194 lines
26 KiB
XML
194 lines
26 KiB
XML
<document id="7B11603754340CF62D99B76944E00E55" ID-CLB-Dataset="31837" ID-DOI="10.5281/zenodo.893462" ID-GBIF-Dataset="13655079-c807-4b16-b3fb-a5bb965f798b" ID-ISSN="0035-418" ID-Zenodo-Dep="893462" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508158438709" checkinUser="plazi" docAuthor="Jochen Martens" docDate="2017" docId="03E57339FFF8FF9510BDF8A3E9F5FBFC" docLanguage="en" docName="rsz_124_2_1_Martens.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Starengovia Snegovaya 2010" docType="treatment" docVersion="7" lastPageNumber="188" masterDocId="FFDC0B41FFF8FF94131EFF9AED6CFFE0" masterDocTitle="On Starengovia Snegovaya, a genus of Asian nemastomatines (Arachnida: Opiliones: Nemastomatidae)" masterLastPageNumber="201" masterPageNumber="187" pageNumber="187" updateTime="1698470539736" updateUser="ExternalLinkService">
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<mods:title id="0DC744F179EDFBDB74D91EC645EFDCFE">On Starengovia Snegovaya, a genus of Asian nemastomatines (Arachnida: Opiliones: Nemastomatidae)</mods:title>
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<mods:namePart id="D613065D00F8907D4B91B805C54B27EE">Jochen Martens</mods:namePart>
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<mods:title id="CF7E776B074F9CE8FEB73739F29401A9">Revue suisse de Zoologie</mods:title>
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<mods:part id="AA7859FCD5E93272A2FBB5774394A179">
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<mods:date id="1A67D955F847DC7D0B9AB080AEDE034B">2017</mods:date>
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<mods:number id="FCD751E70E0D2B6BD7335FACB5BF2505">2017-09-30</mods:number>
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<mods:number id="6DCF840C06F47091F368F4D88835EFDD">124</mods:number>
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<mods:identifier id="8C0A188EC19F75D9644228CBA7B96944" type="DOI">10.5281/zenodo.893462</mods:identifier>
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<treatment id="03E57339FFF8FF9510BDF8A3E9F5FBFC" ID-DOI="http://doi.org/10.5281/zenodo.6030103" ID-GBIF-Taxon="135436958" ID-Zenodo-Dep="6030103" LSID="urn:lsid:plazi:treatment:03E57339FFF8FF9510BDF8A3E9F5FBFC" httpUri="http://treatment.plazi.org/id/03E57339FFF8FF9510BDF8A3E9F5FBFC" lastPageId="1" lastPageNumber="188" pageId="0" pageNumber="187">
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<subSubSection id="C35691A4FFF8FF9410BDF8A3E855F8B0" box="[931,1337,1849,1873]" pageId="0" pageNumber="187" type="nomenclature">
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<paragraph id="8BF3C22FFFF8FF9410BDF8A3E855F8B0" blockId="0.[931,1337,1817,1873]" box="[931,1337,1849,1873]" pageId="0" pageNumber="187">
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<heading id="D0BB7543FFF8FF9410BDF8A3E855F8B0" bold="true" box="[931,1337,1849,1873]" centered="true" fontSize="10" level="3" pageId="0" pageNumber="187" reason="4">
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<emphasis id="B9381E3DFFF8FF9410BDF8A3E855F8B0" bold="true" box="[931,1337,1849,1873]" pageId="0" pageNumber="187">
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Genus
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<taxonomicName id="4C4CB9ACFFF8FF9410ECF8A3E855F8B0" ID-CoL="7Q2PL" authority="Snegovaya, 2010" authorityName="Snegovaya" authorityYear="2010" box="[1010,1337,1849,1873]" class="Arachnida" family="Nemastomatidae" genus="Starengovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF8FF9410ECF8A3E91AF8B1" bold="true" box="[1010,1142,1849,1873]" italics="true" pageId="0" pageNumber="187">Starengovia</emphasis>
