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<document id="B8B64AF65C4F6A2BCF1A9E8238E78394" ID-DOI="10.5281/zenodo.5327198" ID-ISSN="0374-1036" ID-Zenodo-Dep="5327198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630155436148" checkinUser="felipe" docAuthor="Yin, Zi-Wei, Li, Li-Zhen &amp; Zhao, Mei-Jun" docDate="2011" docId="03E587D81951FFF9FE6AFC4EFCD79516" docLanguage="en" docName="ActaEntMusNatPra.2011.51.1.123-135.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 51 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Linan Hlavac 2002" docType="treatment" docVersion="4" lastPageNumber="126" masterDocId="FFDCFFA01950FFFAFFA9FFF1FF92904E" masterDocTitle="A review of the genus Linan (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="135" masterPageNumber="123" pageNumber="124" updateTime="1699167225380" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F3B8DC2756115CA1AB3592F15273B756">A review of the genus Linan (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="05B9B51C87EFAF329ACF7FC282F1B6BA">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="48D2D1E427725E7D0588A547D58B159F">Department of Biology, College of Life and Environment Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China; e-mails: yin _ ziwei @ yahoo. com; lizhenli @ shnu. edu. cn</mods:affiliation>
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<mods:namePart id="1D5AD9939217BB1CDF6305EC97188765">Li, Li-Zhen</mods:namePart>
<mods:affiliation id="CF948710206A17E4F123D1A82121B6EF">Department of Biology, College of Life and Environment Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China; e-mails: yin _ ziwei @ yahoo. com; lizhenli @ shnu. edu. cn</mods:affiliation>
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<mods:namePart id="9795174CEE74F9595536A66A4FEFACE2">Zhao, Mei-Jun</mods:namePart>
<mods:affiliation id="779AC66D25AD5FBBD1178D404DA5EF51">Department of Biology, College of Life and Environment Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China; e-mails: yin _ ziwei @ yahoo. com; lizhenli @ shnu. edu. cn &amp; Corresponding author; e-mail: mjzhao @ shnu. edu. cn</mods:affiliation>
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<mods:title id="E9D70AA496659C6A2D9452E319E65014">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="5A490FFC41C334E8EF7DCBEF76F9D96C">2011</mods:date>
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<mods:number id="40556CEF9F1D9E125E3B4CACA23E019A">51</mods:number>
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Genus
<taxonomicName id="4C4C4D4D1951FFFBFDBAFC4EFD7C9399" ID-CoL="8VYXL" authority="Hlavac, 2002" authorityName="Hlavac" authorityYear="2002" box="[531,750,959,984]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B938EADC1951FFFBFDBAFC4EFDC49399" bold="true" box="[531,598,959,983]" italics="true" pageId="1" pageNumber="124">Linan</emphasis>
<bibRefCitation id="EFDD4B3F1951FFFBFDF5FC4EFD7C9399" author="HLAVAC P." box="[604,750,959,984]" pageId="1" pageNumber="124" pagination="283 - 297" refId="ref5971" refString="HLAVAC P. 2002: A taxonomic revision of the Tyrini of the Oriental region. II. Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae). Annales de la Societe Entomologique de France 38: 283 - 297." type="journal article" year="2002">Hlaváč, 2002</bibRefCitation>
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(
