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<document id="B839761A15F8513BEE29478A634501A0" ID-CLB-Dataset="22479" ID-DOI="10.5281/zenodo.3776222" ID-GBIF-Dataset="c1129e00-0019-4411-9efb-a9d6c83fcf9d" ID-ISSN="0253-116X" ID-Zenodo-Dep="3776222" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588180845528" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2018" docId="03C487B82908B836CE8FF920FE6974AE" docLanguage="en" docName="Linzer.biol.Beitr.50.2.1015-1032.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Myllaena graeca Kraatz 1858" docType="treatment" docVersion="7" lastPageNumber="1018" masterDocId="FFFDFFC0290AB835CE3DF876FFAB7050" masterDocTitle="On some Myllaena species in the East Mediterranean and Caucasus regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1032" masterPageNumber="1015" pageNumber="1017" updateTime="1698838018870" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0029508106F1DF7E0D172B317F97D58B">On some Myllaena species in the East Mediterranean and Caucasus regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="20ACA2975062550B0DDCCF5B3057B810">Assing, Volker</mods:namePart>
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<mods:date id="DC3B700F56B2D169558890901BBB140B">2018</mods:date>
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<mods:number id="BDC2A62E101A19291D8261D8C826453F">2018-12-17</mods:number>
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<treatment id="03C487B82908B836CE8FF920FE6974AE" ID-DOI="http://doi.org/10.5281/zenodo.3796591" ID-GBIF-Taxon="163876662" ID-Zenodo-Dep="3796591" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE" httpUri="http://treatment.plazi.org/id/03C487B82908B836CE8FF920FE6974AE" lastPageId="3" lastPageNumber="1018" pageId="2" pageNumber="1017">
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<paragraph id="8BD236AE2908B837CE8FF920FD13713C" blockId="2.[178,696,342,364]" box="[178,696,342,364]" pageId="2" pageNumber="1017">
<heading id="D09A81C22908B837CE8FF920FD13713C" box="[178,696,342,364]" fontSize="8" level="2" pageId="2" pageNumber="1017" reason="2">
<taxonomicName id="4C6D4D2D2908B837CE8FF920FE54713B" ID-CoL="453HB" authority="KRAATZ, 1858" authorityName="Kraatz" authorityYear="1858" box="[178,511,342,364]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CE8FF920FE6A713B" bold="true" box="[178,449,342,364]" pageId="2" pageNumber="1017">
<emphasis id="B919EABC2908B837CE8FF920FECB713B" bold="true" box="[178,352,342,364]" italics="true" pageId="2" pageNumber="1017">Myllaena graeca</emphasis>
<bibRefCitation id="EFFC4B5F2908B837CF5AF920FE6A713B" author="KRAATZ G" box="[359,449,342,363]" pageId="2" pageNumber="1017" pagination="37 - 67" refId="ref8405" refString="KRAATZ G. (1858): Beitrag zur Kaferfauna Griechenlands. Zweites Stuck: Palpicornia, Silphales, Scydmaenidae, Pselaphidae, Staphylinidae. - Berliner Entomologische Zeitschrift 2: 37 - 67." type="journal article" year="1858">KRAATZ</bibRefCitation>
</emphasis>
,
<emphasis id="B919EABC2908B837CFF1F920FE54713B" bold="true" box="[460,511,342,363]" pageId="2" pageNumber="1017">1858</emphasis>
</taxonomicName>
(
<figureCitation id="13562A2B2908B837CC31F920FDC8713B" box="[524,611,342,364]" captionStart="Figs 1-7" captionStartId="5.[178,218,1523,1545]" captionTargetBox="[180,1058,229,1502]" captionTargetId="figure@5.[178,1085,225,1510]" captionTargetPageId="5" captionText="Figs 1-7: Myllaena graeca: (1) forebody; (2) male tergite VIII; (3) male sternite VIII; (4-6) median lobe of aedeagus in lateral and in ventral view; (7) spermatheca. Scale bars: 1: 1.0 mm; 2-3: 0.2 mm; 4-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776226" httpUri="https://zenodo.org/record/3776226/files/figure.png" pageId="2" pageNumber="1017">Figs 1-7</figureCitation>
,
<figureCitation id="13562A2B2908B837CC53F920FD05713C" box="[622,686,342,364]" captionStart="Map 1" captionStartId="4.[178,223,662,683]" captionTargetBox="[179,1086,212,653]" captionTargetId="figure@4.[179,1086,212,653]" captionTargetPageId="4" captionText="Map 1: Distributions of Myllaena graeca (based on revised records) (circles), M. mutabilis (triangles), and M. cretica (star)." figureDoi="http://doi.org/10.5281/zenodo.3776224" httpUri="https://zenodo.org/record/3776224/files/figure.png" pageId="2" pageNumber="1017">Map 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37765252908B837CE8FF90AFB9671FA" pageId="2" pageNumber="1017" type="reference_group">
<paragraph id="8BD236AE2908B837CE8FF90AFDAA71C2" blockId="2.[178,1086,380,815]" box="[178,513,380,402]" pageId="2" pageNumber="1017">
<treatmentCitationGroup id="AB7D11802908B837CE8FF90AFDAA71C2" box="[178,513,380,402]" pageId="2" pageNumber="1017">
