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<document id="BED7D2B253F9D683AE174C7036C1FA47" ID-DOI="10.1080/00222933.2017.1355491" ID-ISSN="1464-5262" ID-Zenodo-Dep="5183671" ID-ZooBank="C8A19CFD-1329-427A-B380-08542BB25718" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628733935909" checkinUser="felipe" docAuthor="Nguyen, Anh D." docDate="2017" docId="03E587B8FFDDFF87E1E0FADA992FF9EF" docLanguage="en" docName="JNatHist.51.39-40.2331-2343.pdf" docOrigin="Journal of Natural History 51 (39 - 40)" docSource="http://dx.doi.org/10.1080/00222933.2017.1355491" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Vietnamorpha pumatensis Nguyen 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2338" masterDocId="FFDCFFC0FFD8FF8FE069FFF59835FFA5" masterDocTitle="A second species of the genus Vietnamorpha Golovatch, 1984 (Polydesmida: Paradoxosomatidae) and notes on the generic relationship" masterLastPageNumber="2343" masterPageNumber="2331" pageNumber="2335" updateTime="1699050494882" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6E3A78CC34143C77F366B6ECE9C0C66A">A second species of the genus Vietnamorpha Golovatch, 1984 (Polydesmida: Paradoxosomatidae) and notes on the generic relationship</mods:title>
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<mods:namePart id="D9ECA6024769BDBF70659154A12AC0AD">Nguyen, Anh D.</mods:namePart>
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<mods:title id="A7A64AAB36A64011F06A4A0BA719AD55">Journal of Natural History</mods:title>
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<mods:date id="4458773315234B2368EC0B72A58827DB">2017</mods:date>
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<mods:number id="84DC49D9DD543E3CABE6444F85F4580B">2017-08-15</mods:number>
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<mods:number id="4DDB4B47F28A370CB9E7246A243DD193">51</mods:number>
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<mods:number id="88C0BF6FE3203D1725EF0FFD1106F40B">39 - 40</mods:number>
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<treatment id="03E587B8FFDDFF87E1E0FADA992FF9EF" LSID="urn:lsid:plazi:treatment:03E587B8FFDDFF87E1E0FADA992FF9EF" httpUri="http://treatment.plazi.org/id/03E587B8FFDDFF87E1E0FADA992FF9EF" lastPageId="8" lastPageNumber="2338" pageId="5" pageNumber="2335">
<subSubSection id="C3566525FFDDFF8AE1E0FADA9BAEFAEC" box="[393,923,1327,1353]" pageId="5" pageNumber="2335" type="nomenclature">
<paragraph id="8BF336AEFFDDFF8AE1E0FADA9BAEFAEC" blockId="5.[393,923,1327,1388]" box="[393,923,1327,1353]" pageId="5" pageNumber="2335">
<heading id="D0BB81C2FFDDFF8AE1E0FADA9BAEFAEC" box="[393,923,1327,1353]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="2335" reason="5">
<taxonomicName id="4C4C4D2DFFDDFF8AE1E0FADA9B01FAEC" ID-CoL="5BDG2" authority="Nguyen" authorityName="Nguyen" authorityYear="2017" box="[393,820,1327,1353]" class="Diplopoda" family="Paradoxosomatidae" genus="Vietnamorpha" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="2335" phylum="Arthropoda" rank="species" species="pumatensis" status="sp. nov.">
<emphasis id="B938EABCFFDDFF8AE1E0FADA9AFBFAEC" bold="true" box="[393,718,1327,1353]" italics="true" pageId="5" pageNumber="2335">Vietnamorpha pumatensis</emphasis>
Nguyen
</taxonomicName>