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<bibRefCitation id="EFDDBFDEFFF8FF941763F8A3E855F8B0" author="Snegovaya N. Y." box="[1149,1337,1849,1872]" journalOrPublisher="Acta Zoologica Bulgarica" pageId="0" pageNumber="187" pagination="351 - 354" part="62" refString="Snegovaya N. Y. 2010. New harvestman genus and species from Kyrgyz Republic (Kyrgyzstan) (Arachnida: Opiliones: Nemastomatidae). Acta Zoologica Bulgarica 62 (3): 351 - 354." title="New harvestman genus and species from Kyrgyz Republic (Kyrgyzstan) (Arachnida: Opiliones: Nemastomatidae)" type="journal article" year="2010">Snegovaya, 2010</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35691A4FFF8FF941021F8F3E979F841" pageId="0" pageNumber="187" type="reference_group">
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<paragraph id="8BF3C22FFFF8FF941021F8F3E979F841" blockId="0.[831,1437,1896,1953]" pageId="0" pageNumber="187">
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<emphasis id="B9381E3DFFF8FF941021F8F3EEB6F89F" bold="true" box="[831,986,1896,1921]" pageId="0" pageNumber="187">Type species:</emphasis>
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<taxonomicName id="4C4CB9ACFFF8FF9410FBF8F3EE0BF87F" ID-CoL="7Z4S9" authority="Snegovaya, 2010: 351" authorityName="Snegovaya" authorityPageNumber="351" authorityYear="2010" class="Arachnida" family="Nemastomatidae" genus="Starengovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="kirgizica">
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<emphasis id="B9381E3DFFF8FF9410FBF8F3E9A1F89F" box="[997,1229,1896,1920]" italics="true" pageId="0" pageNumber="187">Starengovia kirgizica</emphasis>
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<bibRefCitation id="EFDDBFDEFFF8FF9417C6F8F3E8F4F861" author="Snegovaya N. Y." box="[1240,1432,1897,1921]" journalOrPublisher="Acta Zoologica Bulgarica" pageId="0" pageNumber="187" pagination="351 - 354" part="62" refString="Snegovaya N. Y. 2010. New harvestman genus and species from Kyrgyz Republic (Kyrgyzstan) (Arachnida: Opiliones: Nemastomatidae). Acta Zoologica Bulgarica 62 (3): 351 - 354." title="New harvestman genus and species from Kyrgyz Republic (Kyrgyzstan) (Arachnida: Opiliones: Nemastomatidae)" type="journal article" year="2010">Snegovaya, 2010</bibRefCitation>
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: 351
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</taxonomicName>
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(by monotypy).
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</paragraph>
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</subSubSection>
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<footnote id="E857DE21FFF8FF941389F79BECBAF7D8" pageId="0" pageNumber="187">
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<paragraph id="8BF3C22FFFF8FF941389F79BECBAF7D8" blockId="0.[151,499,2049,2106]" pageId="0" pageNumber="187">
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Manuscript accepted 28.02.
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<docYear id="932A2ED4FFF8FF9412A3F798EC9FF7FA" box="[445,499,2050,2074]" pageId="0" pageNumber="187">2017</docYear>
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DOI: 10.5281/zenodo.