<treatmentCitation id="0ACC10BF2908B837CE8FF90AFE5671C2" author="KRAATZ G" box="[178,509,380,402]" page="54" pageId="2" pageNumber="1017" year="1858">
<taxonomicName id="4C6D4D2D2908B837CE8FF90AFE5671C2" ID-CoL="453HB" authority="KRAATZ, 1858: 54" authorityName="Kraatz" authorityYear="1858" box="[178,509,380,402]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CE8FF90AFEE771C1" box="[178,332,380,401]" italics="true" pageId="2" pageNumber="1017">Myllaena graeca</emphasis>
<bibRefCitation id="EFFC4B5F2908B837CF6FF90AFE5671C2" author="KRAATZ G" box="[338,509,380,402]" pageId="2" pageNumber="1017" pagination="37 - 67" refId="ref8405" refString="KRAATZ G. (1858): Beitrag zur Kaferfauna Griechenlands. Zweites Stuck: Palpicornia, Silphales, Scydmaenidae, Pselaphidae, Staphylinidae. - Berliner Entomologische Zeitschrift 2: 37 - 67." type="journal article" year="1858">KRAATZ, 1858: 54</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD236AE2908B837CE8FF9E3FB9671FA" blockId="2.[178,1086,380,815]" box="[178,1085,401,427]" pageId="2" pageNumber="1017">
<materialsCitation id="3B053CF32908B837CE8FF9E3FB9671FA" ID-GBIF-Occurrence="2609436601" box="[178,1085,401,427]" country="Greece" location="Arta" municipality="Athamanon Oros" pageId="2" pageNumber="1017" specimenCount="1" specimenCount-female="1" typeStatus="neotype">
T y p e m a t e r i a l:
<typeStatus id="54D6880C2908B837CF8DF9E3FE5671FA" box="[432,509,405,426]" pageId="2" pageNumber="1017" type="neotype">Neotype</typeStatus>
<specimenCount id="9D6BFD272908B837CC3BF9E3FDBC71FA" box="[518,535,405,426]" pageId="2" pageNumber="1017" type="female"></specimenCount>
, present designation: &quot;
<collectingCountry id="F37A763E2908B837CCCDF9E3FC9E71FA" box="[752,821,405,426]" name="Greece" pageId="2" pageNumber="1017">Greece</collectingCountry>
:
<location id="8EB260752908B837CD7CF9E3FCC771FA" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CD7CF9E3FCC771FA" box="[833,876,405,426]" country="Greece" municipality="Athamanon Oros" name="Arta" pageId="2" pageNumber="1017">Arta</location>
,
<collectingMunicipality id="6BB6ACD42908B837CD45F9E3FBB271FA" box="[888,1049,401,427]" pageId="2" pageNumber="1017">Athamánon Óros</collectingMunicipality>
, 33
</materialsCitation>
</paragraph>
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<subSubSection id="C37765252908B836CE8FF9DDFE6974AE" lastPageId="3" lastPageNumber="1018" pageId="2" pageNumber="1017" type="description">
<paragraph id="8BD236AE2908B837CE8FF9DDFB967191" blockId="2.[178,1086,380,815]" box="[178,1085,423,449]" pageId="2" pageNumber="1017">
km N Arta, Strasse Arta &lt;Ágnanda, kl. Wasserfall,
<quantity id="4C959B4B2908B837CCB7F9DAFD6D7191" box="[650,710,428,449]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="1017" unit="m" value="340.0">340 m</quantity>
,
<geoCoordinate id="EE5950692908B837CCEDF9DDFC907191" box="[720,827,427,449]" degrees="39" direction="north" minutes="27" orientation="latitude" pageId="2" pageNumber="1017" precision="15" seconds="26" value="39.457222">39°27'26''N</geoCoordinate>
,
<geoCoordinate id="EE5950692908B837CD78F9DAFC067191" box="[837,941,428,449]" degrees="21" direction="east" minutes="02" orientation="longitude" pageId="2" pageNumber="1017" precision="15" seconds="37" value="21.043612">21°02'37''E</geoCoordinate>
, Genist + Streu
</paragraph>
<paragraph id="8BD236AE2908B837CE8FF9B4FB967188" blockId="2.[178,1086,380,815]" box="[178,1085,449,472]" pageId="2" pageNumber="1017">
Gesiebe,
<date id="FFD3106E2908B837CF3AF9B4FEDA7187" box="[263,369,450,471]" pageId="2" pageNumber="1017" value="2018-04-17">17.IV.2018</date>
, leg. M. Schülke [GR18-09] /
<typeStatus id="54D6880C2908B837CCB7F9B5FD4A7188" box="[650,737,451,472]" pageId="2" pageNumber="1017" type="neotype">Neotypus</typeStatus>
<specimenCount id="9D6BFD272908B837CCD5F9B5FD527188" box="[744,761,451,472]" pageId="2" pageNumber="1017" type="female"></specimenCount>
<taxonomicName id="4C6D4D2D2908B837CD3DF9B4FC4B7188" authority="Kraatz" authorityName="Kraatz" authorityYear="1858" box="[768,992,450,472]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CD3DF9B4FC307188" box="[768,923,450,472]" italics="true" pageId="2" pageNumber="1017">Myllaena graeca</emphasis>
Kraatz
</taxonomicName>
, desig. V.
</paragraph>
<paragraph id="8BD236AE2908B837CE8FF9AFFC8171BF" blockId="2.[178,1086,380,815]" box="[178,810,473,495]" pageId="2" pageNumber="1017">
Assing 2018 /
<taxonomicName id="4C6D4D2D2908B837CF0BF9AFFDBF71BE" authority="Kraatz" authorityName="Kraatz" authorityYear="1858" box="[310,532,473,495]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CF0BF9AFFE7B71BF" box="[310,464,473,495]" italics="true" pageId="2" pageNumber="1017">Myllaena graeca</emphasis>
Kraatz
</taxonomicName>
, det. V. Assing 2018&quot; (MNB).
