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<document id="A892259B3E8EA5638EA22BA0F98383E6" ID-DOI="10.5281/zenodo.282509" ID-GBIF-Dataset="fa676e46-f615-4a54-91bf-d097ea18f64c" ID-ISSN="1175-5326" ID-Zenodo-Dep="282509" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460541281962" checkinUser="plazi" docAuthor="Paredes-León, Ricardo, Klompen, Hans &amp; Pérez, Tila M." docDate="2012" docId="03E587D3FB756A73CCCCFCB2FAB5F9D6" docLanguage="en" docName="zt03510p040.pdf" docOrigin="Zootaxa 3510" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Geckobiella pyriformis Newell and Ryckman, new combination" docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="FFDCFFABFB7F6A79CC5BFF80FF93FF8A" masterDocTitle="Systematic revision of the genera Geckobiella Hirst, 1917 and Hirstiella Berlese, 1920 (Acari: Prostigmata: Pterygosomatidae) with description of a new genus for American species parasites on geckos formerly placed in Hirstiella" masterLastPageNumber="40" masterPageNumber="1" pageNumber="11" updateTime="1698317095416" updateUser="plazi">
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<mods:title id="CDE0310C7743EC98FBE54A72916C70E1">Systematic revision of the genera Geckobiella Hirst, 1917 and Hirstiella Berlese, 1920 (Acari: Prostigmata: Pterygosomatidae) with description of a new genus for American species parasites on geckos formerly placed in Hirstiella</mods:title>
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<mods:namePart id="4CF5A9F57F9F2C6F2EC6F0657618EC65">Paredes-León, Ricardo</mods:namePart>
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<mods:namePart id="A4E55BAB12CBA54FFB23A1C4ABA0C5A6">Klompen, Hans</mods:namePart>
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<mods:namePart id="FD7459CDF4D61C9F8E84541D78167535">Pérez, Tila M.</mods:namePart>
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<treatment id="03E587D3FB756A73CCCCFCB2FAB5F9D6" ID-DOI="http://doi.org/10.5281/zenodo.6178817" ID-GBIF-Taxon="119528331" ID-Zenodo-Dep="6178817" LSID="urn:lsid:plazi:treatment:03E587D3FB756A73CCCCFCB2FAB5F9D6" httpUri="http://treatment.plazi.org/id/03E587D3FB756A73CCCCFCB2FAB5F9D6" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="C356654EFB756A73CCCCFCB2FC3EFCC6" box="[151,941,818,844]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BF336C5FB756A73CCCCFCB2FC3EFCC6" blockId="10.[151,941,818,844]" box="[151,941,818,844]" pageId="10" pageNumber="11">
<heading id="D0BB81A9FB756A73CCCCFCB2FC3EFCC6" bold="true" box="[151,941,818,844]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B938EAD7FB756A73CCCCFCB2FC3EFCC6" bold="true" box="[151,941,818,844]" pageId="10" pageNumber="11">
<taxonomicName id="4C4C4D46FB756A73CCCCFCB2FD5CFCC6" authority="Newell and Ryckman" authorityName="Newell and Ryckman" box="[151,719,818,844]" class="Arachnida" family="Pterygosomatidae" genus="Geckobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pyriformis" status="comb. nov.">
<emphasis id="B938EAD7FB756A73CCCCFCB2FE39FCC6" bold="true" box="[151,426,818,844]" italics="true" pageId="10" pageNumber="11">Geckobiella pyriformis</emphasis>
(Newell and Ryckman)
</taxonomicName>
<taxonomicNameLabel id="A20B57ACFB756A73CE8CFCB2FC3EFCC6" box="[727,941,818,844]" pageId="10" pageNumber="11">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356654EFB756A73CCCCFCF9FD51FC05" box="[151,706,889,911]" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BF336C5FB756A73CCCCFCF9FD51FC05" blockId="10.[151,706,889,911]" box="[151,706,889,911]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB5C11EBFB756A73CCCCFCF9FD51FC05" box="[151,706,889,911]" pageId="10" pageNumber="11">