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<docID id="FA692B12FFF8FF94129AF7BBECBAF7D8" box="[388,470,2081,2104]" pageId="0" pageNumber="187">893462</docID>
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</paragraph>
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</footnote>
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<subSubSection id="C35691A4FFF9FF951389FE99EC9BFEBA" pageId="1" pageNumber="188" type="discussion">
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<paragraph id="8BF3C22FFFF9FF951389FE99EC9BFEBA" blockId="1.[151,757,258,348]" pageId="1" pageNumber="188">
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<emphasis id="B9381E3DFFF9FF951389FE99EC64FEFB" bold="true" box="[151,264,259,283]" pageId="1" pageNumber="188">Remarks:</emphasis>
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A genus of
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<taxonomicName id="4C4CB9ACFFF9FF951294FE98EF51FEF9" box="[394,573,258,281]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="family">Nemastomatidae</taxonomicName>
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in the subfamily
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<taxonomicName id="4C4CB9ACFFF9FF951389FEB8EC3CFED9" authorityName="Simon" authorityYear="1872" box="[151,336,290,313]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="subFamily" subFamily="Nemastomatinae">Nemastomatinae</taxonomicName>
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, presently comprising two species. Gender of genus name is female.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35691A4FFF9FF951389FEE9ED93FD2E" pageId="1" pageNumber="188" type="diagnosis">
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<paragraph id="8BF3C22FFFF9FF951389FEE9ED93FD2E" blockId="1.[151,757,370,1075]" pageId="1" pageNumber="188">
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<emphasis id="B9381E3DFFF9FF951389FEE9EC11FE69" bold="true" box="[151,381,370,395]" pageId="1" pageNumber="188">Emended diagnosis:</emphasis>
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Small species (up to
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<quantity id="4CB46FCAFFF9FF95117BFEE9EFDBFE6B" box="[613,695,371,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="188" unit="mm" value="1.7">1.7 mm</quantity>
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body length, females being larger than males), dorsal scutum with lines of anvil-shaped tubercles along margins of all scutal areas. Pairs of marked para-median tubercles on opisthosomal areas of dorsal scutum. Apophysis on basal cheliceral article of male well-marked but low, with a distad-directed hook, discharge area for secretion in a bowl-like excavation on medial side of apophysis. Its inner wall perforated by numerous minute pores, apparently the extrusion area for cheliceral gland secretion.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35691A4FFF9FF951389FD4DEC52FBD3" pageId="1" pageNumber="188" type="description">
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<paragraph id="8BF3C22FFFF9FF951389FD4DEC52FBD3" blockId="1.[151,757,370,1075]" pageId="1" pageNumber="188">Genital morphology of males: Characterized by rather unspecialized truncus penis, moderately slender, not markedly inflated; muscle-containing inflated base of penis large, rather compact, deeply incised medially, occupying about one fourth to one third of truncus length, truncus in straight continuation of inflated base. Glans inconspicuous, small, short, not well differentiated from truncus; armament of glans simple, with few stiff spicules in a dorsally and ventrally symmetrical arrangement, stylus short, straight to slightly curved.</paragraph>
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</subSubSection>
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<subSubSection id="C35691A4FFF9FF951389FBD6EC71FA47" pageId="1" pageNumber="188" type="distribution">
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<paragraph id="8BF3C22FFFF9FF951389FBD6EC71FA47" blockId="1.[151,756,1100,1447]" pageId="1" pageNumber="188">
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<emphasis id="B9381E3DFFF9FF951389FBD6EC41FB83" bold="true" box="[151,301,1100,1123]" pageId="1" pageNumber="188">Distribution:</emphasis>
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Hitherto only two males of one species were known from
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<collectingCountry id="F35B82BFFFF9FF95127DFBF7EC8CFB65" box="[355,480,1133,1157]" name="Kyrgyzstan" pageId="1" pageNumber="188">Kyrgyzstan</collectingCountry>
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, a second species is here described from
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<collectingCountry id="F35B82BFFFF9FF951257FB16ECC4FB43" box="[329,424,1164,1187]" name="Pakistan" pageId="1" pageNumber="188">Pakistan</collectingCountry>
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.