</paragraph>
<paragraph id="8BD236AE2908B837CE8FF98FFC00737F" blockId="2.[178,1086,380,815]" pageId="2" pageNumber="1017">
<materialsCitation id="3B053CF32908B837CE8FF98FFEB0726E" ID-GBIF-Occurrence="2609436621" collectingDate="1998-04-03" collectionCode="MNB" collectorName="Assing &amp; Schulke" country="Greece" elevation="450" latitude="39.416668" location="Pilion Oros" longLatPrecision="1169" longitude="23.083334" pageId="2" pageNumber="1017" specimenCount="2" specimenCount-female="2" stateProvince="Thessalia">
A d d i t i o n a l m a t e r i a l e x a m i n e d:
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:
<specimenCount id="9D6BFD272908B837CCC1F98FFC82725E" box="[764,809,505,526]" pageId="2" pageNumber="1017" type="female">2♀♀</specimenCount>
,
<collectingRegion id="49A9F84C2908B837CD0AF98EFC20725E" box="[823,907,504,526]" country="Greece" name="Thessalia" pageId="2" pageNumber="1017">Thessalía</collectingRegion>
,
<location id="8EB260752908B837CDAAF98EFBAB725E" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CDAAF98EFBAB725E" box="[919,1024,504,526]" country="Greece" latitude="39.416668" longLatPrecision="1169" longitude="23.083334" name="Pilion Oros" pageId="2" pageNumber="1017" stateProvince="Thessalia">Pilion Oros</location>
,
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SW Zagora
</locationDeviation>
,
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,
<geoCoordinate id="EE5950692908B837CF91FA66FE587276" box="[428,499,528,550]" degrees="23" direction="east" minutes="05" orientation="longitude" pageId="2" pageNumber="1017" precision="925" value="23.083334">23°05'E</geoCoordinate>
,
<quantity id="4C959B4B2908B837CFC2FA66FD967276" box="[511,573,528,550]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="1017" unit="m" value="450.0">
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</quantity>
, in wet moss,
<date id="FFD3106E2908B837CCFAFA67FC8A7276" box="[711,801,529,550]" pageId="2" pageNumber="1017" value="1998-04-03">
<collectingDate id="EF97E9862908B837CCFAFA67FC8A7276" box="[711,801,529,550]" pageId="2" pageNumber="1017" value="1998-04-03">3.IV.1998</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CD6AFA66FC3F7276" box="[855,916,528,550]" pageId="2" pageNumber="1017">Assing</collectorName>
&amp;
<collectorName id="269853782908B837CD8BFA66FC577276" box="[950,1020,528,550]" pageId="2" pageNumber="1017">Schülke</collectorName>
(cAss,
<collectionCode id="ED7CAE6B2908B837CEE2FA5FFEBE726E" box="[223,277,553,574]" pageId="2" pageNumber="1017">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053CF32908B837CF15FA5FFEB57207" ID-GBIF-Occurrence="2609436609" collectingDate="2005-03-18" collectingDateMax="2014-03-21" collectingDateMin="2005-03-18" collectorName="Lompe &amp; Meybohm &amp; Hetzel" country="Turkey" county="Lesbos" determinerName="Feldmann" elevation="200" latitude="41.183334" location="Belgrad Ormani" longLatPrecision="1158" longitude="29.0" municipality="Skotino" pageId="2" pageNumber="1017" specimenCount="13" specimenCount-female="1" stateProvince="Istanbul">
<specimenCount id="9D6BFD272908B837CF15FA5FFEEF726E" box="[296,324,553,574]" pageId="2" pageNumber="1017" type="female">1♀</specimenCount>
,
<collectingCounty id="62B34E222908B837CF6FFA5EFE39726E" box="[338,402,552,574]" pageId="2" pageNumber="1017">Lesbos</collectingCounty>
,
<collectingMunicipality id="6BB6ACD42908B837CFA2FA5EFE4E726E" box="[415,485,552,574]" pageId="2" pageNumber="1017">Skotino</collectingMunicipality>
,
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,
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,
<quantity id="4C959B4B2908B837CCA1FA5FFD72726E" box="[668,729,553,574]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="1017" unit="m" value="200.0">
<elevation id="0040D19D2908B837CCA1FA5FFD72726E" box="[668,729,553,574]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="1017" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="FFD3106E2908B837CCDBFA5EFCE2726E" box="[742,841,552,574]" pageId="2" pageNumber="1017" value="2005-03-18">
<collectingDate id="EF97E9862908B837CCDBFA5EFCE2726E" box="[742,841,552,574]" pageId="2" pageNumber="1017" value="2005-03-18">18.III.2005</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CDBDFA5FFC15726E" box="[896,958,553,574]" pageId="2" pageNumber="1017">Lompe</collectorName>
&amp;
<collectorName id="269853782908B837CDDFFA5EFB96726E" box="[994,1085,552,574]" pageId="2" pageNumber="1017">Meybohm</collectorName>
(cAss).
</materialsCitation>
<materialsCitation id="3B053CF32908B837CF19FA36FC0F723E" ID-GBIF-Occurrence="2609436625" collectingDate="2005-03-18" collectingDateMax="2014-03-21" collectingDateMin="2005-03-18" collectorName="Lompe &amp; Meybohm &amp; Hetzel" country="Turkey" county="Lesbos" determinerName="Feldmann" elevation="200" latitude="41.183334" location="Belgrad Ormani" longLatPrecision="1158" longitude="29.0" municipality="Skotino" pageId="2" pageNumber="1017" specimenCount="13" specimenCount-female="1" stateProvince="Istanbul">
<collectingCountry id="F37A763E2908B837CF19FA36FECD7206" box="[292,358,576,598]" name="Turkey" pageId="2" pageNumber="1017">Turkey</collectingCountry>
:
<specimenCount id="9D6BFD272908B837CF4DFA37FE1A7206" box="[368,433,577,598]" pageId="2" pageNumber="1017" type="generic">12 exs.</specimenCount>
[det.
<determinerName id="7E0EDA6A2908B837CFD8FA36FDEA7206" box="[485,577,576,598]" pageId="2" pageNumber="1017">Feldmann</determinerName>
],
<collectingRegion id="49A9F84C2908B837CC6CFA36FD327206" box="[593,665,576,598]" country="Turkey" name="Istanbul" pageId="2" pageNumber="1017">Istanbul</collectingRegion>
,