<figureCitation id="13772A4B1951FFFBFD82FC2FFD1593BF" box="[555,647,990,1009]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStart-6="Fig" captionStart-7="Figs" captionStartId-0="2.[151,190,1554,1573]" captionStartId-1="3.[113,152,875,894]" captionStartId-2="5.[113,152,747,766]" captionStartId-3="5.[113,152,1548,1567]" captionStartId-4="7.[113,152,798,817]" captionStartId-5="8.[151,190,1520,1539]" captionStartId-6="10.[151,182,711,730]" captionStartId-7="11.[113,152,1518,1537]" captionTargetBox-0="[237,1039,224,1535]" captionTargetBox-1="[227,980,232,848]" captionTargetBox-2="[130,1075,219,703]" captionTargetBox-3="[120,1084,883,1493]" captionTargetBox-4="[123,1071,219,775]" captionTargetBox-5="[158,1125,525,1484]" captionTargetBox-6="[274,1005,219,693]" captionTargetBox-7="[114,1089,395,1491]" captionTargetId-0="figure-10@2.[227,1048,217,901]" captionTargetId-1="figure-285@3.[213,989,218,859]" captionTargetId-2="figure-14@5.[113,1088,217,708]" captionTargetId-3="figure-16@5.[113,1088,875,1513]" captionTargetId-4="figure-298@7.[113,1088,219,785]" captionTargetId-5="figure-9@8.[151,1126,514,1496]" captionTargetId-6="figure-279@10.[274,1005,218,694]" captionTargetId-7="figure-14@11.[114,1089,395,1491]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="Figs. 1–4. Male habitus of Linan spp. 1 – L. cardialis Hlaváč, 2002; 2 – L. chinensis (Löbl, 1964); 3 – L. hainanicus Hlaváč, 2002; 4 – L. inornatus sp. nov." captionText-1="Figs. 5–6. Dorsal habitus of Linan spp. 5 – L. megalobus sp. nov., male; 6 – L. sp. indet., female." captionText-2="Figs. 7–11.Antennal segments VIII–XI of Linan spp. in male. 7 – L. cardialis Hlaváč, 2002; 8 – L. chinensis (Löbl, 1964); 9 – L. hainanicus Hlaváč, 2002; 10 – L. inornatus sp. nov.; 11 – L. megalobus sp. nov. Scales: 0.3 mm." captionText-3="Figs. 12–21. Pronotum and metaventral horn-like processes of Linan spp. in male. 12, 17 – L. cardialis Hlaváč, 2002; 13, 18 – L. chinensis (Löbl, 1964); 14, 19 – L. hainanicus Hlaváč, 2002; 15, 20 – L. inornatus sp. nov.; 16, 21 – L. megalobus sp. nov. Scales: 0.2 mm." captionText-4="Figs. 22–26. Details of legs of Linan spp. in male. 22 – protrochanter and profemur of L. cardialis Hlaváč, 2002; 23 – mesotrochanter and mesofemur of L. cardialis; 24 – protibia of L. cardialis; 25 – protibia of L. hainanicus Hlaváč, 2002; 26 – protibia of L. megalobus sp. nov. Scales: 0.3 mm." captionText-5="Figs. 27–41. Aedeagus and sternite IX of Linan spp. 27, 29, 31, 33, 35 – aedeagus, in dorsal view; 28, 30, 32, 34, 36 – aedeagus, in lateral view; 37–41 – sternite IX of Linan spp. 27, 28, 37 – L. cardialis Hlaváč, 2002; 29–30, 38 – L. chinensis (Löbl, 1964); 31–32, 39 – L. hainanicus Hlaváč, 2002; 33–34, 40 – L. inornatus sp. nov.; 35–36, 41 – L. megalobus sp. nov. Scales: 0.2 mm (a, b, c, d, e); 0.1 mm (f, g, h, i, j)." captionText-6="Fig. 42. Distribution of the genus Linan Hlaváč, 2002. Black squares – L. cardialis Hlaváč, 2002; white triangle – L. chinensis (Löbl, 1964); white squares – L. hainanicus Hlaváč, 2002; white circle – L. inornatus sp. nov.; black circle – L. megalobus sp. nov.; black triangle – L. sp. indet." captionText-7="Figs. 43–48. Habitats of the genus Linan Hlaváč, 2002. 43 – Nabanhe Nature Reserve, Yunnan (L. cardialis Hlaváč, 2002); 44 – Longwangshan Mountain, Zhejiang (L. chinensis (Löbl, 1964)); 45 – Bawangling Nature Reserve, Hainan (L. hainanicus Hlaváč, 2002); 46 – Tianzhushan Mountain, Anhui (L. inornatus sp. nov.); 47 – Kuankuoshui N. R., Guizhou (L. megalobus sp. nov.); 48 – Sanqingshan Mountain, Jiangxi (L. sp. indet.)." figureDoi-0="http://doi.org/10.5281/zenodo.5327200" figureDoi-1="http://doi.org/10.5281/zenodo.5327206" figureDoi-2="http://doi.org/10.5281/zenodo.5327208" figureDoi-3="http://doi.org/10.5281/zenodo.5327212" figureDoi-4="http://doi.org/10.5281/zenodo.5327214" figureDoi-5="http://doi.org/10.5281/zenodo.5327218" figureDoi-6="http://doi.org/10.5281/zenodo.5327222" figureDoi-7="http://doi.org/10.5281/zenodo.5327226" httpUri-0="https://zenodo.org/record/5327200/files/figure.png" httpUri-1="https://zenodo.org/record/5327206/files/figure.png" httpUri-2="https://zenodo.org/record/5327208/files/figure.png" httpUri-3="https://zenodo.org/record/5327212/files/figure.png" httpUri-4="https://zenodo.org/record/5327214/files/figure.png" httpUri-5="https://zenodo.org/record/5327218/files/figure.png" httpUri-6="https://zenodo.org/record/5327222/files/figure.png" httpUri-7="https://zenodo.org/record/5327226/files/figure.png" pageId="1" pageNumber="124">Figs. 1–48</figureCitation>