<taxonomicNameLabel id="A20B57C7FFDDFF8AE357FAC59BAEFAEC" box="[830,923,1328,1353]" pageId="5" pageNumber="2335" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3566525FFDDFF8AE25FFAA69AD8FAC9" box="[566,749,1362,1388]" pageId="5" pageNumber="2335" type="description">
<paragraph id="8BF336AEFFDDFF8AE25FFAA69AD8FAC9" blockId="5.[393,923,1327,1388]" box="[566,749,1362,1388]" pageId="5" pageNumber="2335">
(
<figureCitation id="13772A2BFFDDFF8AE257FAA69A9EFAC9" box="[574,683,1363,1388]" captionStart="Figure 1" captionStartId="5.[160,225,1067,1090]" captionTargetBox="[163,1153,142,1036]" captionTargetId="figure-71@5.[163,1153,142,1036]" captionTargetPageId="5" captionText="Figure 1. Collecting sites for the genus Vietnamorpha Golovatch, 1984. 1: Xuan Son National Park, 2: Ba Vi National Park, 3: Thuong Tien Natural Reserve, 4: Cuc Phuong National Park, 5: Pu Mat National Park." figureDoi="http://doi.org/10.5281/zenodo.5183673" httpUri="https://zenodo.org/record/5183673/files/figure.png" pageId="5" pageNumber="2335">Figures 1</figureCitation>
,
<figureCitation id="13772A2BFFDDFF8AE2D3FAA69AD0FAC9" box="[698,741,1362,1388]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStartId-0="7.[160,225,933,956]" captionStartId-1="8.[160,225,905,928]" captionStartId-2="9.[160,225,980,1003]" captionTargetBox-0="[166,1151,145,901]" captionTargetBox-1="[163,1151,142,874]" captionTargetBox-2="[163,1152,142,949]" captionTargetId-0="figure-234@7.[163,1152,142,902]" captionTargetId-1="figure-180@8.[163,1152,142,874]" captionTargetId-2="figure-198@9.[163,1152,142,949]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." captionText-1="Figure 4. Vietnamorpha pumatensis sp.nov., holotype. Right gonopod, mesal (a), dorsal (b), ventral (c), and lateral view (d). Scale bar = 1 mm." captionText-2="Figure 5. Vietnamorpha pumatensis sp. nov., holotype. Right gonopod, dorsal (a) and lateral view (b); postfemoral region, dorsal view (e). Left gonopod, mesal (c) and ventral view (d); postfemoral region, ventral view (f). Scale bars: ad = 1 mm; ef = 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5183677" figureDoi-1="http://doi.org/10.5281/zenodo.5183679" figureDoi-2="http://doi.org/10.5281/zenodo.5183681" httpUri-0="https://zenodo.org/record/5183677/files/figure.png" httpUri-1="https://zenodo.org/record/5183679/files/figure.png" httpUri-2="https://zenodo.org/record/5183681/files/figure.png" pageId="5" pageNumber="2335">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3566525FFDDFF8AE0C9FA589C21F98B" pageId="5" pageNumber="2335" type="materials_examined">
<paragraph id="8BF336AEFFDDFF8AE0C9FA5899B5FA63" blockId="5.[160,1156,1453,1582]" box="[160,384,1453,1478]" pageId="5" pageNumber="2335">
<emphasis id="B938EABCFFDDFF8AE0C9FA5899B5FA63" bold="true" box="[160,384,1453,1478]" italics="true" pageId="5" pageNumber="2335">Material examined</emphasis>
</paragraph>
<paragraph id="8BF336AEFFDDFF8AE0C9FA249B93F9AE" blockId="5.[160,1156,1453,1582]" pageId="5" pageNumber="2335">
<materialsCitation id="3B243CF3FFDDFF8AE0C9FA249B93F9AE" collectingDate="2011-04-04" collectingDateMax="2011-04-10" collectingDateMin="2011-04-04" collectorName="Anh D. Nguyen." elevation="430" latitude="18.9715" location="Chem" longLatPrecision="7" longitude="104.8008" municipality="Pu Mat National Park" pageId="5" pageNumber="2335" specimenCount="1" specimenCount-male="1" stateProvince="Nghe An" typeStatus="holotype">
