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<document id="A7D1B2AA4EFD3B1BAC07DCA2D61F8EF3" ID-CLB-Dataset="54097" ID-DOI="10.11646/megataxa.4.1.1" ID-GBIF-Dataset="7c9bc39d-9910-46f1-9558-c0647b9cbbdb" ID-ISSN="2703-3090" ID-Zenodo-Dep="4591138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614033145335" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2020" docId="03C487CBEFD23AB3FF4DF8BDFD89F903" docLanguage="en" docName="megataxa.4.1.1.pdf.imf" docOrigin="Megataxa 4 (1)" docSource="http://zoobank.org/3fdd970e-62f1-4f67-8cce-10870bdb3c01" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Pachypygus bisetiger Kim &amp; Boxshall 2020, sp. nov." docType="treatment" docVersion="11" lastPageNumber="135" masterDocId="FFFDFFB3EF513A35FFC5FFCEFFC5FFC1" masterDocTitle="Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="132" updateTime="1698930853666" updateUser="ExternalLinkService">
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<mods:title id="9A07D743FE9A5D478EC0B0FDA3D02BD1">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<treatment id="03C487CBEFD23AB3FF4DF8BDFD89F903" ID-DOI="http://doi.org/10.5281/zenodo.5828400" ID-GBIF-Taxon="194475159" ID-Zenodo-Dep="5828400" LSID="urn:lsid:plazi:treatment:03C487CBEFD23AB3FF4DF8BDFD89F903" httpUri="http://treatment.plazi.org/id/03C487CBEFD23AB3FF4DF8BDFD89F903" lastPageId="134" lastPageNumber="135" pageId="131" pageNumber="132">
<subSubSection id="C3776556EFD23AB6FF4DF8BDFE29F84F" box="[136,492,1907,1934]" pageId="131" pageNumber="132" type="nomenclature">
<paragraph id="8BD236DDEFD23AB6FF4DF8BDFE29F84F" blockId="131.[136,492,1907,1968]" box="[136,492,1907,1934]" pageId="131" pageNumber="132">
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<emphasis id="B919EACFEFD23AB6FF4DF8BDFE29F84F" bold="true" box="[136,492,1907,1934]" pageId="131" pageNumber="132">
<taxonomicName id="4C6D4D5EEFD23AB6FF4DF8BDFE42F84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[136,391,1907,1934]" class="Maxillopoda" family="Notodelphyidae" genus="Pachypygus" kingdom="Animalia" order="Cyclopoida" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="bisetiger" status="sp. nov.">
<emphasis id="B919EACFEFD23AB6FF4DF8BDFE42F84F" bold="true" box="[136,391,1907,1934]" italics="true" pageId="131" pageNumber="132">Pachypygusbisetiger</emphasis>
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<taxonomicNameLabel id="A22A57B4EFD23AB6FE4BF8BAFE29F84F" box="[398,492,1908,1934]" pageId="131" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3776556EFD23AB6FF4DF858FEDAF871" box="[136,287,1941,1968]" pageId="131" pageNumber="132" type="description">
<paragraph id="8BD236DDEFD23AB6FF4DF858FEDAF871" blockId="131.[136,492,1907,1968]" box="[136,287,1941,1968]" pageId="131" pageNumber="132">
(
<figureCitation id="13562A58EFD23AB6FF55F858FF2BF86E" box="[144,238,1941,1968]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Figs. 86</figureCitation>
,
<figureCitation id="13562A58EFD23AB6FF3FF858FED2F871" box="[250,279,1942,1968]" captionStart="FIGURE 87" captionStartId="133.[136,245,1907,1933]" captionTargetBox="[242,1356,222,1827]" captionTargetId="figure-18@133.[202,1386,185,1870]" captionTargetPageId="133" captionText="FIGURE 87. Pachypygus bisetiger sp. nov., female. A, paragnath; B, maxilla; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: A, G, 0.02 mm; BF, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591410" httpUri="https://zenodo.org/record/4591410/files/figure.png" pageId="131" pageNumber="132">87</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776556EFD23AB6FF4DF814FA88FECA" pageId="131" pageNumber="132" type="materials_examined">
<paragraph id="8BD236DDEFD23AB6FF4DF814FA88FECA" blockId="131.[136,778,2009,2070]" lastBlockId="131.[810,1452,173,2070]" pageId="131" pageNumber="132">
<emphasis id="B919EACFEFD23AB6FF4DF814FEF3F832" bold="true" box="[136,310,2009,2036]" pageId="131" pageNumber="132">Typematerial</emphasis>
.