)
</paragraph>
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<paragraph id="8BF336CE1951FFFBFFD8FBF0FDF1945D" blockId="1.[113,653,990,1044]" box="[113,611,1024,1044]" pageId="1" pageNumber="124">
<treatmentCitationGroup id="AB5C11E01951FFFBFFD8FBF0FDF1945D" box="[113,611,1024,1044]" pageId="1" pageNumber="124">
<treatmentCitation id="0AED10DF1951FFFBFFD8FBF0FEDB945D" author="HLAVAC P." box="[113,329,1024,1044]" page="294" pageId="1" pageNumber="124" year="2002">
<taxonomicName id="4C4C4D4D1951FFFBFFD8FBF0FEDB945D" ID-CoL="8VYXL" authority="Hlavac, 2002: 294" authorityName="Hlavac" authorityPageNumber="294" authorityYear="2002" box="[113,329,1024,1044]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B938EADC1951FFFBFFD8FBF0FF31945D" box="[113,163,1025,1043]" italics="true" pageId="1" pageNumber="124">Linan</emphasis>
<bibRefCitation id="EFDD4B3F1951FFFBFF00FBF1FEDB945D" author="HLAVAC P." box="[169,329,1024,1044]" pageId="1" pageNumber="124" pagination="283 - 297" refId="ref5971" refString="HLAVAC P. 2002: A taxonomic revision of the Tyrini of the Oriental region. II. Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae). Annales de la Societe Entomologique de France 38: 283 - 297." type="journal article" year="2002">Hlaváč, 2002: 294</bibRefCitation>
</taxonomicName>
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.
<treatmentCitation id="0AED10DF1951FFFBFEF9FBF0FDCD945D" author="HLAVAC P. &amp; CHANDLER D. S." box="[336,607,1024,1044]" page="110" pageId="1" pageNumber="124" year="2005">
<bibRefCitation id="EFDD4B3F1951FFFBFEF9FBF0FDCD945D" author="HLAVAC P. &amp; CHANDLER D. S." box="[336,607,1024,1044]" pageId="1" pageNumber="124" pagination="81 - 139" refId="ref6020" refString="HLAVAC P. &amp; CHANDLER D. S. 2005: World catalog of the species of Tyrini with a key to the genera (Coleoptera: Staphylindae: Pselaphinae). Folia Heyrovskyana 13: 81 - 139." type="journal article" year="2005">HLAVÁČ &amp; CHANDLER 2005: 110</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BF336CE1951FFFBFFD8FBDDFCDD940D" blockId="1.[113,847,1067,1092]" box="[113,847,1067,1092]" pageId="1" pageNumber="124">
<emphasis id="B938EADC1951FFFBFFD8FBDDFE97940D" bold="true" box="[113,261,1067,1092]" pageId="1" pageNumber="124">
<typeStatus id="54F7886C1951FFFBFFD8FBDDFF3B940A" box="[113,169,1068,1092]" pageId="1" pageNumber="124">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C4C4D4D1951FFFBFEA5FBDCFDD5940D" ID-CoL="3SCW9" authority="Lobl, 1964" authorityName="Lobl" authorityYear="1964" box="[268,583,1067,1092]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B938EADC1951FFFBFEA5FBDCFE5A940A" box="[268,456,1069,1092]" italics="true" pageId="1" pageNumber="124">Lasinus chinensis</emphasis>
<bibRefCitation id="EFDD4B3F1951FFFBFE66FBDAFDD5940D" author="LOBL I." box="[463,583,1067,1091]" pageId="1" pageNumber="124" pagination="45 - 48" refId="ref6060" refString="LOBL I. 1964: Eine neue Art der Gattung Lasinus Sharp aus China (Col., Pselaphidae). Acta Rerum Naturalium Musei Nationalis Slovaci (Bratislava) 10: 45 - 48." type="journal article" year="1964">Löbl, 1964</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
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<subSubSection id="C35665451951FFFBFFD8FBABFB87955D" pageId="1" pageNumber="124" type="diagnosis">