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<bibRefCitation id="EFDDBFDEFFF9FF9512A9FB17EF37FB43" author="Starega W." box="[439,603,1164,1190]" journalOrPublisher="Bulletin of the Polish Academy of Sciences, Biological Sciences" pageId="1" pageNumber="188" pagination="301 - 305" part="34" refString="Starega W. 1986. Eine neueArt der Nemastomatidae (Opiliones) aus dem Pamir, nebst nomenklatorisch-taxonomischen Anmerkungen. Bulletin of the Polish Academy of Sciences, Biological Sciences 34: 301 - 305." title="Eine neueArt der Nemastomatidae (Opiliones) aus dem Pamir, nebst nomenklatorisch-taxonomischen Anmerkungen" type="journal article" year="1986">Staręga (1986)</bibRefCitation>
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provisionally named a single female from the Tadjik Republic as “
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<taxonomicName id="4C4CB9ACFFF9FF9513BDFB57EC44FB04" authorityName="Kratochvil" authorityYear="1958" box="[163,296,1229,1252]" class="Arachnida" family="Nemastomatidae" genus="Mediostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF9FF9513BDFB57EC44FB04" box="[163,296,1229,1252]" italics="true" pageId="1" pageNumber="188">Mediostoma</emphasis>
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</taxonomicName>
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”
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<taxonomicName id="4C4CB9ACFFF9FF95125EFB55EF32FB04" authority="Starega, 1986" authorityName="Starega" authorityYear="1986" box="[320,606,1229,1255]" class="Arachnida" family="Nemastomatidae" genus="Mediostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="pamiricum">
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<emphasis id="B9381E3DFFF9FF95125EFB55ECD9FB06" box="[320,437,1231,1254]" italics="true" pageId="1" pageNumber="188">pamiricum</emphasis>
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<bibRefCitation id="EFDDBFDEFFF9FF9512DFFB54EF32FB04" author="Starega W." box="[449,606,1229,1255]" journalOrPublisher="Bulletin of the Polish Academy of Sciences, Biological Sciences" pageId="1" pageNumber="188" pagination="301 - 305" part="34" refString="Starega W. 1986. Eine neueArt der Nemastomatidae (Opiliones) aus dem Pamir, nebst nomenklatorisch-taxonomischen Anmerkungen. Bulletin of the Polish Academy of Sciences, Biological Sciences 34: 301 - 305." title="Eine neueArt der Nemastomatidae (Opiliones) aus dem Pamir, nebst nomenklatorisch-taxonomischen Anmerkungen" type="journal article" year="1986">Staręga, 1986</bibRefCitation>
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</taxonomicName>
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. The locality of this specimen is about half way between and approximately
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<quantity id="4CB46FCAFFF9FF95125DFA94ECF0FAC5" box="[323,412,1294,1317]" metricMagnitude="5" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="188" unit="km" value="360.0">360 km</quantity>
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from
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<taxonomicName id="4C4CB9ACFFF9FF9512F1FA95EF1DFAC6" authorityName="Snegovaya" authorityYear="2010" box="[495,625,1295,1318]" class="Arachnida" family="Nemastomatidae" genus="Starengovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF9FF9512F1FA95EF1DFAC6" box="[495,625,1295,1318]" italics="true" pageId="1" pageNumber="188">Starengovia</emphasis>
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</taxonomicName>
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records in
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<collectingCountry id="F35B82BFFFF9FF951389FAAAEC7EFAA8" box="[151,274,1328,1352]" name="Kyrgyzstan" pageId="1" pageNumber="188">Kyrgyzstan</collectingCountry>
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and in
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<collectingCountry id="F35B82BFFFF9FF95126EFAB5ECA7FAA6" box="[368,459,1327,1350]" name="Pakistan" pageId="1" pageNumber="188">Pakistan</collectingCountry>
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and thus this female may also belong to
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<taxonomicName id="4C4CB9ACFFF9FF951220FACAECD2FA87" authorityName="Snegovaya" authorityYear="2010" box="[318,446,1360,1383]" class="Arachnida" family="Nemastomatidae" genus="Starengovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF9FF951220FACAECD2FA87" box="[318,446,1360,1383]" italics="true" pageId="1" pageNumber="188">Starengovia</emphasis>
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</taxonomicName>
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. This assumption is backed by characters of external morphology (