<emphasis id="B938EABCFFDDFF8AE0C9FA249934FA4C" box="[160,257,1489,1513]" italics="true" pageId="5" pageNumber="2335">
<typeStatus id="54F7880CFFDDFF8AE0C9FA249934FA4C" box="[160,257,1489,1513]" pageId="5" pageNumber="2335" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D4AFD27FFDDFF8AE167FA24996FFA4C" box="[270,346,1489,1513]" pageId="5" pageNumber="2335" type="male">1 male</specimenCount>
(
<emphasis id="B938EABCFFDDFF8AE100FA249A29FA4C" bold="true" box="[361,540,1489,1513]" pageId="5" pageNumber="2335">IEBR-Myr 158H</emphasis>
)
<collectingRegion id="4988F84CFFDDFF8AE242FA249AE4FA4C" box="[555,721,1489,1513]" country="Vietnam" name="Nghe An" pageId="5" pageNumber="2335">Nghe An Prov.</collectingRegion>
,
<collectingMunicipality id="6B97ACD4FFDDFF8AE2B3FA249BFFFA4C" box="[730,970,1489,1513]" pageId="5" pageNumber="2335">Pu Mat National Park</collectingMunicipality>
,
<location id="8E936075FFDDFF8AE3BDFA249C23FA4C" LSID="urn:lsid:plazi:treatment:03E587B8FFDDFF87E1E0FADA992FF9EF:8E936075FFDDFF8AE3BDFA249C23FA4C" box="[980,1046,1489,1513]" latitude="18.9715" longLatPrecision="7" longitude="104.8008" municipality="Pu Mat National Park" name="Chem" pageId="5" pageNumber="2335" stateProvince="Nghe An">Chem</location>
waterfall,
<quantity id="4CB49B4BFFDDFF8AE0C9FA0698DFF9AE" box="[160,234,1523,1547]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="2335" unit="m" value="430.0">
<elevation id="0061D19DFFDDFF8AE0C9FA0698DFF9AE" box="[160,234,1523,1547]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="2335" unit="m" value="430.0">430 m</elevation>
</quantity>
,
<geoCoordinate id="EE785069FFDDFF8AE09FFA069945F9AE" box="[246,368,1523,1547]" degrees="18.9715" direction="north" orientation="latitude" pageId="5" pageNumber="2335" precision="5" value="18.9715">18.9715°N-</geoCoordinate>
<geoCoordinate id="EE785069FFDDFF8AE119FA0699DBF9AE" box="[368,494,1523,1547]" degrees="104.8008" direction="east" orientation="longitude" pageId="5" pageNumber="2335" precision="5" value="104.8008">104.8008°E</geoCoordinate>
,
<date id="FFF2106EFFDDFF8AE190FA069A9AF9AE" box="[505,687,1523,1547]" pageId="5" pageNumber="2335" value="2011-04-04" valueMax="2011-04-10" valueMin="2011-04-04">
<collectingDate id="EFB6E986FFDDFF8AE190FA069A9AF9AE" box="[505,687,1523,1547]" pageId="5" pageNumber="2335" value="2011-04-04" valueMax="2011-04-10" valueMin="2011-04-04">410 April 2011</collectingDate>
</date>
, coll.
<collectorName id="26B95378FFDDFF8AE298FA069B93F9AE" box="[753,934,1523,1547]" pageId="5" pageNumber="2335">Anh D. Nguyen.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BF336AEFFDDFF8AE0A9F9E39C21F98B" blockId="5.[160,1156,1453,1582]" box="[192,1044,1558,1582]" pageId="5" pageNumber="2335">
<materialsCitation id="3B243CF3FFDDFF8AE0A9F9E39C24F98B" box="[192,1041,1558,1582]" collectingDate="2011-04-04" collectingDateMax="2011-04-10" collectingDateMin="2011-04-04" collectorName="Anh D. Nguyen." elevation="430" latitude="18.9715" location="Chem" longLatPrecision="7" longitude="104.8008" municipality="Pu Mat National Park" pageId="5" pageNumber="2335" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Nghe An" typeStatus="paratype">
<emphasis id="B938EABCFFDDFF8AE0A9F9E3991FF98B" box="[192,298,1558,1582]" italics="true" pageId="5" pageNumber="2335">
<typeStatus id="54F7880CFFDDFF8AE0A9F9E3991FF98B" box="[192,298,1558,1582]" pageId="5" pageNumber="2335" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="9D4AFD27FFDDFF8AE151F9E399A6F98B" box="[312,403,1558,1582]" pageId="5" pageNumber="2335" type="male">2 males</specimenCount>
