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<document id="E0D393915B1858DC5CC017CAF68AAF02" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105362570" checkinUser="felipe" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel-Angel" docDate="2005" docId="03C487D0C454FFB4174FF94D20A9A4D4" docLanguage="en" docName="zt01012p037.pdf" docOrigin="Zootaxa 1012 (1012)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrodactylus hondurensis Arce-Perez and Moron 2005, new species" docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="FFFDFFA8C45DFFB81647FFEA2418A64A" masterDocTitle="New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua" masterLastPageNumber="37" masterPageNumber="23" pageNumber="32" updateTime="1698980141271" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1312615D197724C657687B08C6822619">New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua</mods:title>
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<treatment id="03C487D0C454FFB4174FF94D20A9A4D4" ID-DOI="http://doi.org/10.5281/zenodo.5049200" ID-Zenodo-Dep="5049200" LSID="urn:lsid:plazi:treatment:03C487D0C454FFB4174FF94D20A9A4D4" httpUri="http://treatment.plazi.org/id/03C487D0C454FFB4174FF94D20A9A4D4" lastPageId="12" lastPageNumber="35" pageId="9" pageNumber="32">
<subSubSection id="C377654DC454FFB1174FF94D2005A08B" box="[264,1053,1703,1729]" pageId="9" pageNumber="32" type="nomenclature">
<paragraph id="8BD236C6C454FFB1174FF94D2005A08B" blockId="9.[264,1053,1703,1764]" box="[264,1053,1703,1729]" pageId="9" pageNumber="32">
<heading id="D09A81AAC454FFB1174FF94D2005A08B" bold="true" box="[264,1053,1703,1729]" fontSize="11" level="1" pageId="9" pageNumber="32" reason="1">
<emphasis id="B919EAD4C454FFB1174FF94D2005A08B" bold="true" box="[264,1053,1703,1729]" pageId="9" pageNumber="32">
<taxonomicName id="4C6D4D45C454FFB1174FF94D2798A08B" authority="Arce-Perez and Moron" authorityName="Arce-Perez and Moron" authorityYear="2005" box="[264,896,1703,1729]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="hondurensis" status="sp. nov.">
<emphasis id="B919EAD4C454FFB1174FF94D2640A08B" bold="true" box="[264,600,1703,1729]" italics="true" pageId="9" pageNumber="32">Macrodactylus hondurensis</emphasis>
Arce­Pérez and Morón
</taxonomicName>
,
<taxonomicNameLabel id="A22A57AFC454FFB115CAF94D2005A08B" box="[909,1053,1703,1729]" pageId="9" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377654DC454FFB1174FF92025F6A0AE" box="[264,494,1738,1764]" pageId="9" pageNumber="32" type="description">
<paragraph id="8BD236C6C454FFB1174FF92025F6A0AE" blockId="9.[264,1053,1703,1764]" box="[264,494,1738,1764]" pageId="9" pageNumber="32">
(
<figureCitation id="13562A43C454FFB11756F9202596A0AE" box="[273,398,1738,1764]" captionStart="FIGURES 712" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 712. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="9" pageNumber="32">Figs. 8, 11</figureCitation>
,
<figureCitation id="13562A43C454FFB117DCF92025FDA0AE" box="[411,485,1738,1764]" captionStart="FIGURES 1321" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 1321. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="9" pageNumber="32">1618</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C377654DC454FFB2174FF8F02747A7CC" lastPageId="10" lastPageNumber="33" pageId="9" pageNumber="32" type="diagnosis">
<paragraph id="8BD236C6C454FFB2174FF8F02747A7CC" blockId="9.[264,1323,1818,1884]" lastBlockId="10.[264,1325,284,1870]" lastPageId="10" lastPageNumber="33" pageId="9" pageNumber="32">
<emphasis id="B919EAD4C454FFB1174FF8F02590A17E" bold="true" box="[264,392,1818,1844]" pageId="9" pageNumber="32">Diagnosis.</emphasis>
This species is distinguished by the following combination of characters: antennal club bicolored; pronotum, elytra, and pygidium with dense vestiture of scale­like, mustard yellow setae; abdomen and legs black; tarsi with rings of white setae; parameres with lanceolate apex and long setae on the external borders; female with scale­like, mustard yellow setae only at the sides of the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC457FFB3177FFE79210AA0E4" lastPageId="11" lastPageNumber="34" pageId="10" pageNumber="33" type="description">
<paragraph id="8BD236C6C457FFB2177FFE7920B7A014" blockId="10.[264,1325,284,1870]" pageId="10" pageNumber="33">
<emphasis id="B919EAD4C457FFB2177FFE792683A7E7" bold="true" box="[312,667,403,429]" pageId="10" pageNumber="33">
Description.
<typeStatus id="54D68864C457FFB2179DFE792656A7E7" box="[474,590,403,429]" pageId="10" pageNumber="33" type="holotype">Holotype</typeStatus>
. Male
</emphasis>
: Body length 10.0 mm; humeral width
<quantity id="4C959B23C457FFB2123DFE7E20C1A7E4" box="[1146,1241,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="33" unit="mm" value="3.5">3.5 mm</quantity>
.
<emphasis id="B919EAD4C457FFB212A1FE7E2132A7E7" box="[1254,1322,404,429]" italics="true" pageId="10" pageNumber="33">Color</emphasis>
(
<figureCitation id="13562A43C457FFB21756FE562540A79C" box="[273,344,444,470]" captionStart="FIGURES 712" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 712. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="10" pageNumber="33">Fig. 8</figureCitation>
): clypeus, most of antennae, mouthparts, elytra, pygidium, last sternite, femora, and most of tibiae reddish­yellow; head, distal half of antennal club, pronotum, scutellum, pterothorax, abdomen, apex of tibiae, and tarsi black.
<emphasis id="B919EAD4C457FFB2153CFDE627CCA46F" box="[891,980,524,549]" italics="true" pageId="10" pageNumber="33">Surface</emphasis>
: clypeus and frons with scattered scale­like, yellow setae; pronotum with dense vestiture of scale­like, yellow, short setae that cover most of integument; elytra weakly striated without scale­like setae along sutural and lateral borders; elytral disc with recumbent scale­like, yellow, short setae that do not completely cover the integument; ventrally with short, white setae. Disc of pygidium with scale­like, yellow setae, surrounded by glabrous, wide margins.
<emphasis id="B919EAD4C457FFB21238FD3E20D0A4A7" box="[1151,1224,724,749]" italics="true" pageId="10" pageNumber="33">Head:</emphasis>
clypeus trapezoidal, anterior border nearly straight; antennal club yellow with apical half black; mentum elongate with scattered, longitudinally furrowed setae; anterior border straight; anterolateral emargination exposing basal segment of labial palpus.
<emphasis id="B919EAD4C457FFB21217FCA620BCA52F" box="[1104,1188,844,869]" italics="true" pageId="10" pageNumber="33">Thorax</emphasis>
: pronotum hexagonal, convex,
<quantity id="4C959B23C457FFB217BBFC9E2642A5C4" box="[508,602,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="33" unit="mm" value="3.2">3.2 mm</quantity>
length,
<quantity id="4C959B23C457FFB21485FC9E2738A5C4" box="[706,800,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="33" unit="mm" value="2.9">2.9 mm</quantity>
width; prosternal process short, narrowed, slightly depressed, with acute apex hidden in frontal view; scutellum elongate with rounded apex. Metasternum with two longitudinal, irregular rows of long, yellow setae between meso­ and metacoxae. Procoxa conical, prominent. Protibia with two rounded teeth at the apex of external border. Protibial articulated spur absent. Protarsus shortened with spine­like setae at the apex of each segment and rings of white setae. Meso­ and metafemur with short, spine­like setae scattered on ventral surface. Mesotibia widened toward the apex with two long, narrow, weakly curved, acute spurs, and long spine­like setae around the apex. Metatibia widened toward the apex with numerous preapical long, black setae on the inner surface and long, spine­like setae around the apex. Meso­ and metatarsi enlarged, each segment with rings of white setae on basal half and long spinelike setae around the apex. All tarsal claws narrowly cleft.
<emphasis id="B919EAD4C457FFB21587FAC62036A30F" box="[960,1070,1324,1349]" italics="true" pageId="10" pageNumber="33">Abdomen</emphasis>
: sternites 2­5 moderately concave at middle, each with 45 spine­like, erect, reddish, short setae at the sides of midline; length of sternite 5 twice the length of sternite 4, without setae near the posterior border; sternite 6 ovate with scattered white setae on the surface and numerous long, black setae on the apical border. Pygidium elongate, strongly convex.
<emphasis id="B919EAD4C457FFB21266FA262088A3AF" box="[1057,1168,1484,1509]" italics="true" pageId="10" pageNumber="33">Genitalia</emphasis>
: total length
<quantity id="4C959B23C457FFB2174FFA1E256DA044" box="[264,373,1524,1550]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="10" pageNumber="33" unit="mm" value="2.95">2.95 mm</quantity>
; parameres stout with long setae on the distal half of lateral borders, completely fused dorsobasally, widely curved toward the apex, horse­shoe shape in distal view with
<quantity id="4C959B23C457FFB2174FF9AE2568A014" box="[264,368,1604,1630]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="10" pageNumber="33" unit="mm" value="1.14">1.14 mm</quantity>
length and
<quantity id="4C959B23C457FFB217BDF9AE267AA014" box="[506,610,1604,1630]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="10" pageNumber="33" unit="mm" value="0.96">0.96 mm</quantity>
width; each apex clearly lanceolate (
<figureCitation id="13562A43C457FFB21250F9AE20B9A014" box="[1047,1185,1604,1630]" captionStart="FIGURES 1321" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 1321. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="10" pageNumber="33">Figs. 16­17</figureCitation>
).
</paragraph>
<paragraph id="8BD236C6C457FFB2177FF98627AEA104" blockId="10.[264,1325,284,1870]" pageId="10" pageNumber="33">
<emphasis id="B919EAD4C457FFB2177FF9862617A0CC" bold="true" box="[312,527,1644,1670]" pageId="10" pageNumber="33">
<typeStatus id="54D68864C457FFB2177FF98625BCA0CC" box="[312,420,1644,1670]" pageId="10" pageNumber="33" type="allotype">Allotype</typeStatus>
. Female
</emphasis>
: Body length
<quantity id="4C959B23C457FFB21485F9862702A0CC" box="[706,794,1644,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="33" unit="mm" value="11.0">11 mm</quantity>
; humeral width
<quantity id="4C959B23C457FFB215A4F9862032A0CC" box="[995,1066,1644,1670]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="33" unit="mm" value="4.0">4 mm</quantity>
(
<figureCitation id="13562A43C457FFB21278F9862081A0CC" box="[1087,1177,1644,1670]" captionStart="FIGURES 712" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 712. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="10" pageNumber="33">Fig. 11</figureCitation>
). Similar to male except as follows: pronotum
<quantity id="4C959B23C457FFB214D8F97E26E1A0E4" box="[671,761,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="33" unit="mm" value="2.8">2.8 mm</quantity>
in length,
<quantity id="4C959B23C457FFB2153FF97E27C9A0E4" box="[888,977,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="33" unit="mm" value="2.9">2.9 mm</quantity>
in width with scale­like dark yellow setae only at the sides of the pronotum; protibia with preapical articulated spur; abdominal sternites 24 convex at middle without erect, long setae; pygidium short, nearly triangular, weakly convex, slightly projected before the apex. Genital plates narrowed, apex rounded and external border briefly sinuate (
<figureCitation id="13562A43C457FFB21516F8DE27B0A104" box="[849,936,1844,1870]" captionStart="FIGURES 1321" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 1321. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="10" pageNumber="33">Fig. 18</figureCitation>
).
</paragraph>
<caption id="DF12664EC456FFB3174FFA7C258AA072" pageId="11" pageNumber="34" startId="11.[264,383,1430,1454]" targetBox="[319,1270,326,1406]" targetPageId="11">
<paragraph id="8BD236C6C456FFB3174FFA7C258AA072" blockId="11.[264,1323,1430,1592]" pageId="11" pageNumber="34">
<emphasis id="B919EAD4C456FFB3174FFA7C25C8A3E4" bold="true" box="[264,464,1430,1454]" pageId="11" pageNumber="34">FIGURES 1321.</emphasis>
Diagnostic structures of species of
<taxonomicName id="4C6D4D45C456FFB31513FA7D27EBA3E4" authorityName="Dejean" authorityYear="1821" box="[852,1011,1431,1454]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B919EAD4C456FFB31513FA7D27EBA3E4" box="[852,1011,1431,1454]" italics="true" pageId="11" pageNumber="34">Macrodactylus</emphasis>
</taxonomicName>
:
<taxonomicName id="4C6D4D45C456FFB31245FA7D2069A3E4" box="[1026,1137,1431,1454]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B919EAD4C456FFB31245FA7D2069A3E4" box="[1026,1137,1431,1454]" italics="true" pageId="11" pageNumber="34">M. tibialis</emphasis>
</taxonomicName>
, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view;
<taxonomicName id="4C6D4D45C456FFB31238FA50213CA39B" box="[1151,1316,1466,1489]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="hondurensis">
<emphasis id="B919EAD4C456FFB31238FA50213CA39B" box="[1151,1316,1466,1489]" italics="true" pageId="11" pageNumber="34">M. hondurensis</emphasis>
</taxonomicName>
, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view;
<taxonomicName id="4C6D4D45C456FFB3174FFA15258AA05C" box="[264,402,1535,1558]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B919EAD4C456FFB3174FFA15258AA05C" box="[264,402,1535,1558]" italics="true" pageId="11" pageNumber="34">M. montanus</emphasis>
</taxonomicName>
, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view.
</paragraph>
</caption>
<paragraph id="8BD236C6C456FFB3177FF986210AA0E4" blockId="11.[264,1322,1644,1870]" pageId="11" pageNumber="34">
<emphasis id="B919EAD4C456FFB3177FF98625B6A0CC" bold="true" box="[312,430,1644,1670]" pageId="11" pageNumber="34">Variation</emphasis>
(
<specimenCount id="9D6BFD4FC456FFB317FAF9862651A0CC" box="[445,585,1644,1670]" pageId="11" pageNumber="34" type="generic" typeStatus="paratypes">6 paratypes</specimenCount>
). Body length
<quantity id="4C959B23C456FFB314B5F9862792A0CC" box="[754,906,1644,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.15" metricValueMin="0.98" pageId="11" pageNumber="34" unit="mm" value="10.65" valueMax="11.5" valueMin="9.8">9.8­11.5 mm</quantity>
; humeral width
<quantity id="4C959B23C456FFB31201F98620CEA0CC" box="[1094,1238,1644,1670]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.1" metricValueMin="3.4" pageId="11" pageNumber="34" unit="mm" value="3.75" valueMax="4.1" valueMin="3.4">3.44.1 mm</quantity>
; antennal club dark and abdominal sternites reddish­yellow to dark yellow in some specimens.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC456FFB3177FF95626ACA104" pageId="11" pageNumber="34" type="materials_examined">
<paragraph id="8BD236C6C456FFB3177FF95626ACA104" blockId="11.[264,1322,1644,1870]" pageId="11" pageNumber="34">
<emphasis id="B919EAD4C456FFB3177FF9562634A09C" bold="true" box="[312,556,1724,1750]" pageId="11" pageNumber="34">Material examined</emphasis>
(
<specimenCount id="9D6BFD4FC456FFB31402F95626B2A09C" box="[581,682,1724,1750]" pageId="11" pageNumber="34" type="male">2 males</specimenCount>
,
<specimenCount id="9D6BFD4FC456FFB314F9F9562725A09C" box="[702,829,1724,1750]" pageId="11" pageNumber="34" type="female">6 females</specimenCount>
).
<materialsCitation id="3B053C9BC456FFB3151FF95625F8A16C" collectingDate="1995-03-02" collectionCode="R, CMNC" country="Honduras" county="San Pedro Sula" location="Danto Trail" municipality="Parque Nacional Cusuco" pageId="11" pageNumber="34" specimenCount="1" stateProvince="Cortes" typeStatus="holotype">
<typeStatus id="54D68864C456FFB3151FF95627DDA09C" box="[856,965,1724,1750]" pageId="11" pageNumber="34" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F37A7656C456FFB31263F95620D3A09C" box="[1060,1227,1724,1750]" name="Honduras" pageId="11" pageNumber="34">HONDURAS</collectingCountry>
.
<collectingRegion id="49A9F824C456FFB31299F9562597A0B4" country="Honduras" name="Cortes" pageId="11" pageNumber="34">Cortes department</collectingRegion>
,
<collectingCounty id="62B34E4AC456FFB317DCF90E2649A0B4" box="[411,593,1764,1790]" pageId="11" pageNumber="34">San Pedro Sula</collectingCounty>
,
<collectingMunicipality id="6BB6ACBCC456FFB3141AF90E2799A0B4" box="[605,897,1764,1790]" pageId="11" pageNumber="34">Parque Nacional Cusuco</collectingMunicipality>
,
<location id="8EB2601DC456FFB315CAF90E200FA0B4" LSID="urn:lsid:plazi:treatment:03C487D0C454FFB4174FF94D20A9A4D4:8EB2601DC456FFB315CAF90E200FA0B4" box="[909,1047,1764,1790]" country="Honduras" county="San Pedro Sula" municipality="Parque Nacional Cusuco" name="Danto Trail" pageId="11" pageNumber="34" stateProvince="Cortes">Danto Trail</location>
,
<date id="FFD31006C456FFB31264F90E20D2A0B4" box="[1059,1226,1764,1790]" pageId="11" pageNumber="34" value="1995-03-02">
<collectingDate id="EF97E9EEC456FFB31264F90E20D2A0B4" box="[1059,1226,1764,1790]" pageId="11" pageNumber="34" value="1995-03-02">2 March 1995</collectingDate>
</date>
, Col.
<collectionCode id="ED7CAE03C456FFB31348F90E213EA0B4" box="[1295,1318,1764,1790]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="34">R</collectionCode>
. Cordero (
<collectionCode id="ED7CAE03C456FFB3173CF8E625C0A16C" box="[379,472,1804,1830]" pageId="11" pageNumber="34">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B053C9BC456FFB317A9F8E626A8A104" collectingDate="1995-03-02" collectionCode="CMNC, MXAL" country="Honduras" county="San Pedro Sula" location="Danto Trail" municipality="Parque Nacional Cusuco" pageId="11" pageNumber="34" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Cortes" typeStatus="allotype">
<typeStatus id="54D68864C456FFB317A9F8E6264DA16C" box="[494,597,1804,1830]" pageId="11" pageNumber="34" type="allotype">Allotype</typeStatus>
female (
<collectionCode id="ED7CAE03C456FFB31484F8E62707A16C" box="[707,799,1804,1830]" pageId="11" pageNumber="34">CMNC</collectionCode>
),
<specimenCount id="9D6BFD4FC456FFB31571F8E62791A16C" box="[822,905,1804,1830]" pageId="11" pageNumber="34" type="male">1 male</specimenCount>
and
<specimenCount id="9D6BFD4FC456FFB3158FF8E62025A16C" box="[968,1085,1804,1830]" pageId="11" pageNumber="34" type="female">5 females</specimenCount>
<typeStatus id="54D68864C456FFB3120FF8E620A0A16C" box="[1096,1208,1804,1830]" pageId="11" pageNumber="34" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED7CAE03C456FFB3128CF8E6213EA16C" box="[1227,1318,1804,1830]" pageId="11" pageNumber="34">CMNC</collectionCode>
,
<collectionCode id="ED7CAE03C456FFB3174FF8DE2578A104" box="[264,352,1844,1870]" pageId="11" pageNumber="34">MXAL</collectionCode>
) with same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC451FFB4177FFEF127DFA43C" pageId="12" pageNumber="35" type="biology_ecology">
<paragraph id="8BD236C6C451FFB4177FFEF1213CA714" blockId="12.[264,1323,283,670]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFEF1258CA77F" bold="true" box="[312,404,283,309]" pageId="12" pageNumber="35">Biology</emphasis>
. This species inhabits secondary montane moist forest of
<taxonomicName id="4C6D4D45C451FFB4127BFEF62549A717" authority=", Liquidambar, Trema" authorityName="Liquidambar, Trema" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="12" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B919EAD4C451FFB4127BFEF62549A717" italics="true" pageId="12" pageNumber="35">Pinus, Liquidambar, Trema</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C6D4D45C451FFB417D6FEAE2657A714" box="[401,591,324,350]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="35" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B919EAD4C451FFB417D6FEAE25EBA717" box="[401,499,324,349]" italics="true" pageId="12" pageNumber="35">Quercus</emphasis>
species
</taxonomicName>
located at
<quantity id="4C959B23C451FFB4148AFEAE273EA714" box="[717,806,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="35" unit="m" value="1800.0">1800 m</quantity>
in altitude. Month of collection: March (8).
</paragraph>
<paragraph id="8BD236C6C451FFB4177FFE812102A7CC" blockId="12.[264,1323,283,670]" box="[312,1306,363,390]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFE812664A7CF" bold="true" box="[312,636,363,389]" pageId="12" pageNumber="35">Geographical distribution</emphasis>
.
<materialsCitation id="3B053C9BC451FFB414CCFE86210EA7CC" box="[651,1302,364,390]" country="Honduras" latitude="15.5" location="Mountains" longLatPrecision="1284" longitude="-88.083336" pageId="12" pageNumber="35" specimenCount="1">
<location id="8EB2601DC451FFB414CCFE862710A7CC" LSID="urn:lsid:plazi:treatment:03C487D0C454FFB4174FF94D20A9A4D4:8EB2601DC451FFB414CCFE862710A7CC" box="[651,776,364,390]" country="Honduras" latitude="15.5" longLatPrecision="1284" longitude="-88.083336" name="Mountains" pageId="12" pageNumber="35">Mountains</location>
in western
<collectingCountry id="F37A7656C451FFB415D6FE86201BA7CC" box="[913,1027,364,390]" name="Honduras" pageId="12" pageNumber="35">Honduras</collectingCountry>
(
<geoCoordinate id="EE595001C451FFB41253FE86209EA7CC" box="[1044,1158,364,390]" degrees="15" direction="north" minutes="30" orientation="latitude" pageId="12" pageNumber="35" precision="925" value="15.5">15° 30' N</geoCoordinate>
,
<geoCoordinate id="EE595001C451FFB412D5FE862115A7CC" box="[1170,1293,364,390]" degrees="88" direction="west" minutes="05" orientation="longitude" pageId="12" pageNumber="35" precision="925" value="-88.083336">88° 05' W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD236C6C451FFB4177FFE7927DFA43C" blockId="12.[264,1323,283,670]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFE792651A7E7" bold="true" box="[312,585,403,429]" pageId="12" pageNumber="35">Taxonomic comments</emphasis>
.
<taxonomicName id="4C6D4D45C451FFB41426FE7E27BEA7E7" box="[609,934,404,429]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="hondurensis">
<emphasis id="B919EAD4C451FFB41426FE7E27BEA7E7" box="[609,934,404,429]" italics="true" pageId="12" pageNumber="35">Macrodactylus hondurensis</emphasis>
</taxonomicName>
is included in the “
<taxonomicName id="4C6D4D45C451FFB412D7FE7E2535A79F" authorityName="&quot; (sensu Arce-Perez and Moron" authorityYear="2000" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="subspinosus">
<emphasis id="B919EAD4C451FFB412D7FE7E20B0A7E7" box="[1168,1192,404,429]" italics="true" pageId="12" pageNumber="35">M</emphasis>
.
<emphasis id="B919EAD4C451FFB412FEFE7E2535A79F" italics="true" pageId="12" pageNumber="35">subspinosus</emphasis>
</taxonomicName>
” group (
<emphasis id="B919EAD4C451FFB417DEFE5625C2A79F" box="[409,474,444,469]" italics="true" pageId="12" pageNumber="35">sensu</emphasis>
Arce­Pérez and
<bibRefCitation id="EFFC4B37C451FFB414D8FE562723A79C" author="Moron, M. A." box="[671,827,444,470]" pageId="12" pageNumber="35" pagination="123 - 239" refId="ref6864" refString="Arce-Perez R. &amp; Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) de Mexico y America del Norte. Acta Zoologica Mexicana (n. s), 79, 123 - 239." type="journal article" year="2000">Morón, 2000</bibRefCitation>
) because the dorsal surface and pygidium have scale­like setae, the metasternum has rows of long setae between meso­ and metacoxae, the largest mesotibial apical spur is curved, the tarsal segments have rings of white setae, the parameres have setae on the external borders, and the genital plates of the females are narrowed with the lateral borders not expanded.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC451FFB4177FFD6E20A9A4D4" box="[312,1201,644,670]" pageId="12" pageNumber="35" type="etymology">
<paragraph id="8BD236C6C451FFB4177FFD6E20A9A4D4" blockId="12.[264,1323,283,670]" box="[312,1201,644,670]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFD6E25A5A4D4" bold="true" box="[312,445,644,670]" pageId="12" pageNumber="35">Etymology</emphasis>
. Latinized name of the country where the species was collected.
</paragraph>
</subSubSection>
</treatment>
</document>