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<figureCitation id="1377DEAAFFF9FF95117CFAEAEFD3FA67" box="[610,703,1392,1415]" captionStart="Figs 5 - 6" captionStartId="3.[151,192,2025,2046]" captionTargetBox="[161,1428,1231,1972]" captionTargetId="figure@3.[151,1436,1213,1972]" captionTargetPageId="3" captionText="Figs 5 - 6 B. Body of male in lateral view (5 - 6 A) and in dorsal view (6 B). (5) Starengovia kirgizica. (6 A) S. ivanloebli sp. n. (6 B) Mediostoma pamiricum Staręga, 1986; reproduced from Staręga (1986). Scale: 1 mm." httpUri="https://zenodo.org/record/1012867/files/figure.png" pageId="1" pageNumber="188">Fig. 6B</figureCitation>
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; see Discussion).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35691A4FFF9FF951389FA5AEF63F9D8" pageId="1" pageNumber="188" type="etymology">
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<paragraph id="8BF3C22FFFF9FF951389FA5AEF63F9D8" blockId="1.[151,756,1472,1594]" pageId="1" pageNumber="188">
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<emphasis id="B9381E3DFFF9FF951389FA5AED8FFA38" bold="true" box="[151,227,1472,1496]" pageId="1" pageNumber="188">Name:</emphasis>
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It refers to Wojciech Krzysztof Staręga (
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<date id="FFF2E4EFFFF9FF9511AAFA5AEDBCFA17" pageId="1" pageNumber="188" value="1939" valueMax="2015">1939- 2015</date>
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), a distinguished Polish arachnologist who worked on systematics and taxonomy of European, Asian and African harvestmen (
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<bibRefCitation id="EFDDBFDEFFF9FF951264F9BBEF68F9D8" author="Zabka M." box="[378,516,1569,1592]" journalOrPublisher="Arachnologische Mitteilungen" pageId="1" pageNumber="188" part="50" refString="Zabka M. 2015. Prof. dr hab. Wojciech Krzysztof Starega (1939 - 2015). Arachnologische Mitteilungen 50: vi-viii." title="Prof. dr hab. Wojciech Krzysztof Starega (1939 - 2015)" type="journal volume" year="2015">Żabka, 2015</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35691A4FFF9FF951389F9CBE9F5FBFC" pageId="1" pageNumber="188" type="discussion">
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<paragraph id="8BF3C22FFFF9FF951389F9CBEEFCFD9A" blockId="1.[151,756,1617,1738]" lastBlockId="1.[831,1437,258,1052]" pageId="1" pageNumber="188">
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<emphasis id="B9381E3DFFF9FF951389F9CBEC50F988" bold="true" box="[151,316,1617,1640]" pageId="1" pageNumber="188">Relationships:</emphasis>
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Rows of anvil-like tubercles forming lines and encircling the dorsal scutum magnum or parts of it and all or some of the dorsal scutal areas (probably equivalents to “somites”) are present in several nemastomatine genera and are documented for
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<taxonomicName id="4C4CB9ACFFF9FF951021FEB9E907FED9" authority="Roewer, 1951" authorityName="Roewer" authorityYear="1951" box="[831,1131,290,314]" class="Arachnida" family="Nemastomatidae" genus="Mitostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF9FF951021FEB9EEDFFEDA" box="[831,947,291,314]" italics="true" pageId="1" pageNumber="188">Mitostoma</emphasis>
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<bibRefCitation id="EFDDBFDEFFF9FF9510D8FEB9E907FED9" author="Roewer C. - F." box="[966,1131,290,314]" journalOrPublisher="Senckenbergiana biologica" pageId="1" pageNumber="188" pagination="95 - 153" part="32" refString="Roewer C. - F. 1951. Uber Nemastomatiden. Weitere Weberknechte XVI. Senckenbergiana biologica 32: 95 - 153." title="Uber Nemastomatiden. Weitere Weberknechte XVI" type="journal article" year="1951">Roewer, 1951</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C4CB9ACFFF9FF95179FFEB9E925FEBB" authority="Kratochvil, 1958" authorityName="Kratochvil" authorityYear="1958" class="Arachnida" family="Nemastomatidae" genus="Carinostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF9FF95179FFEB9E863FEDA" box="[1153,1295,291,314]" italics="true" pageId="1" pageNumber="188">Carinostoma</emphasis>
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<bibRefCitation id="EFDDBFDEFFF9FF95163FFEB8EE19FEB9" author="Kratochvil J." journalOrPublisher="Prace Brnenske zakladny Ceskoslovenske akademie ved, Brno" pageId="1" pageNumber="188" pagination="523 - 576" part="30" refString="Kratochvil J. 1958. Hohlenweberknechte Bulgariens (Palpatores - Nemastomatidae). Prace Brnenske zakladny Ceskoslovenske akademie ved, Brno 30 (12): 523 - 576." title="Hohlenweberknechte Bulgariens (Palpatores - Nemastomatidae)" type="journal article" year="1958">Kratochvíl, 1958</bibRefCitation>