<paragraph id="8BF336CE1951FFFBFFD8FBABFB87955D" blockId="1.[113,1089,1114,1299]" pageId="1" pageNumber="124">
<emphasis id="B938EADC1951FFFBFFD8FBABFF74943C" bold="true" box="[113,230,1114,1138]" pageId="1" pageNumber="124">Diagnosis.</emphasis>
The genus
<taxonomicName id="4C4C4D4D1951FFFBFEC8FBADFE0D943D" authorityName="Hlavac" authorityYear="2002" box="[353,415,1116,1139]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B938EADC1951FFFBFEC8FBADFE0D943D" box="[353,415,1116,1139]" italics="true" pageId="1" pageNumber="124">Linan</emphasis>
</taxonomicName>
could be readily separated from allied genera within the
<taxonomicName id="4C4C4D4D1951FFFBFC55FBADFF2F94DD" authorityName="Westwood" authorityYear="1870" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B938EADC1951FFFBFC55FBADFF2F94DD" italics="true" pageId="1" pageNumber="124">Pselaphodes</emphasis>
</taxonomicName>
complex by the following combination of characters: Head with vertexal and frontal foveae indistinct; maxillary palpi with segments III–IV asymmetrical, roundly expanded or with external protuberance; pronotum with small but well-defined median and lateral antebasal foveae; head, pronotum and legs densely and roughly punctate; median metaventral fovea present; metaventral horn-like processes present in male; elytra simple without carinae.
</paragraph>
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<paragraph id="8BF336CE1951FFFBFFD8FAD8FC8D952C" blockId="1.[113,1089,1321,1602]" pageId="1" pageNumber="124">
<emphasis id="B938EADC1951FFFBFFD8FAD8FE87950F" bold="true" box="[113,277,1321,1345]" pageId="1" pageNumber="124">Redescription.</emphasis>
Length
<quantity id="4CB49B2B1951FFFBFEC0FADBFD96950F" box="[361,516,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="2.99" metricValueMax="3.63" metricValueMin="2.35" pageId="1" pageNumber="124" unit="mm" value="2.99" valueMax="3.63" valueMin="2.35">2.35–3.63 mm</quantity>
. Head, antennae, pronotum, abdomen and legs reddishbrown, elytra lighter, maxillary palpi and tarsi yellowish-brown.
</paragraph>
<paragraph id="8BF336CE1951FFFBFF39FA9BFC6295AC" blockId="1.[113,1089,1321,1602]" pageId="1" pageNumber="124">Head nearly triangular, roughly punctate, sparsely covered with pubescence; frontal fovea and pair of vertexal foveae very indistinct; antennae elongate, pubescent; scape roughly punctate, about as long as segments II–V combined; antennal club three-segmented; antennomeres IX–X strongly modified or not in male, not modified and weekly clubbed in female.</paragraph>
<paragraph id="8BF336CE1951FFF9FF39FA1BFD3D9467" blockId="1.[113,1089,1321,1602]" lastBlockId="3.[113,1089,945,1194]" lastPageId="3" lastPageNumber="126" pageId="1" pageNumber="124">
Pronotum about as long as wide, roughly punctate, pubescent. Width of the elytra wider than long, pubescent; each elytron with two basal foveae; discal stria reaching apical twothirds of elytral length. Legs elongate and slender with femora roughly punctate, usually simple without spine (except for
<taxonomicName id="4C4C4D4D1953FFF9FE7FFC43FDC39387" authorityName="Hlavac" authorityYear="2002" box="[470,593,946,969]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="cardialis">
<emphasis id="B938EADC1953FFF9FE7FFC43FE779387" box="[470,485,946,969]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FE5AFC43FDC39387" box="[499,593,946,969]" italics="true" pageId="3" pageNumber="126">cardialis</emphasis>
</taxonomicName>
); protibiae in some species expanded mesally at apex as a spur or lobe in male; tarsi three-segmented. Venter with metaventrite depressed, smooth; median metaventral horn-like processes paired and well-defined in male, various in length and shape; median metaventral foveae present.