<materialsCitation id="3B053C80EFD23AB6FE89F814FF1FF7D7" ID-GBIF-Occurrence="3748446343" pageId="131" pageNumber="132" specimenCode="MNHN-IU-2014- 21243" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54D6887FEFD23AB6FE89F814FE71F835" box="[332,436,2010,2036]" pageId="131" pageNumber="132" type="holotype">Holotype</typeStatus>
(intact
<specimenCount id="9D6BFD54EFD23AB6FDDDF814FDE8F835" box="[536,557,2010,2036]" pageId="131" pageNumber="132" type="female"></specimenCount>
,
<specimenCode id="DBCB9EA6EFD23AB6FD86F814FF17F7D7" pageId="131" pageNumber="132">MNHN-IU-2014- 21243</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B053C80EFD23AB6FF34F832FA88FECA" ID-GBIF-Occurrence="3748446367" pageId="131" pageNumber="132" typeStatus="paratype">
<typeStatus id="54D6887FEFD23AB6FF34F832FE94F7D7" box="[241,337,2044,2070]" pageId="131" pageNumber="132" type="paratype">paratype</typeStatus>
(intact
<specimenCount id="9D6BFD54EFD23AB6FE7BF832FE16F7D7" box="[446,467,2044,2070]" pageId="131" pageNumber="132"></specimenCount>
, MNHN-IU-2014-21244), and
<typeStatus id="54D6887FEFD23AB6FC9EFF63FBEFFF06" box="[859,1066,173,199]" pageId="131" pageNumber="132">dissectedparatype</typeStatus>
(
<specimenCount id="9D6BFD54EFD23AB6FBFAFF63FB97FF06" box="[1087,1106,173,199]" pageId="131" pageNumber="132"></specimenCount>
, figured), from the compound ascidian
<taxonomicName id="4C6D4D5EEFD23AB6FC55FF01FACFFF28" authority="(Sluiter, 1904)" baseAuthorityName="Sluiter" baseAuthorityYear="1904" box="[912,1290,207,233]" class="Ascidiacea" family="Clavelinidae" genus="Clavelina" kingdom="Animalia" order="Enterogona" pageId="131" pageNumber="132" phylum="Chordata" rank="species" species="detorta">
<emphasis id="B919EACFEFD23AB6FC55FF01FB9DFF28" box="[912,1112,207,233]" italics="true" pageId="131" pageNumber="132">Clavelina detorta</emphasis>
(Sluiter, 1904)
</taxonomicName>
, lagoon,
<collectingCountry id="F37A764DEFD23AB6FAB3FF01FC5BFECA" name="New Caledonia" pageId="131" pageNumber="132">New Caledonia</collectingCountry>
, B. Richer de Forges -
<collectionCode id="ED7CAE18EFD23AB6FB65FF3FFAD1FECA" box="[1184,1300,241,267]" country="Bolivia" httpUri="http://grbio.org/cool/mdr4-z094" name="Office de la Recherche scientifique et Technique Outre-mer" pageId="131" pageNumber="132">ORSTOM</collectionCode>
coll.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3776556EFD23AB6FC9AFEDDFA50FE8E" pageId="131" pageNumber="132" type="etymology">
<paragraph id="8BD236DDEFD23AB6FC9AFEDDFA50FE8E" blockId="131.[810,1452,173,2070]" pageId="131" pageNumber="132">
<emphasis id="B919EACFEFD23AB6FC9AFEDDFC25FEEC" bold="true" box="[863,992,275,301]" pageId="131" pageNumber="132">Etymology</emphasis>
. The specific name refers to the presence of the two setae on the third segment of the maxilliped.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EFD23AB3FC9AFE99FEBBFA8A" lastPageId="134" lastPageNumber="135" pageId="131" pageNumber="132" type="description">
<paragraph id="8BD236DDEFD23AB6FC9AFE99FA48FC14" blockId="131.[810,1452,173,2070]" pageId="131" pageNumber="132">
<emphasis id="B919EACFEFD23AB6FC9AFE99FBA7FEB0" bold="true" box="[863,1122,343,369]" pageId="131" pageNumber="132">Descriptionoffemale</emphasis>
. Body (
<figureCitation id="13562A58EFD23AB6FB04FE99FAEDFEB3" box="[1217,1320,343,370]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86A</figureCitation>
) bilaterally compressed. Dorsal shield of cephalosome with short, broad posterolateral processes and convex dorsal protrusion. Dorsal tergites of first to third prosomites incompletely developed, with unsclerotized gaps between them. Third and fourth pedigerous somites forming brood pouch; fourth pedigeroussomite suboval. Urosome (
<figureCitation id="13562A58EFD23AB6FABBFDEDFC9CFDA1" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86B</figureCitation>
) 6-segmented, graduallynarrowing distally. Fifth pedigerous somite short, 90×242 μm, but well-defined from fourth pedigerous somite. Genital and 4 free abdominalsomites 155×215, 181×176, 167×158, and 121×124, and 67×94 μm, respectively. Anal somite (
<figureCitation id="13562A58EFD23AB6FABBFD03FC9CFCCB" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86C</figureCitation>
) with highly sclerotized posteroventral protrusion and short dorsal margin. Caudal ramus (
<figureCitation id="13562A58EFD23AB6FB27FCDFFA80FCED" box="[1250,1349,785,812]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86C</figureCitation>
) inserted dorsolaterally on anal somite, about 2.5 times longer than wide (114×45 μm), narrowing distally, and armed with 4 distal claws (1 large and 3 small) and 2 setae; lengths of claws 35, 20, 17, and 14 μm; setae as long as distal width of ramus, positioned at 56% and 75% of ramus length.