,
<specimenCount id="9D4AFD27FFDDFF8AE1F7F9E39A3EF98B" box="[414,523,1558,1582]" pageId="5" pageNumber="2335" type="female">3 females</specimenCount>
(
<emphasis id="B938EABCFFDDFF8AE275F9E39AFBF98B" bold="true" box="[540,718,1558,1582]" pageId="5" pageNumber="2335">IEBR-Myr 158P</emphasis>
); same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF336626FFDEFF89E0C9FC9E9A68FC3A" ID-DOI="http://doi.org/10.5281/zenodo.5183675" ID-Zenodo-Dep="5183675" httpUri="https://zenodo.org/record/5183675/files/figure.png" pageId="6" pageNumber="2336" startId="6.[160,225,875,898]" targetBox="[163,1152,142,844]" targetPageId="6">
<paragraph id="8BF336AEFFDEFF89E0C9FC9E9A68FC3A" blockId="6.[160,1155,875,927]" pageId="6" pageNumber="2336">
<emphasis id="B938EABCFFDEFF89E0C9FC9E98C8FC27" bold="true" box="[160,253,875,898]" pageId="6" pageNumber="2336">Figure 2.</emphasis>
<taxonomicName id="4C4C4D2DFFDEFF89E16FFC9E9AB0FC27" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[262,645,875,898]" class="Diplopoda" family="Paradoxosomatidae" genus="Vietnamorpha" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="2336" phylum="Arthropoda" rank="species" species="spiralis">
<emphasis id="B938EABCFFDEFF89E16FFC9E99EFFC27" box="[262,474,875,898]" italics="true" pageId="6" pageNumber="2336">Vietnamorpha spiralis</emphasis>
<bibRefCitation id="EFDD4B5FFFDEFF89E18AFC9E9AB0FC27" author="Golovatch SI" box="[483,645,875,898]" pageId="6" pageNumber="2336" pagination="5 - 77" refId="ref4654" refString="Golovatch SI. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica. 30 (1 - 2): 5 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
from Cuc Phuong NP. Left gonopod, lateral view (a) and mesal view (b). Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3566525FFDEFF89E0C9FC2F9BD1FBFF" pageId="6" pageNumber="2336" type="diagnosis">
<paragraph id="8BF336AEFFDEFF89E0C9FC2F9921FC56" blockId="6.[160,1155,986,1114]" box="[160,276,986,1011]" pageId="6" pageNumber="2336">
<emphasis id="B938EABCFFDEFF89E0C9FC2F9921FC56" bold="true" box="[160,276,986,1011]" italics="true" pageId="6" pageNumber="2336">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF336AEFFDEFF89E0C9FC089BD1FBFF" blockId="6.[160,1155,986,1114]" pageId="6" pageNumber="2336">
The species differs from its congener,
<taxonomicName id="4C4C4D2DFFDEFF89E21EFC089B85FBB0" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[631,944,1021,1045]" class="Diplopoda" family="Paradoxosomatidae" genus="Vietnamorpha" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="2336" phylum="Arthropoda" rank="species" species="spiralis">
<emphasis id="B938EABCFFDEFF89E21EFC089AD0FBB0" box="[631,741,1021,1045]" italics="true" pageId="6" pageNumber="2336">V. spiralis</emphasis>
<bibRefCitation id="EFDD4B5FFFDEFF89E29AFC089B85FBB0" author="Golovatch SI" box="[755,944,1021,1045]" pageId="6" pageNumber="2336" pagination="5 - 77" refId="ref4654" refString="Golovatch SI. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica. 30 (1 - 2): 5 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
</taxonomicName>
, in the gonopod femorite being erect and cylindrically elongate; and the solenomere being almost completely sheathed by solenophore, and being only exposed at the tip.