</paragraph>
<caption id="DF3366461952FFF8FF3EF9E3FE77960E" ID-DOI="http://doi.org/10.5281/zenodo.5327200" ID-Zenodo-Dep="5327200" httpUri="https://zenodo.org/record/5327200/files/figure.png" pageId="2" pageNumber="125" startId="2.[151,190,1554,1573]" targetBox="[237,1039,224,1535]" targetPageId="2">
<paragraph id="8BF336CE1952FFF8FF3EF9E3FE77960E" blockId="2.[151,1127,1554,1600]" pageId="2" pageNumber="125">
Figs. 1–4. Male habitus of
<taxonomicName id="4C4C4D4D1952FFF8FED0F9E2FE70966B" box="[377,482,1554,1573]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B938EADC1952FFF8FED0F9E2FE39966B" box="[377,427,1555,1573]" italics="true" pageId="2" pageNumber="125">Linan</emphasis>
spp. 1
</taxonomicName>
–
<taxonomicName id="4C4C4D4D1952FFF8FE5CF9E2FD5C966B" authority="Hlavac, 2002" authorityName="Hlavac" authorityYear="2002" box="[501,718,1554,1573]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="cardialis">
<emphasis id="B938EADC1952FFF8FE5CF9E2FD93966B" box="[501,513,1555,1573]" italics="true" pageId="2" pageNumber="125">L</emphasis>
.
<emphasis id="B938EADC1952FFF8FDA2F9E2FDC4966B" box="[523,598,1555,1573]" italics="true" pageId="2" pageNumber="125">cardialis</emphasis>
<bibRefCitation id="EFDD4B3F1952FFF8FDF2F9E3FD5C966B" author="HLAVAC P." box="[603,718,1554,1573]" pageId="2" pageNumber="125" pagination="283 - 297" refId="ref5971" refString="HLAVAC P. 2002: A taxonomic revision of the Tyrini of the Oriental region. II. Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae). Annales de la Societe Entomologique de France 38: 283 - 297." type="journal article" year="2002">Hlaváč, 2002</bibRefCitation>
</taxonomicName>
; 2 –
<taxonomicName id="4C4C4D4D1952FFF8FD5CF9E2FC5E966B" authority="(Lobl, 1964)" baseAuthorityName="Lobl" baseAuthorityYear="1964" box="[757,972,1554,1573]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B938EADC1952FFF8FD5CF9E2FC93966B" box="[757,769,1555,1573]" italics="true" pageId="2" pageNumber="125">L</emphasis>
.
<emphasis id="B938EADC1952FFF8FCA2F9E2FCC8966B" box="[779,858,1555,1573]" italics="true" pageId="2" pageNumber="125">chinensis</emphasis>
(
<bibRefCitation id="EFDD4B3F1952FFF8FCCCF9E3FC54966B" author="LOBL I." box="[869,966,1554,1573]" pageId="2" pageNumber="125" pagination="45 - 48" refId="ref6060" refString="LOBL I. 1964: Eine neue Art der Gattung Lasinus Sharp aus China (Col., Pselaphidae). Acta Rerum Naturalium Musei Nationalis Slovaci (Bratislava) 10: 45 - 48." type="journal article" year="1964">Löbl, 1964</bibRefCitation>
)
</taxonomicName>
; 3 –
<taxonomicName id="4C4C4D4D1952FFF8FC5AF9E2FE999671" authority="Hlavac, 2002" authorityName="Hlavac" authorityYear="2002" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="hainanicus">
<emphasis id="B938EADC1952FFF8FC5AF9E2FC6D966B" box="[1011,1023,1555,1573]" italics="true" pageId="2" pageNumber="125">L</emphasis>
.