<taxonomicName id="4C4C4D46FB756A73CCCCFCF9FD2DFC05" authority="Newell and Ryckman, 1964: 164" authorityName="Newell and Ryckman" authorityPageNumber="164" authorityYear="1964" box="[151,702,889,911]" class="Arachnida" family="Tetranychidae" genus="Hirstiella" higherTaxonomySource="CoL" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="B938EAD7FB756A73CCCCFCF9FEFBFC05" box="[151,360,889,911]" italics="true" pageId="10" pageNumber="11">Hirstiella pyriformis</emphasis>
<treatmentCitation id="0AED10D4FB756A73CD35FCF9FD2DFC05" author="Newell" box="[366,702,889,911]" page="164" pageId="10" pageNumber="11" year="1964">
<bibRefCitation id="EFDD4B34FB756A73CD35FCF9FD1FFC05" author="Newell" box="[366,652,889,911]" pageId="10" pageNumber="11" refString="Newell, I. M. &amp; R. E. Ryckman. (1964) Hirstiella pyriformis sp. n. (Acari, Pterygosomidae), a new parasite of lizards from Baja California. Journal of Parasitology, 50, 163 - 171." type="journal article" year="1964">Newell and Ryckman, 1964</bibRefCitation>
: 164
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C356654EFB756A73CCCCFC3BFD7AFC7D" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BF336C5FB756A73CCCCFC3BFD7AFC7D" blockId="10.[151,1436,955,1628]" pageId="10" pageNumber="11">
<emphasis id="B938EAD7FB756A73CCCCFC3BFEE4FC5E" bold="true" box="[151,375,955,980]" pageId="10" pageNumber="11">Diagnosis. Female.</emphasis>
Prodorsal shield pyriform (anterior margin not sharply defined) with 1 pair of setae (
<emphasis id="B938EAD7FB756A73C96DFC3DFAC6FC5E" box="[1334,1365,957,980]" italics="true" pageId="10" pageNumber="11">sci</emphasis>
) (
<figureCitation id="13772A40FB756A73C935FC3CFF35FC7D" captionStart="FIGURE 2" captionStartId="12.[151,250,1955,1978]" captionTargetBox="[375,1198,222,1909]" captionTargetId="figure@12.[351,1235,195,1934]" captionTargetPageId="12" captionText="FIGURE 2. Prodorsal shields of the species of Geckobiella. A, G. trombidiiformis; B, G. p e l a e z i; C, G. boneti; D, G. bakeri; E, G. pyriformis; F, G. javieri; G, G. diolii; H, G. donnae sp. nov. Scale bar 100 µm." httpUri="https://zenodo.org/record/282511/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
E). Tarsus III with residual alveolus or minute pore.
</paragraph>
</subSubSection>
<subSubSection id="C356654EFB756A73CC9CFB84FDEBFBEE" pageId="10" pageNumber="11" type="description">
<paragraph id="8BF336C5FB756A73CC9CFB84FDEBFBEE" blockId="10.[151,1436,955,1628]" pageId="10" pageNumber="11">
<emphasis id="B938EAD7FB756A73CC9CFB84FE98FB96" bold="true" box="[199,267,1028,1052]" pageId="10" pageNumber="11">Male.</emphasis>
Well-defined prodorsal shield absent. Trochanter IV without setae (1 seta present in female); genua IV with a large, hollow ensiform seta (
<emphasis id="B938EAD7FB756A73CE73FBA8FDADFBB5" box="[552,574,1064,1087]" italics="true" pageId="10" pageNumber="11">v</emphasis>
) and tibia with 1 solid spine-like seta (
<emphasis id="B938EAD7FB756A73CFA2FBA8FB86FBB5" box="[1017,1045,1064,1087]" italics="true" pageId="10" pageNumber="11">v”</emphasis>
), both covered with short, spinose ornamentation (
<bibRefCitation id="EFDD4B34FB756A73CD1CFBCCFDF8FBEE" author="Newell" box="[327,619,1100,1124]" pageId="10" pageNumber="11" refString="Newell, I. M. &amp; R. E. Ryckman. (1964) Hirstiella pyriformis sp. n. (Acari, Pterygosomidae), a new parasite of lizards from Baja California. Journal of Parasitology, 50, 163 - 171." type="journal article" year="1964">Newell &amp; Ryckman 1964</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C356654EFB756A73CC9CFBF0FC2EFA0E" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BF336C5FB756A73CC9CFBF0FF6EFB26" blockId="10.[151,1436,955,1628]" pageId="10" pageNumber="11">
<emphasis id="B938EAD7FB756A73CC9CFBF0FE95FB02" bold="true" box="[199,262,1136,1160]" pageId="10" pageNumber="11">
<typeStatus id="54F78867FB756A73CC9CFBF0FE93FB02" box="[199,256,1136,1160]" pageId="10" pageNumber="11">Type</typeStatus>
.
