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<document id="C4BFE59563471ACDC6E0E3078661F157" ID-DOI="10.11646/zootaxa.5519.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13915579" ID-ZooBank="C12023C8-FF3D-4058-A590-E1E2DC38B4FA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728584281405" checkinUser="plazi" docAuthor="Huang, Tian, Janák, Jiří &amp; Zhou, Hong-Zhang" docDate="2024" docId="03E587A3FFFCFFD5B9C08B29FE85FD69" docLanguage="en" docName="zootaxa.5519.1.5.pdf" docOrigin="Zootaxa 5519 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diochus chambrieri Huang, Janak &amp; Zhou 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="115" masterDocId="FFDCFFDBFFF4FFD9B9578E0CFFB2FC0C" masterDocTitle="Taxonomy of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) of Papua New Guinea, Vanuatu and Solomon Islands" masterLastPageNumber="118" masterPageNumber="103" pageNumber="111" updateTime="1728585392433" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F9EA9725DECEA19038703AC11D4C7D4D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C69FD6C6C6E9FA307A849E704702CF7">
<mods:title id="2F8E87A3DB2EA12F3CD1934D9944D70C">Taxonomy of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) of Papua New Guinea, Vanuatu and Solomon Islands</mods:title>
</mods:titleInfo>
<mods:name id="0409FB6E1CE97D344C91E7D0BA939A43" type="personal">
<mods:role id="2E5860EA6E3C4E6C55C7C414C7A308C4">
<mods:roleTerm id="ADD4D43ACAFD8E0DF9BABBCB91541BF3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E972BFE2074A0377098CB3E6BE98AA2">Huang, Tian</mods:namePart>
<mods:affiliation id="293D2E40034629E0031A8E046E82F1E1">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. &amp; University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
</mods:name>
<mods:name id="BD758DFE33351D3B4D8A82568DB025D4" type="personal">
<mods:role id="B625639EEB6ADA76B742B4D9890DDAFB">
<mods:roleTerm id="A0A0ACCC8791EB42295062ED90F107A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B75926F0580F77D55CDAEE992C91319">Janák, Jiří</mods:namePart>
<mods:affiliation id="1A5A702DC29D94AC23754FC050064D45">CZ- 41762 Rtyně nad Bílinou, č. 4, Czech Republic.</mods:affiliation>
</mods:name>
<mods:name id="E224D0AF49FACE91029AFC9F528EC072" type="personal">
<mods:role id="C6BA49A6EB4601E48E36280B4608E32C">
<mods:roleTerm id="2FB8688125E1FEF60C2C52E710853A63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70544DBB9407DFC7EF75331C3B314F87">Zhou, Hong-Zhang</mods:namePart>
<mods:affiliation id="7733B4EA53650E309839E1D9CF47D2B9">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. &amp; University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F33437E7038B07A660B2F855BEA9781C">text</mods:typeOfResource>
<mods:relatedItem id="79C96FF18B373DE07D4C518E1E03EADB" type="host">
<mods:titleInfo id="FF21359C7997E5D887411975EC197BBF">
<mods:title id="5993F0DA94EDF925220A85C4377D5117">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B8794BA9889B53858D625454428E5AB4">
<mods:date id="62FD9D7A0C2823E0E0F6D957501F8A75">2024</mods:date>
<mods:detail id="63907E64C4DCC68804DED36478C7A6CB" type="pubDate">
<mods:number id="D1A3FCA3D1F1340355223922EE8C684D">2024-10-07</mods:number>
</mods:detail>
<mods:detail id="A79B774E6D763209DCF0D1B4ABACFBFB" type="volume">
<mods:number id="110DBC8457BBFF7D8276375FA6CD86BF">5519</mods:number>
</mods:detail>
<mods:detail id="960AF4A439FA0F56862A3B65DAAFEDC7" type="issue">
<mods:number id="04EE0D29F45ADB679DAB97D31A95ED9D">1</mods:number>
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<mods:start id="41B72183D6106C73B844CD5592C4DB1B">103</mods:start>
<mods:end id="7F5C6A3661ABE07BFFCC349F14FADF10">118</mods:end>
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<mods:location id="3C11DC594554A5A64723AE6DC03CD53A">
<mods:url id="5371A7C51AA74F3633D0CB244D3B9771">http://dx.doi.org/10.11646/zootaxa.5519.1.5</mods:url>
</mods:location>
<mods:classification id="8C46AFBD5B2804186A560202029B8E4D">journal article</mods:classification>
<mods:identifier id="17E9E65C6903CB91A793AC49931C5A6D" type="DOI">10.11646/zootaxa.5519.1.5</mods:identifier>
<mods:identifier id="441A871E8C3FC08D15872D6121FD663D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5170FC4871577D156707B4A5599A2255" type="Zenodo-Dep">13915579</mods:identifier>
<mods:identifier id="E6C506DE039344A16938B7D259620CB7" type="ZooBank">C12023C8-FF3D-4058-A590-E1E2DC38B4FA</mods:identifier>
</mods:mods>
<treatment id="03E587A3FFFCFFD5B9C08B29FE85FD69" LSID="urn:lsid:plazi:treatment:03E587A3FFFCFFD5B9C08B29FE85FD69" httpUri="http://treatment.plazi.org/id/03E587A3FFFCFFD5B9C08B29FE85FD69" lastPageId="12" lastPageNumber="115" pageId="8" pageNumber="111">
<subSubSection id="C356653EFFFCFFD1B9C08B29FCF5F94C" box="[151,839,1317,1344]" pageId="8" pageNumber="111" type="nomenclature">
<paragraph id="8BF336B5FFFCFFD1B9C08B29FCF5F94C" blockId="8.[151,839,1317,1380]" box="[151,839,1317,1344]" pageId="8" pageNumber="111">
<heading id="D0BB81D9FFFCFFD1B9C08B29FCF5F94C" bold="true" box="[151,839,1317,1344]" fontSize="11" level="1" pageId="8" pageNumber="111" reason="1">
<emphasis id="B938EAA7FFFCFFD1B9C08B29FCF5F94C" bold="true" box="[151,839,1317,1344]" pageId="8" pageNumber="111">
3.