</paragraph>
<paragraph id="8BD236DDEFD23AB6FC9AFC10FB95FA73" blockId="131.[810,1452,173,2070]" pageId="131" pageNumber="132">
Rostrum (
<figureCitation id="13562A58EFD23AB6FC15FC13FBF2FC39" box="[976,1079,989,1016]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86D</figureCitation>
) bulbous, narrowing proximally, widest in middle, rounded distally and with large pore at apex. Antennule (
<figureCitation id="13562A58EFD23AB6FC28FBEFFB8AFBFD" box="[1005,1103,1057,1084]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86E</figureCitation>
) 9-segmented, secondsegment widest, more distal segments narrowing gradually; fifth segment subdivided by incomplete suture line on posterior side; armatureformula 3, 16, 5, 2+ 2 aesthetascs, 4, 2+aesthetasc, 2, 2+aesthetasc, and 7+aesthetasc; all setae naked and most setae long. Antenna (
<figureCitation id="13562A58EFD23AB6FB34FB05FA9CFB27" box="[1265,1369,1227,1254]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86F</figureCitation>
) stout, 4-segmented; coxaandbasisunarmed; firstendopodal segment slightly shorter than basis, with 1 seta on inner margin; compound distal endopodal segment slightly shorter than first, twice as long as wide (61×32 μm); armedwith 8 setae (grouped as 3, 2, and 3) plusterminal claw longerthan segment.
</paragraph>
<paragraph id="8BD236DDEFD23AB3FC9AFA77FD3EFE52" blockId="131.[810,1452,173,2070]" lastBlockId="134.[136,778,173,846]" lastPageId="134" lastPageNumber="135" pageId="131" pageNumber="132">
Labrum (
<figureCitation id="13562A58EFD23AB6FC18FA77FB9CFA15" box="[989,1113,1465,1492]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86G</figureCitation>
) with large semicircular posterolateral lobes ornamented with setules on outer marginand inner side; convex posteriormargin setulose. Mandible (
<figureCitation id="13562A58EFD23AB6FC69F9D1FBDDF9FB" box="[940,1048,1567,1594]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86H</figureCitation>
) with 6 teeth, including 4 blunt, indistinct proximal ones, and 2 smallsetae on coxal gnathobase; basiswith 1 setaonmedial margin; exopod with 5 setae, outer 2 distinctly larger than 3 inner setae; endopod with 4 and 9 setaeon first and second segments, respectively; 2 outer distal setae on second endopodal segment subequal in length, third much larger than other 8. Paragnath (
<figureCitation id="13562A58EFD23AB6FC15F8C3FBF9F8E9" box="[976,1084,1805,1832]" captionStart="FIGURE 87" captionStartId="133.[136,245,1907,1933]" captionTargetBox="[242,1356,222,1827]" captionTargetId="figure-18@133.[202,1386,185,1870]" captionTargetPageId="133" captionText="FIGURE 87. Pachypygus bisetiger sp. nov., female. A, paragnath; B, maxilla; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: A, G, 0.02 mm; BF, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591410" httpUri="https://zenodo.org/record/4591410/files/figure.png" pageId="131" pageNumber="132">Fig. 87A</figureCitation>
) as usual for genus. Maxillule (
<figureCitation id="13562A58EFD23AB6FCF7F8E1FC4BF88B" box="[818,910,1839,1866]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="131" pageNumber="132">Fig. 86I</figureCitation>