<location id="8EB260752908B837CC9EFA36FC9F7206" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CC9EFA36FC9F7206" box="[675,820,576,598]" country="Turkey" county="Lesbos" latitude="41.183334" longLatPrecision="1158" longitude="29.0" municipality="Skotino" name="Belgrad Ormani" pageId="2" pageNumber="1017" stateProvince="Istanbul">Belgrad Ormanı</location>
,
<locationDeviation id="9A602F8C2908B837CD03FA37FC4E7206" box="[830,997,576,598]" location="Bahcekoy" pageId="2" pageNumber="1017">
<quantity id="4C959B4B2908B837CD03FA37FCC77206" box="[830,876,576,598]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="1017" unit="km" value="1.0">1 km</quantity>
N Bahçeköy
</locationDeviation>
,
<geoCoordinate id="EE5950692908B837CDD2FA37FB927206" box="[1007,1081,577,598]" degrees="41" direction="north" minutes="11" orientation="latitude" pageId="2" pageNumber="1017" precision="925" value="41.183334">41°11'N</geoCoordinate>
,
<geoCoordinate id="EE5950692908B837CEE2FA2FFE8D723E" box="[223,294,601,622]" degrees="29" direction="east" minutes="00" orientation="longitude" pageId="2" pageNumber="1017" precision="925" value="29.0">29°00'E</geoCoordinate>
,
<quantity id="4C959B4B2908B837CF0DFA2EFEC1723E" box="[304,362,600,622]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="1017" unit="m" value="160.0">
<elevation id="0040D19D2908B837CF0DFA2EFEC1723E" box="[304,362,600,622]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="1017" unit="m" value="160.0">160 m</elevation>
</quantity>
, mixed broadleaved forest, sifted,
<date id="FFD3106E2908B837CCA6FA2FFD55723E" box="[667,766,601,622]" pageId="2" pageNumber="1017" value="2014-03-21">
<collectingDate id="EF97E9862908B837CCA6FA2FFD55723E" box="[667,766,601,622]" pageId="2" pageNumber="1017" value="2014-03-21">21.III.2014</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CD12FA2EFCC3723E" box="[815,872,600,622]" pageId="2" pageNumber="1017">Hetzel</collectorName>
(cFel)
</materialsCitation>
;
<materialsCitation id="3B053CF32908B837CD93FA2FFBAA72D7" ID-GBIF-Occurrence="2609436608" collectingDate="2008-07-21" collectorName="Assing" country="Turkey" elevation="220" latitude="41.316666" location="Bafra" longLatPrecision="1157" longitude="35.85" pageId="2" pageNumber="1017" specimenCount="1" specimenCount-female="1" stateProvince="Samsun">
<specimenCount id="9D6BFD272908B837CD93FA2FFC61723E" box="[942,970,601,622]" pageId="2" pageNumber="1017" type="female">1♀</specimenCount>
,
<collectingRegion id="49A9F84C2908B837CDEBFA2FFBB6723E" box="[982,1053,601,622]" country="Turkey" name="Samsun" pageId="2" pageNumber="1017">Samsun</collectingRegion>
,
<locationDeviation id="9A602F8C2908B837CA1AFA2FFECF72D6" location="Samsun" pageId="2" pageNumber="1017">
<quantity id="4C959B4B2908B837CA1AFA2FFF5072D6" metricMagnitude="4" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="1017" unit="km" value="41.0">41 km</quantity>
W
<collectingRegion id="49A9F84C2908B837CF20FA07FECF72D6" box="[285,356,625,646]" country="Turkey" name="Samsun" pageId="2" pageNumber="1017">Samsun</collectingRegion>
</locationDeviation>
,
<quantity id="4C959B4B2908B837CF52FA07FE0C72D6" box="[367,423,624,646]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="1017" unit="km" value="27.0">27 km</quantity>
S
<location id="8EB260752908B837CFFCFA06FE5872D6" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CFFCFA06FE5872D6" box="[449,499,624,646]" country="Turkey" latitude="41.316666" longLatPrecision="1157" longitude="35.85" name="Bafra" pageId="2" pageNumber="1017" stateProvince="Samsun">Bafra</location>
,
<geoCoordinate id="EE5950692908B837CFC0FA07FDED72D6" box="[509,582,625,646]" degrees="41" direction="north" minutes="19" orientation="latitude" pageId="2" pageNumber="1017" precision="925" value="41.316666">41°19'N</geoCoordinate>
,
<geoCoordinate id="EE5950692908B837CC6CFA06FD3C72D6" box="[593,663,624,646]" degrees="35" direction="east" minutes="51" orientation="longitude" pageId="2" pageNumber="1017" precision="925" value="35.85">35°51'E</geoCoordinate>
,
<quantity id="4C959B4B2908B837CC9FFA07FD7672D6" box="[674,733,625,646]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="1017" unit="m" value="220.0">
<elevation id="0040D19D2908B837CC9FFA07FD7672D6" box="[674,733,625,646]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="1017" unit="m" value="220.0">220 m</elevation>
</quantity>
,
<date id="FFD3106E2908B837CCDAFA07FCF872D6" box="[743,851,625,646]" pageId="2" pageNumber="1017" value="2008-07-21">
<collectingDate id="EF97E9862908B837CCDAFA07FCF872D6" box="[743,851,625,646]" pageId="2" pageNumber="1017" value="2008-07-21">21.VII.2008</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CDBEFA06FC1472D6" box="[899,959,624,646]" pageId="2" pageNumber="1017">Assing</collectorName>
(cAss)
</materialsCitation>
;
<materialsCitation id="3B053CF32908B837CA36FA07FD3172E6" ID-GBIF-Occurrence="2609436618" collectingDate="2006-07-30" collectionCode="MNB" collectorName="Assing &amp; Schulke" country="Turkey" elevation="990" latitude="40.816666" location="Kabaduz" longLatPrecision="1160" longitude="37.9" pageId="2" pageNumber="1017" specimenCount="12" specimenCount-female="3" specimenCount-male="7" stateProvince="Ordu">
<specimenCount id="9D6BFD272908B837CA36FA07FB9372D6" box="[1035,1080,625,646]" pageId="2" pageNumber="1017" type="male">7♂♂</specimenCount>
,
<specimenCount id="9D6BFD272908B837CEE2FAFFFEA072CE" box="[223,267,649,670]" pageId="2" pageNumber="1017" type="female">3♀♀</specimenCount>
,
<specimenCount id="9D6BFD272908B837CF26FAFFFEFD72CE" box="[283,342,649,670]" pageId="2" pageNumber="1017" type="generic">2 exs.</specimenCount>
,
<collectingRegion id="49A9F84C2908B837CF5EFAFEFE3972CE" box="[355,402,648,670]" country="Turkey" name="Ordu" pageId="2" pageNumber="1017">Ordu</collectingRegion>
,
<locationDeviation id="9A602F8C2908B837CF9DFAFEFD8572CE" box="[416,558,648,670]" location="Ordu" pageId="2" pageNumber="1017">