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(
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<bibRefCitation id="EFDDBFDEFFF9FF951093FED9E92DFEBB" author="Martens J." box="[909,1089,323,347]" journalOrPublisher="Die Tierwelt Deutschlands" pageId="1" pageNumber="188" pagination="1 - 464" part="64" refString="Martens J. 1978 a. Spinnentiere, Arachnida: Weberknechte, Opiliones. Die Tierwelt Deutschlands 64: 1 - 464." title="Spinnentiere, Arachnida: Weberknechte, Opiliones" type="journal article" year="1978" yearSuffix="a">Martens, 1978a</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="4C4CB9ACFFF9FF951791FED9E941FE9B" authority="Roewer, 1951" authorityName="Roewer" authorityYear="1951" class="Arachnida" family="Nemastomatidae" genus="Acromitostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF9FF951791FED9E85FFEBA" box="[1167,1331,323,346]" italics="true" pageId="1" pageNumber="188">Acromitostoma</emphasis>
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<bibRefCitation id="EFDDBFDEFFF9FF95165DFED9EE18FE99" author="Roewer C. - F." journalOrPublisher="Senckenbergiana biologica" pageId="1" pageNumber="188" pagination="95 - 153" part="32" refString="Roewer C. - F. 1951. Uber Nemastomatiden. Weitere Weberknechte XVI. Senckenbergiana biologica 32: 95 - 153." title="Uber Nemastomatiden. Weitere Weberknechte XVI" type="journal article" year="1951">Roewer, 1951</bibRefCitation>
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(
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<bibRefCitation id="EFDDBFDEFFF9FF951098FEF8E949FE9B" author="Rambla M." box="[902,1061,354,379]" journalOrPublisher="Speleon 26" pageId="1" pageNumber="188" pagination="21 - 27" part="27" refString="Rambla M. 1983. Sobre los Nemastomatidae (Arachnida, Opiliones) de la Peninsula Iberica. VI. Acromitostoma rhinocerus y Acromitostoma hispanum (nueva combinacion). Speleon 26 / 27: 21 - 27." title="Sobre los Nemastomatidae (Arachnida, Opiliones) de la Peninsula Iberica. VI. Acromitostoma rhinocerus y Acromitostoma hispanum (nueva combinacion)" type="journal article" year="1983">Rambla, 1983</bibRefCitation>
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)
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</taxonomicName>
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. These structures are excessively developed in all New World ortholasmatines. According to the (still incomplete) molecular-based results of
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<bibRefCitation id="EFDDBFDEFFF9FF951021FE59E9FDFE3B" author="Schonhofer A. L. & Martens J." box="[831,1169,451,475]" journalOrPublisher="Organisms Diversity and Evolution" pageId="1" pageNumber="188" pagination="409 - 419" part="12" refString="Schonhofer A. L., Martens J. 2012. The enigmatic Alpine opilionid Saccarella schilleri gen. n., sp. n. (Arachnida: Nemastomatidae) - isolated systematic placement inferred from comparative genital morphology. Organisms Diversity and Evolution 12 (4): 409 - 419." title="The enigmatic Alpine opilionid Saccarella schilleri gen. n., sp. n. (Arachnida: Nemastomatidae) - isolated systematic placement inferred from comparative genital morphology" type="journal article" year="2012">Schönhofer & Martens (2012)</bibRefCitation>
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, these dorsal ornaments developed independently in at least two different lineages. Two of the relevant genera analyzed, namely
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<taxonomicName id="4C4CB9ACFFF9FF951021FDBEEEA0FDDB" authorityName="Kratochvil" authorityYear="1958" box="[831,972,548,571]" class="Arachnida" family="Nemastomatidae" genus="Carinostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B9381E3DFFF9FF951021FDBEEEA0FDDB" box="[831,972,548,571]" italics="true" pageId="1" pageNumber="188">Carinostoma</emphasis>
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</taxonomicName>
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and
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<emphasis id="B9381E3DFFF9FF951710FDBEE9E4FDDB" box="[1038,1160,548,571]" italics="true" pageId="1" pageNumber="188">
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<taxonomicName id="4C4CB9ACFFF9FF951710FDBEE9E8FDDB" authorityName="Roewer" authorityYear="1951" box="[1038,1156,548,571]" class="Arachnida" family="Nemastomatidae" genus="Mitostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">Mitostoma</taxonomicName>
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,
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</emphasis>
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are placed on different branches of the molecular-based tree and are not closely related.