</emphasis>
<typeStatus id="54F78867FB756A73CD43FBEFFE3FFB0D" box="[280,428,1135,1159]" pageId="10" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
depositary unknown; female, male and deutonymphal
<typeStatus id="54F78867FB756A73C815FBEFFB7BFB0D" box="[1102,1256,1135,1159]" pageId="10" pageNumber="11" type="paratype">PARATYPES</typeStatus>
(
<collectionCode id="ED5DAE00FB756A73C8A4FBEFFAF7FB0D" box="[1279,1380,1135,1159]" pageId="10" pageNumber="11">USNMC</collectionCode>
and
<collectionCode id="ED5DAE00FB756A73CCCCFB14FF61FB26" LSID="urn:lsid:biocol.org:col:34925" box="[151,242,1172,1196]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="10" pageNumber="11">AMNH</collectionCode>
).
</paragraph>
<paragraph id="8BF336C5FB756A73CC9CFB38FBADFB45" blockId="10.[151,1436,955,1628]" box="[199,1086,1207,1232]" pageId="10" pageNumber="11">
<emphasis id="B938EAD7FB756A73CC9CFB38FEF6FB5A" bold="true" box="[199,357,1207,1232]" pageId="10" pageNumber="11">
<typeStatus id="54F78867FB756A73CC9CFB38FE91FB5A" box="[199,258,1208,1232]" pageId="10" pageNumber="11">Type</typeStatus>
locality.
</emphasis>
San Esteban Island, Gulf of California, Baja California,
<collectingCountry id="F35B7655FB756A73CFBAFB37FBA9FB45" box="[993,1082,1207,1231]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="8BF336C5FB756A73CC9CFB5CFD0FFB7E" blockId="10.[151,1436,955,1628]" box="[199,668,1244,1268]" pageId="10" pageNumber="11">
<emphasis id="B938EAD7FB756A73CC9CFB5CFED2FB7E" bold="true" box="[199,321,1244,1268]" pageId="10" pageNumber="11">
<typeStatus id="54F78867FB756A73CC9CFB5CFE91FB7E" box="[199,258,1244,1268]" pageId="10" pageNumber="11">Type</typeStatus>
host.
</emphasis>
<taxonomicName id="4C4C4D46FB756A73CD13FB5CFD0FFB7E" authority="Dickerson." authorityName="Dickerson." box="[328,668,1244,1268]" class="Reptilia" family="Iguanidae" genus="Sauromalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="varius">
<emphasis id="B938EAD7FB756A73CD13FB5CFD89FB7E" box="[328,538,1244,1268]" italics="true" pageId="10" pageNumber="11">Sauromalus varius</emphasis>
Dickerson.