<quantity id="4C959B4B2908B837CF9DFAFEFE7672CE" box="[416,477,648,670]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="1017" unit="km" value="15.0">15 km</quantity>
S
<collectingRegion id="49A9F84C2908B837CFC2FAFEFD8572CE" box="[511,558,648,670]" country="Turkey" name="Ordu" pageId="2" pageNumber="1017">Ordu</collectingRegion>
</locationDeviation>
, S
<location id="8EB260752908B837CC6EFAFEFD0872CE" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CC6EFAFEFD0872CE" box="[595,675,648,670]" country="Turkey" latitude="40.816666" longLatPrecision="1160" longitude="37.9" name="Kabaduz" pageId="2" pageNumber="1017" stateProvince="Ordu">Kabaduz</location>
,
<geoCoordinate id="EE5950692908B837CC8CFAFFFD5072CE" box="[689,763,649,670]" degrees="40" direction="north" minutes="49" orientation="latitude" pageId="2" pageNumber="1017" precision="925" value="40.816666">40°49'N</geoCoordinate>
,
<geoCoordinate id="EE5950692908B837CD34FAFEFCFA72CE" box="[777,849,648,670]" degrees="37" direction="east" minutes="54" orientation="longitude" pageId="2" pageNumber="1017" precision="925" value="37.9">37°54'E</geoCoordinate>
,
<quantity id="4C959B4B2908B837CD62FAFFFC3672CE" box="[863,925,649,670]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="2" pageNumber="1017" unit="m" value="990.0">
<elevation id="0040D19D2908B837CD62FAFFFC3672CE" box="[863,925,649,670]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="2" pageNumber="1017" unit="m" value="990.0">990 m</elevation>
</quantity>
, grassy roadside,
<date id="FFD3106E2908B837CEE2FAD6FEE072E6" box="[223,331,672,694]" pageId="2" pageNumber="1017" value="2006-07-30">
<collectingDate id="EF97E9862908B837CEE2FAD6FEE072E6" box="[223,331,672,694]" pageId="2" pageNumber="1017" value="2006-07-30">30.VII.2006</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CF46FAD6FE1372E6" box="[379,440,672,694]" pageId="2" pageNumber="1017">Assing</collectorName>
&amp;
<collectorName id="269853782908B837CFE8FAD6FDB072E6" box="[469,539,672,694]" pageId="2" pageNumber="1017">Schülke</collectorName>
(cAss,
<collectionCode id="ED7CAE6B2908B837CC62FAD7FD3E72E6" box="[607,661,673,694]" pageId="2" pageNumber="1017">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053CF32908B837CC99FAD7FC75729F" ID-GBIF-Occurrence="2609436660" collectingDate="2008-07-22" collectionCode="MNB" collectorName="Schulke" country="Turkey" determinerName="Schulke" elevation="1040" latitude="40.683334" location="11 km W Golkoy" longLatPrecision="1161" longitude="37.5" pageId="2" pageNumber="1017" specimenCount="7" stateProvince="Ordu">
<specimenCount id="9D6BFD272908B837CC99FAD7FD7272E6" box="[676,729,673,694]" pageId="2" pageNumber="1017" type="generic">7 exs.</specimenCount>
[det.
<determinerName id="7E0EDA6A2908B837CD36FAD6FCFE72E6" box="[779,853,672,694]" pageId="2" pageNumber="1017">Schülke</determinerName>
],
<collectingRegion id="49A9F84C2908B837CD59FAD6FC3872E6" box="[868,915,672,694]" country="Turkey" name="Ordu" pageId="2" pageNumber="1017">Ordu</collectingRegion>
,
<location id="8EB260752908B837CDA1FAD7FB9172E6" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CDA1FAD7FB9172E6" box="[924,1082,672,694]" country="Turkey" latitude="40.683334" longLatPrecision="1161" longitude="37.5" name="11 km W Golkoy" pageId="2" pageNumber="1017" stateProvince="Ordu">
<quantity id="4C959B4B2908B837CDA1FAD7FC7F72E6" box="[924,980,672,694]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="1017" unit="km" value="11.0">11 km</quantity>
W Gölköy
</location>
,
<geoCoordinate id="EE5950692908B837CEE2FACFFE82729E" box="[223,297,697,718]" degrees="40" direction="north" minutes="41" orientation="latitude" pageId="2" pageNumber="1017" precision="925" value="40.683334">40°41'N</geoCoordinate>
,
<geoCoordinate id="EE5950692908B837CF05FACFFE2B729E" box="[312,384,697,718]" degrees="37" direction="east" minutes="30" orientation="longitude" pageId="2" pageNumber="1017" precision="925" value="37.5">37°30'E</geoCoordinate>
,
<quantity id="4C959B4B2908B837CFB2FACFFE71729E" box="[399,474,697,718]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="1017" unit="m" value="1040.0">
<elevation id="0040D19D2908B837CFB2FACFFE71729E" box="[399,474,697,718]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="1017" unit="m" value="1040.0">1040 m</elevation>
</quantity>
, wet stream valley,
<date id="FFD3106E2908B837CC9FFACFFCA5729E" box="[674,782,697,718]" pageId="2" pageNumber="1017" value="2008-07-22">
<collectingDate id="EF97E9862908B837CC9FFACFFCA5729E" box="[674,782,697,718]" pageId="2" pageNumber="1017" value="2008-07-22">22.VII.2008</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CD74FACEFC24729E" box="[841,911,696,718]" pageId="2" pageNumber="1017">Schülke</collectorName>
(
<collectionCode id="ED7CAE6B2908B837CD9FFACFFC73729F" box="[930,984,697,719]" pageId="2" pageNumber="1017">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053CF32908B837CDD3FACFFCAF72AF" ID-GBIF-Occurrence="2609436630" collectingDate="2006-07-29" collectionCode="MNB" collectorName="Schulke" country="Turkey" determinerName="Schulke" elevation="1250" latitude="40.583332" location="35 km S Giresun" longLatPrecision="1162" longitude="38.45" pageId="2" pageNumber="1017" specimenCount="1" specimenCount-male="1" stateProvince="Giresun">
<specimenCount id="9D6BFD272908B837CDD3FACFFBA1729E" box="[1006,1034,697,718]" pageId="2" pageNumber="1017" type="male">1♂</specimenCount>
[det.
<determinerName id="7E0EDA6A2908B837CEE2FAA6FE8272B6" box="[223,297,720,742]" pageId="2" pageNumber="1017">Schülke</determinerName>
],
<collectingRegion id="49A9F84C2908B837CF02FAA6FE2D72B6" box="[319,390,720,742]" country="Turkey" name="Giresun" pageId="2" pageNumber="1017">Giresun</collectingRegion>
, ca.