</paragraph>
</subSubSection>
<subSubSection id="C3566525FFDEFF89E0C9FB699ADCFB7D" box="[160,745,1180,1240]" pageId="6" pageNumber="2336" type="etymology">
<paragraph id="8BF336AEFFDEFF89E0C9FB699928FB10" blockId="6.[160,745,1180,1240]" box="[160,285,1180,1205]" pageId="6" pageNumber="2336">
<emphasis id="B938EABCFFDEFF89E0C9FB699928FB10" bold="true" box="[160,285,1180,1205]" italics="true" pageId="6" pageNumber="2336">Etymology</emphasis>
</paragraph>
<paragraph id="8BF336AEFFDEFF89E0C9FB359ADCFB7D" blockId="6.[160,745,1180,1240]" box="[160,745,1216,1240]" pageId="6" pageNumber="2336">
<materialsCitation id="3B243CF3FFDEFF89E0C9FB359AD0FB7D" box="[160,741,1216,1240]" location="Pu Mat National Park" pageId="6" pageNumber="2336" specimenCount="1">
Named after its
<typeStatus id="54F7880CFFDEFF89E130FB3599BEFB7D" box="[345,395,1216,1240]" pageId="6" pageNumber="2336">type</typeStatus>
locality,
<location id="8E936075FFDEFF89E198FB359AD0FB7D" LSID="urn:lsid:plazi:treatment:03E587B8FFDDFF87E1E0FADA992FF9EF:8E936075FFDEFF89E198FB359AD0FB7D" box="[497,741,1216,1240]" name="Pu Mat National Park" pageId="6" pageNumber="2336">Pu Mat National Park</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566525FFDEFF87E0C9FAEC9AA9FA0F" lastPageId="8" lastPageNumber="2338" pageId="6" pageNumber="2336" type="description">
<paragraph id="8BF336AEFFDEFF89E0C9FAEC9913FA97" blockId="6.[160,1156,1305,1677]" box="[160,294,1305,1330]" pageId="6" pageNumber="2336">
<emphasis id="B938EABCFFDEFF89E0C9FAEC9913FA97" bold="true" box="[160,294,1305,1330]" italics="true" pageId="6" pageNumber="2336">Description</emphasis>
</paragraph>
<paragraph id="8BF336AEFFDEFF89E0C9FAC89A5AFA18" blockId="6.[160,1156,1305,1677]" pageId="6" pageNumber="2336">
Size: length ca.
<quantity id="4CB49B4BFFDEFF89E127FAC899DEFAF0" box="[334,491,1341,1365]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="1.97" metricValueMin="1.93" pageId="6" pageNumber="2336" unit="mm" value="19.5" valueMax="19.7" valueMin="19.3">19.319.7 mm</quantity>
(male), 18.120.0 mm (female); width of midbody pro- and metazonae
<quantity id="4CB49B4BFFDEFF89E14DFA959992FADD" box="[292,423,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="6" pageNumber="2336" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
(male),
<quantity id="4CB49B4BFFDEFF89E26BFA959AB0FADD" box="[514,645,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="6" pageNumber="2336" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.71.8 mm</quantity>
(female) and
<quantity id="4CB49B4BFFDEFF89E34BFA959B90FADD" box="[802,933,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="6" pageNumber="2336" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 mm</quantity>
(male),
<quantity id="4CB49B4BFFDEFF89E469FA959CB6FADD" box="[1024,1155,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="6" pageNumber="2336" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.92.1 mm</quantity>
(female), respectively.
<typeStatus id="54F7880CFFDEFF89E1F7FA779A30FA3F" box="[414,517,1410,1434]" pageId="6" pageNumber="2336" type="holotype">Holotype</typeStatus>
length ca.
<quantity id="4CB49B4BFFDEFF89E2E0FA779AC5FA3F" box="[649,752,1410,1434]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="6" pageNumber="2336" unit="mm" value="19.7">19.7 mm</quantity>
, width of midbody pro- and metazonae
<quantity id="4CB49B4BFFDEFF89E085FA509977FA18" box="[236,322,1445,1469]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="2336" unit="mm" value="1.3">1.3 mm</quantity>
and
<quantity id="4CB49B4BFFDEFF89E117FA5099E2FA18" box="[382,471,1445,1469]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="2336" unit="mm" value="1.5">1.5 mm</quantity>
, respectively.
</paragraph>
<paragraph id="8BF336AEFFDEFF89E0A9FA3D9A2EF9A7" blockId="6.[160,1156,1305,1677]" pageId="6" pageNumber="2336">Body uniformly darkish brown with a broad median yellowish area on metaterga. Sterna and legs brownish yellow.</paragraph>
<paragraph id="8BF336AEFFDEFF89E0A9F9F89CB1F928" blockId="6.[160,1156,1305,1677]" pageId="6" pageNumber="2336">
Head slightly broader than collum in width (
<figureCitation id="13772A2BFFDEFF89E2D5F9F89B2FF980" box="[700,794,1549,1573]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="6" pageNumber="2336">Figure 3</figureCitation>
(a)); clypeolabral region densely setose. Epicranial suture clearly distinct. Antennae long, reaching segment 4 if stretched posteriorly; antennomeres 24 slender while antennomeres 56 claviform; antennomeres 3&gt; 4&gt; 2 = 5&gt; 6&gt; 7&gt;
<quantity id="4CB49B4BFFDEFF89E19DF9809A2BF928" box="[500,542,1653,1677]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="2336" unit="in" value="1.0">1 in</quantity>
length; antennomere 7 with four sensory cones at tip.
</paragraph>
<caption id="DF336626FFDFFF88E0C9FC509CB1FC53" ID-DOI="http://doi.org/10.5281/zenodo.5183677" ID-Zenodo-Dep="5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="7" pageNumber="2337" startId="7.[160,225,933,956]" targetBox="[166,1151,145,901]" targetPageId="7">
<paragraph id="8BF336AEFFDFFF88E0C9FC509CB1FC53" blockId="7.[160,1156,933,1014]" pageId="7" pageNumber="2337">
<emphasis id="B938EABCFFDFFF88E0C9FC5098CFFC19" bold="true" box="[160,250,933,956]" pageId="7" pageNumber="2337">Figure 3.</emphasis>
<taxonomicName id="4C4C4D2DFFDFFF88E16AFC5099C8FC19" authority="Nguyen, 2017" authorityName="Nguyen" authorityYear="2017" box="[259,509,933,956]" class="Diplopoda" family="Paradoxosomatidae" genus="Vietnamorpha" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="2337" phylum="Arthropoda" rank="species" species="pumatensis" status="sp. nov.">
<emphasis id="B938EABCFFDFFF88E16AFC5099C8FC19" box="[259,509,933,956]" italics="true" pageId="7" pageNumber="2337">Vietnamorpha pumatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57C7FFDFFF88E26AFC509A64FC19" box="[515,593,933,956]" pageId="7" pageNumber="2337" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF336AEFFDFFF88E0A9FBB79C64FB3A" blockId="7.[160,1156,1089,1668]" pageId="7" pageNumber="2337">
Collum (
<figureCitation id="13772A2BFFDFFF88E149FBB79948FBFF" box="[288,381,1090,1114]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="7" pageNumber="2337">Figure 3</figureCitation>
(ab)) subequal to segment
<quantity id="4CB49B4BFFDFFF88E2D6FBB79ADEFBFF" box="[703,747,1090,1114]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="2337" unit="in" value="2.0">2 in</quantity>
width, oval-shaped; surface shining and smooth, neither wrinkled nor granulated, with three rows of setae: 6 + 6 anterior, 2 + 2 intermediate and 6 + 6 posterior. Axial line distinct. Paraterga totally absent.
</paragraph>
<paragraph id="8BF336AEFFDFFF88E0A9FB5F9B36FACA" blockId="7.[160,1156,1089,1668]" pageId="7" pageNumber="2337">
Body (
<figureCitation id="13772A2BFFDFFF88E160FB5F9952FB67" box="[265,359,1194,1218]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="7" pageNumber="2337">Figure 3</figureCitation>
(af)) submoniliform, segment 4 &lt;3 &lt;2 =
<quantity id="4CB49B4BFFDFFF88E33DFB5F9B88FB67" box="[852,957,1193,1218]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="4.318" metricValueMin="1.27" pageId="7" pageNumber="2337" unit="in" value="11.0" valueMax="17.0" valueMin="5.0">5 17 in</quantity>
width, thereafter tapering towards telson. Prozonae and metazonae smooth and shining, neither granulated nor wrinkled. Metaterga 218 with a row of 2 + 2 setae near anterior margin. Transverse sulcus absent. Axial line distinctly present. Pleurosternal carinae present as a keel with a caudal denticle on segments 217. Waist between pro- and metazonae clearly broad, neither striolate nor beaded at bottom.
</paragraph>
<paragraph id="8BF336AEFFDFFF88E0A9FA8F9A3DFA72" blockId="7.[160,1156,1089,1668]" pageId="7" pageNumber="2337">
Paraterga (
<figureCitation id="13772A2BFFDFFF88E152FA8F99ADFA37" box="[315,408,1402,1426]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="7" pageNumber="2337">Figure 3</figureCitation>
(cd)) totally absent on poreless segments, but very tiny on porebearing segments. Ozopores normal, lying inside white hallows on lateral side of segments 5, 7, 910, 1213, 1519.
</paragraph>
<paragraph id="8BF336AEFFDFFF88E0A9FA179904F99A" blockId="7.[160,1156,1089,1668]" pageId="7" pageNumber="2337">
Epiproct (
<figureCitation id="13772A2BFFDFFF88E147FA1799B9FA5F" box="[302,396,1506,1530]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="7" pageNumber="2337">Figure 3</figureCitation>
(fg)) broadly truncated, dorsally inflattened. Tip with four spinnerets. Hypoproct trapeziform, with two well separated, distolateral, setiferous knobs (
<figureCitation id="13772A2BFFDFFF88E0C1F9D29933F99A" box="[168,262,1575,1599]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="7" pageNumber="2337">Figure 3</figureCitation>
(h)).
</paragraph>
<paragraph id="8BF336AEFFDFFF87E0A9F9BF9B43FBEA" blockId="7.[160,1156,1089,1668]" lastBlockId="8.[160,1156,1044,1450]" lastPageId="8" lastPageNumber="2338" pageId="7" pageNumber="2337">
Sterna (
<figureCitation id="13772A2BFFDFFF88E17EF9BF9941F9C7" box="[279,372,1610,1634]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="7" pageNumber="2337">Figure 3</figureCitation>
(e)) densely setose with four long sternal spines, each next to a coxa; posterior spines much more developed and longer than anterior ones. Cross impressions clearly distinct. Sternum 5 without modifications, but sternum 4 with two long, anteriorly curved, setiferous tubercles between coxae 3 (
<figureCitation id="13772A2BFFD0FF87E29CFBC29B63FBEA" box="[757,854,1079,1103]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="8" pageNumber="2338">Figure 3</figureCitation>
(i)).
</paragraph>
<caption id="DF336626FFD0FF87E0C9FC7C9A17FC18" ID-DOI="http://doi.org/10.5281/zenodo.5183679" ID-Zenodo-Dep="5183679" httpUri="https://zenodo.org/record/5183679/files/figure.png" pageId="8" pageNumber="2338" startId="8.[160,225,905,928]" targetBox="[163,1151,142,874]" targetPageId="8">
<paragraph id="8BF336AEFFD0FF87E0C9FC7C9A17FC18" blockId="8.[160,1156,905,957]" pageId="8" pageNumber="2338">
<emphasis id="B938EABCFFD0FF87E0C9FC7C98CFFC05" bold="true" box="[160,250,905,928]" pageId="8" pageNumber="2338">Figure 4.</emphasis>
<taxonomicName id="4C4C4D2DFFD0FF87E16AFC7C99C8FC05" authority="Nguyen, 2017" authorityName="Nguyen" authorityYear="2017" box="[259,509,905,928]" class="Diplopoda" family="Paradoxosomatidae" genus="Vietnamorpha" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="2338" phylum="Arthropoda" rank="species" species="pumatensis" status="sp. nov.">
<emphasis id="B938EABCFFD0FF87E16AFC7C99C8FC05" box="[259,509,905,928]" italics="true" pageId="8" pageNumber="2338">Vietnamorpha pumatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57C7FFD0FF87E26BFC7C9A7AFC05" box="[514,591,905,928]" pageId="8" pageNumber="2338" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. Right gonopod, mesal (a), dorsal (b), ventral (c), and lateral view (d). Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BF336AEFFD0FF87E0A9FBAF999EFB12" blockId="8.[160,1156,1044,1450]" pageId="8" pageNumber="2338">
Legs (
<figureCitation id="13772A2BFFD0FF87E16DFBAF9954FBD7" box="[260,353,1114,1138]" captionStart="Figure 3" captionStartId="7.[160,225,933,956]" captionTargetBox="[166,1151,145,901]" captionTargetId="figure-234@7.[163,1152,142,902]" captionTargetPageId="7" captionText="Figure 3. Vietnamorpha pumatensis sp. nov., holotype. Head, lateral view (a), dorsal view (b); midbody segments, lateral view (c), dorsal view (d); sterna 1215, ventral view (e); telson, dorsal view (f), lateral view (g), ventral view (h); sterna 45, ventral view (i). Scale bars: ag = 1 mm; h = 0.5 mm; i = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5183677" httpUri="https://zenodo.org/record/5183677/files/figure.png" pageId="8" pageNumber="2338">Figure 3</figureCitation>
(cd)) slender, ca. 2.02.2 times as long as midbody height. Prefemora strongly swollen ventrally. Femora without modifications. Tarsal brushes, if present, only on pre-gonopodal legs.
</paragraph>
<paragraph id="8BF336AEFFD0FF87E0A9FB379AA9FA0F" blockId="8.[160,1156,1044,1450]" pageId="8" pageNumber="2338">
Gonopod simple (
<figureCitation id="13772A2BFFD0FF87E1E3FB379A39FB7F" box="[394,524,1218,1242]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStartId-0="8.[160,225,905,928]" captionStartId-1="9.[160,225,980,1003]" captionTargetBox-0="[163,1151,142,874]" captionTargetBox-1="[163,1152,142,949]" captionTargetId-0="figure-180@8.[163,1152,142,874]" captionTargetId-1="figure-198@9.[163,1152,142,949]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figure 4. Vietnamorpha pumatensis sp.nov., holotype. Right gonopod, mesal (a), dorsal (b), ventral (c), and lateral view (d). Scale bar = 1 mm." captionText-1="Figure 5. Vietnamorpha pumatensis sp. nov., holotype. Right gonopod, dorsal (a) and lateral view (b); postfemoral region, dorsal view (e). Left gonopod, mesal (c) and ventral view (d); postfemoral region, ventral view (f). Scale bars: ad = 1 mm; ef = 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5183679" figureDoi-1="http://doi.org/10.5281/zenodo.5183681" httpUri-0="https://zenodo.org/record/5183679/files/figure.png" httpUri-1="https://zenodo.org/record/5183681/files/figure.png" pageId="8" pageNumber="2338">Figures 45</figureCitation>
). Coxite about 1/31/2 femorite length; distovental part sparsely setose. Prefemorite densely setose, separated laterally from femorite by an oblique sulcus. Femorite (
<emphasis id="B938EABCFFD0FF87E1ACFAF299E9FABA" bold="true" box="[453,476,1287,1311]" pageId="8" pageNumber="2338">fe</emphasis>
) erect, not curved, cylindrically elongate, without modifications, but with a distal small spine (
<emphasis id="B938EABCFFD0FF87E22AFADF9A6BFAE7" bold="true" box="[579,606,1322,1346]" pageId="8" pageNumber="2338">ds</emphasis>
). Postfemoral region consisting only of solenophore (
<emphasis id="B938EABCFFD0FF87E09BFAB89928FAC0" bold="true" box="[242,285,1357,1381]" pageId="8" pageNumber="2338">sph</emphasis>
) and solenomere (
<emphasis id="B938EABCFFD0FF87E19BFAB89A30FAC0" bold="true" box="[498,517,1357,1381]" pageId="8" pageNumber="2338">sl</emphasis>
). Solenomere completely sheathed by solenophore, and only exposed at the tip. Solenophore strongly coiled, almost twice. Tip with a small denticle (
<emphasis id="B938EABCFFD0FF87E163FA679910FA0F" bold="true" box="[266,293,1426,1450]" pageId="8" pageNumber="2338">sd</emphasis>
) and a large pointed lobule (
<emphasis id="B938EABCFFD0FF87E211FA679ABAFA0F" bold="true" box="[632,655,1426,1450]" pageId="8" pageNumber="2338">pl</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C3566525FFD0FF87E0C9FA19992FF9EF" pageId="8" pageNumber="2338" type="discussion">
<paragraph id="8BF336AEFFD0FF87E0C9FA199933F9A0" blockId="8.[160,1155,1516,1610]" box="[160,262,1516,1541]" pageId="8" pageNumber="2338">
<emphasis id="B938EABCFFD0FF87E0C9FA199933F9A0" bold="true" box="[160,262,1516,1541]" italics="true" pageId="8" pageNumber="2338">Remarks</emphasis>
</paragraph>
<paragraph id="8BF336AEFFD0FF87E0C9F9FA992FF9EF" blockId="8.[160,1155,1516,1610]" pageId="8" pageNumber="2338">
The species was found in the Pu Mat National Park (
<collectingCountry id="F35B763EFFD0FF87E284F9FA9B78F982" box="[749,845,1551,1575]" name="Vietnam" pageId="8" pageNumber="2338">Vietnam</collectingCountry>
) where the forests are best conserved.
</paragraph>
</subSubSection>
</treatment>
</document>