</taxonomicName>
</paragraph>
<paragraph id="8BF336C5FB756A73CC9CFB7FFC2EFA0E" blockId="10.[151,1436,955,1628]" pageId="10" pageNumber="11">
<emphasis id="B938EAD7FB756A73CC9CFB7FFE23FA92" bold="true" box="[199,432,1279,1304]" pageId="10" pageNumber="11">Material examined.</emphasis>
<specimenCount id="9D4AFD4CFB756A73CDECFB7FFDBAFA9D" box="[439,553,1279,1303]" pageId="10" pageNumber="11" type="female">15 female</specimenCount>
,
<specimenCount id="9D4AFD4CFB756A73CE6FFB7FFD11FA9D" box="[564,642,1279,1303]" pageId="10" pageNumber="11" type="male">4 male</specimenCount>
, 1 deutonymphal
<typeStatus id="54F78867FB756A73CF11FB7FFC77FA9D" box="[842,996,1279,1303]" pageId="10" pageNumber="11" type="paratype">PARATYPES</typeStatus>
ex
<taxonomicName id="4C4C4D46FB756A73C857FA80FB4DFA92" box="[1036,1246,1280,1304]" class="Reptilia" family="Iguanidae" genus="Sauromalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="varius">
<emphasis id="B938EAD7FB756A73C857FA80FB4DFA92" box="[1036,1246,1280,1304]" italics="true" pageId="10" pageNumber="11">Sauromalus varius</emphasis>
</taxonomicName>
,
<collectingCountry id="F35B7655FB756A73C8B7FB7FFACEFA9D" box="[1260,1373,1279,1303]" name="Mexico" pageId="10" pageNumber="11">MEXICO</collectingCountry>
, Baja California, San Esteban Island,
<geoCoordinate id="EE785002FB756A73CDA2FAA4FDF8FAB6" box="[505,619,1316,1340]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="28.333334">28° 20 N</geoCoordinate>
, -
<geoCoordinate id="EE785002FB756A73CE25FAA4FC91FAB6" box="[638,770,1316,1340]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="-112.61667">112° 37 W</geoCoordinate>
,
<date id="FFF21005FB756A73CF56FAA4FC36FAB6" box="[781,933,1316,1341]" pageId="10" pageNumber="11" value="1963-05-23">23 May 1963</date>
, coll. R. E. Ryckman, A. E. Ryckman and C. P. Christianson (
<collectionCode id="ED5DAE00FB756A73CD0AFAC7FE28FAD5" box="[337,443,1351,1375]" pageId="10" pageNumber="11">USNMC</collectionCode>
).
<specimenCount id="9D4AFD4CFB756A73CD8BFAC7FDD2FAD5" box="[464,577,1351,1375]" pageId="10" pageNumber="11" type="female">5 females</specimenCount>
,
<specimenCount id="9D4AFD4CFB756A73CE15FAC7FD35FAD5" box="[590,678,1351,1375]" pageId="10" pageNumber="11" type="male">5 males</specimenCount>
ex
<taxonomicName id="4C4C4D46FB756A73CE89FAC8FC47FAEA" box="[722,980,1352,1376]" class="Reptilia" family="Iguanidae" genus="Ctenosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="hemilopha">
<emphasis id="B938EAD7FB756A73CE89FAC8FC47FAEA" box="[722,980,1352,1376]" italics="true" pageId="10" pageNumber="11">Ctenosaura hemilopha</emphasis>
</taxonomicName>
,
<collectingCountry id="F35B7655FB756A73CFBFFAC7FBC6FAD5" box="[996,1109,1351,1375]" name="Mexico" pageId="10" pageNumber="11">MEXICO</collectingCountry>
, Sonora, Hermosillo, Centro Ecológico de Sonora,
<date id="FFF21005FB756A73CDD5FAECFDA8FA0E" box="[398,571,1388,1412]" pageId="10" pageNumber="11" value="1986-03-11">11 March 1986</date>
, coll. G. Lara (
<collectionCode id="ED5DAE00FB756A73CE85FAECFCBCFA0E" box="[734,815,1388,1412]" pageId="10" pageNumber="11">CNAC</collectionCode>
00703342).
</paragraph>
</subSubSection>
<subSubSection id="C356654EFB756A73CC9CFA10FAB5F9D6" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BF336C5FB756A73CC9CFA10FAB5F9D6" blockId="10.[151,1436,955,1628]" pageId="10" pageNumber="11">
<emphasis id="B938EAD7FB756A73CC9CFA10FEA8FA22" bold="true" box="[199,315,1424,1448]" pageId="10" pageNumber="11">Remarks.</emphasis>
This species appears to be a specific ectoparasite of
<taxonomicName id="4C4C4D46FB756A73CFC2FA0FFB94FA2D" box="[921,1031,1423,1447]" class="Reptilia" family="Iguanidae" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Iguanidae</taxonomicName>
lizards (see
<tableCitation id="C6CE037EFB756A73C8C2FA0FFB63FA2D" box="[1177,1264,1423,1447]" captionStart="TABLE 2" captionStartId="6.[151,235,514,535]" captionTargetBox="[151,1424,605,1799]" captionTargetPageId="6" captionText="TABLE 2. Species included in Geckobiella with a summary of host and distribution data." httpUri="http://table.plazi.org/id/DF33664DFB796A7FCCCCFD82FBB0FD9D" pageId="10" pageNumber="11" tableUuid="DF33664DFB796A7FCCCCFD82FBB0FD9D">Table 2</tableCitation>
).
<bibRefCitation id="EFDD4B34FB756A73C95DFA0FFA0FFA2D" author="Baker" box="[1286,1436,1423,1447]" pageId="10" pageNumber="11" refString="Baker, A. (1998) A new species of Hirstiella Berlese (Acari: Pterygosomatidae) from captive rhinoceros iguanas, Cyclura cornuta Bonnaterre (Reptilia: Iguanidae). Systematic and Applied Acarology, 3, 183 - 192." type="journal article" year="1998">
<collectingCountry id="F35B7655FB756A73C95DFA0FFADAFA2D" box="[1286,1353,1423,1447]" name="United States Pacific Island Wildlife Refuges" pageId="10" pageNumber="11">Baker</collectingCountry>
(1998)
</bibRefCitation>
reports this species from
<collectingCountry id="F35B7655FB756A73CDE2FA34FE62FA46" box="[441,497,1460,1484]" name="United States of America" pageId="10" pageNumber="11">USA</collectingCountry>
based on
<typeStatus id="54F78867FB756A73CE32FA35FD47FA47" box="[617,724,1461,1485]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
deposited at the Natural History Museum, London, but in the original description
<bibRefCitation id="EFDD4B34FB756A73CD20FA57FD52FA65" author="Newell" box="[379,705,1495,1519]" pageId="10" pageNumber="11" refString="Newell, I. M. &amp; R. E. Ryckman. (1964) Hirstiella pyriformis sp. n. (Acari, Pterygosomidae), a new parasite of lizards from Baja California. Journal of Parasitology, 50, 163 - 171." type="journal article" year="1964">Newell and Ryckman (1964)</bibRefCitation>
mentioned this species only from Baja California,
<collectingCountry id="F35B7655FB756A73C95DFA57FACCFA65" box="[1286,1375,1495,1519]" name="Mexico" pageId="10" pageNumber="11">México</collectingCountry>
. This confusion must be due to the fact that the microscope slides are labeled as Loma Linda, California,
<collectingCountry id="F35B7655FB756A73C8B1FA7CFAB5F99E" box="[1258,1318,1532,1556]" name="United States of America" pageId="10" pageNumber="11">USA</collectingCountry>
. However,
<bibRefCitation id="EFDD4B34FB756A73CCCCF9A0FE48F9B2" author="Newell" box="[151,475,1568,1592]" pageId="10" pageNumber="11" refString="Newell, I. M. &amp; R. E. Ryckman. (1964) Hirstiella pyriformis sp. n. (Acari, Pterygosomidae), a new parasite of lizards from Baja California. Journal of Parasitology, 50, 163 - 171." type="journal article" year="1964">Newell and Ryckman (1964)</bibRefCitation>
mentioned that the description was based from mites collected in a laboratory colony of
<taxonomicName id="4C4C4D46FB756A73CCEEF9C4FE14F9D6" box="[181,391,1604,1628]" class="Reptilia" family="Iguanidae" genus="Sauromalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="varius">
<emphasis id="B938EAD7FB756A73CCEEF9C4FE14F9D6" box="[181,391,1604,1628]" italics="true" pageId="10" pageNumber="11">Sauromalus varius</emphasis>
</taxonomicName>
at Loma Linda University, but that the lizards originated from San Esteban Island.
</paragraph>
</subSubSection>
</treatment>
</document>