<location id="8EB260752908B837CF83FAA6FDC672B6" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CF83FAA6FDC672B6" box="[446,621,720,742]" country="Turkey" latitude="40.583332" longLatPrecision="1162" longitude="38.45" name="35 km S Giresun" pageId="2" pageNumber="1017" stateProvince="Giresun">
<quantity id="4C959B4B2908B837CF83FAA6FE5572B6" box="[446,510,720,742]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="1017" unit="km" value="35.0">35 km</quantity>
S
<collectingRegion id="49A9F84C2908B837CC1BFAA6FDC672B6" box="[550,621,720,742]" country="Turkey" name="Giresun" pageId="2" pageNumber="1017">Giresun</collectingRegion>
</location>
,
<geoCoordinate id="EE5950692908B837CC43FAA6FD6272B6" box="[638,713,720,742]" degrees="40" direction="north" minutes="35" orientation="latitude" pageId="2" pageNumber="1017" precision="925" value="40.583332">40°35'N</geoCoordinate>
,
<geoCoordinate id="EE5950692908B837CCE7FAA7FC8A72B6" box="[730,801,721,742]" degrees="38" direction="east" minutes="27" orientation="longitude" pageId="2" pageNumber="1017" precision="925" value="38.45">38°27'E</geoCoordinate>
,
<quantity id="4C959B4B2908B837CD0FFAA6FCD472B6" box="[818,895,720,742]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="1017" unit="m" value="1250.0">
<elevation id="0040D19D2908B837CD0FFAA6FCD472B6" box="[818,895,720,742]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="1017" unit="m" value="1250.0">1250 m</elevation>
</quantity>
, spruce forest with rhododendron undergrowth,
<date id="FFD3106E2908B837CFE7FA9EFDEE72AE" box="[474,581,744,766]" pageId="2" pageNumber="1017" value="2006-07-29">
<collectingDate id="EF97E9862908B837CFE7FA9EFDEE72AE" box="[474,581,744,766]" pageId="2" pageNumber="1017" value="2006-07-29">29.VII.2006</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CC48FA9EFD1072AE" box="[629,699,744,766]" pageId="2" pageNumber="1017">Schülke</collectorName>
(
<collectionCode id="ED7CAE6B2908B837CCFAFA9FFD5672AF" box="[711,765,745,767]" pageId="2" pageNumber="1017">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B053CF32908B837CD33FA9FFC0C737F" ID-GBIF-Occurrence="2609436643" collectingDate="2006-07-26" collectionCode="MNB, NHMW" collectorName="Schulke &amp; Identification &amp; v. Oertzen" country="Turkey" determinerName="Schulke" elevation="1540" latitude="40.666668" location="Altindere Milli Park" longLatPrecision="1161" longitude="39.666668" pageId="2" pageNumber="1017" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Trabzon">
<specimenCount id="9D6BFD272908B837CD33FA9FFC8172AE" box="[782,810,745,766]" pageId="2" pageNumber="1017" type="female">1♀</specimenCount>
[det.
<determinerName id="7E0EDA6A2908B837CD61FA9EFC0D72AE" box="[860,934,744,766]" pageId="2" pageNumber="1017">Schülke</determinerName>
],
<collectingRegion id="49A9F84C2908B837CD89FA9EFC5472AE" box="[948,1023,744,766]" country="Turkey" name="Trabzon" pageId="2" pageNumber="1017">Trabzon</collectingRegion>
, ca.
<locationDeviation id="9A602F8C2908B837CA1AFA9FFEC37346" location="Trabzon" pageId="2" pageNumber="1017">
<quantity id="4C959B4B2908B837CA1AFA9FFF507346" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="1017" unit="km" value="40.0">40 km</quantity>
S
<collectingRegion id="49A9F84C2908B837CF20FB76FEC37346" box="[285,360,768,790]" country="Turkey" name="Trabzon" pageId="2" pageNumber="1017">Trabzon</collectingRegion>
</locationDeviation>
,
<location id="8EB260752908B837CF4BFB76FD997346" LSID="urn:lsid:plazi:treatment:03C487B82908B836CE8FF920FE6974AE:8EB260752908B837CF4BFB76FD997346" box="[374,562,768,790]" country="Turkey" latitude="40.666668" longLatPrecision="1161" longitude="39.666668" name="Altindere Milli Park" pageId="2" pageNumber="1017" stateProvince="Trabzon">Altındere Milli Park</location>
,
<quantity id="4C959B4B2908B837CC7CFB76FD217346" box="[577,650,768,790]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="1017" unit="m" value="1540.0">
<elevation id="0040D19D2908B837CC7CFB76FD217346" box="[577,650,768,790]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="1017" unit="m" value="1540.0">1540 m</elevation>
</quantity>
,
<geoCoordinate id="EE5950692908B837CCA4FB77FD497346" box="[665,738,769,790]" degrees="40" direction="north" minutes="40" orientation="latitude" pageId="2" pageNumber="1017" precision="925" value="40.666668">40°40'N</geoCoordinate>
,
<geoCoordinate id="EE5950692908B837CCCCFB77FC937346" box="[753,824,769,790]" degrees="39" direction="east" minutes="40" orientation="longitude" pageId="2" pageNumber="1017" precision="925" value="39.666668">39°40'E</geoCoordinate>
,
<date id="FFD3106E2908B837CD7BFB76FC167346" box="[838,957,768,790]" pageId="2" pageNumber="1017" value="2006-07-26">
<collectingDate id="EF97E9862908B837CD7BFB76FC167346" box="[838,957,768,790]" pageId="2" pageNumber="1017" value="2006-07-26">26. VII.2006</collectingDate>
</date>
, leg.
<collectorName id="269853782908B837CDCBFB76FB977346" box="[1014,1084,768,790]" pageId="2" pageNumber="1017">Schülke</collectorName>
(
<collectionCode id="ED7CAE6B2908B837CED8FB6FFEB0737F" box="[229,283,793,815]" pageId="2" pageNumber="1017">MNB</collectionCode>
).
<collectorName id="269853782908B837CF17FB6EFE35737E" box="[298,414,792,814]" pageId="2" pageNumber="1017">Identification</collectorName>
doubtful:
<specimenCount id="9D6BFD272908B837CFC4FB6FFD8E737E" box="[505,549,793,814]" pageId="2" pageNumber="1017" type="male">2♂♂</specimenCount>
[teneral], &quot;Creta&quot;, leg.