) with 10 setae on arthrite, 1 on coxal endite, 2 on epipodite; basiswith 4 setaeon medial margin, second proximal seta larger than other 3; exopod with 4 setae distally; endopod 2-segmented, with 3 smallsetae on first, and 2 (1 long, naked and 1 short pinnate) setae on second segment. Maxilla (
<figureCitation id="13562A58EFD23AB6FBC5F814FBA3F835" box="[1024,1126,2010,2036]" captionStart="FIGURE 87" captionStartId="133.[136,245,1907,1933]" captionTargetBox="[242,1356,222,1827]" captionTargetId="figure-18@133.[202,1386,185,1870]" captionTargetPageId="133" captionText="FIGURE 87. Pachypygus bisetiger sp. nov., female. A, paragnath; B, maxilla; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: A, G, 0.02 mm; BF, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591410" httpUri="https://zenodo.org/record/4591410/files/figure.png" pageId="131" pageNumber="132">Fig. 87B</figureCitation>
) 6-segmented; precoxa with 4 and 1 setae on first and second endites, respectively; coxa incompletely articulated from precoxa, with 2 and 3 setae on first and second endites, respectively; basis with slender, smooth claw plus 2 setae; endopod slender with 1, 1, and 4 setae on first to third segments, respectively. Maxilliped (
<figureCitation id="13562A58EFD73AB3FED4FEFBFEA9FE91" box="[273,364,309,336]" captionStart="FIGURE 86" captionStartId="132.[136,245,1907,1933]" captionTargetBox="[245,1344,214,1837]" captionTargetId="figure-19@132.[202,1386,185,1870]" captionTargetPageId="132" captionText="FIGURE 86. Pachypygus bisetiger sp. nov., female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="134" pageNumber="135">Fig. 86J</figureCitation>
) 3-segmented, armedwith 9, 1, and 2 setae onfirst to third segments, respectively; thirdsegment small; articulation incomplete between first 2 segments.
</paragraph>
<caption id="DF126655EFD53AB1FF4DF8BDFE3AF813" ID-DOI="http://doi.org/10.5281/zenodo.4591406" ID-Zenodo-Dep="4591406" httpUri="https://zenodo.org/record/4591406/files/figure.png" pageId="132" pageNumber="133" startId="132.[136,245,1907,1933]" targetBox="[245,1344,214,1837]" targetPageId="132">
<paragraph id="8BD236DDEFD53AB1FF4DF8BDFE3AF813" blockId="132.[136,1451,1907,2002]" pageId="132" pageNumber="133">
<emphasis id="B919EACFEFD53AB1FF4DF8BDFEDAF84F" bold="true" box="[136,287,1907,1934]" pageId="132" pageNumber="133">FIGURE 86.</emphasis>
<taxonomicName id="4C6D4D5EEFD53AB1FEE3F8BAFDD2F84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[294,535,1908,1934]" class="Maxillopoda" family="Notodelphyidae" genus="Pachypygus" kingdom="Animalia" order="Cyclopoida" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="bisetiger" status="sp. nov.">
<emphasis id="B919EACFEFD53AB1FEE3F8BAFDD2F84F" box="[294,535,1908,1934]" italics="true" pageId="132" pageNumber="133">Pachypygus bisetiger</emphasis>
</taxonomicName>
<emphasis id="B919EACFEFD53AB1FDDBF8BAFDBDF84F" bold="true" box="[542,632,1908,1934]" pageId="132" pageNumber="133">
<taxonomicNameLabel id="A22A57B4EFD53AB1FDDBF8BAFDBDF84F" box="[542,632,1908,1934]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, habitus, right; B, urosome, ventral C, distal part of abdomen, right; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilliped. A, 0.5 mm; B, 0.1 mm; CF, HJ, 0.05 mm; G, 0.02 mm.