<taxonomicName id="4C4C4D36FFFCFFD1B9908B29FD6FF94C" authority="Huang, Janak &amp; Zhou" authorityName="Huang, Janak &amp; Zhou" authorityYear="2024" box="[199,733,1317,1344]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="111" phylum="Arthropoda" rank="species" species="chambrieri" status="sp. nov.">
<emphasis id="B938EAA7FFFCFFD1B9908B29FE08F94C" bold="true" box="[199,442,1317,1344]" italics="true" pageId="8" pageNumber="111">Diochus chambrieri</emphasis>
Huang, Janák &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="A20B57DCFFFCFFD1BBBE8B2AFCF5F94C" box="[745,839,1318,1344]" pageId="8" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356653EFFFCFFD1B9C08B45FE24F968" box="[151,406,1353,1380]" pageId="8" pageNumber="111" type="description">
<paragraph id="8BF336B5FFFCFFD1B9C08B45FE24F968" blockId="8.[151,839,1317,1380]" box="[151,406,1353,1380]" pageId="8" pageNumber="111">
(
<figureCitation id="13772A30FFFCFFD1B9C88B45FF5DF968" box="[159,239,1353,1380]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[154,1432,744,1846]" captionTargetId="figure-232@2.[151,1435,741,1850]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of four species from Papua New Guinea, Vanuatu and Solomon Islands in dorsal view:A. Diochus elongatus sp. nov.; B. D. chambrieri sp. nov.; C. D. papuanus Cameron, 1939, D. D. loebli sp. nov. (Scale bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915581" httpUri="https://zenodo.org/record/13915581/files/figure.png" pageId="8" pageNumber="111">Fig 1B</figureCitation>
; Figs 4-1; 4-2)
</paragraph>
</subSubSection>
<subSubSection id="C356653EFFFCFFD1B9C08B9EFA36FA6C" pageId="8" pageNumber="111" type="materials_examined">
<paragraph id="8BF336B5FFFCFFD1B9C08B9EFD2DF9A7" blockId="8.[151,1437,1425,2028]" box="[151,671,1425,1452]" pageId="8" pageNumber="111">
<materialsCitation id="3B243CE8FFFCFFD1B9C08B9EFD29F9A7" box="[151,667,1425,1452]" country="Vanuatu" location="Tanna" municipality="Ipenkangien" pageId="8" pageNumber="111" specimenCount="1" typeStatus="holotype">
<emphasis id="B938EAA7FFFCFFD1B9C08B9EFDB4F9A0" bold="true" box="[151,518,1425,1452]" pageId="8" pageNumber="111">
<typeStatus id="54F78817FFFCFFD1B9C08B9EFF60F9A0" box="[151,210,1426,1452]" pageId="8" pageNumber="111">Type</typeStatus>
locality.
<collectingCountry id="F35B7625FFFCFFD1B86B8B9EFE14F9A0" box="[316,422,1426,1452]" name="Vanuatu" pageId="8" pageNumber="111">Vanuatu</collectingCountry>
:
<location id="8E93606EFFFCFFD1B8E68B9EFDB3F9A0" LSID="urn:lsid:plazi:treatment:03E587A3FFFCFFD5B9C08B29FE85FD69:8E93606EFFFCFFD1B8E68B9EFDB3F9A0" box="[433,513,1426,1452]" country="Vanuatu" municipality="Ipenkangien" name="Tanna" pageId="8" pageNumber="111">Tanna</location>
:
</emphasis>
<collectingMunicipality id="6B97ACCFFFFCFFD1BB5A8B9DFD29F9A7" box="[525,667,1425,1451]" pageId="8" pageNumber="111">Ipenkangien</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336B5FFFCFFD1B9908BBAFA36FA6C" blockId="8.[151,1437,1425,2028]" pageId="8" pageNumber="111">
<emphasis id="B938EAA7FFFCFFD1B9908BBAFEC6F9C3" bold="true" box="[199,372,1461,1488]" pageId="8" pageNumber="111">Type material.</emphasis>
<materialsCitation id="3B243CE8FFFCFFD1B82D8BBAFC73F9FF" collectingDate="1979-05-24" collectingDateMax="1979-05-27" collectingDateMin="1979-05-24" collectionCode="MHNG" collectorName="A. de Chambrier &amp; Nlles &amp; de Tanna Ipenkangien White Sands" country="Vanuatu" location="Tanna" municipality="Ipenkangien" pageId="8" pageNumber="111" specimenCount="1" typeStatus="holotype">
<emphasis id="B938EAA7FFFCFFD1B82D8BBAFE42F9DC" bold="true" box="[378,496,1462,1488]" pageId="8" pageNumber="111">
<typeStatus id="54F78817FFFCFFD1B82D8BBAFE59F9DC" box="[378,491,1462,1488]" pageId="8" pageNumber="111" type="holotype">Holotype</typeStatus>
:
</emphasis>
male,
<emphasis id="B938EAA7FFFCFFD1BB6D8BB9FC96F9DC" bold="true" box="[570,804,1461,1488]" pageId="8" pageNumber="111">
<collectingCountry id="F35B7625FFFCFFD1BB6D8BB9FD77F9C3" box="[570,709,1461,1487]" name="Vanuatu" pageId="8" pageNumber="111">VANUATU</collectingCountry>
:
<location id="8E93606EFFFCFFD1BB988BBAFCADF9DC" LSID="urn:lsid:plazi:treatment:03E587A3FFFCFFD5B9C08B29FE85FD69:8E93606EFFFCFFD1BB988BBAFCADF9DC" box="[719,799,1462,1488]" country="Vanuatu" municipality="Ipenkangien" name="Tanna" pageId="8" pageNumber="111">Tanna</location>
:
</emphasis>
<collectingMunicipality id="6B97ACCFFFFCFFD1BA7D8BB9FC05F9C3" box="[810,951,1461,1487]" pageId="8" pageNumber="111">Ipenkangien</collectingMunicipality>
,
<date id="FFF21075FFFCFFD1BA958BBAFBDDF9DC" box="[962,1135,1462,1488]" pageId="8" pageNumber="111" value="1979-05-24" valueMax="1979-05-27" valueMin="1979-05-24">
<collectingDate id="EFB6E99DFFFCFFD1BA958BBAFBDDF9DC" box="[962,1135,1462,1488]" pageId="8" pageNumber="111" value="1979-05-24" valueMax="1979-05-27" valueMin="1979-05-24">24−27. V. 1979</collectingDate>
</date>
,
<collectorName id="26B95363FFFCFFD1BD208BBAFA83F9C3" box="[1143,1329,1461,1488]" pageId="8" pageNumber="111">A. de Chambrier</collectorName>
collected (
<collectionCode id="ED5DAE70FFFCFFD1B9C88BD6FF49F9F8" box="[159,251,1498,1524]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="111" type="Museum">MHNG</collectionCode>
). [
<collectorName id="26B95363FFFCFFD1B8408BD5FEE7F9FF" box="[279,341,1497,1523]" pageId="8" pageNumber="111">Nlles</collectorName>
HEBRIDES Ile
<collectorName id="26B95363FFFCFFD1BB4C8BD5FC08F9FF" box="[539,954,1497,1524]" pageId="8" pageNumber="111">de Tanna Ipenkangien White Sands</collectorName>
]
</materialsCitation>
.
<materialsCitation id="3B243CE8FFFCFFD1BA988BD6FE2DFA1B" collectingDate="1979-05-24" collectingDateMax="1979-05-27" collectingDateMin="1979-05-24" collectionCode="MHNG" collectorName="A. de Chambrier &amp; Nlles &amp; de Tanna Ipenkangien White Sands" country="Vanuatu" location="Tanna" municipality="Ipenkangien" pageId="8" pageNumber="111" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B938EAA7FFFCFFD1BA988BD6FB58F9FF" bold="true" box="[975,1258,1497,1524]" pageId="8" pageNumber="111">
<typeStatus id="54F78817FFFCFFD1BA988BD6FBF9F9F8" box="[975,1099,1498,1524]" pageId="8" pageNumber="111" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F35B7625FFFCFFD1BD0D8BD5FB57F9FF" box="[1114,1253,1497,1523]" name="Vanuatu" pageId="8" pageNumber="111">VANUATU</collectingCountry>
:
</emphasis>
<specimenCount id="9D4AFD3CFFFCFFD1BDA38BD6FAEEF9FF" box="[1268,1372,1497,1524]" count="1" pageId="8" pageNumber="111" type="female">1 female</specimenCount>
, with same data as holotype
</materialsCitation>
;
<materialsCitation id="3B243CE8FFFCFFD1B8F98BF1FB07FA30" collectingDate="1965-02-16" collectionCode="NHMUK, JJRC" collectorName="P. Greenslade" country="Solomon Islands" location="Malaita" pageId="8" pageNumber="111" specimenCode="BM 1966-477" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Malaita" typeStatus="paratype">
<emphasis id="B938EAA7FFFCFFD1B8F98BF1FD7DFA1B" bold="true" box="[430,719,1533,1559]" pageId="8" pageNumber="111">
<collectingCountry id="F35B7625FFFCFFD1B8F98BF1FD78FA1B" box="[430,714,1533,1559]" name="Solomon Islands" pageId="8" pageNumber="111">SOLOMON ISLANDS</collectingCountry>
:
</emphasis>
<specimenCount id="9D4AFD3CFFFCFFD1BB8D8BF2FC9FFA1B" box="[730,813,1533,1560]" count="1" pageId="8" pageNumber="111" type="male">1 male</specimenCount>
,
<specimenCount id="9D4AFD3CFFFCFFD1BA6A8BF2FC10FA1B" box="[829,930,1533,1560]" count="1" pageId="8" pageNumber="111" type="female">1 female</specimenCount>
“SOLOMON IS.