<collectorName id="269853782908B837CCD3FB6FFCE2737E" box="[750,841,793,814]" pageId="2" pageNumber="1017">v. Oertzen</collectorName>
(
<collectionCode id="ED7CAE6B2908B837CD69FB6FFC0A737F" box="[852,929,793,815]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="1017" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD236AE2908B837CE8FFB4BFC197453" blockId="2.[178,1085,829,1027]" pageId="2" pageNumber="1017">
C o m m e n t: The original description is based on an unspecified number of
<typeStatus id="54D6880C2908B837CDD9FB4BFB967302" box="[996,1085,829,850]" pageId="2" pageNumber="1017" type="syntype">syntypes</typeStatus>
from &quot;Zante (v. Kiesenwetter),
<collectingRegion id="49A9F84C2908B837CFCBFB2CFD86733F" box="[502,557,858,879]" country="Greece" name="Kriti" pageId="2" pageNumber="1017">Creta</collectingRegion>
(Zebe)&quot; (
<bibRefCitation id="EFFC4B5F2908B837CCAFFB2CFC8E7320" author="KRAATZ G" box="[658,805,858,880]" pageId="2" pageNumber="1017" pagination="37 - 67" refId="ref8405" refString="KRAATZ G. (1858): Beitrag zur Kaferfauna Griechenlands. Zweites Stuck: Palpicornia, Silphales, Scydmaenidae, Pselaphidae, Staphylinidae. - Berliner Entomologische Zeitschrift 2: 37 - 67." type="journal article" year="1858">KRAATZ 1858</bibRefCitation>
). According to Lutz Behne (e-mail 22 Aug., 2018) there is no type material in the Kraatz collection at the SDEI. The collection of Gustav Zebe was moved to &quot;Hartmann/Arnswalde&quot; around 1870 (
<bibRefCitation id="EFFC4B5F2908B837CDD8FBE3FEA17398" author="HORN W &amp; KAHLE I &amp; FRIESE G" pageId="2" pageNumber="1017" refId="ref8357" refString="HORN W., KAHLE I., FRIESE G. &amp; R. GAEDIKE (1990): Collectiones entomologicae. Ein Kompendium uber den Verbleib entomologischer Sammlungen der Welt bis 1960. - Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin: 573 pp." type="book" year="1990">HORN et al. 1990</bibRefCitation>
); the Kiesenwetter collection is deposited in the Zoologische Staatssammlung München. It does not seem likely that the former collection will ever be located, and no type material was found in München (Michael Balke, e-mail 7 Sept., 2018).
</paragraph>
<paragraph id="8BD236AE2908B837CE8FFC64FE8B74EB" blockId="2.[178,1085,1041,1211]" pageId="2" pageNumber="1017">
According to SCHÜLKE &amp; SMETANA (2015),
<taxonomicName id="4C6D4D2D2908B837CCBFFC65FD467478" authorityName="Kraatz" authorityYear="1858" box="[642,749,1043,1064]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CCBFFC65FD467478" box="[642,749,1043,1064]" italics="true" pageId="2" pageNumber="1017">M. graeca</emphasis>
</taxonomicName>
has a trans-Palaearctic distribution and been recorded from North Africa (
<collectingCountry id="F37A763E2908B837CC4CFC46FD7B7415" box="[625,720,1072,1093]" name="Morocco" pageId="2" pageNumber="1017">Morocco</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CCE1FC46FC817416" box="[732,810,1072,1094]" name="Algeria" pageId="2" pageNumber="1017">Algeria</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CD0BFC46FC2D7416" box="[822,902,1072,1094]" name="Tunisia" pageId="2" pageNumber="1017">Tunisia</collectingCountry>
), West and South Europe (
<collectingCountry id="F37A763E2908B837CF2AFC3BFE1B7433" box="[279,432,1101,1123]" name="United Kingdom" pageId="2" pageNumber="1017">Great Britain</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CFF4FC3BFDBA7432" box="[457,529,1101,1122]" name="France" pageId="2" pageNumber="1017">France</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CC14FC3BFDF17433" box="[553,602,1101,1123]" name="Italy" pageId="2" pageNumber="1017">Italy</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CC49FC3BFD657433" box="[628,718,1101,1123]" name="Bulgaria" pageId="2" pageNumber="1017">Bulgaria</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CCDAFC3BFC987433" box="[743,819,1101,1123]" name="Greece" pageId="2" pageNumber="1017">Greece</collectingCountry>
), the Caucasus region (
<collectingCountry id="F37A763E2908B837CE84FC1CFE8574D0" box="[185,302,1130,1152]" name="Azerbaijan" pageId="2" pageNumber="1017">Azerbaijan</collectingCountry>
), Middle Asia (&quot;Turkestan&quot;),
<collectingCountry id="F37A763E2908B837CC59FC1CFD0874D0" box="[612,675,1130,1152]" name="China" pageId="2" pageNumber="1017">China</collectingCountry>
, and
<collectingCountry id="F37A763E2908B837CCE0FC1DFCB174D0" box="[733,794,1131,1152]" name="Japan" pageId="2" pageNumber="1017">Japan</collectingCountry>
. The identity of
<taxonomicName id="4C6D4D2D2908B837CDF6FC1DFB9D74D0" authorityName="Kraatz" authorityYear="1858" box="[971,1078,1131,1152]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CDF6FC1DFB9D74D0" box="[971,1078,1131,1152]" italics="true" pageId="2" pageNumber="1017">M. graeca</emphasis>
</taxonomicName>
, however, has never been clarified based on a study (and illustrations) of the sexual characters.