</paragraph>
</caption>
<caption id="DF126655EFD43AB0FF4DF8BDFD6EF871" ID-DOI="http://doi.org/10.5281/zenodo.4591410" ID-Zenodo-Dep="4591410" httpUri="https://zenodo.org/record/4591410/files/figure.png" pageId="133" pageNumber="134" startId="133.[136,245,1907,1933]" targetBox="[242,1356,222,1827]" targetPageId="133">
<paragraph id="8BD236DDEFD43AB0FF4DF8BDFD6EF871" blockId="133.[136,1452,1907,1968]" pageId="133" pageNumber="134">
<emphasis id="B919EACFEFD43AB0FF4DF8BDFEDAF84F" bold="true" box="[136,287,1907,1934]" pageId="133" pageNumber="134">FIGURE 87.</emphasis>
<taxonomicName id="4C6D4D5EEFD43AB0FEE0F8BAFDD3F84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[293,534,1908,1934]" class="Maxillopoda" family="Notodelphyidae" genus="Pachypygus" kingdom="Animalia" order="Cyclopoida" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="bisetiger" status="sp. nov.">
<emphasis id="B919EACFEFD43AB0FEE0F8BAFDD3F84F" box="[293,534,1908,1934]" italics="true" pageId="133" pageNumber="134">Pachypygusbisetiger</emphasis>
</taxonomicName>
<emphasis id="B919EACFEFD43AB0FDD9F8BAFDB3F84F" bold="true" box="[540,630,1908,1934]" pageId="133" pageNumber="134">
<taxonomicNameLabel id="A22A57B4EFD43AB0FDD9F8BAFDB3F84F" box="[540,630,1908,1934]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, paragnath; B, maxilla; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: A, G, 0.02 mm; BF, 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BD236DDEFD73AB3FF78FE55FEC6FC8C" blockId="134.[136,778,173,846]" pageId="134" pageNumber="135">
Leg 1 (
<figureCitation id="13562A58EFD73AB3FED7FE55FEBDFE74" box="[274,376,411,437]" captionStart="FIGURE 87" captionStartId="133.[136,245,1907,1933]" captionTargetBox="[242,1356,222,1827]" captionTargetId="figure-18@133.[202,1386,185,1870]" captionTargetPageId="133" captionText="FIGURE 87. Pachypygus bisetiger sp. nov., female. A, paragnath; B, maxilla; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: A, G, 0.02 mm; BF, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591410" httpUri="https://zenodo.org/record/4591410/files/figure.png" pageId="134" pageNumber="135">Fig. 87C</figureCitation>
) with 3-segmented rami. Legs 24 (
<figureCitation id="13562A58EFD73AB3FF55FE73FF2CFE16" box="[144,233,445,471]" captionStart="FIGURE 87" captionStartId="133.[136,245,1907,1933]" captionTargetBox="[242,1356,222,1827]" captionTargetId="figure-18@133.[202,1386,185,1870]" captionTargetPageId="133" captionText="FIGURE 87. Pachypygus bisetiger sp. nov., female. A, paragnath; B, maxilla; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: A, G, 0.02 mm; BF, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591410" httpUri="https://zenodo.org/record/4591410/files/figure.png" pageId="134" pageNumber="135">Fig. 87</figureCitation>
DF) each with 3-segmented exopod and 2- segmented endopod. Inner seta on coxa well-developed in leg 1, rudimentary in leg 3, but apparently absent in legs 2 and 4. Outer seta on basis broad and pinnate in leg 1, but small and naked in legs 24. Inner distal spine on basis of leg 1 acutely pointed at tip, 39 μm long, slightly longer than first endopodal segment. Outer spines on exopods of legs 24 slender, but swollen distally. First exopodal segment of legs 24 elongated, about 3.3 times longer than wide in legs 2 and 3 and 2.5 times in leg 4. Setae on legs 24 sparsely pinnate. Armature formula for legs 14 as follows:
</paragraph>
<paragraph id="8BD236DDEFD73AB3FF2DFCB9FD1CFBD8" pageId="134" pageNumber="135">
<table id="F96DC47DEFD7C5CAFF4DFCB9FCCDFBDB" box="[136,776,887,1050]" gridcols="5" gridrows="5" inLine="true" pageId="134" pageNumber="135">
<tr id="355D349FEFD7C5CAFF4DFCB9FCCDFC50" box="[136,776,887,913]" gridrow="0" pageId="134" pageNumber="135" rowspan-0="1">
<th id="768C5DE3EFD7C5CAFF2DFCB9FEE6FC50" box="[232,291,887,913]" gridcol="1" gridrow="0" pageId="134" pageNumber="135">Coxa</th>
<th id="768C5DE3EFD7C5CAFEFBFCB9FEB9FC50" box="[318,380,887,913]" gridcol="2" gridrow="0" pageId="134" pageNumber="135">Basis</th>
<th id="768C5DE3EFD7C5CAFE50FCB9FD80FC50" box="[405,581,887,913]" gridcol="3" gridrow="0" pageId="134" pageNumber="135">Exopod</th>
<th id="768C5DE3EFD7C5CAFD9FFCB9FCCDFC50" box="[602,776,887,913]" gridcol="4" gridrow="0" pageId="134" pageNumber="135">Endopod</th>
</tr>
<tr id="355D349FEFD7C5CAFF4DFC57FCCDFC75" box="[136,776,921,948]" gridrow="1" pageId="134" pageNumber="135">
<th id="768C5DE3EFD7C5CAFF4DFC57FF0CFC75" box="[136,201,921,948]" gridcol="0" gridrow="1" pageId="134" pageNumber="135">Leg 1</th>
<td id="768C5DE3EFD7C5CAFF2DFC57FEE6FC75" box="[232,291,921,948]" gridcol="1" gridrow="1" pageId="134" pageNumber="135">0-1</td>
<td id="768C5DE3EFD7C5CAFEFBFC57FEB9FC75" box="[318,380,921,948]" gridcol="2" gridrow="1" pageId="134" pageNumber="135">1-I</td>
<td id="768C5DE3EFD7C5CAFE50FC57FD80FC75" box="[405,581,921,948]" gridcol="3" gridrow="1" pageId="134" pageNumber="135">I-1; I-1; III, I, 4</td>
<td id="768C5DE3EFD7C5CAFD9FFC57FCCDFC75" box="[602,776,921,948]" gridcol="4" gridrow="1" pageId="134" pageNumber="135">0-1; 0-1; 1, 2, 3</td>
</tr>
<tr id="355D349FEFD7C5CAFF4DFC75FCCDFC17" box="[136,776,955,982]" gridrow="2" pageId="134" pageNumber="135">
<th id="768C5DE3EFD7C5CAFF4DFC75FF0CFC17" box="[136,201,955,982]" gridcol="0" gridrow="2" pageId="134" pageNumber="135">Leg 2</th>
<td id="768C5DE3EFD7C5CAFF2DFC75FEE6FC17" box="[232,291,955,982]" gridcol="1" gridrow="2" pageId="134" pageNumber="135">0-0</td>
<td id="768C5DE3EFD7C5CAFEFBFC75FEB9FC17" box="[318,380,955,982]" gridcol="2" gridrow="2" pageId="134" pageNumber="135">1-0</td>
<td id="768C5DE3EFD7C5CAFE50FC75FD80FC17" box="[405,581,955,982]" gridcol="3" gridrow="2" pageId="134" pageNumber="135">I-1; I-1; II, II, 0</td>
<td id="768C5DE3EFD7C5CAFD9FFC75FCCDFC17" box="[602,776,955,982]" gridcol="4" gridrow="2" pageId="134" pageNumber="135">0-1; 1, 2, 5</td>
</tr>
<tr id="355D349FEFD7C5CAFF4DFC13FCCDFC39" box="[136,776,989,1016]" gridrow="3" pageId="134" pageNumber="135">
<th id="768C5DE3EFD7C5CAFF4DFC13FF0CFC39" box="[136,201,989,1016]" gridcol="0" gridrow="3" pageId="134" pageNumber="135">Leg 3</th>
<td id="768C5DE3EFD7C5CAFF2DFC13FEE6FC39" box="[232,291,989,1016]" gridcol="1" gridrow="3" pageId="134" pageNumber="135">0-1</td>
<td id="768C5DE3EFD7C5CAFEFBFC13FEB9FC39" box="[318,380,989,1016]" gridcol="2" gridrow="3" pageId="134" pageNumber="135">1-0</td>
<td id="768C5DE3EFD7C5CAFE50FC13FD80FC39" box="[405,581,989,1016]" gridcol="3" gridrow="3" pageId="134" pageNumber="135">I-1; I-0; II, II, 0</td>
<td id="768C5DE3EFD7C5CAFD9FFC13FCCDFC39" box="[602,776,989,1016]" gridcol="4" gridrow="3" pageId="134" pageNumber="135">0-1; 1, 2, 5</td>
</tr>
<tr id="355D349FEFD7C5CAFF4DFC31FCCDFBDB" box="[136,776,1023,1050]" gridrow="4" pageId="134" pageNumber="135">
<th id="768C5DE3EFD7C5CAFF4DFC31FF0CFBDB" box="[136,201,1023,1050]" gridcol="0" gridrow="4" pageId="134" pageNumber="135">Leg 4</th>
<td id="768C5DE3EFD7C5CAFF2DFC31FEE6FBDB" box="[232,291,1023,1050]" gridcol="1" gridrow="4" pageId="134" pageNumber="135">0-0</td>
<td id="768C5DE3EFD7C5CAFEFBFC31FEB9FBDB" box="[318,380,1023,1050]" gridcol="2" gridrow="4" pageId="134" pageNumber="135">1-0</td>
<td id="768C5DE3EFD7C5CAFE50FC31FD80FBDB" box="[405,581,1023,1050]" gridcol="3" gridrow="4" pageId="134" pageNumber="135">I-1; I-0; II, II, 0</td>
<td id="768C5DE3EFD7C5CAFD9FFC31FCCDFBDB" box="[602,776,1023,1050]" gridcol="4" gridrow="4" pageId="134" pageNumber="135">0-1; 1, 2, 4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDEFD73AB3FF78FB8AFD7DFAE8" blockId="134.[136,778,1091,1730]" pageId="134" pageNumber="135">
Leg 5 (
<figureCitation id="13562A58EFD73AB3FEC8FB8DFEB7FB9C" box="[269,370,1091,1117]" captionStart="FIGURE 87" captionStartId="133.[136,245,1907,1933]" captionTargetBox="[242,1356,222,1827]" captionTargetId="figure-18@133.[202,1386,185,1870]" captionTargetPageId="133" captionText="FIGURE 87. Pachypygus bisetiger sp. nov., female. A, paragnath; B, maxilla; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: A, G, 0.02 mm; BF, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591410" httpUri="https://zenodo.org/record/4591410/files/figure.png" pageId="134" pageNumber="135">Fig. 87G</figureCitation>
) 2-segmented. Protopod longer than wide, fused with somite, armed with 1 naked seta at outer distal corner and ornamented with row of several minute spinules near inner distal corner; free exopodal segment about 3.4 timeslongerthan wide (65×19 μm), armed distally with 1 small spine and 1 naked seta; ornamented with 3 rows of minute spinules on medial surface.
</paragraph>
<paragraph id="8BD236DDEFD73AB3FC71FB05FA49FA8A" pageId="134" pageNumber="135">
<table id="F96DC47DEFD7C5CAFCEFFB05FA4BFA8D" box="[810,1422,1227,1356]" gridcols="5" gridrows="4" inLine="true" pageId="134" pageNumber="135">
<tr id="355D349FEFD7C5CAFCEFFB05FA4BFB24" box="[810,1422,1227,1253]" gridrow="0" pageId="134" pageNumber="135" rowspan-0="1">
<th id="768C5DE3EFD7C5CAFC71FB05FC2AFB24" box="[948,1007,1227,1253]" gridcol="1" gridrow="0" pageId="134" pageNumber="135">Coxa</th>
<th id="768C5DE3EFD7C5CAFC39FB05FBFFFB24" box="[1020,1082,1227,1253]" gridcol="2" gridrow="0" pageId="134" pageNumber="135">Basis</th>
<th id="768C5DE3EFD7C5CAFB82FB05FB3AFB24" box="[1095,1279,1227,1253]" gridcol="3" gridrow="0" pageId="134" pageNumber="135">Exopod</th>
<th id="768C5DE3EFD7C5CAFAC9FB05FA4BFB24" box="[1292,1422,1227,1253]" gridcol="4" gridrow="0" pageId="134" pageNumber="135">Endopod</th>
</tr>
<tr id="355D349FEFD7C5CAFCEFFB23FA4BFAC9" box="[810,1422,1261,1288]" gridrow="1" pageId="134" pageNumber="135">
<th id="768C5DE3EFD7C5CAFCEFFB23FC62FAC9" box="[810,935,1261,1288]" gridcol="0" gridrow="1" pageId="134" pageNumber="135">Leg 1</th>
<td id="768C5DE3EFD7C5CAFC71FB23FC2AFAC9" box="[948,1007,1261,1288]" gridcol="1" gridrow="1" pageId="134" pageNumber="135">0-1</td>
<td id="768C5DE3EFD7C5CAFC39FB23FBFFFAC9" box="[1020,1082,1261,1288]" gridcol="2" gridrow="1" pageId="134" pageNumber="135">1-I</td>
<td id="768C5DE3EFD7C5CAFB82FB23FB3AFAC9" box="[1095,1279,1261,1288]" gridcol="3" gridrow="1" pageId="134" pageNumber="135">I-1; I-1; III, I, 4</td>
<td id="768C5DE3EFD7C5CAFAC9FB23FA4BFAC9" box="[1292,1422,1261,1288]" gridcol="4" gridrow="1" pageId="134" pageNumber="135">0-1; 1, 2, 4</td>
</tr>
<tr id="355D349FEFD7C5CAFCEFFAC1FA4BFAEB" box="[810,1422,1295,1322]" gridrow="2" pageId="134" pageNumber="135">
<th id="768C5DE3EFD7C5CAFCEFFAC1FC62FAEB" box="[810,935,1295,1322]" gridcol="0" gridrow="2" pageId="134" pageNumber="135">
Legs 2 &amp;
<td id="768C5DE3EFD7C5CAFC71FAC1FC2AFAEB" box="[948,1007,1295,1322]" gridcol="1" gridrow="2" pageId="134" pageNumber="135">30-1</td>
</th>
<td id="768C5DE3EFD7C5CAFC39FAC1FBFFFAEB" box="[1020,1082,1295,1322]" gridcol="2" gridrow="2" pageId="134" pageNumber="135">1-0</td>
<td id="768C5DE3EFD7C5CAFB82FAC1FB3AFAEB" box="[1095,1279,1295,1322]" gridcol="3" gridrow="2" pageId="134" pageNumber="135">I-1; I-1; III, I, 0</td>
<td id="768C5DE3EFD7C5CAFAC9FAC1FA4BFAEB" box="[1292,1422,1295,1322]" gridcol="4" gridrow="2" pageId="134" pageNumber="135">0-1; 1, 2, 5</td>
</tr>
<tr id="355D349FEFD7C5CAFCEFFAFFFA4BFA8D" box="[810,1422,1329,1356]" gridrow="3" pageId="134" pageNumber="135">
<th id="768C5DE3EFD7C5CAFCEFFAFFFC62FA8D" box="[810,935,1329,1356]" gridcol="0" gridrow="3" pageId="134" pageNumber="135">Leg 4</th>
<td id="768C5DE3EFD7C5CAFC71FAFFFC2AFA8D" box="[948,1007,1329,1356]" gridcol="1" gridrow="3" pageId="134" pageNumber="135">0-1</td>
<td id="768C5DE3EFD7C5CAFC39FAFFFBFFFA8D" box="[1020,1082,1329,1356]" gridcol="2" gridrow="3" pageId="134" pageNumber="135">1-0</td>
<td id="768C5DE3EFD7C5CAFB82FAFFFB3AFA8D" box="[1095,1279,1329,1356]" gridcol="3" gridrow="3" pageId="134" pageNumber="135">I-0; I-1; II, I,0</td>
<td id="768C5DE3EFD7C5CAFAC9FAFFFA4BFA8D" box="[1292,1422,1329,1356]" gridcol="4" gridrow="3" pageId="134" pageNumber="135">0-1; 1, 2, 4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDEFD73AB3FF78FAFFFEBBFA8A" blockId="134.[136,778,1091,1730]" box="[189,382,1329,1356]" pageId="134" pageNumber="135">
<emphasis id="B919EACFEFD73AB3FF78FAFFFF3FFA8A" bold="true" box="[189,250,1329,1355]" pageId="134" pageNumber="135">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EFD73AB3FF78FA9DFD89F903" pageId="134" pageNumber="135" type="discussion">
<paragraph id="8BD236DDEFD73AB3FF78FA9DFD89F903" blockId="134.[136,778,1091,1730]" pageId="134" pageNumber="135">
<emphasis id="B919EACFEFD73AB3FF78FA9DFEEFFAAC" bold="true" box="[189,298,1363,1389]" pageId="134" pageNumber="135">Remarks</emphasis>
.
<taxonomicName id="4C6D4D5EEFD73AB3FEFEFA9DFDEAFAAC" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[315,559,1363,1389]" class="Maxillopoda" family="Notodelphyidae" genus="Pachypygus" kingdom="Animalia" order="Cyclopoida" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="bisetiger" status="sp. nov.">
<emphasis id="B919EACFEFD73AB3FEFEFA9DFDEAFAAC" box="[315,559,1363,1389]" italics="true" pageId="134" pageNumber="135">Pachypygus bisetiger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EFD73AB3FDFCFA9DFD52FAAF" box="[569,663,1363,1390]" pageId="134" pageNumber="135" rank="species">
<emphasis id="B919EACFEFD73AB3FDFCFA9DFD55FAAC" bold="true" box="[569,656,1363,1389]" pageId="134" pageNumber="135">sp. nov</emphasis>
.
</taxonomicNameLabel>
is unique in the genus in having only 2 setae on the third segment of the maxilliped; all other species have either 3 or 4 setae. In addition, there are only 2 setaeon the second endopodal segment of the maxillule in the new species, whereas there are 3 setae on this segment in all other described species. The 2-segmented endopod of legs
<quantity id="4C959B38EFD73AB3FD25F9EEFF5BF99A" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="134" pageNumber="135" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">24 in</quantity>
the new species is also unique: all other species have 3-segmented endopods on these legs. The distal segments of the 2-segmented endopods are compound, representing the second and third ancestral segments.
</paragraph>
</subSubSection>
</treatment>
</document>