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</paragraph>
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<paragraph id="8BF3C22FFFF9FF951021FD19E9F5FBFC" blockId="1.[831,1437,258,1052]" pageId="1" pageNumber="188">
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Alternatively, this dorsal armament may have been lost at various stages of nemastomatine evolution. This assumption is backed by the fact that nearly all ortholasmatine genera and species exhibit rich and elaborate scutal ornamentations. It is reduced in the few known Asian species (
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<bibRefCitation id="EFDDBFDEFFF9FF95175CFCBEE82BFCDD" author="Shear W. A. & Gruber J." box="[1090,1351,804,829]" journalOrPublisher="American Museum Novitates" pageId="1" pageNumber="188" pagination="1 - 65" part="2757" refString="Shear W. A., Gruber J. 1983. The opilionid subfamily Ortholasmatinae (Opiliones: Troguloidea, Nemastomatidae). American Museum Novitates 2757: 1 - 65." title="The opilionid subfamily Ortholasmatinae (Opiliones: Troguloidea, Nemastomatidae)" type="journal article" year="1983">Shear & Gruber, 1983</bibRefCitation>
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;
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<bibRefCitation id="EFDDBFDEFFF9FF951644FCBEEE14FCBD" author="Shear W. A." journalOrPublisher="ZooKeys" pageId="1" pageNumber="188" pagination="9 - 45" part="52" refString="Shear W. A. 2010. New species and records of ortholasmatine harvestmen from Mexico, Honduras, and the western United States (Opiliones, Nemastomatidae Ortholasmatinae). ZooKeys 52: 9 - 45." title="New species and records of ortholasmatine harvestmen from Mexico, Honduras, and the western United States (Opiliones, Nemastomatidae Ortholasmatinae)" type="journal article" year="2010">Shear, 2010</bibRefCitation>
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;
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<bibRefCitation id="EFDDBFDEFFF9FF95109CFCDEE903FCBD" author="Zhang C. & Zhang F." box="[898,1135,836,861]" journalOrPublisher="Zootaxa 3691" pageId="1" pageNumber="188" pagination="443 - 452" part="4" refString="Zhang C., Zhang F. 2013: Description of a new Cladolasma (Opiliones: Nemastomatidae: Ortholasmatinae) species from China. Zootaxa 3691 (4): 443 - 452." title="Description of a new Cladolasma (Opiliones: Nemastomatidae: Ortholasmatinae) species from China" type="journal article" year="2013">Zhang & Zhang, 2013</bibRefCitation>
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). Thus, lines of anvil-shaped tubercles may represent a plesiomorphic character within the
|
|
<taxonomicName id="4C4CB9ACFFF9FF951072FC1EE94EFC7B" box="[876,1058,900,923]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="family">Nemastomatidae</taxonomicName>
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, an element of the nemastomatid “Bauplan”. In addition, equally shaped thorns turn up as coxal-rim denticles in most species of
|
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<taxonomicName id="4C4CB9ACFFF9FF9517FCFC5FE8F5FC3C" box="[1250,1433,965,988]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="188" phylum="Arthropoda" rank="family">Nemastomatidae</taxonomicName>
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. If plesiomorphic, this character would not be useful to infer phylogenetic relationships.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |