png" pageId="4" pageNumber="511">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BF336E6624BFFBBFF5A379CBF492FB1" blockId="4.[151,1437,150,2011]" pageId="4" pageNumber="511">
Right (major) cheliped (
<figureCitation id="13772A63624BFFBBFE4F379CBF3C2AC6" box="[466,532,463,489]" captionStart="FIGURE 4" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[163,1424,190,1794]" captionTargetId="figure-17@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 4. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344, chelipeds (setae omitted except for A, F). A, right (major) cheliped, ischium and merus, lateral view; B, same, carpus and chela, lateral view; C, same, ischium and merus, mesial view; D, same, carpus and chela, mesial view; E, left (minor) cheliped, ischium and merus, lateral view; F, same, carpus and chela, lateral view; G, same, ischium and merus, mesial view; H, same, carpus and chela, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4412678" httpUri="https://zenodo.org/record/4412678/files/figure.png" pageId="4" pageNumber="511">Fig. 4</figureCitation>
A–D) coxa bearing 1 spinule on distolateral margin. Basis unarmed. Ischium with 1 small subdistal spine on dorsal margin; ventral margin also with 1 subdistal spine followed by 2 small tubercles. Merus widening distally; dorsal margin strongly arcuate, sharply carinate, with 1 small subdistal spine followed by 1 somewhat distant small spine and row of setae in distal half; lateral face glabrous, generally convex, distoventrally with deep excavation delimited by distinct ridge and fitting flexor margin of carpus, when cheliped is flexed; distolateral angle with prominent articular condyle; mesial face smooth, somewhat flattened; ventral margin carinate, bearing 1 small but conspicuous spine subterminally, followed by 6 small spines decreasing in size proximally. Carpus short, much higher than long, dorsal margin sharply carinate, with 1 small subdistal spine and numerous tufts of long setae; lateral face with several tufts of short to long setae on dorsal half, almost glabrous on ventral half, with sharp keel along ventral margin; mesial and ventral surfaces glabrous; mesial surface deeply excavated proximally to accommodate distoventral margin of merus when cheliped is flexed. Palm 1.3 times as long as high; dorsal surface non-carinate, armed with 4 widely spaced, relatively small spines, obscured by numerous tufts of long plumose setae; lateral surface flattened, with 3 small spines adjacent to dorsal margin, obscured by numerous tufts of short to long plumose setae; finger base unarmed; ventrolateral margin sharply keeled, this keel not extending onto fixed finger; mesial face inflated, almost glabrous except for some tufts of short setae on distal part. Fingers crossing, each terminating in acute tip; leaving narrow hiatus proximally when closed. Fixed finger with slightly arcuate ventral margin; occlusal margin with 3 obtuse teeth; lateral surface partially obscured by numerous tufts of long plumose setae; mesial face with tufts of short setae adjacent to ventral margin. Dactylus 0.8 times as long as palm, gently curved, with numerous tufts of long plumose setae on non-carinate, spineless dorsal margin and lateral surface; both lateral and mesial surfaces slightly elevated in midline, appearing as blunt ridge in proximal half; occlusal margin without conspicuous teeth.
</paragraph>
<paragraph id="8BF336E6624BFFBBFF5A32FBB8062EB5" blockId="4.[151,1437,150,2011]" pageId="4" pageNumber="511">
Left (minor) cheliped (
<figureCitation id="13772A63624BFFBBFE5E32FBBF2C2FED" box="[451,516,1192,1218]" captionStart="FIGURE 4" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[163,1424,190,1794]" captionTargetId="figure-17@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 4. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344, chelipeds (setae omitted except for A, F). A, right (major) cheliped, ischium and merus, lateral view; B, same, carpus and chela, lateral view; C, same, ischium and merus, mesial view; D, same, carpus and chela, mesial view; E, left (minor) cheliped, ischium and merus, lateral view; F, same, carpus and chela, lateral view; G, same, ischium and merus, mesial view; H, same, carpus and chela, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4412678" httpUri="https://zenodo.org/record/4412678/files/figure.png" pageId="4" pageNumber="511">Fig. 4</figureCitation>
E–H) with coxa to carpus similar to those of major cheliped; ischium with 1 subdistal spine on dorsal margin, ventral margin with 1 subdistal spine followed by 2 small tubercles. Merus with 1 small subdistal spine and 1 additional spine on dorsal margin; ventral margin with 1 large distal spine followed by row of 5 small spines diminishing in size proximally. Carpus with 1 small subdistal spine on dorsal margin. Palm and fingers slightly narrower than those of major cheliped, otherwise generally similar; palm 1.3 times as long as wide, dorsal margin with 5 well-spaced spines; lateral face with 4 small spines adjacent to dorsal margin. Fixed finger with 2 widely spaced small subacute teeth. Dactylus 1.2 times as long as palm; occlusal margin without tooth.
</paragraph>
<paragraph id="8BF336E6624BFFBBFF5A33F7BFF12D29" blockId="4.[151,1437,150,2011]" pageId="4" pageNumber="511">
Pereopod 2 (
<figureCitation id="13772A63624BFFBBFECE33F7BC802E90" box="[339,424,1444,1471]" captionStart="FIGURE 5" captionStartId="8.[151,250,1812,1836]" captionTargetBox="[167,1401,216,1758]" captionTargetId="figure-19@8.[164,1423,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4412680" httpUri="https://zenodo.org/record/4412680/files/figure.png" pageId="4" pageNumber="511">Fig. 5C</figureCitation>
) moderately slender. Ischium unarmed. Merus with 1 small spine on ventral margin located distally to mid-length. Chela 1.1 times as long as carpus; fixed finger straight, occlusal margin with row of minute spiniform setae; dactylus 1.2 times as long as palm.
</paragraph>
<paragraph id="8BF336E6624BFFBBFF5A3043B9D22DF1" blockId="4.[151,1437,150,2011]" pageId="4" pageNumber="511">
Pereopod 3 (
<figureCitation id="13772A63624BFFBBFEC73043BC9B2D04" box="[346,435,1552,1579]" captionStart="FIGURE 5" captionStartId="8.[151,250,1812,1836]" captionTargetBox="[167,1401,216,1758]" captionTargetId="figure-19@8.[164,1423,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4412680" httpUri="https://zenodo.org/record/4412680/files/figure.png" pageId="4" pageNumber="511">Fig. 5D</figureCitation>
) moderately slender. Ischium unarmed. Merus with 2 minute spines on ventral margin. Propodus with 5 transverse rows of spiniform setae (each consisting of 2 or 3 unequal spiniform setae) on flexor lateral surface; distal margin with row of several spiniform setae, longest seta on flexor side reaching mid-length of dactylus; flexor margin with cluster of stiff grooming setae in its distal half. Dactylus 0.6 times as long as propodus, gently curving, terminating in minute, basally demarcated unguis; lateral surface with longitudinal row of small spiniform setae adjacent to extensor margin and row of minute spiniform setae on flexor margin; mesial face unarmed.
</paragraph>
<paragraph id="8BF336E6624BFFBBFF5A30BBBE2C2C99" blockId="4.[151,1437,150,2011]" pageId="4" pageNumber="511">
Pereopod 4 (
<figureCitation id="13772A63624BFFBBFEC830BBBC812C2C" box="[341,425,1768,1795]" captionStart="FIGURE 5" captionStartId="8.[151,250,1812,1836]" captionTargetBox="[167,1401,216,1758]" captionTargetId="figure-19@8.[164,1423,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4412680" httpUri="https://zenodo.org/record/4412680/files/figure.png" pageId="4" pageNumber="511">Fig. 5E</figureCitation>
) noticeably slenderer than pereopod 3. Ischium to carpus unarmed. Propodus with 4 transverse rows of spiniform setae on flexor lateral face (each consisting of 2 or 3 unequal spiniform setae) and grooming setae on distal half of flexor margin; distal margin with row of several spiniform setae, longest seta on flexor side reaching mid-length of dactylus. Dactylus 0.5 times as long as propodus, gently arcuate, terminating in basally demarcated unguis; lateral surface with row of 3 spiniform setae adjacent to extensor margin and row of minute spiniform setae on flexor margin; mesial face unarmed.
</paragraph>
<paragraph id="8BF336E6624BFFBAFF5A3193B9032A6E" blockId="4.[151,1437,150,2011]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="512" pageId="4" pageNumber="511">
Pereopod 5 (
<figureCitation id="13772A63624BFFBBFECB3193BCBF2CF4" box="[342,407,1984,2011]" captionStart="FIGURE 5" captionStartId="8.[151,250,1812,1836]" captionTargetBox="[167,1401,216,1758]" captionTargetId="figure-19@8.[164,1423,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4412680" httpUri="https://zenodo.org/record/4412680/files/figure.png" pageId="4" pageNumber="511">Fig. 5</figureCitation>
F–H) most slender and shortest among pereopods, neither chelate nor semichelate. Ischium to carpus unarmed. Propodus with grooming apparatus consisting of 3 subterminal spiniform setae and field of stiff setae extending to mid-length of propodus. Dactylus lanceolate, somewhat twisted, terminating in basally demarcate, acute unguis; extensor and outer surface with thin setae, but without spiniform setae; inner surface excavated proximally; flexor margin with 3 minute spiniform setae followed by some microscopic spiniform setae; mesial surface excavated proximally, with 3 spiniform setae adjacent to extensor margin.
</paragraph>
<paragraph id="8BF336E6624AFFBAFF5A3718BF642AA6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">Pleopod 1 uniramous, consisting of 2 articles; pleopods 2–5 narrow, each with well-developed, rod-like appendix interna, 0.3–0.4 length of endopod.</paragraph>
<paragraph id="8BF336E6624AFFBAFF5A37C0BCEF294E" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
Uropod (
<figureCitation id="13772A63624AFFBAFEB137C0BCA92A82" box="[300,385,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1831,1855]" captionTargetBox="[163,1424,539,1797]" captionTargetId="figure-172@3.[151,1436,525,1808]" captionTargetPageId="3" captionText="FIGURE 2. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, carapace, dorsal view (setae omitted); B, anterior part of carapace and cephalic appendages, dorsal view; C, coxae of pereopods 3 and 4 and thoracic sternite 7; D, telson, dorsal view (setae omitted); E, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.4412674" httpUri="https://zenodo.org/record/4412674/files/figure.png" pageId="5" pageNumber="512">Fig. 2E</figureCitation>
) with endopod about twice as long as wide, lateral margin with row of 6 small spines on posterior half of lateral margin (posterolateral spine largest); median ridge on dorsal surface with row of 4 spines, posterior marginal spine strongest. Exopod subequal in length to endopod; lateral margin slightly convex, with 5 small spines on posterior 0.6 of exopod length (including posterolateral spine) and with 1 strong spiniform seta mesial to posterolateral spine; mid-dorsal ridge unarmed, obsolete lateral ridge with 2 small spines; transverse suture serrate, with row of 6 small spines.
</paragraph>
<paragraph id="8BF336E6624AFFBAFF5A3438B9C12F1A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
<emphasis id="B938EAF4624AFFBAFF5A3438BC4329AA" box="[199,363,619,645]" italics="true" pageId="5" pageNumber="512">
<typeStatus id="54F78844624AFFBAFF5A3438BC0629AB" box="[199,302,619,644]" pageId="5" pageNumber="512" type="paratype">Paratype</typeStatus>
male
</emphasis>
. Generally similar to 
<typeStatus id="54F78844624AFFBAFDFD3438BFEE29AA" box="[608,710,619,645]" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
. Rostrum as in 
<typeStatus id="54F78844624AFFBAFCEE3438BEF129AA" box="[883,985,619,645]" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
. Carapace median gastric carina bearing 2 small spines, 1 obsolete tubercle and 1 minute tubercle; submedian carinae each with 1 small spine followed by 2 minute tubercles; posterior part of lateral gastric carinae each with 1 spinule and 1 minute tubercle. Telson almost as in 
<typeStatus id="54F78844624AFFBAFF493484BC1129DE" box="[212,313,727,753]" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
, with 2 pairs of posterolateral spiniform setae. Maxilliped 3 armature as in 
<typeStatus id="54F78844624AFFBAFB113484B9DA29DE" box="[1164,1266,727,753]" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
. Cheliped meri with ventral margins each armed with 1 strong subdistal spine followed by smaller 5 or 6 spines; occlusal margin of fixed finger with 2 rounded teeth, that of dactylus with 1 rounded tooth proximally; dorsal margins of palm each armed with 4 or 5 moderately large spines. Pereopodal armature almost as in 
<typeStatus id="54F78844624AFFBAFB903510B95B2872" box="[1037,1139,835,861]" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
; pereopod 2 merus with 2 small spines on ventral margin; pereopod 3 merus with 2 or 3 small spines on ventral margin; pereopod 5 coxae each with gonopore. Pleopod 1 absent; pleopod 2 devoid of appendix masculina, perhaps not yet differentiated. Uropodal endopod with row of 5 or 6 small spines on lateral margin; median ridge on dorsal surface with row of 4 spines, posterior marginal spine strongest. Exopod lateral margin with 5 small spines on posterior 0.6 of its length (including posterolateral spine) and with 1 strong spiniform seta mesial to posterolateral spine; mid-dorsal ridge unarmed; dorsolateral ridge obsolete, with 2 or 3 spinules; transverse suture serrate, with 6 or 7 small spines.
</paragraph>
<paragraph id="8BF336E6624AFFBAFF5A326CBE0F2E22" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
Gill formula examined on 
<typeStatus id="54F78844624AFFBAFE60326CBF492F76" box="[509,609,1087,1113]" pageId="5" pageNumber="512" type="paratype">paratype</typeStatus>
. Arthrobranch on maxilliped 2 rudimentary, simple; 2 arthrobranchs on maxilliped 3 to pereopod 4, all well lamellate. Pleurobranch above each base of pereopods 2–4 small, lamellate, partially hidden by arthrobranchs. Epipods present on maxilliped 1 to pereopod 4, those on maxilliped 3 to pereopod 3 slender, located inside gill chamber, each with simple or poorly lamellate (with few papillae) podobranch, those on maxillipeds 1 and 2 and pereopod 4 devoid of podobranch. Setobranchs on pereopods 1–4 well developed, each consisting of tuft of long setae located inside gill chamber.
</paragraph>
<paragraph id="8BF336E6624AFFBAFF5A3345BFE52D01" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
<emphasis id="B938EAF4624AFFBAFF5A3345BC422E1F" bold="true" box="[199,362,1302,1328]" pageId="5" pageNumber="512">Colour in life</emphasis>
. Anterior part of carapace, including rostrum, whitish semi-translucent; dorsal part brownish, becoming paler laterally, with scattered red-brown spots on cardiac and branchial regions (spots on cardiac area being larger and with more intense red-brown colour); pleon semi-translucent, with tinge of brown laterally, including pleura, pleuron 2 with 2 brown spots anteriorly and posteriorly, pleura 3–5 each with brown spot posteriorly; ocular peduncle whitish semi-translucent, cornea darky pigmented; articles 1 and 2 of antennular peduncle whitish, article 3 brown, flagella pale brown; articles 1–4 of antennal peduncle whitish, article 5 brown, flagellum pale brown; chelipeds generally whitish, with tinge of yellow-brown on basal half of fingers, ventrolateral keel on palm conspicuous yellow-brown; pereopods 2–5 pale brown (
<figureCitation id="13772A63624AFFBAFDE63040BF972D01" box="[635,703,1555,1582]" captionStart="FIGURE 6" captionStartId="9.[151,250,1856,1880]" captionTargetBox="[183,1405,181,1832]" captionTargetId="figure-17@9.[183,1405,181,1832]" captionTargetPageId="9" captionText="FIGURE 6. Manaxius paullus n. sp., holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, habitus, dorsolateral view; B, same, lateral view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412682" httpUri="https://zenodo.org/record/4412682/files/figure.png" pageId="5" pageNumber="512">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C356656D624AFFBAFF5A3064BC8B2D5A" pageId="5" pageNumber="512" type="etymology">
<paragraph id="8BF336E6624AFFBAFF5A3064BC8B2D5A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
<emphasis id="B938EAF4624AFFBAFF5A3064BC662D7E" bold="true" box="[199,334,1591,1617]" pageId="5" pageNumber="512">Etymology.</emphasis>
From the alternative form (
<taxonomicName id="4C4C4D65624AFFBAFD143064BFF32D7E" authorityName="Komai &amp; Anker" authorityYear="2020" box="[649,731,1591,1617]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="512" phylum="Arthropoda" rank="species" species="paullus">
<emphasis id="B938EAF4624AFFBAFD143064BFF32D7E" box="[649,731,1591,1617]" italics="true" pageId="5" pageNumber="512">paullus</emphasis>
</taxonomicName>
) of the Latin adjective 
<emphasis id="B938EAF4624AFFBAFC793064B9072D7E" box="[996,1071,1591,1617]" italics="true" pageId="5" pageNumber="512">paulus</emphasis>
, for small, little, referring to the new species’ small size.
</paragraph>
</subSubSection>
<subSubSection id="C356656D624AFFBAFF5A302CBE7D2DB6" box="[199,853,1662,1689]" pageId="5" pageNumber="512" type="materials_examined">
<paragraph id="8BF336E6624AFFBAFF5A302CBE7D2DB6" blockId="5.[151,1437,151,2013]" box="[199,853,1662,1689]" pageId="5" pageNumber="512">
<emphasis id="B938EAF4624AFFBAFF5A302CBC482DB7" bold="true" box="[199,352,1662,1689]" pageId="5" pageNumber="512">Type locality</emphasis>
. Moorea, Society Islands, 
<collectingCountry id="F35B7676624AFFBAFD16302CBE792DB6" box="[651,849,1663,1689]" name="French Polynesia" pageId="5" pageNumber="512">French Polynesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C356656D624AFFBAFF5A30F1B9AE2DCE" pageId="5" pageNumber="512" type="distribution">
<paragraph id="8BF336E6624AFFBAFF5A30F1B9AE2DCE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
<emphasis id="B938EAF4624AFFBAFF5A30F1BC712D93" bold="true" box="[199,345,1698,1724]" pageId="5" pageNumber="512">Distribution</emphasis>
. Presently known only from two distant localities, viz. Society Islands, 
<collectingCountry id="F35B7676624AFFBAFB0330F0B84E2D92" box="[1182,1382,1699,1725]" name="French Polynesia" pageId="5" pageNumber="512">French Polynesia</collectingCountry>
, and Ryukyu Islands, 
<collectingCountry id="F35B7676624AFFBAFECE3094BCBE2DCE" box="[339,406,1735,1761]" name="Japan" pageId="5" pageNumber="512">Japan</collectingCountry>
; presumably more widely distributed in the western Pacific Ocean.
</paragraph>
</subSubSection>
<subSubSection id="C356656D624AFFBAFF5A30B8BE092C06" pageId="5" pageNumber="512" type="biology_ecology">
<paragraph id="8BF336E6624AFFBAFF5A30B8BE092C06" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
<emphasis id="B938EAF4624AFFBAFF5A30B8BC0E2C2A" bold="true" box="[199,294,1771,1797]" pageId="5" pageNumber="512">Ecology</emphasis>
. Shallow coral reefs, both on outer slope and reef flats, presently known depth range: 
<quantity id="4CB49B03624AFFBAFB7930B8B8052C2A" box="[1252,1325,1771,1797]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="5" pageNumber="512" unit="m" value="4.5" valueMax="7.0" valueMin="2.0">2–7 m</quantity>
; the 
<typeStatus id="54F78844624AFFBAFAFC30B8BDEF2C06" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
was extracted from a dead coral head (
<taxonomicName id="4C4C4D65624AFFBAFDE2315CBFC12C06" box="[639,745,1807,1833]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="5" pageNumber="512" phylum="Cnidaria" rank="genus">
<emphasis id="B938EAF4624AFFBAFDE2315CBFC12C06" box="[639,745,1807,1833]" italics="true" pageId="5" pageNumber="512">Acropora</emphasis>
</taxonomicName>
sp.).
</paragraph>
</subSubSection>
<subSubSection id="C356656D624AFFB3FF5A3160BCB82E1E" lastPageId="12" lastPageNumber="519" pageId="5" pageNumber="512" type="discussion">
<paragraph id="8BF336E6624AFFBAFF5A3160BE2A2CBA" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="512">
<emphasis id="B938EAF4624AFFBAFF5A3160BC1C2C62" bold="true" box="[199,308,1843,1869]" pageId="5" pageNumber="512">Remarks</emphasis>
. The two presently available specimens of 
<taxonomicName id="4C4C4D65624AFFBAFCB13160BEDE2C62" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[812,1014,1843,1869]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="512" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF4624AFFBAFCB13160BEDE2C62" box="[812,1014,1843,1869]" italics="true" pageId="5" pageNumber="512">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF4624AFFBAFB9D3160B9692C62" bold="true" box="[1024,1089,1843,1869]" pageId="5" pageNumber="512">
<taxonomicNameLabel id="A20B578F624AFFBAFB9D3160B9692C62" box="[1024,1089,1843,1869]" pageId="5" pageNumber="512" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
agree well with each other in almost every diagnostic detail. The minor differences between the 
<typeStatus id="54F78844624AFFBAFC173104BEC42C5E" box="[906,1004,1879,1905]" pageId="5" pageNumber="512" type="holotype">holotype</typeStatus>
and the 
<typeStatus id="54F78844624AFFBAFBD23104B9872C5E" box="[1103,1199,1879,1905]" pageId="5" pageNumber="512" type="paratype">paratype</typeStatus>
mentioned above all appear to be within the new species’ range of variation.
</paragraph>
<paragraph id="8BF336E6624AFFB3FF5A31CCB9BC29A9" blockId="5.[151,1437,151,2013]" lastBlockId="12.[151,1437,150,1329]" lastPageId="12" lastPageNumber="519" pageId="5" pageNumber="512">
As discussed by 
<bibRefCitation id="EFDD4B17624AFFBAFE0931CCBFF02C96" author="Kensley, B. &amp; Hickman, C. P. Jr." box="[404,728,1951,1977]" pageId="5" pageNumber="512" pagination="484 - 488" refId="ref6797" refString="Kensley, B. &amp; Hickman, C. P. Jr. (2001) A new species of Calaxius Sakai &amp; de Saint Laurent, 1989, from the Galapagos Islands (Crustacea: Decapoda: Axiidae). Proceedings of the Biological Society of Washington, 114, 484 - 488." type="journal article" year="2001">Kensley &amp; Hickman (2001</bibRefCitation>
; under 
<taxonomicName id="4C4C4D65624AFFBAFCAB31CCBEB22C96" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[822,922,1951,1977]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAF4624AFFBAFCAB31CCBEB22C96" box="[822,922,1951,1977]" italics="true" pageId="5" pageNumber="512">Calaxius</emphasis>
</taxonomicName>
), species of 
<taxonomicName id="4C4C4D65624AFFBAFBA431CCB98F2C97" authorityName="Kensley" authorityYear="2003" box="[1081,1191,1951,1976]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAF4624AFFBAFBA431CCB98F2C97" box="[1081,1191,1951,1976]" italics="true" pageId="5" pageNumber="512">Manaxius</emphasis>
</taxonomicName>
can be divided into two groups, viz. those with ventrally or posteroventrally acute pleura and those with ventrally rounded pleura. However, in 
<taxonomicName id="4C4C4D656243FFB3FEB836C4BC8B2B9E" baseAuthorityName="Clark, Galil &amp; Poore" baseAuthorityYear="2007" box="[293,419,151,177]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="B938EAF46243FFB3FEB836C4BC8B2B9E" box="[293,419,151,177]" italics="true" pageId="12" pageNumber="519">M. kensleyi</emphasis>
</taxonomicName>
, only the pleuron 2 is posteroventrally acutely pointed, whereas the pleura 3–5 are rounded; thus, this species appears to be somewhat intermediate between the two groups. 
<taxonomicName id="4C4C4D656243FFB3FB8636E8B9CA2BFA" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[1051,1250,187,213]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46243FFB3FB8636E8B9CA2BFA" box="[1051,1250,187,213]" italics="true" pageId="12" pageNumber="519">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46243FFB3FB7736E8B8012BFA" bold="true" box="[1258,1321,187,213]" pageId="12" pageNumber="519">
<taxonomicNameLabel id="A20B578F6243FFB3FB7736E8B8012BFA" box="[1258,1321,187,213]" pageId="12" pageNumber="519" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is characterised by the rounded pleura 2–5 and in therefore can be grouped together with 
<taxonomicName id="4C4C4D656243FFB3FB99368CB9E02BD6" baseAuthorityName="Sakai" baseAuthorityYear="2015" box="[1028,1224,223,249]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="andamanensis">
<emphasis id="B938EAF46243FFB3FB99368CB9E02BD6" box="[1028,1224,223,249]" italics="true" pageId="12" pageNumber="519">M. andamanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FB49368CB8BD2BD6" baseAuthorityName="de Man" baseAuthorityYear="1905" box="[1236,1429,223,249]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="euophthalmus">
<emphasis id="B938EAF46243FFB3FB49368CB8BD2BD6" box="[1236,1429,223,249]" italics="true" pageId="12" pageNumber="519">M. euophthalmus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FF0A3750BC7A2A32" authorityName="Kensley &amp; Hickman" authorityYear="2001" box="[151,338,259,285]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="galapagensis">
<emphasis id="B938EAF46243FFB3FF0A3750BC7A2A32" box="[151,338,259,285]" italics="true" pageId="12" pageNumber="519">M. galapagensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FEFC3750BCD42A32" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[353,508,259,285]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B938EAF46243FFB3FEFC3750BCD42A32" box="[353,508,259,285]" italics="true" pageId="12" pageNumber="519">M. inaequalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FD963750BF7A2A33" authorityName="Poore" authorityYear="2020" box="[523,594,259,284]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="iro">
<emphasis id="B938EAF46243FFB3FD963750BF7A2A33" box="[523,594,259,284]" italics="true" pageId="12" pageNumber="519">M. iro</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FDFC3750BFC92A33" baseAuthorityName="Komai" baseAuthorityYear="2011" box="[609,737,259,284]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="B938EAF46243FFB3FDFC3750BFC92A33" box="[609,737,259,284]" italics="true" pageId="12" pageNumber="519">M. izuensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FD723750BE4D2A33" baseAuthorityName="Williams" baseAuthorityYear="1974" box="[751,869,259,284]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="jenneri">
<emphasis id="B938EAF46243FFB3FD723750BE4D2A33" box="[751,869,259,284]" italics="true" pageId="12" pageNumber="519">M. jenneri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FCE83750B9342A33" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[885,1052,259,284]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="mimasensis">
<emphasis id="B938EAF46243FFB3FCE83750B9342A33" box="[885,1052,259,284]" italics="true" pageId="12" pageNumber="519">M. mimasensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FBB13750B9F12A32" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[1068,1241,259,285]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="pailoloensis">
<emphasis id="B938EAF46243FFB3FBB13750B9F12A32" box="[1068,1241,259,285]" italics="true" pageId="12" pageNumber="519">M. pailoloensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FB743750B8BD2A33" baseAuthorityName="de Man" baseAuthorityYear="1925" box="[1257,1429,259,284]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="B938EAF46243FFB3FB743750B8BD2A33" box="[1257,1429,259,284]" italics="true" pageId="12" pageNumber="519">M. pitatucensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FF0A3774BC3B2A6F" baseAuthorityName="Komai" baseAuthorityYear="2016" box="[151,275,295,320]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="poupini">
<emphasis id="B938EAF46243FFB3FF0A3774BC3B2A6F" box="[151,275,295,320]" italics="true" pageId="12" pageNumber="519">M. poupini</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C4C4D656243FFB3FECC3774BC9B2A6F" authorityName="Poore" authorityYear="2020" box="[337,435,295,320]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="supia">
<emphasis id="B938EAF46243FFB3FECC3774BC9B2A6F" box="[337,435,295,320]" italics="true" pageId="12" pageNumber="519">M. supia</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFDD4B176243FFB3FE753774BE392A6E" author="Kensley, B. &amp; Hickman, C. P. Jr." box="[488,785,295,321]" pageId="12" pageNumber="519" pagination="484 - 488" refId="ref6797" refString="Kensley, B. &amp; Hickman, C. P. Jr. (2001) A new species of Calaxius Sakai &amp; de Saint Laurent, 1989, from the Galapagos Islands (Crustacea: Decapoda: Axiidae). Proceedings of the Biological Society of Washington, 114, 484 - 488." type="journal article" year="2001">Kensley &amp; Hickman 2001</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FC803774BEF52A6E" author="Clark, P. F. &amp; Galil, B. &amp; Poore, G. C. B." box="[797,989,295,321]" pageId="12" pageNumber="519" pagination="63 - 73" refId="ref6526" refString="Clark, P. F., Galil, B. &amp; Poore, G. C. B. (2007) A new species of Calaxius Sakai &amp; de Saint Laurent, 1989, from West Africa (Crustacea, Decapoda, Axiidae) and synonymy of Manaxius Kensley, 2003. Proceedings of the Biological Society of Washington, 120, 63 - 73. https: // doi. org / 10.2988 / 0006 - 324 X (2007) 120 [63: ANSOCS] 2.0. CO; 2" type="journal article" year="2007">
Clark 
<emphasis id="B938EAF46243FFB3FCFF3774BEBC2A6E" box="[866,916,295,321]" italics="true" pageId="12" pageNumber="519">et al</emphasis>
. 2007
</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FC743774B9902A6E" author="Komai, T. &amp; Ohtsuka, S. &amp; Nakaguchi, K. &amp; Go, A." box="[1001,1208,295,321]" pageId="12" pageNumber="519" pagination="19 - 73" refId="ref6964" refString="Komai, T., Ohtsuka, S., Nakaguchi, K. &amp; Go, A. (2002) Decapod crustaceans collected from the southern part of the Sea of Japan in 2000 - 2001 using TRV Toyoshio-maru. Natural History Research, 7, 19 - 73." type="journal article" year="2002">
Komai 
<emphasis id="B938EAF46243FFB3FBA03774B9462A6E" box="[1085,1134,295,321]" italics="true" pageId="12" pageNumber="519">et al</emphasis>
. 2002
</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FB593774B8792A6E" author="Komai, T." box="[1220,1361,295,321]" pageId="12" pageNumber="519" pagination="279 - 337" refId="ref6852" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011">Komai 2011</bibRefCitation>
, 
<bibRefCitation id="EFDD4B176243FFB3FAC13774B8B02A6E" author="Komai, T." box="[1372,1432,295,321]" pageId="12" pageNumber="519" pagination="498 - 510" refId="ref6901" refString="Komai, T. (2016) Reappraisal of the two axiid genera Manaxius Kensley, 2003 and Calaxidium Sakai, 2014, and description of a new species of Calaxius Sakai &amp; de Saint Laurent, 1989 from French Polynesia, Southwest Pacific (Crustacea: Decapoda: Axiidea). Zootaxa, 4098 (3), 498 - 510." type="journal article" year="2016">2016</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FF0A3718BC332A4A" author="Poore, G. C. B." box="[151,283,331,357]" pageId="12" pageNumber="519" pagination="261 - 365" refId="ref7181" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds.), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum National d'Histoire Naturelle, Paris, 213, pp. 261 - 365." type="journal article" year="2020">Poore 2020</bibRefCitation>
). 
<taxonomicName id="4C4C4D656243FFB3FEB13718BF3D2A4A" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[300,533,331,357]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B938EAF46243FFB3FEB13718BF3D2A4A" box="[300,533,331,357]" italics="true" pageId="12" pageNumber="519">Manaxius inaequalis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C4D656243FFB3FDD53718BF932A4B" baseAuthorityName="Williams" baseAuthorityYear="1974" box="[584,699,331,356]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="jenneri">
<emphasis id="B938EAF46243FFB3FDD53718BF932A4B" box="[584,699,331,356]" italics="true" pageId="12" pageNumber="519">M. jenneri</emphasis>
</taxonomicName>
are very different from the other species in this group by their long ocular peduncles reaching the rostral apex (cf. 
<bibRefCitation id="EFDD4B176243FFB3FD01373CBE142AA6" author="Rathbun, M. J." box="[668,828,367,393]" pageId="12" pageNumber="519" pagination="1 - 137" refId="ref7321" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Porto Rico. Bulletin of the U. S. Fish Commission, 20, 1 - 137." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FCDB373CBEC62AA6" author="Williams, A. B." box="[838,1006,367,393]" pageId="12" pageNumber="519" pagination="451 - 464" refId="ref7670" refString="Williams, A. B. (1974) Two new axiids (Crustacea: Decapoda: Thalassinidea: Calocaris) from North Carolina and the Straits of Florida. Proceedings of the Biological Society of Washington, 87, 451 - 464." type="journal article" year="1974">Williams 1974</bibRefCitation>
); in addition, 
<taxonomicName id="4C4C4D656243FFB3FB1B373CB9D12AA7" baseAuthorityName="Williams" baseAuthorityYear="1974" box="[1158,1273,367,392]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="jenneri">
<emphasis id="B938EAF46243FFB3FB1B373CB9D12AA7" box="[1158,1273,367,392]" italics="true" pageId="12" pageNumber="519">M. jenneri</emphasis>
</taxonomicName>
is distinguishable by its markedly dissimilar chelipeds (
<bibRefCitation id="EFDD4B176243FFB3FDFA37C0BE382A82" author="Williams, A. B." box="[615,784,403,429]" pageId="12" pageNumber="519" pagination="451 - 464" refId="ref7670" refString="Williams, A. B. (1974) Two new axiids (Crustacea: Decapoda: Thalassinidea: Calocaris) from North Carolina and the Straits of Florida. Proceedings of the Biological Society of Washington, 87, 451 - 464." type="journal article" year="1974">Williams 1974</bibRefCitation>
). 
<taxonomicName id="4C4C4D656243FFB3FCBF37C0B91E2A82" baseAuthorityName="Sakai" baseAuthorityYear="2015" box="[802,1078,403,429]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="andamanensis">
<emphasis id="B938EAF46243FFB3FCBF37C0B91E2A82" box="[802,1078,403,429]" italics="true" pageId="12" pageNumber="519">Manaxius andamanensis</emphasis>
</taxonomicName>
is diagnostic in having unarmed gastric carinae, of which the median one extends posteriorly to the cervical groove (
<bibRefCitation id="EFDD4B176243FFB3FBA937E4B99E2AFE" author="Sakai, K." box="[1076,1206,438,465]" pageId="12" pageNumber="519" pagination="867 - 880" refId="ref7540" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (Infraorder Axiidea de Saint Laurent, 1979). Crustaceana, 88 (7 / 8), 867 - 880. https: // doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai 2015</bibRefCitation>
). 
<taxonomicName id="4C4C4D656243FFB3FB5537E4B8742AFF" authorityName="Poore" authorityYear="2020" box="[1224,1372,439,464]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="iro">
<emphasis id="B938EAF46243FFB3FB5537E4B8742AFF" box="[1224,1372,439,464]" italics="true" pageId="12" pageNumber="519">Manaxius iro</emphasis>
</taxonomicName>
is the only species in the genus characterised by the absence of spines on the dorsal (upper) margin of the cheliped palms (
<bibRefCitation id="EFDD4B176243FFB3FF0237ACBC002936" author="Poore, G. C. B." box="[159,296,511,537]" pageId="12" pageNumber="519" pagination="261 - 365" refId="ref7181" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds.), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum National d'Histoire Naturelle, Paris, 213, pp. 261 - 365." type="journal article" year="2020">Poore 2020</bibRefCitation>
). 
<taxonomicName id="4C4C4D656243FFB3FEA137ACBF602936" authorityName="Kensley &amp; Hickman" authorityYear="2001" box="[316,584,511,537]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="galapagensis">
<emphasis id="B938EAF46243FFB3FEA137ACBF602936" box="[316,584,511,537]" italics="true" pageId="12" pageNumber="519">Manaxius galapagensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C4D656243FFB3FD1E37ACBE182937" baseAuthorityName="de Man" baseAuthorityYear="1925" box="[643,816,511,536]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="B938EAF46243FFB3FD1E37ACBE182937" box="[643,816,511,536]" italics="true" pageId="12" pageNumber="519">M. pitatucensis</emphasis>
</taxonomicName>
are characteristic in the rostrum presenting a strongly tridentate apex, as well as in the highly spinose gastric carinae on the carapace in the former (
<bibRefCitation id="EFDD4B176243FFB3FB293470BDFA294E" author="Kensley, B. &amp; Hickman, C. P. Jr." pageId="12" pageNumber="519" pagination="484 - 488" refId="ref6797" refString="Kensley, B. &amp; Hickman, C. P. Jr. (2001) A new species of Calaxius Sakai &amp; de Saint Laurent, 1989, from the Galapagos Islands (Crustacea: Decapoda: Axiidae). Proceedings of the Biological Society of Washington, 114, 484 - 488." type="journal article" year="2001">Kensley &amp; Hickman 2001</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FF403414BC76294E" author="Sakai, K." box="[221,350,583,609]" pageId="12" pageNumber="519" pagination="1 - 616" refId="ref7436" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011</bibRefCitation>
). 
<taxonomicName id="4C4C4D656243FFB3FEED3414BF44294E" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[368,620,583,609]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="pailoloensis">
<emphasis id="B938EAF46243FFB3FEED3414BF44294E" box="[368,620,583,609]" italics="true" pageId="12" pageNumber="519">Manaxius pailoloensis</emphasis>
</taxonomicName>
is also unique within the genus, for instance, in having a pubescent body, a granulate carapace and only two distodorsal spines on the cheliped palm (
<bibRefCitation id="EFDD4B176243FFB3FC7E3438B9AF29A9" author="Rathbun, M. J." box="[995,1159,619,646]" pageId="12" pageNumber="519" pagination="827 - 930" refId="ref7354" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fisheries Commission, 23, 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
).
</paragraph>
<caption id="DF33666E6249FFB9FF0A3128B99A2C97" ID-DOI="http://doi.org/10.5281/zenodo.4412676" ID-Zenodo-Dep="4412676" httpUri="https://zenodo.org/record/4412676/files/figure.png" pageId="6" pageNumber="513" startId="6.[151,250,1915,1939]" targetBox="[162,1426,199,1883]" targetPageId="6">
<paragraph id="8BF336E66249FFB9FF0A3128B99A2C97" blockId="6.[151,1436,1915,1976]" pageId="6" pageNumber="513">
<emphasis id="B938EAF46249FFB9FF0A3128BC3C2CBB" bold="true" box="[151,276,1915,1940]" pageId="6" pageNumber="513">FIGURE 3.</emphasis>
<taxonomicName id="4C4C4D656249FFB9FE86312FBCFA2CBB" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[283,466,1916,1940]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="513" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46249FFB9FE86312FBCFA2CBB" box="[283,466,1916,1940]" italics="true" pageId="6" pageNumber="513">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46249FFB9FE44312FBF3B2CBB" bold="true" box="[473,531,1916,1940]" pageId="6" pageNumber="513">
<taxonomicNameLabel id="A20B578F6249FFB9FE44312FBF3B2CBB" box="[473,531,1916,1940]" pageId="6" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 3.5 mm), FLMNH-UF 16344, showing setation of chelae. A, right major chela, lateral view; B, same, mesial view; C, left minor chela, lateral view; D, same, mesial view.
</paragraph>
</caption>
<caption id="DF33666E6248FFB8FF0A3177B8302C87" ID-DOI="http://doi.org/10.5281/zenodo.4412678" ID-Zenodo-Dep="4412678" httpUri="https://zenodo.org/record/4412678/files/figure.png" pageId="7" pageNumber="514" startId="7.[151,250,1828,1852]" targetBox="[163,1424,190,1794]" targetPageId="7">
<paragraph id="8BF336E66248FFB8FF0A3177B8302C87" blockId="7.[151,1437,1828,1961]" pageId="7" pageNumber="514">
<emphasis id="B938EAF46248FFB8FF0A3177BC3B2C13" bold="true" box="[151,275,1828,1852]" pageId="7" pageNumber="514">FIGURE 4.</emphasis>
<taxonomicName id="4C4C4D656248FFB8FE843176BCE62C13" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[281,462,1828,1852]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="514" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46248FFB8FE843176BCE62C13" box="[281,462,1828,1852]" italics="true" pageId="7" pageNumber="514">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46248FFB8FE493176BF242C12" bold="true" box="[468,524,1829,1853]" pageId="7" pageNumber="514">
<taxonomicNameLabel id="A20B578F6248FFB8FE493176BF242C12" box="[468,524,1829,1853]" pageId="7" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 3.5 mm), FLMNH-UF 16344, chelipeds (setae omitted except for A, F). A, right (major) cheliped, ischium and merus, lateral view; B, same, carpus and chela, lateral view; C, same, ischium and merus, mesial view; D, same, carpus and chela, mesial view; E, left (minor) cheliped, ischium and merus, lateral view; F, same, carpus and chela, lateral view; G, same, ischium and merus, mesial view; H, same, carpus and chela, mesial view.
</paragraph>
</caption>
<caption id="DF33666E6247FFB7FF0A3147B8A72C5B" ID-DOI="http://doi.org/10.5281/zenodo.4412680" ID-Zenodo-Dep="4412680" httpUri="https://zenodo.org/record/4412680/files/figure.png" pageId="8" pageNumber="515" startId="8.[151,250,1812,1836]" targetBox="[167,1401,216,1758]" targetPageId="8">
<paragraph id="8BF336E66247FFB7FF0A3147B8A72C5B" blockId="8.[151,1437,1812,1908]" pageId="8" pageNumber="515">
<emphasis id="B938EAF46247FFB7FF0A3147BC3C2C03" bold="true" box="[151,276,1812,1836]" pageId="8" pageNumber="515">FIGURE 5.</emphasis>
<taxonomicName id="4C4C4D656247FFB7FE863146BCF92C03" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[283,465,1812,1836]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="515" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46247FFB7FE863146BCF92C03" box="[283,465,1812,1836]" italics="true" pageId="8" pageNumber="515">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46247FFB7FE4A3147BF392C03" bold="true" box="[471,529,1812,1836]" pageId="8" pageNumber="515">
<taxonomicNameLabel id="A20B578F6247FFB7FE4A3147BF392C03" box="[471,529,1812,1836]" pageId="8" pageNumber="515" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, left maxilliped 3, lateral view; B, same, crista dentata on ischium, ventral view; C, left pereopod 2, lateral view; D, left pereopod 3, lateral view; E, left pereopod 4, lateral view; F, left pereopod 5, lateral view; G, same, propodus and dactylus, ventral view; H, same, dactylus, dorsal view.
</paragraph>
</caption>
<caption id="DF33666E6246FFB6FF0A3113BF372C53" ID-DOI="http://doi.org/10.5281/zenodo.4412682" ID-Zenodo-Dep="4412682" httpUri="https://zenodo.org/record/4412682/files/figure.png" pageId="9" pageNumber="516" startId="9.[151,250,1856,1880]" targetBox="[183,1405,181,1832]" targetPageId="9">
<paragraph id="8BF336E66246FFB6FF0A3113BF372C53" blockId="9.[151,1436,1856,1917]" pageId="9" pageNumber="516">
<emphasis id="B938EAF46246FFB6FF0A3113BC3A2C77" bold="true" box="[151,274,1856,1880]" pageId="9" pageNumber="516">FIGURE 6.</emphasis>
<taxonomicName id="4C4C4D656246FFB6FE853112BCE42C77" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[280,460,1856,1880]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="516" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46246FFB6FE853112BCE42C77" box="[280,460,1856,1880]" italics="true" pageId="9" pageNumber="516">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46246FFB6FE4C3112BF222C76" bold="true" box="[465,522,1857,1881]" pageId="9" pageNumber="516">
<taxonomicNameLabel id="A20B578F6246FFB6FE4C3112BF222C76" box="[465,522,1857,1881]" pageId="9" pageNumber="516" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 3.5 mm), FLMNH-UF 16344. A, habitus, dorsolateral view; B, same, lateral view. Photographs by A. Anker.
</paragraph>
</caption>
<caption id="DF33666E6245FFB5FF0A3418B9AB294A" ID-Table-UUID="DF33666E6245FFB5FF0A3418B9AB294A" box="[151,1155,587,613]" httpUri="http://table.plazi.org/id/DF33666E6245FFB5FF0A3418B9AB294A" pageId="10" pageNumber="517" targetBox="[165,2021,628,1577]" targetIsTable="true" targetPageId="10">
<paragraph id="8BF336E66245FFB5FF0A3418B9AB294A" blockId="10.[151,1155,587,613]" box="[151,1155,587,613]" pageId="10" pageNumber="517">
<emphasis id="B938EAF46245FFB5FF0A3418BC22294A" bold="true" box="[151,266,587,613]" pageId="10" pageNumber="517">TABLE 1.</emphasis>
Differentiating characters between 
<taxonomicName id="4C4C4D656245FFB5FD153418BE6E294A" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[648,838,587,613]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="517" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46245FFB5FD153418BE6E294A" box="[648,838,587,613]" italics="true" pageId="10" pageNumber="517">Manaxius paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46245FFB5FCD03418BEA1294A" bold="true" box="[845,905,587,613]" pageId="10" pageNumber="517">
<taxonomicNameLabel id="A20B578F6245FFB5FCD03418BEA1294A" box="[845,905,587,613]" pageId="10" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and five allied species.
</paragraph>
</caption>
<paragraph id="8BF336E66245FFB5FF383427BAB42D06" pageId="10" pageNumber="517">
<table id="F94CC44662450040FF383427BACD2D06" box="[165,2021,628,1577]" gridcols="4" gridrows="15" pageId="10" pageNumber="517" rowsContinueIn="11.[151,2008,605,1625]">
<tr id="357C34A462450040FF383427BACD29A2" box="[165,2021,628,653]" gridrow="0" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383427BF2429A2" box="[165,524,628,653]" gridcol="0" gridrow="0" pageId="10" pageNumber="517">Characters/species</th>
<th id="76AD5DD862450040FDA03427B92D29A2" box="[573,1029,628,653]" gridcol="1" gridrow="0" pageId="10" pageNumber="517">
<taxonomicName id="4C4C4D656245FFB5FD563427BE1029A2" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[715,824,628,653]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="517" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46245FFB5FD563427BE1029A2" box="[715,824,628,653]" italics="true" pageId="10" pageNumber="517">M. paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46245FFB5FCA33427BE5F29A2" bold="true" box="[830,887,628,653]" pageId="10" pageNumber="517">
<taxonomicNameLabel id="A20B578F6245FFB5FCA33427BE5F29A2" box="[830,887,628,653]" pageId="10" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76AD5DD862450040FBBB3427B8D829A2" box="[1062,1520,628,653]" gridcol="2" gridrow="0" pageId="10" pageNumber="517">
<taxonomicName id="4C4C4D656245FFB5FB2E3427B84B29A2" baseAuthorityName="de Man" baseAuthorityYear="1905" box="[1203,1379,628,653]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="517" phylum="Arthropoda" rank="species" species="euophthalmus">
<emphasis id="B938EAF46245FFB5FB2E3427B84B29A2" box="[1203,1379,628,653]" italics="true" pageId="10" pageNumber="517">M. euophthalmus</emphasis>
</taxonomicName>
</th>
<th id="76AD5DD862450040F98C3427BACD29A2" box="[1553,2021,628,653]" gridcol="3" gridrow="0" pageId="10" pageNumber="517">
<taxonomicName id="4C4C4D656245FFB5F95F3427BA1C29A2" baseAuthorityName="Komai" baseAuthorityYear="2011" box="[1730,1844,628,653]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="517" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="B938EAF46245FFB5F95F3427BA1C29A2" box="[1730,1844,628,653]" italics="true" pageId="10" pageNumber="517">M. izuensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="357C34A462450040FF3834CEBACD2999" box="[165,2021,669,694]" gridrow="1" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF3834CEBF242999" box="[165,524,669,694]" gridcol="0" gridrow="1" pageId="10" pageNumber="517">Rostral shape</th>
<td id="76AD5DD862450040FDA034CEB92D2999" box="[573,1029,669,694]" gridcol="1" gridrow="1" pageId="10" pageNumber="517">Relatively broad, triangular, straight</td>
<td id="76AD5DD862450040FBBB34CEB8D82999" box="[1062,1520,669,694]" gridcol="2" gridrow="1" pageId="10" pageNumber="517">Narrowly triangular, nearly straight</td>
<td id="76AD5DD862450040F98C34CEBACD2999" box="[1553,2021,669,694]" gridcol="3" gridrow="1" pageId="10" pageNumber="517">Narrowly triangular, slightly upturned</td>
</tr>
<tr id="357C34A462450040FF383496BACD282D" box="[165,2021,709,770]" gridrow="2" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383496BF24282D" box="[165,524,709,770]" gridcol="0" gridrow="2" pageId="10" pageNumber="517">Rostral length</th>
<td id="76AD5DD862450040FDA03496B92D282D" box="[573,1029,709,770]" gridcol="1" gridrow="2" pageId="10" pageNumber="517">Reaching distal end of article 2 of antennular peduncle</td>
<td id="76AD5DD862450040FBBB3496B8D8282D" box="[1062,1520,709,770]" gridcol="2" gridrow="2" pageId="10" pageNumber="517">Reaching distal end of article 2 of antennular peduncle</td>
<td id="76AD5DD862450040F98C3496BACD282D" box="[1553,2021,709,770]" gridcol="3" gridrow="2" pageId="10" pageNumber="517">Reaching distal end of article 3 of antennular peduncle</td>
</tr>
<tr id="357C34A462450040FF383542BACD2805" box="[165,2021,785,810]" gridrow="3" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383542BF242805" box="[165,524,785,810]" gridcol="0" gridrow="3" pageId="10" pageNumber="517">Rostral armature</th>
<td id="76AD5DD862450040FDA03542B92D2805" box="[573,1029,785,810]" gridcol="1" gridrow="3" pageId="10" pageNumber="517">With 1 pair of spines on lateral margins</td>
<td id="76AD5DD862450040FBBB3542B8D82805" box="[1062,1520,785,810]" gridcol="2" gridrow="3" pageId="10" pageNumber="517">With 2 or 3 spines on each lateral margin</td>
<td id="76AD5DD862450040F98C3542BACD2805" box="[1553,2021,785,810]" gridcol="3" gridrow="3" pageId="10" pageNumber="517">With 2 pairs of spines on lateral margins</td>
</tr>
<tr id="357C34A462450040FF38356ABACD287D" box="[165,2021,825,850]" gridrow="4" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF38356ABF24287D" box="[165,524,825,850]" gridcol="0" gridrow="4" pageId="10" pageNumber="517">Carapace ornamentation</th>
<td id="76AD5DD862450040FDA0356AB92D287D" box="[573,1029,825,850]" gridcol="1" gridrow="4" pageId="10" pageNumber="517">Smooth</td>
<td id="76AD5DD862450040FBBB356AB8D8287D" box="[1062,1520,825,850]" gridcol="2" gridrow="4" pageId="10" pageNumber="517">Smooth</td>
<td id="76AD5DD862450040F98C356ABACD287D" box="[1553,2021,825,850]" gridcol="3" gridrow="4" pageId="10" pageNumber="517">Smooth</td>
</tr>
<tr id="357C34A462450040FF383531BACD28B0" box="[165,2021,866,927]" gridrow="5" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383531BF2428B0" box="[165,524,866,927]" gridcol="0" gridrow="5" pageId="10" pageNumber="517">Carapace, gastric median carina</th>
<td id="76AD5DD862450040FDA03531B92D28B0" box="[573,1029,866,927]" gridcol="1" gridrow="5" pageId="10" pageNumber="517">With 1 anterior spine followed by 2 small denticles and 1 obsolescent tubercle</td>
<td id="76AD5DD862450040FBBB3531B8D828B0" box="[1062,1520,866,927]" gridcol="2" gridrow="5" pageId="10" pageNumber="517">With 1 anterior spine followed by 2 small tubercles</td>
<td id="76AD5DD862450040F98C3531BACD28B0" box="[1553,2021,866,927]" gridcol="3" gridrow="5" pageId="10" pageNumber="517">With 1–3 tubercles and 1 low convexity</td>
</tr>
<tr id="357C34A462450040FF3835FDBACD28E8" box="[165,2021,942,967]" gridrow="6" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF3835FDBF2428E8" box="[165,524,942,967]" gridcol="0" gridrow="6" pageId="10" pageNumber="517">Carapace, gastric submedian carina</th>
<td id="76AD5DD862450040FDA035FDB92D28E8" box="[573,1029,942,967]" gridcol="1" gridrow="6" pageId="10" pageNumber="517">With 3 small spines</td>
<td id="76AD5DD862450040FBBB35FDB8D828E8" box="[1062,1520,942,967]" gridcol="2" gridrow="6" pageId="10" pageNumber="517">With 2 spines</td>
<td id="76AD5DD862450040F98C35FDBACD28E8" box="[1553,2021,942,967]" gridcol="3" gridrow="6" pageId="10" pageNumber="517">With 3–6 tubercles or denticles</td>
</tr>
<tr id="357C34A462450040FF383585BACD2F3C" box="[165,2021,982,1043]" gridrow="7" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383585BF242F3C" box="[165,524,982,1043]" gridcol="0" gridrow="7" pageId="10" pageNumber="517">Antennal scaphocerite</th>
<td id="76AD5DD862450040FDA03585B92D2F3C" box="[573,1029,982,1043]" gridcol="1" gridrow="7" pageId="10" pageNumber="517">Reaching mid-length of article 4 of antennal peduncle</td>
<td id="76AD5DD862450040FBBB3585B8D82F3C" box="[1062,1520,982,1043]" gridcol="2" gridrow="7" pageId="10" pageNumber="517">Overreaching mid-length of article 4 of antennal peduncle</td>
<td id="76AD5DD862450040F98C3585BACD2F3C" box="[1553,2021,982,1043]" gridcol="3" gridrow="7" pageId="10" pageNumber="517">Overreaching mid-length of article 4 of antennal peduncle</td>
</tr>
<tr id="357C34A462450040FF383271BACD2F70" box="[165,2021,1058,1119]" gridrow="8" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383271BF242F70" box="[165,524,1058,1119]" gridcol="0" gridrow="8" pageId="10" pageNumber="517">Pereopod 1 ischium armature</th>
<td id="76AD5DD862450040FDA03271B92D2F70" box="[573,1029,1058,1119]" gridcol="1" gridrow="8" pageId="10" pageNumber="517">Dorsal margin with 1 spine, ventral margin with 1 subdistal spine and some tubercles</td>
<td id="76AD5DD862450040FBBB3271B8D82F70" box="[1062,1520,1058,1119]" gridcol="2" gridrow="8" pageId="10" pageNumber="517">Dorsal margin unarmed, ventral margin with 1 spine</td>
<td id="76AD5DD862450040F98C3271BACD2F70" box="[1553,2021,1058,1119]" gridcol="3" gridrow="8" pageId="10" pageNumber="517">Dorsal margin unarmed, ventral margin with 1 spine</td>
</tr>
<tr id="357C34A462450040FF38323CBACD2FDB" box="[165,2021,1135,1268]" gridrow="9" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF38323CBF242FDB" box="[165,524,1135,1268]" gridcol="0" gridrow="9" pageId="10" pageNumber="517">Pereopod 1 merus armature</th>
<td id="76AD5DD862450040FDA0323CB92D2FDB" box="[573,1029,1135,1268]" gridcol="1" gridrow="9" pageId="10" pageNumber="517">Dorsal margin with 2 subdistal spines; ventral margin with 1 strong subdistal spine followed by 5 or 6 spines</td>
<td id="76AD5DD862450040FBBB323CB8D82FDB" box="[1062,1520,1135,1268]" gridcol="2" gridrow="9" pageId="10" pageNumber="517">Dorsal margin unarmed; ventral margin with 1 subdistal spine followed by 8 blunt teeth or tubercles</td>
<td id="76AD5DD862450040F98C323CBACD2FDB" box="[1553,2021,1135,1268]" gridcol="3" gridrow="9" pageId="10" pageNumber="517">Dorsal margin with 1 subdistal spine; ventral margin with 1 conspicuous subdistal spine followed by several smaller spines or tubercles</td>
</tr>
<tr id="357C34A462450040FF383350BACD2E33" box="[165,2021,1283,1308]" gridrow="10" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383350BF242E33" box="[165,524,1283,1308]" gridcol="0" gridrow="10" pageId="10" pageNumber="517">Pereopod 1 carpus armature</th>
<td id="76AD5DD862450040FDA03350B92D2E33" box="[573,1029,1283,1308]" gridcol="1" gridrow="10" pageId="10" pageNumber="517">With 1 small dorsodistal spine</td>
<td id="76AD5DD862450040FBBB3350B8D82E33" box="[1062,1520,1283,1308]" gridcol="2" gridrow="10" pageId="10" pageNumber="517">With 1 subdistal spine on dorsal margin</td>
<td id="76AD5DD862450040F98C3350BACD2E33" box="[1553,2021,1283,1308]" gridcol="3" gridrow="10" pageId="10" pageNumber="517">With 1 or 2 subdistal spines on dorsal margin</td>
</tr>
<tr id="357C34A462450040FF383378BACD2E47" box="[165,2021,1323,1384]" gridrow="11" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383378BF242E47" box="[165,524,1323,1384]" gridcol="0" gridrow="11" pageId="10" pageNumber="517">Pereopod 1 palm, dorsal margin armature</th>
<td id="76AD5DD862450040FDA03378B92D2E47" box="[573,1029,1323,1384]" gridcol="1" gridrow="11" pageId="10" pageNumber="517">With 3 or 4 spines</td>
<td id="76AD5DD862450040FBBB3378B8D82E47" box="[1062,1520,1323,1384]" gridcol="2" gridrow="11" pageId="10" pageNumber="517">With 2 spines</td>
<td id="76AD5DD862450040F98C3378BACD2E47" box="[1553,2021,1323,1384]" gridcol="3" gridrow="11" pageId="10" pageNumber="517">With 3 or 4 spines</td>
</tr>
<tr id="357C34A462450040FF383324BACD2EF7" box="[165,2021,1399,1496]" gridrow="12" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383324BF242EF7" box="[165,524,1399,1496]" gridcol="0" gridrow="12" pageId="10" pageNumber="517">Pereopod 1 palm, lateral surface armature</th>
<td id="76AD5DD862450040FDA03324B92D2EF7" box="[573,1029,1399,1496]" gridcol="1" gridrow="12" pageId="10" pageNumber="517">With 3 or 4 small spines adjacent to dorsal margin; lateral surface without spine near base of fingers</td>
<td id="76AD5DD862450040FBBB3324B8D82EF7" box="[1062,1520,1399,1496]" gridcol="2" gridrow="12" pageId="10" pageNumber="517">No spines adjacent to dorsal margin; lateral face with spine near base of fingers</td>
<td id="76AD5DD862450040F98C3324BACD2EF7" box="[1553,2021,1399,1496]" gridcol="3" gridrow="12" pageId="10" pageNumber="517">No spines adjacent to dorsal margin; lateral face with spine near base of fingers at least on minor side</td>
</tr>
<tr id="357C34A462450040FF3833BBBACD2D2E" box="[165,2021,1512,1537]" gridrow="13" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF3833BBBF242D2E" box="[165,524,1512,1537]" gridcol="0" gridrow="13" pageId="10" pageNumber="517">Bathymetric range</th>
<td id="76AD5DD862450040FDA033BBB92D2D2E" box="[573,1029,1512,1537]" gridcol="1" gridrow="13" pageId="10" pageNumber="517">Subtidal</td>
<td id="76AD5DD862450040FBBB33BBB8D82D2E" box="[1062,1520,1512,1537]" gridcol="2" gridrow="13" pageId="10" pageNumber="517">397 m</td>
<td id="76AD5DD862450040F98C33BBBACD2D2E" box="[1553,2021,1512,1537]" gridcol="3" gridrow="13" pageId="10" pageNumber="517">440–465 m</td>
</tr>
<tr id="357C34A462450040FF383043BACD2D06" box="[165,2021,1552,1577]" gridrow="14" pageId="10" pageNumber="517">
<th id="76AD5DD862450040FF383043BF242D06" box="[165,524,1552,1577]" gridcol="0" gridrow="14" pageId="10" pageNumber="517">References</th>
<td id="76AD5DD862450040FDA03043B92D2D06" box="[573,1029,1552,1577]" gridcol="1" gridrow="14" pageId="10" pageNumber="517">Present study</td>
<td id="76AD5DD862450040FBBB3043B8D82D06" box="[1062,1520,1552,1577]" gridcol="2" gridrow="14" pageId="10" pageNumber="517">
de 
<bibRefCitation id="EFDD4B176245FFB5FBC53043B8202D06" author="Man, J. G. de" box="[1112,1288,1552,1577]" pageId="10" pageNumber="517" pagination="1 - 127" refId="ref7052" refString="Man, J. G. de (1925 a) The Decapoda of the Siboga-Expedition. Part VI. The Axiidae collected by the Siboga-Expedition. Siboga Expeditie Monographie, 39 a 5, 1 - 127." type="journal article" year="1925">Man (1925a: 122</bibRefCitation>
, pl. 10, fig. 22–22g.)
</td>
<td id="76AD5DD862450040F98C3043BACD2D06" box="[1553,2021,1552,1577]" gridcol="3" gridrow="14" pageId="10" pageNumber="517">
<bibRefCitation id="EFDD4B176245FFB5F9C63043BA3F2D06" author="Komai, T." box="[1627,1815,1552,1577]" pageId="10" pageNumber="517" pagination="279 - 337" refId="ref6852" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011">Komai (2011: 320</bibRefCitation>
, figs. 20–24)
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF336E66245FFB5F92C306BBACF2D7E" blockId="10.[1713,2023,1592,1617]" box="[1713,2023,1592,1617]" pageId="10" pageNumber="517">
<tableNote id="76AA37686245FFB5F92C306BBACF2D7E" box="[1713,2023,1592,1617]" pageId="10" pageNumber="517" targetBox="[165,2021,628,1577]" targetPageId="10">......continued on the next page</tableNote>
</paragraph>
<caption id="DF33666E6244FFB4FF0A3467BCB82962" box="[151,400,564,589]" pageId="11" pageNumber="518" targetBox="[151,2008,605,1625]" targetIsTable="true" targetPageId="11">
<paragraph id="8BF336E66244FFB4FF0A3467BCB82962" blockId="11.[151,400,564,589]" box="[151,400,564,589]" pageId="11" pageNumber="518">TABLE 1. (Continued)</paragraph>
</caption>
<paragraph id="8BF336E66244FFB4FF0A340EBA822D76" pageId="11" pageNumber="518">
<table id="F94CC44662440040FF0A340EBAF02D76" box="[151,2008,605,1625]" gridcols="4" gridrows="15" pageId="11" pageNumber="518" rowsContinueFrom="10.[165,2021,628,1577]">
<tr id="357C34A462440040FF0A340EBAF02959" box="[151,2008,605,630]" gridrow="0" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A340EBCD62959" box="[151,510,605,630]" gridcol="0" gridrow="0" pageId="11" pageNumber="518">Characters/species</th>
<th id="76AD5DD862440040FDA1340EB9292959" box="[572,1025,605,630]" gridcol="1" gridrow="0" pageId="11" pageNumber="518">
<taxonomicName id="4C4C4D656244FFB4FD4E340EBE422959" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[723,874,605,630]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="518" phylum="Arthropoda" rank="species" species="mimasensis">
<emphasis id="B938EAF46244FFB4FD4E340EBE422959" box="[723,874,605,630]" italics="true" pageId="11" pageNumber="518">M. mimasensis</emphasis>
</taxonomicName>
</th>
<th id="76AD5DD862440040FBB8340EB8C52959" box="[1061,1517,605,630]" gridcol="2" gridrow="0" pageId="11" pageNumber="518">
<taxonomicName id="4C4C4D656244FFB4FB4C340EB8682959" baseAuthorityName="Komai" baseAuthorityYear="2016" box="[1233,1344,605,630]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="518" phylum="Arthropoda" rank="species" species="poupini">
<emphasis id="B938EAF46244FFB4FB4C340EB8682959" box="[1233,1344,605,630]" italics="true" pageId="11" pageNumber="518">M. poupini</emphasis>
</taxonomicName>
</th>
<th id="76AD5DD862440040F992340EBAF02959" box="[1551,2008,605,630]" gridcol="3" gridrow="0" pageId="11" pageNumber="518">
<taxonomicName id="4C4C4D656244FFB4F95A340EBA082959" authorityName="Poore" authorityYear="2020" box="[1735,1824,605,630]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="518" phylum="Arthropoda" rank="species" species="supia">
<emphasis id="B938EAF46244FFB4F95A340EBA082959" box="[1735,1824,605,630]" italics="true" pageId="11" pageNumber="518">M. supia</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="357C34A462440040FF0A34D6BAF029B1" box="[151,2008,645,670]" gridrow="1" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A34D6BCD629B1" box="[151,510,645,670]" gridcol="0" gridrow="1" pageId="11" pageNumber="518">Rostral shape</th>
<td id="76AD5DD862440040FDA134D6B92929B1" box="[572,1025,645,670]" gridcol="1" gridrow="1" pageId="11" pageNumber="518">Narrowly triangular, nearly straight</td>
<td id="76AD5DD862440040FBB834D6B8C529B1" box="[1061,1517,645,670]" gridcol="2" gridrow="1" pageId="11" pageNumber="518">Narrowly triangular, nearly straight</td>
<td id="76AD5DD862440040F99234D6BAF029B1" box="[1551,2008,645,670]" gridcol="3" gridrow="1" pageId="11" pageNumber="518">Narrowly triangular, nearly straight</td>
</tr>
<tr id="357C34A462440040FF0A34FEBAF029C5" box="[151,2008,685,746]" gridrow="2" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A34FEBCD629C5" box="[151,510,685,746]" gridcol="0" gridrow="2" pageId="11" pageNumber="518">Rostral length</th>
<td id="76AD5DD862440040FDA134FEB92929C5" box="[572,1025,685,746]" gridcol="1" gridrow="2" pageId="11" pageNumber="518">Reaching mid-length of article 3 of antennular peduncle</td>
<td id="76AD5DD862440040FBB834FEB8C529C5" box="[1061,1517,685,746]" gridcol="2" gridrow="2" pageId="11" pageNumber="518">Reaching mid-length of article 3 of antennular peduncle</td>
<td id="76AD5DD862440040F99234FEBAF029C5" box="[1551,2008,685,746]" gridcol="3" gridrow="2" pageId="11" pageNumber="518">Reaching distal end of article 2 of antennular peduncle</td>
</tr>
<tr id="357C34A462440040FF0A34A9BAF02818" box="[151,2008,762,823]" gridrow="3" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A34A9BCD62818" box="[151,510,762,823]" gridcol="0" gridrow="3" pageId="11" pageNumber="518">Rostral armature</th>
<td id="76AD5DD862440040FDA134A9B9292818" box="[572,1025,762,823]" gridcol="1" gridrow="3" pageId="11" pageNumber="518">With 2 spines on either lateral margin</td>
<td id="76AD5DD862440040FBB834A9B8C52818" box="[1061,1517,762,823]" gridcol="2" gridrow="3" pageId="11" pageNumber="518">With 1–3 lateral spines on either lateral margin</td>
<td id="76AD5DD862440040F99234A9BAF02818" box="[1551,2008,762,823]" gridcol="3" gridrow="3" pageId="11" pageNumber="518">With 2 or 3 spines on either lateral margin</td>
</tr>
<tr id="357C34A462440040FF0A3515BAF02870" box="[151,2008,838,863]" gridrow="4" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A3515BCD62870" box="[151,510,838,863]" gridcol="0" gridrow="4" pageId="11" pageNumber="518">Carapace ornamentation</th>
<td id="76AD5DD862440040FDA13515B9292870" box="[572,1025,838,863]" gridcol="1" gridrow="4" pageId="11" pageNumber="518">Dorsally granulate</td>
<td id="76AD5DD862440040FBB83515B8C52870" box="[1061,1517,838,863]" gridcol="2" gridrow="4" pageId="11" pageNumber="518">Smooth</td>
<td id="76AD5DD862440040F9923515BAF02870" box="[1551,2008,838,863]" gridcol="3" gridrow="4" pageId="11" pageNumber="518">Smooth</td>
</tr>
<tr id="357C34A462440040FF0A353DBAF02884" box="[151,2008,878,939]" gridrow="5" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A353DBCD62884" box="[151,510,878,939]" gridcol="0" gridrow="5" pageId="11" pageNumber="518">Carapace, gastric median carina</th>
<td id="76AD5DD862440040FDA1353DB9292884" box="[572,1025,878,939]" gridcol="1" gridrow="5" pageId="11" pageNumber="518">With 7–10 small tubercles or denticles</td>
<td id="76AD5DD862440040FBB8353DB8C52884" box="[1061,1517,878,939]" gridcol="2" gridrow="5" pageId="11" pageNumber="518">With 2–4 tubercles followed by 1 low con- vexity</td>
<td id="76AD5DD862440040F992353DBAF02884" box="[1551,2008,878,939]" gridcol="3" gridrow="5" pageId="11" pageNumber="518">With 3 tubercles</td>
</tr>
<tr id="357C34A462440040FF0A35E9BAF028FC" box="[151,2008,954,979]" gridrow="6" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A35E9BCD628FC" box="[151,510,954,979]" gridcol="0" gridrow="6" pageId="11" pageNumber="518">Carapace, gastric submedian carina</th>
<td id="76AD5DD862440040FDA135E9B92928FC" box="[572,1025,954,979]" gridcol="1" gridrow="6" pageId="11" pageNumber="518">With 9–11 spinules</td>
<td id="76AD5DD862440040FBB835E9B8C528FC" box="[1061,1517,954,979]" gridcol="2" gridrow="6" pageId="11" pageNumber="518">With 2–4 tubercles or denticles</td>
<td id="76AD5DD862440040F99235E9BAF028FC" box="[1551,2008,954,979]" gridcol="3" gridrow="6" pageId="11" pageNumber="518">With 4 tubercles</td>
</tr>
<tr id="357C34A462440040FF0A35B1BAF02F30" box="[151,2008,994,1055]" gridrow="7" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A35B1BCD62F30" box="[151,510,994,1055]" gridcol="0" gridrow="7" pageId="11" pageNumber="518">Antennal scaphocerite</th>
<td id="76AD5DD862440040FDA135B1B9292F30" box="[572,1025,994,1055]" gridcol="1" gridrow="7" pageId="11" pageNumber="518">Overreaching mid-length of article 4 of antennal peduncle</td>
<td id="76AD5DD862440040FBB835B1B8C52F30" box="[1061,1517,994,1055]" gridcol="2" gridrow="7" pageId="11" pageNumber="518">Overreaching mid-length of article 4 of antennal peduncle</td>
<td id="76AD5DD862440040F99235B1BAF02F30" box="[1551,2008,994,1055]" gridcol="3" gridrow="7" pageId="11" pageNumber="518">Overreaching mid-length of article 4 of antennal peduncle</td>
</tr>
<tr id="357C34A462440040FF0A327CBAF02F43" box="[151,2008,1071,1132]" gridrow="8" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A327CBCD62F43" box="[151,510,1071,1132]" gridcol="0" gridrow="8" pageId="11" pageNumber="518">Pereopod 1 ischium armature</th>
<td id="76AD5DD862440040FDA1327CB9292F43" box="[572,1025,1071,1132]" gridcol="1" gridrow="8" pageId="11" pageNumber="518">Dorsal margin unarmed; ventral margin with 3 or 4 ventral spines</td>
<td id="76AD5DD862440040FBB8327CB8C52F43" box="[1061,1517,1071,1132]" gridcol="2" gridrow="8" pageId="11" pageNumber="518">Dorsal margin unarmed; ventral margin with 1 spine</td>
<td id="76AD5DD862440040F992327CBAF02F43" box="[1551,2008,1071,1132]" gridcol="3" gridrow="8" pageId="11" pageNumber="518">Dorsal margin unarmed; ventral margin with 1 spine followed by small tubercle(s)</td>
</tr>
<tr id="357C34A462440040FF0A32CCBAF02E2F" box="[151,2008,1183,1280]" gridrow="9" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A32CCBCD62E2F" box="[151,510,1183,1280]" gridcol="0" gridrow="9" pageId="11" pageNumber="518">Pereopod 1 merus armature</th>
<td id="76AD5DD862440040FDA132CCB9292E2F" box="[572,1025,1183,1280]" gridcol="1" gridrow="9" pageId="11" pageNumber="518">Dorsal margin with 2–4 subdistal spines; ventral margin with 5 or 6 conspicuous spines</td>
<td id="76AD5DD862440040FBB832CCB8C52E2F" box="[1061,1517,1183,1280]" gridcol="2" gridrow="9" pageId="11" pageNumber="518">Dorsal margin with 1 subdistal spine; ventral margin unarmed or armed with 1 subdistal spine</td>
<td id="76AD5DD862440040F99232CCBAF02E2F" box="[1551,2008,1183,1280]" gridcol="3" gridrow="9" pageId="11" pageNumber="518">Dorsal margin with 2 or 3 spines; ventral margin with 1 or 2 spines near midlength</td>
</tr>
<tr id="357C34A462440040FF0A335CBAF02E63" box="[151,2008,1295,1356]" gridrow="10" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A335CBCD62E63" box="[151,510,1295,1356]" gridcol="0" gridrow="10" pageId="11" pageNumber="518">Pereopod 1 carpus armature</th>
<td id="76AD5DD862440040FDA1335CB9292E63" box="[572,1025,1295,1356]" gridcol="1" gridrow="10" pageId="11" pageNumber="518">With 3 spines on dorsal margin on major cheliped side</td>
<td id="76AD5DD862440040FBB8335CB8C52E63" box="[1061,1517,1295,1356]" gridcol="2" gridrow="10" pageId="11" pageNumber="518">With 4 or 5 small spines on dorsal margin</td>
<td id="76AD5DD862440040F992335CBAF02E63" box="[1551,2008,1295,1356]" gridcol="3" gridrow="10" pageId="11" pageNumber="518">With 1 subdistal spine on dorsal margin</td>
</tr>
<tr id="357C34A462440040FF0A3308BAF02EB7" box="[151,2008,1371,1432]" gridrow="11" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A3308BCD62EB7" box="[151,510,1371,1432]" gridcol="0" gridrow="11" pageId="11" pageNumber="518">Pereopod 1 palm, dorsal margin armature</th>