<emphasis id="B938EADC1952FFF8FBA0F9E2FBF5966B" box="[1033,1127,1555,1573]" italics="true" pageId="2" pageNumber="125">hainanicus</emphasis>
<bibRefCitation id="EFDD4B3F1952FFF8FF3EF9DDFE999671" author="HLAVAC P." box="[151,267,1580,1600]" pageId="2" pageNumber="125" pagination="283 - 297" refId="ref5971" refString="HLAVAC P. 2002: A taxonomic revision of the Tyrini of the Oriental region. II. Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae). Annales de la Societe Entomologique de France 38: 283 - 297." type="journal article" year="2002">Hlaváč, 2002</bibRefCitation>
</taxonomicName>
; 4 –
<taxonomicName id="4C4C4D4D1952FFF8FE9DF9DCFE0F9671" authority="Yin &amp; Li &amp; Zhao, 2011" authorityName="Yin &amp; Li &amp; Zhao" authorityYear="2011" box="[308,413,1581,1600]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="inornatus" status="sp. nov.">
<emphasis id="B938EADC1952FFF8FE9DF9DCFED29671" box="[308,320,1581,1599]" italics="true" pageId="2" pageNumber="125">L</emphasis>
.
<emphasis id="B938EADC1952FFF8FEE2F9DCFE0F9671" box="[331,413,1581,1599]" italics="true" pageId="2" pageNumber="125">inornatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57A71952FFF8FE0BF9DCFE77960E" box="[418,485,1581,1600]" pageId="2" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF3366461953FFF9FFD8FC9AFC3E9330" ID-DOI="http://doi.org/10.5281/zenodo.5327206" ID-Zenodo-Dep="5327206" box="[113,940,875,895]" httpUri="https://zenodo.org/record/5327206/files/figure.png" pageId="3" pageNumber="126" startId="3.[113,152,875,894]" targetBox="[227,980,232,848]" targetPageId="3">
<paragraph id="8BF336CE1953FFF9FFD8FC9AFC3E9330" blockId="3.[113,940,875,895]" box="[113,940,875,895]" pageId="3" pageNumber="126">
Figs. 5–6. Dorsal habitus of
<taxonomicName id="4C4C4D4D1953FFF9FECDFC9DFE5C9330" box="[356,462,875,895]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="undefined-5">
<emphasis id="B938EADC1953FFF9FECDFC9DFE049330" box="[356,406,876,894]" italics="true" pageId="3" pageNumber="126">Linan</emphasis>
spp. 5
</taxonomicName>
–
<taxonomicName id="4C4C4D4D1953FFF9FE4AFC9DFDC49330" authority="Yin &amp; Li &amp; Zhao, 2011" authorityName="Yin &amp; Li &amp; Zhao" authorityYear="2011" box="[483,598,876,895]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="megalobus" status="sp. nov.">
<emphasis id="B938EADC1953FFF9FE4AFC9DFE7D9330" box="[483,495,876,894]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FE53FC9DFDC49330" box="[506,598,876,894]" italics="true" pageId="3" pageNumber="126">megalobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57A71953FFF9FDF2FC9DFD329331" box="[603,672,876,895]" pageId="3" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
, male; 6 –
<emphasis id="B938EADC1953FFF9FD55FC9DFC9A9330" box="[764,776,876,894]" italics="true" pageId="3" pageNumber="126">L</emphasis>
. sp. indet., female.
</paragraph>
</caption>
<paragraph id="8BF336CE1953FFF9FF39FBC0FEBE94E7" blockId="3.[113,1089,945,1194]" pageId="3" pageNumber="126">
Abdomen with first visible tergite (morphological tergite IV) largest, discal ridges present or absent; following segments successively shorter and narrower; paratergites well-defined. Aedeagus with median lobe large; parameres paired; endophallus provided with various
<typeStatus id="54F7886C1953FFF9FFD8FB63FF3B94E4" box="[113,169,1170,1194]" pageId="3" pageNumber="126">types</typeStatus>
of sclerites.