<collectingRegion id="4988F857FFFCFFD1BD2C8BF1FB61FA1B" box="[1147,1235,1533,1559]" country="Solomon Islands" name="Malaita" pageId="8" pageNumber="111">Malaita</collectingRegion>
, Dala,
<date id="FFF21075FFFCFFD1BC7C8BF1FA2EFA14" box="[1323,1436,1533,1560]" pageId="8" pageNumber="111" value="1965-02-16">
<collectingDate id="EFB6E99DFFFCFFD1BC7C8BF1FA2EFA14" box="[1323,1436,1533,1560]" pageId="8" pageNumber="111" value="1965-02-16">16/2/1965</collectingDate>
</date>
PJM Greenslade 16688.”, “SOLOMON IS
<collectorName id="26B95363FFFCFFD1BBDB882EFC95FA37" box="[652,807,1569,1596]" pageId="8" pageNumber="111">P. Greenslade</collectorName>
<specimenCode id="DBEA9ECEFFFCFFD1BA66882EFC66FA37" box="[817,980,1569,1596]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="8" pageNumber="111">BM 1966-477</specimenCode>
” (
<collectionCode id="ED5DAE70FFFCFFD1BAA6882EFBEDFA30" box="[1009,1119,1570,1596]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="111" type="Museum">NHMUK</collectionCode>
,
<collectionCode id="ED5DAE70FFFCFFD1BD3A882EFB1FFA30" box="[1133,1197,1570,1596]" pageId="8" pageNumber="111">JJRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B243CE8FFFCFFD1BD93882EFA32FA6C" collectionCode="NHMUK" collectorName="P. Greenslade" country="Solomon Islands" pageId="8" pageNumber="111" specimenCode="BM 1966-477" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D4AFD3CFFFCFFD1BD93882EFA9FFA37" box="[1220,1325,1569,1596]" count="1" pageId="8" pageNumber="111" type="female">1 female</specimenCount>
: “18/3/65 SOLOMON IS., Russel Is. PJM Greenslade 17102.”, “SOLOMON IS
<collectorName id="26B95363FFFCFFD1BAF8884AFBF5FA53" box="[943,1095,1605,1632]" pageId="8" pageNumber="111">P. Greenslade</collectorName>
<specimenCode id="DBEA9ECEFFFCFFD1BD19884AFB5CFA53" box="[1102,1262,1605,1632]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="8" pageNumber="111">BM 1966-477</specimenCode>
” (
<collectionCode id="ED5DAE70FFFCFFD1BC50884AFACAFA6C" box="[1287,1400,1606,1632]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="111" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C356653EFFFCFFD3B9908866FEE5FCD9" lastPageId="10" lastPageNumber="113" pageId="8" pageNumber="111" type="description">
<paragraph id="8BF336B5FFFCFFD1B9908866FC67FAC0" blockId="8.[151,1437,1425,2028]" pageId="8" pageNumber="111">
<emphasis id="B938EAA7FFFCFFD1B9908866FECCFA88" bold="true" box="[199,382,1642,1668]" pageId="8" pageNumber="111">Measurements.</emphasis>
BL =
<quantity id="4CB49B50FFFCFFD1B8968866FD94FA88" box="[449,550,1642,1668]" metricMagnitude="-3" metricUnit="m" metricValue="4.82" pageId="8" pageNumber="111" unit="mm" value="4.82">4.82 mm</quantity>
, FL =
<quantity id="4CB49B50FFFCFFD1BB3B8866FD62FA88" box="[620,720,1642,1668]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="8" pageNumber="111" unit="mm" value="2.17">2.17 mm</quantity>
, HL =
<quantity id="4CB49B50FFFCFFD1BA4D8865FCCDFA88" box="[794,895,1641,1668]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="8" pageNumber="111" unit="mm" value="0.59">0.59 mm</quantity>
, HW =
<quantity id="4CB49B50FFFCFFD1BA848865FB8AFA88" box="[979,1080,1641,1668]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="111" unit="mm" value="0.5">0.50 mm</quantity>
, EyL =
<quantity id="4CB49B50FFFCFFD1BDDA8866FB43FA88" box="[1165,1265,1642,1668]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="111" unit="mm" value="0.19">0.19 mm</quantity>
,TL =
<quantity id="4CB49B50FFFCFFD1BC648866FA25FA88" box="[1331,1431,1642,1668]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="111" unit="mm" value="0.34">0.34 mm</quantity>
, ANL =
<quantity id="4CB49B50FFFCFFD1B9BA8882FEE1FAA4" box="[237,339,1678,1704]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="111" unit="mm" value="1.3">1.30 mm</quantity>
, ANT1 =
<quantity id="4CB49B50FFFCFFD1B8958881FD9AFAA4" box="[450,552,1677,1704]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="111" unit="mm" value="0.16">0.16 mm</quantity>
, ANT2 = 0.12, ANT3 =
<quantity id="4CB49B50FFFCFFD1BA6E8881FC2DFAA4" box="[825,927,1677,1704]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="111" unit="mm" value="0.15">0.15 mm</quantity>
, ANT4 =
<quantity id="4CB49B50FFFCFFD1BD598882FBC0FAA4" box="[1038,1138,1678,1704]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="111" unit="mm" value="0.11">0.11 mm</quantity>
, ANT 11 =
<quantity id="4CB49B50FFFCFFD1BDA28882FAE8FAA4" box="[1269,1370,1678,1704]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="111" unit="mm" value="0.13">0.13 mm</quantity>
, PL =
<quantity id="4CB49B50FFFCFFD1B9C088BEFF4FFAC0" box="[151,253,1714,1740]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="8" pageNumber="111" unit="mm" value="0.81">0.81 mm</quantity>
, PW =
<quantity id="4CB49B50FFFCFFD1B80788BDFE04FAC0" box="[336,438,1713,1740]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="111" unit="mm" value="0.65">0.65 mm</quantity>
, EL =
<quantity id="4CB49B50FFFCFFD1BB5688BDFDD5FAC0" box="[513,615,1713,1740]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="111" unit="mm" value="0.76">0.76 mm</quantity>
, EW =
<quantity id="4CB49B50FFFCFFD1BBEC88BEFC93FAC0" box="[699,801,1714,1740]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="111" unit="mm" value="0.93">0.93 mm</quantity>
, SL =
<quantity id="4CB49B50FFFCFFD1BA3D88BDFC63FAC0" box="[874,977,1713,1740]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="8" pageNumber="111" unit="mm" value="0.57">0.57 mm</quantity>
.
</paragraph>
<paragraph id="8BF336B5FFFCFFD1B99088D9FC55FB3B" blockId="8.[151,1437,1425,2028]" pageId="8" pageNumber="111">
<emphasis id="B938EAA7FFFCFFD1B99088D9FEEBFAE3" bold="true" box="[199,345,1749,1775]" pageId="8" pageNumber="111">Description.</emphasis>
Body (
<figureCitation id="13772A30FFFCFFD1B8E288D9FDBBFAFC" box="[437,521,1749,1776]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[154,1432,744,1846]" captionTargetId="figure-232@2.[151,1435,741,1850]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of four species from Papua New Guinea, Vanuatu and Solomon Islands in dorsal view:A. Diochus elongatus sp. nov.; B. D. chambrieri sp. nov.; C. D. papuanus Cameron, 1939, D. D. loebli sp. nov. (Scale bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915581" httpUri="https://zenodo.org/record/13915581/files/figure.png" pageId="8" pageNumber="111">Fig 1B</figureCitation>
) long, narrowed anteriorly and posteriorly, medium sized. Body dark brown, abdominal intersegmental membrane lighter. Legs brown, tarsi distinctly lighter. Antennae dark brown to yellowish brown, apex of each antennomere lighter. Maxillary and labial palpi brown.
</paragraph>
<paragraph id="8BF336B5FFFCFFD0B990894EFF62FD85" blockId="8.[151,1437,1425,2028]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="112" pageId="8" pageNumber="111">
<emphasis id="B938EAA7FFFCFFD1B990894EFEB5FB50" bold="true" box="[199,263,1858,1884]" pageId="8" pageNumber="111">Head</emphasis>
(Fig 4-1A). Round square in shape, 1.18 times as long as wide. Tempora slightly widened backwards, posterior angles rounded. Dorsal surface shiny, bearing distinct and transverse microstriae, also with coarse and sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of tempus and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye large-sized, distinctly protruding laterad, longitudinal diameter about 1/2 the length of tempora (eye: tempora = 0.55). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions (
<quantity id="4CB49B50FFFDFFD0B9C88F2BFEB6FD4D" box="[159,260,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="112" unit="mm" value="0.11">0.11 mm</quantity>
) distinctly longer than distance from antenna to eye (
<quantity id="4CB49B50FFFDFFD0BA0E8F2BFC71FD4D" box="[857,963,295,321]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="9" pageNumber="112" unit="mm" value="0.07">0.07 mm</quantity>
). Ventral surface with scattered punctures, interspaces between them about 4 puncture diameters. Gular sutures deep, confluent at basal 1/5, then extended to base.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908F9EFDDBFE15" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908F9EFE88FDA0" bold="true" box="[199,314,402,428]" pageId="9" pageNumber="112">Antennae</emphasis>
(Fig 4-1B). Scape rod-shaped, slightly thickened apically, distinctly shorter than two subsequent antennomeres combined; antennomere 2 slightly elongate; antennomere 3 slightly longer than 2; antennomere 4 about thrice of width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908C2FFD7FFE89" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908C2FFEE4FE31" bold="true" box="[199,342,547,573]" pageId="9" pageNumber="112">Mouthparts</emphasis>
(Fig 4-1A). Labrum nearly hexagonal, lateral margin straight, widest at basal 1/3 to 1/2; width of anterior margin is about 1/3 the width at the widest point. Mandibles (Figs 4-1D) falciform, left one with three small teeth on inner margin, whereas right one with two.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908C83FD37FEFD" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908C83FEB6FEA5" bold="true" box="[199,260,655,681]" pageId="9" pageNumber="112">Neck</emphasis>
(Fig 4-1A). Cylindrical, shiny, only bearing shallow microsculpture, width
<quantity id="4CB49B50FFFDFFD0BD268C83FB6BFEA5" box="[1137,1241,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="112" unit="mm" value="0.11">0.11 mm</quantity>
, shorter than 1/3 of head width. Dorsal surface with distinct groove; ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908CF7FCF9FFE1" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908CF7FEF3FF19" bold="true" box="[199,321,763,789]" pageId="9" pageNumber="112">Prothorax</emphasis>
(Fig 4-1A). Pronotum dark brown, elongate (PL to PW ratio 1.25), distinctly longer and wider than head. Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/4 (widest at 1/4), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composing of 3 large punctures and 2 additional punctures outside of puncture row, 13–15 punctures scattered near each margin (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908DFBFB7EF855" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908DFBFEE5F81D" bold="true" box="[199,343,1015,1041]" pageId="9" pageNumber="112">Pterothorax</emphasis>
(Fig 4-1A). Mesoscutellum triangular, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; the paired oblique furrows short but deep; the carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908A6FFB6AF8E5" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908A6FFEA1F871" bold="true" box="[199,275,1123,1149]" pageId="9" pageNumber="112">Elytra</emphasis>
(Fig 4-1A). Elytra dark brown, short and transverse (EL to EW ratio 0.82), distinctly longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal surface shiny, flattened, without microsculpture; each elytron with row of 5 punctures along suture, row of 5 punctures in median, also with 4−5 rows of punctures on deflexed portion.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908AFFFE62F959" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908AFFFF4DF901" bold="true" box="[199,255,1267,1293]" pageId="9" pageNumber="112">Legs</emphasis>
(Fig 4-1C). First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi slightly shorter than 2−4 together, that of meso-, meta- slightly shorter than the length of each 2−3.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B9908B52FE5AFA21" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908B52FE8EF974" bold="true" box="[199,316,1374,1400]" pageId="9" pageNumber="112">Abdomen</emphasis>
(
<figureCitation id="13772A30FFFDFFD0B8078B53FE14F975" box="[336,422,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[154,1432,744,1846]" captionTargetId="figure-232@2.[151,1435,741,1850]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of four species from Papua New Guinea, Vanuatu and Solomon Islands in dorsal view:A. Diochus elongatus sp. nov.; B. D. chambrieri sp. nov.; C. D. papuanus Cameron, 1939, D. D. loebli sp. nov. (Scale bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915581" httpUri="https://zenodo.org/record/13915581/files/figure.png" pageId="9" pageNumber="112">Fig 1B</figureCitation>
). Broadest at segment V. Tergites III–VII covered with dense and obvious transverse microstriae; each tergite with dense brown pubescence, also with row of darker and coarser setae at posterior margin of III–VI. Tergites III–VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with surface shiny, bearing polygonal reticulum, with dense punctures. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
</paragraph>
<paragraph id="8BF336B5FFFDFFD0B990883BFA96FB7D" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B990883BFEB6FA5D" bold="true" box="[199,260,1591,1617]" pageId="9" pageNumber="112">Male</emphasis>
(Figs 4-1E–J; 4-2A–E). Head with a small round elevation between eyes. Posterior margin of sternite VIII emarginated deeply and arcuately (Figs 4-1E, L; 4-2B). Posterior margin of tergite VIII arcuate (Figs 4-1F; -2C). Tergite IX (Figs 4-1H; 4-2D) symmetrical, connected mediobasally, narrowed apically. Sternite IX (Figs 4-1G; 4- 2E) asymmetrical, widest near basal 1/2; basal margin straight, apical margin normally emarginated; width of apical margin distinctly shorter than of basal margin. Tergite X (Figs 4-1H; 4-2D) symmetrical, triangular. Aedeagus (Figs 4-1I−K; 4-2A) symmetrical, medium-sized, ca.
<quantity id="4CB49B50FFFDFFD0BBE788E7FCA6FB09" box="[688,788,1771,1797]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="9" pageNumber="112" unit="mm" value="0.61">0.61 mm</quantity>
long, normally sclerotized. Parameres symmetrical, thick and short, about 1/7 length of median lobe. Internal structures sclerotized, composed of four parts: paired trapezoidal sclerites on upper middle, brown color; paired strip shaped sclerites on both sides, black colored; paired rectangular sclerites and one irregular sclerite in the middle, brown color. Sperm pump coiled, thinned toward apex.
</paragraph>
<paragraph id="8BF336B5FFFDFFD3B9908977FEE5FCD9" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1436,151,249]" lastPageId="10" lastPageNumber="113" pageId="9" pageNumber="112">
<emphasis id="B938EAA7FFFDFFD0B9908977FEACFB99" bold="true" box="[199,286,1915,1941]" pageId="9" pageNumber="112">Female</emphasis>
(Figs 4-2F−I). Head without elevation between eyes. Sternite VIII (Fig 4-2F) with posterior margin rounded. Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX (Fig 4-2H) symmetrical, with deep concaved margin in middle. Tergite X broad, basal margin curved. Sternite X (Fig 4-2G) slender and rhomboidal, with numerous setae in lower half. Tube of spermatheca sclerotized, very long and strongly coiled (Fig 4-2I).
</paragraph>
</subSubSection>
<subSubSection id="C356653EFFFEFFD3B9908ED2FE79FCF5" box="[199,459,222,249]" pageId="10" pageNumber="113" type="distribution">
<paragraph id="8BF336B5FFFEFFD3B9908ED2FE79FCF5" blockId="10.[151,1436,151,249]" box="[199,459,222,249]" pageId="10" pageNumber="113">
<emphasis id="B938EAA7FFFEFFD3B9908ED2FED3FCF4" bold="true" box="[199,353,222,248]" pageId="10" pageNumber="113">Distribution.</emphasis>
<collectingCountry id="F35B7625FFFEFFD3B8308ED3FE75FCF5" box="[359,455,223,249]" name="Vanuatu" pageId="10" pageNumber="113">Vanuatu</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF33663DFFFEFFD3B9C08934FB24FB95" ID-DOI="http://doi.org/10.5281/zenodo.13915591" ID-Zenodo-Dep="13915591" httpUri="https://zenodo.org/record/13915591/files/figure.png" pageId="10" pageNumber="113" startId="10.[151,250,1848,1872]" targetBox="[151,1430,277,1821]" targetPageId="10" targetType="figure">
<paragraph id="8BF336B5FFFEFFD3B9C08934FB24FB95" blockId="10.[151,1436,1848,1945]" pageId="10" pageNumber="113">
<emphasis id="B938EAA7FFFEFFD3B9C08934FE9BFB5C" bold="true" box="[151,297,1848,1872]" pageId="10" pageNumber="113">FIGURE 4-1.</emphasis>
<taxonomicName id="4C4C4D36FFFEFFD3B8668934FE4FFB5C" authorityName="Huang &amp; Janák &amp; Zhou" authorityYear="2024" box="[305,509,1848,1872]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="113" phylum="Arthropoda" rank="species" species="chambrieri" status="sp. nov.">
<emphasis id="B938EAA7FFFEFFD3B8668934FE4FFB5C" box="[305,509,1848,1872]" italics="true" pageId="10" pageNumber="113">Diochus chambrieri</emphasis>
</taxonomicName>
<emphasis id="B938EAA7FFFEFFD3BB538935FDE4FB5D" bold="true" box="[516,598,1849,1873]" pageId="10" pageNumber="113">
<taxonomicNameLabel id="A20B57DCFFFEFFD3BB538935FDE4FB5D" box="[516,598,1849,1873]" pageId="10" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: holotype: A. forebody, dorsal view; B. antenna; C. proleg; D. mandible; E. male sternite VIII; F. male tergite VIII; G. male sternite IX; H. male tergites IX–X; I. aedeagus, dorsal view; J. aedeagus, ventral view; paratypes: K. aedeagus, ventral view; L. male sternite VIII. (Scale bars: A = 0.5 mm, B–L= 0.1 mm).
</paragraph>
</caption>
<caption id="DF33663DFFFFFFD2B9C0899BFE4CFBF4" ID-DOI="http://doi.org/10.5281/zenodo.13915596" ID-Zenodo-Dep="13915596" httpUri="https://zenodo.org/record/13915596/files/figure.png" pageId="11" pageNumber="114" startId="11.[151,250,1943,1967]" targetBox="[189,1397,185,1913]" targetPageId="11" targetType="figure">
<paragraph id="8BF336B5FFFFFFD2B9C0899BFE4CFBF4" blockId="11.[151,1436,1943,2040]" pageId="11" pageNumber="114">
<emphasis id="B938EAA7FFFFFFD2B9C0899BFE99FBA3" bold="true" box="[151,299,1943,1967]" pageId="11" pageNumber="114">FIGURE 4-2.</emphasis>
<taxonomicName id="4C4C4D36FFFFFFD2B8638994FDB0FBBC" authorityName="Huang &amp; Janák &amp; Zhou" authorityYear="2024" box="[308,514,1944,1968]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="114" phylum="Arthropoda" rank="species" species="chambrieri" status="sp. nov.">
<emphasis id="B938EAA7FFFFFFD2B8638994FDB0FBBC" box="[308,514,1944,1968]" italics="true" pageId="11" pageNumber="114">Diochus chambrieri</emphasis>
</taxonomicName>
<emphasis id="B938EAA7FFFFFFD2BB5D8994FDECFBBC" bold="true" box="[522,606,1944,1968]" pageId="11" pageNumber="114">
<taxonomicNameLabel id="A20B57DCFFFFFFD2BB5D8994FDECFBBC" box="[522,606,1944,1968]" pageId="11" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A. aedeagus, dorsal view; B. male sternite VIII; C. male tergite VIII; D. male tergites IX–X; E. male sternite IX; F. female sternite VIII; G. female sternite X; H. female sternite IX; I. spermatheca.(Scale bars: A = 0.05 mm, B–I = 0.1 mm).
</paragraph>
</caption>
<subSubSection id="C356653EFFF8FFD5B9908E9AFB32FD11" pageId="12" pageNumber="115" type="diagnosis">
<paragraph id="8BF336B5FFF8FFD5B9908E9AFB32FD11" blockId="12.[151,1437,150,357]" pageId="12" pageNumber="115">
<emphasis id="B938EAA7FFF8FFD5B9908E9AFEF0FCBC" bold="true" box="[199,322,150,176]" pageId="12" pageNumber="115">Diagnosis.</emphasis>
This species is similar to
<taxonomicName id="4C4C4D36FFF8FFD5BB3E8E9BFD4BFCBC" authorityName="Cameron" authorityYear="1939" box="[617,761,151,176]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B938EAA7FFF8FFD5BB3E8E9BFD4BFCBC" box="[617,761,151,176]" italics="true" pageId="12" pageNumber="115">D. papuanus</emphasis>
</taxonomicName>
in body size and coloration, male sexual structures are also similar. The main difference is the length of elytra. In
<taxonomicName id="4C4C4D36FFF8FFD5BBAF8EB7FC35FCD8" authorityName="Cameron" authorityYear="1939" box="[760,903,187,212]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B938EAA7FFF8FFD5BBAF8EB7FC35FCD8" box="[760,903,187,212]" italics="true" pageId="12" pageNumber="115">D. papuanus</emphasis>
</taxonomicName>
the elytra are distinctly shorter. In addition, for
<taxonomicName id="4C4C4D36FFF8FFD5B9C08ED3FE97FCF4" authorityName="Cameron" authorityYear="1939" box="[151,293,223,248]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B938EAA7FFF8FFD5B9C08ED3FE97FCF4" box="[151,293,223,248]" italics="true" pageId="12" pageNumber="115">D. papuanus</emphasis>
</taxonomicName>
, male sternite VIII is provided with numerous setae on posterior emargination, while
<taxonomicName id="4C4C4D36FFF8FFD5BD808ED3FAC6FCF5" authorityName="Huang &amp; Janák &amp; Zhou" authorityYear="2024" box="[1239,1396,223,249]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="chambrieri" status="sp. nov.">
<emphasis id="B938EAA7FFF8FFD5BD808ED3FAC6FCF5" box="[1239,1396,223,249]" italics="true" pageId="12" pageNumber="115">D. chambrieri</emphasis>
</taxonomicName>
<emphasis id="B938EAA7FFF8FFD5BC2C8ED3FF7AFD11" bold="true" pageId="12" pageNumber="115">
<taxonomicNameLabel id="A20B57DCFFF8FFD5BC2C8ED3FF7AFD11" pageId="12" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is without setae, and the parameres of
<taxonomicName id="4C4C4D36FFF8FFD5BB2B8F0FFCBEFD10" authorityName="Cameron" authorityYear="1939" box="[636,780,259,284]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B938EAA7FFF8FFD5BB2B8F0FFCBEFD10" box="[636,780,259,284]" italics="true" pageId="12" pageNumber="115">D. papuanus</emphasis>
</taxonomicName>
are distinctly longer and slender.
</paragraph>
</subSubSection>
<subSubSection id="C356653EFFF8FFD5B9908F2BFE85FD69" pageId="12" pageNumber="115" type="etymology">
<paragraph id="8BF336B5FFF8FFD5B9908F2BFE85FD69" blockId="12.[151,1437,150,357]" pageId="12" pageNumber="115">
<emphasis id="B938EAA7FFF8FFD5B9908F2BFEFCFD4D" bold="true" box="[199,334,295,321]" pageId="12" pageNumber="115">Etymology.</emphasis>
The species epithet is dedicated to the collector of the
<typeStatus id="54F78817FFF8FFD5BD498F2BFB31FD4D" box="[1054,1155,295,321]" pageId="12" pageNumber="115" type="holotype">holotype</typeStatus>
, Mr. A. de Chambrier (
<collectingCountry id="F35B7625FFF8FFD5B9C88F47FE99FD69" box="[159,299,331,357]" name="Switzerland" pageId="12" pageNumber="115">Switzerland</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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