<td id="76AD5DD862440040FDA13308B9292EB7" box="[572,1025,1371,1432]" gridcol="1" gridrow="11" pageId="11" pageNumber="518">With 3 or 4 spines</td>
<td id="76AD5DD862440040FBB83308B8C52EB7" box="[1061,1517,1371,1432]" gridcol="2" gridrow="11" pageId="11" pageNumber="518">With 3 or 4 spines</td>
<td id="76AD5DD862440040F9923308BAF02EB7" box="[1551,2008,1371,1432]" gridcol="3" gridrow="11" pageId="11" pageNumber="518">With 3 or 4 spines</td>
</tr>
<tr id="357C34A462440040FF0A33FBBAF02D26" box="[151,2008,1448,1545]" gridrow="12" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A33FBBCD62D26" box="[151,510,1448,1545]" gridcol="0" gridrow="12" pageId="11" pageNumber="518">Pereopod 1 palm, lateral surface armature</th>
<td id="76AD5DD862440040FDA133FBB9292D26" box="[572,1025,1448,1545]" gridcol="1" gridrow="12" pageId="11" pageNumber="518">No spines adjacent to dorsal margin; lateral face with spine near base of fingers at least on minor cheliped side</td>
<td id="76AD5DD862440040FBB833FBB8C52D26" box="[1061,1517,1448,1545]" gridcol="2" gridrow="12" pageId="11" pageNumber="518">No spines adjacent to dorsal margin; lateral face with spine near base of fingers at least on minor cheliped side</td>
<td id="76AD5DD862440040F99233FBBAF02D26" box="[1551,2008,1448,1545]" gridcol="3" gridrow="12" pageId="11" pageNumber="518">No spines adjacent to dorsal margin; lateral face with spine near base of fingers</td>
</tr>
<tr id="357C34A462440040FF0A304BBAF02D1E" box="[151,2008,1560,1585]" gridrow="13" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A304BBCD62D1E" box="[151,510,1560,1585]" gridcol="0" gridrow="13" pageId="11" pageNumber="518">Bathymetric range</th>
<td id="76AD5DD862440040FDA1304BB9292D1E" box="[572,1025,1560,1585]" gridcol="1" gridrow="13" pageId="11" pageNumber="518">133–200 m</td>
<td id="76AD5DD862440040FBB8304BB8C52D1E" box="[1061,1517,1560,1585]" gridcol="2" gridrow="13" pageId="11" pageNumber="518">646–737 m</td>
<td id="76AD5DD862440040F992304BBAF02D1E" box="[1551,2008,1560,1585]" gridcol="3" gridrow="13" pageId="11" pageNumber="518">340–430 m</td>
</tr>
<tr id="357C34A462440040FF0A3013BAF02D76" box="[151,2008,1600,1625]" gridrow="14" pageId="11" pageNumber="518">
<th id="76AD5DD862440040FF0A3013BCD62D76" box="[151,510,1600,1625]" gridcol="0" gridrow="14" pageId="11" pageNumber="518">References</th>
<td id="76AD5DD862440040FDA13013B9292D76" box="[572,1025,1600,1625]" gridcol="1" gridrow="14" pageId="11" pageNumber="518">
<bibRefCitation id="EFDD4B176244FFB4FDE83013BE762D76" author="Komai, T. &amp; Ohtsuka, S. &amp; Nakaguchi, K. &amp; Go, A." box="[629,862,1600,1625]" pageId="11" pageNumber="518" pagination="19 - 73" refId="ref6964" refString="Komai, T., Ohtsuka, S., Nakaguchi, K. &amp; Go, A. (2002) Decapod crustaceans collected from the southern part of the Sea of Japan in 2000 - 2001 using TRV Toyoshio-maru. Natural History Research, 7, 19 - 73." type="journal article" year="2002">
Komai 
<emphasis id="B938EAF46244FFB4FD5D3013BFC52D76" box="[704,749,1600,1625]" italics="true" pageId="11" pageNumber="518">et al</emphasis>
. (2002: 30
</bibRefCitation>
, figs. 4–6)
</td>
<td id="76AD5DD862440040FBB83013B8C52D76" box="[1061,1517,1600,1625]" gridcol="2" gridrow="14" pageId="11" pageNumber="518">
<bibRefCitation id="EFDD4B176244FFB4FBE83013B8192D76" author="Komai, T." box="[1141,1329,1600,1625]" pageId="11" pageNumber="518" pagination="498 - 510" refId="ref6901" refString="Komai, T. (2016) Reappraisal of the two axiid genera Manaxius Kensley, 2003 and Calaxidium Sakai, 2014, and description of a new species of Calaxius Sakai &amp; de Saint Laurent, 1989 from French Polynesia, Southwest Pacific (Crustacea: Decapoda: Axiidea). Zootaxa, 4098 (3), 498 - 510." type="journal article" year="2016">Komai (2016: 502</bibRefCitation>
, figs. 1–3)
</td>
<td id="76AD5DD862440040F9923013BAF02D76" box="[1551,2008,1600,1625]" gridcol="3" gridrow="14" pageId="11" pageNumber="518">
<bibRefCitation id="EFDD4B176244FFB4F9A13013BBD82D76" author="Poore, G. C. B." box="[1596,1776,1600,1625]" pageId="11" pageNumber="518" pagination="261 - 365" refId="ref7181" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds.), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum National d'Histoire Naturelle, Paris, 213, pp. 261 - 365." type="journal article" year="2020">Poore (2020: 315</bibRefCitation>
: figs. 22C, 27, 28)
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF336E66243FFB3FF5A34DCBEE72F3E" blockId="12.[151,1437,150,1329]" pageId="12" pageNumber="519">
Differentiating characters among 
<taxonomicName id="4C4C4D656243FFB3FDDB34DCBF972986" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[582,703,655,681]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46243FFB3FDDB34DCBF972986" box="[582,703,655,681]" italics="true" pageId="12" pageNumber="519">M. paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46243FFB3FD5534DCBE212986" bold="true" box="[712,777,655,681]" pageId="12" pageNumber="519">
<taxonomicNameLabel id="A20B578F6243FFB3FD5534DCBE212986" box="[712,777,655,681]" pageId="12" pageNumber="519" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and the remaining five species, i.e. 
<taxonomicName id="4C4C4D656243FFB3FB3734DCB8462986" baseAuthorityName="de Man" baseAuthorityYear="1905" box="[1194,1390,655,681]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="euophthalmus">
<emphasis id="B938EAF46243FFB3FB3734DCB8462986" box="[1194,1390,655,681]" italics="true" pageId="12" pageNumber="519">M. euophthalmus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FAE334DCBDD829E3" baseAuthorityName="Komai" baseAuthorityYear="2011" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="B938EAF46243FFB3FAE334DCBDD829E3" italics="true" pageId="12" pageNumber="519">M. izuensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FE9D34E0BC8129E3" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[256,425,691,716]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="mimasensis">
<emphasis id="B938EAF46243FFB3FE9D34E0BC8129E3" box="[256,425,691,716]" italics="true" pageId="12" pageNumber="519">M. mimasensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FE2434E0BF1E29E3" baseAuthorityName="Komai" baseAuthorityYear="2016" box="[441,566,691,716]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="poupini">
<emphasis id="B938EAF46243FFB3FE2434E0BF1E29E3" box="[441,566,691,716]" italics="true" pageId="12" pageNumber="519">M. poupini</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C4D656243FFB3FDEC34E0BFFD29E3" authorityName="Poore" authorityYear="2020" box="[625,725,691,716]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="supia">
<emphasis id="B938EAF46243FFB3FDEC34E0BFFD29E3" box="[625,725,691,716]" italics="true" pageId="12" pageNumber="519">M. supia</emphasis>
</taxonomicName>
, are summarised in 
<tableCitation id="C6CE035D6243FFB3FC5D34E0B93E29E2" box="[960,1046,691,717]" captionStart="TABLE 1" captionStartId="10.[151,239,587,613]" captionTargetPageId="10" captionText="TABLE 1. Differentiating characters between Manaxius paullus n. sp. and five allied species." httpUri="http://table.plazi.org/id/DF33666E6245FFB5FF0A3418B9AB294A" pageId="12" pageNumber="519" tableUuid="DF33666E6245FFB5FF0A3418B9AB294A">Table 1</tableCitation>
. Interspecific differences are seen in the rostral shape and armature, the armature of the gastric median and submedian carinae, the development of the antennal scaphocerite and the armature of the chelipeds. The main distinguishing characters of the new species are the relatively broad rostrum, with one pair of lateral spines in addition to supraocular spines (
<emphasis id="B938EAF46243FFB3FB4B354CB9DD2817" box="[1238,1269,799,824]" italics="true" pageId="12" pageNumber="519">vs.</emphasis>
usually two or three pairs of lateral spines, often arranged asymmetrically, in other species); the presence of a subdistal spine on the dorsal margin of the cheliped ischium (
<emphasis id="B938EAF46243FFB3FDDA3534BF4D28AF" box="[583,613,871,896]" italics="true" pageId="12" pageNumber="519">vs.</emphasis>
spineless on the dorsal margin in other species); the presence of additional three or four small spines on each cheliped palm adjacent to the dorsal margin (
<emphasis id="B938EAF46243FFB3FC6235D8B935288B" box="[1023,1053,907,932]" italics="true" pageId="12" pageNumber="519">vs.</emphasis>
without spines in this area in other species); and the lack of a spine near the base of fingers of each cheliped palm (
<emphasis id="B938EAF46243FFB3FBBF35FCB91128E7" box="[1058,1081,943,968]" italics="true" pageId="12" pageNumber="519">vs</emphasis>
. with one conspicuous spine in other species). Furthermore, the new species occurs in shallow waters on coral reefs, whereas the other five species are found at lower continental shelf to upper bathyal depths (see 
<tableCitation id="C6CE035D6243FFB3FCF135A4BE962F3E" box="[876,958,1015,1041]" captionStart="TABLE 1" captionStartId="10.[151,239,587,613]" captionTargetPageId="10" captionText="TABLE 1. Differentiating characters between Manaxius paullus n. sp. and five allied species." httpUri="http://table.plazi.org/id/DF33666E6245FFB5FF0A3418B9AB294A" pageId="12" pageNumber="519" tableUuid="DF33666E6245FFB5FF0A3418B9AB294A">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BF336E66243FFB3FF5A3248BCB82E1E" blockId="12.[151,1437,150,1329]" pageId="12" pageNumber="519">
Only one species of 
<taxonomicName id="4C4C4D656243FFB3FE2E3248BF092F1B" authorityName="Kensley" authorityYear="2003" box="[435,545,1051,1076]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAF46243FFB3FE2E3248BF092F1B" box="[435,545,1051,1076]" italics="true" pageId="12" pageNumber="519">Manaxius</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FDAC3248BF862F1B" baseAuthorityName="Komai" baseAuthorityYear="2016" box="[561,686,1051,1076]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="poupini">
<emphasis id="B938EAF46243FFB3FDAC3248BF862F1B" box="[561,686,1051,1076]" italics="true" pageId="12" pageNumber="519">M. poupini</emphasis>
</taxonomicName>
, has been hitherto recorded from 
<collectingCountry id="F35B76766243FFB3FBAF3248B9DE2F1A" box="[1074,1270,1051,1077]" name="French Polynesia" pageId="12" pageNumber="519">French Polynesia</collectingCountry>
(
<bibRefCitation id="EFDD4B176243FFB3FA9B3248B8B02F1A" author="Komai, T." box="[1286,1432,1051,1077]" pageId="12" pageNumber="519" pagination="498 - 510" refId="ref6901" refString="Komai, T. (2016) Reappraisal of the two axiid genera Manaxius Kensley, 2003 and Calaxidium Sakai, 2014, and description of a new species of Calaxius Sakai &amp; de Saint Laurent, 1989 from French Polynesia, Southwest Pacific (Crustacea: Decapoda: Axiidea). Zootaxa, 4098 (3), 498 - 510." type="journal article" year="2016">Komai 2016</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FF0A326CBC372F76" author="Poore, G. C. B." box="[151,287,1087,1113]" pageId="12" pageNumber="519" pagination="261 - 365" refId="ref7181" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds.), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum National d'Histoire Naturelle, Paris, 213, pp. 261 - 365." type="journal article" year="2020">Poore 2020</bibRefCitation>
), with the present new species being the second species of the genus known from the area. 
<bibRefCitation id="EFDD4B176243FFB3FAC6326CBDCA2F52" author="Poore, G. C. B." pageId="12" pageNumber="519" pagination="261 - 365" refId="ref7181" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds.), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum National d'Histoire Naturelle, Paris, 213, pp. 261 - 365." type="journal article" year="2020">Poore (2020)</bibRefCitation>
suggested that specimens from 
<collectingCountry id="F35B76766243FFB3FDD63230BE252F52" box="[587,781,1123,1149]" name="French Polynesia" pageId="12" pageNumber="519">French Polynesia</collectingCountry>
identified as 
<taxonomicName id="4C4C4D656243FFB3FC3B3230B8032F52" authority="(de Man, 1925 b)" baseAuthorityName="de Man" baseAuthorityYear="1925" box="[934,1323,1122,1149]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="B938EAF46243FFB3FC3B3230B9402F52" box="[934,1128,1123,1149]" italics="true" pageId="12" pageNumber="519">Calaxius sibogae</emphasis>
(de 
<bibRefCitation id="EFDD4B176243FFB3FB073230B80B2F52" author="Man, J. G. de" box="[1178,1315,1122,1149]" pageId="12" pageNumber="519" pagination="218 - 219" refId="ref7091" refString="Man, J. G. de (1925 b) Axiopsis (Eiconaxius) sibogae n. sp. Zoologische Mededelingen, Leiden, 8, 218 - 219." type="journal article" year="1925">Man, 1925b</bibRefCitation>
)
</taxonomicName>
by Ngoc- Ho (2005) might actually represent 
<taxonomicName id="4C4C4D656243FFB3FDAB32D4BF9C2F8F" baseAuthorityName="Komai" baseAuthorityYear="2016" box="[566,692,1159,1184]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="poupini">
<emphasis id="B938EAF46243FFB3FDAB32D4BF9C2F8F" box="[566,692,1159,1184]" italics="true" pageId="12" pageNumber="519">M. poupini</emphasis>
</taxonomicName>
. Four species of 
<taxonomicName id="4C4C4D656243FFB3FCE232D4BEC52F8F" authorityName="Kensley" authorityYear="2003" box="[895,1005,1159,1184]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAF46243FFB3FCE232D4BEC52F8F" box="[895,1005,1159,1184]" italics="true" pageId="12" pageNumber="519">Manaxius</emphasis>
</taxonomicName>
, viz. 
<emphasis id="B938EAF46243FFB3FBAF32D4B8452F8F" box="[1074,1389,1159,1184]" italics="true" pageId="12" pageNumber="519">
<taxonomicName id="4C4C4D656243FFB3FBAF32D4B99D2F8F" baseAuthorityName="Komai" baseAuthorityYear="2011" box="[1074,1205,1159,1184]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="izuensis">M. izuensis</taxonomicName>
, 
<taxonomicName id="4C4C4D656243FFB3FB5932D4B8452F8F" baseAuthorityName="Sakai" baseAuthorityYear="1967" box="[1220,1389,1159,1184]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="mimasensis">M. mimasensis</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C4C4D656243FFB3FAE332D4BC082FEA" authorityName="Sakai" authorityYear="2011" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="ohsumiensis">
<emphasis id="B938EAF46243FFB3FAE332D4BC082FEA" italics="true" pageId="12" pageNumber="519">M. ohsumiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C4D656243FFB3FECA32F8BF2B2FEB" baseAuthorityName="de Man" baseAuthorityYear="1925" box="[343,515,1195,1220]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="B938EAF46243FFB3FECA32F8BF2B2FEB" box="[343,515,1195,1220]" italics="true" pageId="12" pageNumber="519">M. pitatucensis</emphasis>
</taxonomicName>
, have been recorded so far from Japanese waters (
<bibRefCitation id="EFDD4B176243FFB3FBA632F8B9952FEA" author="Sakai, K." box="[1083,1213,1195,1221]" pageId="12" pageNumber="519" pagination="39 - 51" refId="ref7392" refString="Sakai, K. (1967) Three new species of Thalassinidea (Decapoda, Crustacea) from Japan. Researches on Crustacea, 3, 39 - 51. https: // doi. org / 10.18353 / rcustacea. 3.0 _ 39" type="journal article" year="1967">Sakai 1967</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FB5432F8B8B02FEA" author="Komai, T. &amp; Ohtsuka, S. &amp; Nakaguchi, K. &amp; Go, A." box="[1225,1432,1195,1221]" pageId="12" pageNumber="519" pagination="19 - 73" refId="ref6964" refString="Komai, T., Ohtsuka, S., Nakaguchi, K. &amp; Go, A. (2002) Decapod crustaceans collected from the southern part of the Sea of Japan in 2000 - 2001 using TRV Toyoshio-maru. Natural History Research, 7, 19 - 73." type="journal article" year="2002">
Komai 
<emphasis id="B938EAF46243FFB3FA8032F8B8662FEA" box="[1309,1358,1195,1221]" italics="true" pageId="12" pageNumber="519">et al</emphasis>
. 2002
</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FF0A329CBC0D2FC6" author="Komai, T." box="[151,293,1231,1257]" pageId="12" pageNumber="519" pagination="279 - 337" refId="ref6852" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011">Komai 2011</bibRefCitation>
; 
<bibRefCitation id="EFDD4B176243FFB3FEAF329CBC9F2FC6" author="Poore, G. C. B." box="[306,439,1231,1257]" pageId="12" pageNumber="519" pagination="261 - 365" refId="ref7181" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds.), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum National d'Histoire Naturelle, Paris, 213, pp. 261 - 365." type="journal article" year="2020">Poore 2020</bibRefCitation>
), with 
<taxonomicName id="4C4C4D656243FFB3FD9B329CBF562FC6" authority="Komai &amp; Anker, 2020" authorityName="Komai &amp; Anker" authorityYear="2020" box="[518,638,1231,1257]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="paullus" status="sp. nov.">
<emphasis id="B938EAF46243FFB3FD9B329CBF562FC6" box="[518,638,1231,1257]" italics="true" pageId="12" pageNumber="519">M. paullus</emphasis>
</taxonomicName>
<emphasis id="B938EAF46243FFB3FD1B329CBFEE2FC6" bold="true" box="[646,710,1231,1257]" pageId="12" pageNumber="519">
<taxonomicNameLabel id="A20B578F6243FFB3FD1B329CBFEE2FC6" box="[646,710,1231,1257]" pageId="12" pageNumber="519" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
now representing the fifth species. 
<bibRefCitation id="EFDD4B176243FFB3FBC6329CB9C72FC6" author="Poore, G. C. B." box="[1115,1263,1231,1257]" pageId="12" pageNumber="519" pagination="261 - 365" refId="ref7181" refString="Poore, G. C. B. (2020) Axiid and micheleid lobsters from Indo-West Pacific deep-sea environments (Crustacea: Decapoda: Axiidea: Axiidae, Micheleidae). In: Corbari, L., Ahyong, S. T. &amp; Chan, T. - Y. (Eds.), Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos 31. Memoires du Museum National d'Histoire Naturelle, Paris, 213, pp. 261 - 365." type="journal article" year="2020">Poore (2020)</bibRefCitation>
concluded that 
<taxonomicName id="4C4C4D656243FFB3FF0A32A0BC6E2E22" authorityName="Sakai" authorityYear="2011" box="[151,326,1267,1293]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="ohsumiensis">
<emphasis id="B938EAF46243FFB3FF0A32A0BC6E2E22" box="[151,326,1267,1293]" italics="true" pageId="12" pageNumber="519">M. ohsumiensis</emphasis>
</taxonomicName>
was conspecific with 
<taxonomicName id="4C4C4D656243FFB3FDA232A0B91D2E22" authority="sensu Kensley (2003)" authorityName="sensu Kensley" authorityYear="2003" box="[575,1077,1267,1293]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="B938EAF46243FFB3FDA232A0BEA82E23" box="[575,896,1267,1292]" italics="true" pageId="12" pageNumber="519">Manaxius pitatucensis sensu</emphasis>
<bibRefCitation id="EFDD4B176243FFB3FC1A32A0B91D2E22" author="Kensley, B." box="[903,1077,1267,1293]" pageId="12" pageNumber="519" pagination="359 - 384" refId="ref6768" refString="Kensley, B. (2003) Axioid shrimps from Guam (Crustacea, Decapoda, Thalassinidea). Micronesica, 35 - 36, 359 - 384." type="journal article" year="2003">Kensley (2003)</bibRefCitation>
</taxonomicName>
(= 
<taxonomicName id="4C4C4D656243FFB3FBC132A0BDF92E1E" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="519" phylum="Arthropoda" rank="species" species="aganaensis">
<emphasis id="B938EAF46243FFB3FBC132A0B8782E23" box="[1116,1360,1267,1292]" italics="true" pageId="12" pageNumber="519">Manaxius aganaensis</emphasis>
<bibRefCitation id="EFDD4B176243FFB3FACA32A0BDF92E1E" author="Sakai, K." pageId="12" pageNumber="519" pagination="1 - 616" refId="ref7436" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. https: // doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B938EAF46243FFB3FF403344BC572E1E" box="[221,383,1303,1329]" italics="true" pageId="12" pageNumber="519">Nomen nudum</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>