</paragraph>
</subSubSection>
<subSubSection id="C35665451953FFF9FFD8FB31FCD79516" pageId="3" pageNumber="126" type="discussion">
<paragraph id="8BF336CE1953FFF9FFD8FB31FCD79516" blockId="3.[113,1089,1216,1369]" pageId="3" pageNumber="126">
<emphasis id="B938EADC1953FFF9FFD8FB31FF629496" bold="true" box="[113,240,1216,1240]" pageId="3" pageNumber="126">Discussion.</emphasis>
Species of the genus
<taxonomicName id="4C4C4D4D1953FFF9FE77FB33FD8E9497" authorityName="Hlavac" authorityYear="2002" box="[478,540,1218,1241]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B938EADC1953FFF9FE77FB33FD8E9497" box="[478,540,1218,1241]" italics="true" pageId="3" pageNumber="126">Linan</emphasis>
</taxonomicName>
can be classified into two groups (here proposed): the
<taxonomicName id="4C4C4D4D1953FFF9FF3EFB13FE8294B7" authorityName="Hlavac" authorityYear="2002" box="[151,272,1249,1273]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="cardialis">
<emphasis id="B938EADC1953FFF9FF3EFB13FF3494B7" box="[151,166,1250,1273]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FF1BFB13FE8294B7" box="[178,272,1250,1273]" italics="true" pageId="3" pageNumber="126">cardialis</emphasis>
</taxonomicName>
species-group, with strongly modified antennal segments IX–X and modified protibiae at apex in male, including
<taxonomicName id="4C4C4D4D1953FFF9FE5EFAF3FDE09557" authorityName="Hlavac" authorityYear="2002" box="[503,626,1281,1305]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="cardialis">
<emphasis id="B938EADC1953FFF9FE5EFAF3FD949557" box="[503,518,1282,1305]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FDBDFAF3FDE09557" box="[532,626,1282,1305]" italics="true" pageId="3" pageNumber="126">cardialis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D4D1953FFF9FD29FAF3FC879557" authorityName="Hlavac" authorityYear="2002" box="[640,789,1281,1305]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="hainanicus">
<emphasis id="B938EADC1953FFF9FD29FAF3FD1D9557" box="[640,655,1282,1305]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FD34FAF3FC879557" box="[669,789,1282,1305]" italics="true" pageId="3" pageNumber="126">hainanicus</emphasis>
</taxonomicName>
<emphasis id="B938EADC1953FFF9FCBCFAF3FC8B9557" box="[789,793,1282,1305]" italics="true" pageId="3" pageNumber="126">,</emphasis>
and
<taxonomicName id="4C4C4D4D1953FFF9FCE6FAF3FC729557" authority="Yin &amp; Li &amp; Zhao, 2011" authorityName="Yin &amp; Li &amp; Zhao" authorityYear="2011" box="[847,992,1281,1305]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="megalobus" status="sp. nov.">
<emphasis id="B938EADC1953FFF9FCE6FAF3FCCC9557" box="[847,862,1282,1305]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FCC5FAF3FC729557" box="[876,992,1282,1305]" italics="true" pageId="3" pageNumber="126">megalobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57A71953FFF9FC4EFAF0FBAF9556" box="[999,1085,1281,1304]" pageId="3" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
, and the
<taxonomicName id="4C4C4D4D1953FFF9FF6BFAD3FEAD9577" baseAuthorityName="Lobl" baseAuthorityYear="1964" box="[194,319,1314,1337]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B938EADC1953FFF9FF6BFAD3FEAD9577" box="[194,319,1314,1337]" italics="true" pageId="3" pageNumber="126">L. chinensis</emphasis>
</taxonomicName>
species-group, lacking obvious sexual modifications of the male antennal club and protibiae, containing
<taxonomicName id="4C4C4D4D1953FFF9FE1FFAB3FDA49517" baseAuthorityName="Lobl" baseAuthorityYear="1964" box="[438,566,1345,1369]" class="Insecta" family="Staphylinidae" genus="Lasinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B938EADC1953FFF9FE1FFAB3FE579517" box="[438,453,1346,1369]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FE7AFAB3FDA49517" box="[467,566,1346,1369]" italics="true" pageId="3" pageNumber="126">chinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D4D1953FFF9FDC3FAB3FD7E9517" authority="Yin &amp; Li &amp; Zhao, 2011" authorityName="Yin &amp; Li &amp; Zhao" authorityYear="2011" box="[618,748,1345,1369]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="inornatus" status="sp. nov.">
<emphasis id="B938EADC1953FFF9FDC3FAB3FDEB9517" box="[618,633,1346,1369]" italics="true" pageId="3" pageNumber="126">L</emphasis>
.
<emphasis id="B938EADC1953FFF9FD2FFAB3FD7E9517" box="[646,748,1346,1369]" italics="true" pageId="3" pageNumber="126">inornatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57A71953FFF9FD5AFAB0FCD79516" box="[755,837,1345,1368]" pageId="3" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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