</paragraph>
<paragraph id="8BD236AE2908B836CE8FFCBCFD7771F0" blockId="2.[178,1086,1226,1600]" lastBlockId="3.[178,1086,219,417]" lastPageId="3" lastPageNumber="1018" pageId="2" pageNumber="1017">
Six specimens labelled as
<taxonomicName id="4C6D4D2D2908B837CFFAFCBCFDDD74B0" authorityName="Kraatz" authorityYear="1858" box="[455,630,1226,1248]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CFFAFCBCFDDD74B0" box="[455,630,1226,1248]" italics="true" pageId="2" pageNumber="1017">Myllaena graeca</emphasis>
</taxonomicName>
from
<collectingCountry id="F37A763E2908B837CC86FCBCFCBD74B0" box="[699,790,1226,1248]" name="Romania" pageId="2" pageNumber="1017">Romania</collectingCountry>
(one specimen),
<collectingCountry id="F37A763E2908B837CDF0FCBCFB9774B0" box="[973,1084,1226,1248]" name="Azerbaijan" pageId="2" pageNumber="1017">Azerbaijan</collectingCountry>
(2),
<collectingCountry id="F37A763E2908B837CEDDFC91FE8774AD" box="[224,300,1255,1277]" name="Algeria" pageId="2" pageNumber="1017">Algeria</collectingCountry>
(1) and
<collectingCountry id="F37A763E2908B837CFB2FC91FE7074AD" box="[399,475,1255,1277]" name="Tunisia" pageId="2" pageNumber="1017">Tunisia</collectingCountry>
(2) were found in the collections of the SDEI. They all belong to
<taxonomicName id="4C6D4D2D2908B837CF1FFD70FE1D754A" authorityName="Erichson" authorityYear="1837" box="[290,438,1285,1307]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B919EABC2908B837CF1FFD70FE1D754A" box="[290,438,1285,1307]" italics="true" pageId="2" pageNumber="1017">M. intermedia</emphasis>
</taxonomicName>
ERICHSON, 1837 (
<collectingCountry id="F37A763E2908B837CCBDFD73FD74754B" box="[640,735,1285,1307]" name="Romania" pageId="2" pageNumber="1017">Romania</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CCD2FD73FCCB754B" box="[751,864,1285,1307]" name="Azerbaijan" pageId="2" pageNumber="1017">Azerbaijan</collectingCountry>
) and
<emphasis id="B919EABC2908B837CD9FFD70FB96754A" box="[930,1085,1285,1307]" italics="true" pageId="2" pageNumber="1017">M. brevicornis</emphasis>
(MATTHEWS, 1838) (
<collectingCountry id="F37A763E2908B837CFB4FD54FE727568" box="[393,473,1314,1336]" name="Algeria" pageId="2" pageNumber="1017">Algeria</collectingCountry>
,
<collectingCountry id="F37A763E2908B837CFD9FD54FD9E7568" box="[484,565,1314,1336]" name="Tunisia" pageId="2" pageNumber="1017">Tunisia</collectingCountry>
). Only two specimens labelled as
<taxonomicName id="4C6D4D2D2908B837CDAAFD55FBAA7568" authorityName="Kraatz" authorityYear="1858" box="[919,1025,1315,1336]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CDAAFD55FBAA7568" box="[919,1025,1315,1336]" italics="true" pageId="2" pageNumber="1017">M. graeca</emphasis>
</taxonomicName>
, both of them teneral males, originally on one pin, and collected in Crete, were found in the collections of the NHMW. They belong to the same species group as the species here interpreted as
<taxonomicName id="4C6D4D2D2908B837CF74FD0DFE1E75C0" authorityName="Kraatz" authorityYear="1858" box="[329,437,1403,1424]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CF74FD0DFE1E75C0" box="[329,437,1403,1424]" italics="true" pageId="2" pageNumber="1017">M. graeca</emphasis>
</taxonomicName>
and as
<taxonomicName id="4C6D4D2D2908B837CC30FD0DFD3A75DF" authorityName="Assing" authorityYear="2018" box="[525,657,1402,1424]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B919EABC2908B837CC30FD0DFD3A75DF" box="[525,657,1402,1424]" italics="true" pageId="2" pageNumber="1017">M. mutabilis</emphasis>
</taxonomicName>
. However, whether they are conspecific with the original type material of
<taxonomicName id="4C6D4D2D2908B837CC1FFDEEFD2575FD" authorityName="Kraatz" authorityYear="1858" box="[546,654,1432,1453]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2908B837CC1FFDEEFD2575FD" box="[546,654,1432,1453]" italics="true" pageId="2" pageNumber="1017">M. graeca</emphasis>
</taxonomicName>
from Zakinthos, with
<taxonomicName id="4C6D4D2D2908B837CDB9FDEEFBA275FD" authorityName="Assing" authorityYear="2018" box="[900,1033,1432,1453]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1017" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B919EABC2908B837CDB9FDEEFBA275FD" box="[900,1033,1432,1453]" italics="true" pageId="2" pageNumber="1017">M. mutabilis</emphasis>
</taxonomicName>
(see below), or whether they belong to yet another (unnamed) species can be clarified only when females are available. In view of the previous confusion regarding the identity of this species, a
<typeStatus id="54D6880C2908B837CF74FD86FE327655" box="[329,409,1520,1541]" pageId="2" pageNumber="1017" type="neotype">neotype</typeStatus>
designation is deemed indispensable. Therefore, a female from a locality in Arta, which is geographically not far from Zakinthos (Zante), is designated as the
<typeStatus id="54D6880C2908B837CEE2FE5DFE987610" box="[223,307,1579,1600]" pageId="2" pageNumber="1017" type="neotype">neotype</typeStatus>
. It is in agreement with the details given in the original description. R e d e s c r i p t i o n: Body length
<quantity id="4C959B4B2909B836CC6EF8ADFD7970A0" box="[595,722,219,241]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.8" metricValueMin="2.6" pageId="3" pageNumber="1018" unit="mm" value="3.2" valueMax="3.8" valueMin="2.6">2.6-3.8 mm</quantity>
; length of forebody
<quantity id="4C959B4B2909B836CD87F8ADFB9170A0" box="[954,1082,219,240]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="3" pageNumber="1018" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4-1.7 mm</quantity>
. Forebody as in
<figureCitation id="13562A2B2909B836CF6FF88EFE3B715D" box="[338,400,248,270]" captionStart="Figs 1-7" captionStartId="5.[178,218,1523,1545]" captionTargetBox="[180,1058,229,1502]" captionTargetId="figure@5.[178,1085,225,1510]" captionTargetPageId="5" captionText="Figs 1-7: Myllaena graeca: (1) forebody; (2) male tergite VIII; (3) male sternite VIII; (4-6) median lobe of aedeagus in lateral and in ventral view; (7) spermatheca. Scale bars: 1: 1.0 mm; 2-3: 0.2 mm; 4-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776226" httpUri="https://zenodo.org/record/3776226/files/figure.png" pageId="3" pageNumber="1018">Fig. 1</figureCitation>
. Coloration variable: head dark-reddish to blackish; pronotum and elytra bright-reddish to dark-brown with paler margins; abdomen reddish to blackishbrown, with the posterior margins of tergites III-VI, the posterior portion of tergite VII, and segments VIII-X reddish; legs yellow; antennae yellow to yellowish-brown, often with the middle more or less extensively and more or less distinctly darker. Other external characters similar to those of
<taxonomicName id="4C6D4D2D2909B836CC3FF9FDFD7D71F0" authorityName="Erichson" authorityYear="1837" box="[514,726,395,416]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1018" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B919EABC2909B836CC3FF9FDFD7D71F0" box="[514,726,395,416]" italics="true" pageId="3" pageNumber="1018">Myllaena intermedia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD236AE2909B836CE8FF9C7FBA47250" blockId="3.[178,1085,432,513]" pageId="3" pageNumber="1018">
<specimenCount id="9D6BFD272909B836CE8FF9C7FF637196" box="[178,200,433,454]" pageId="3" pageNumber="1018" type="male"></specimenCount>
: tergite VIII (
<figureCitation id="13562A2B2909B836CF66F9C6FE307196" box="[347,411,432,454]" captionStart="Figs 1-7" captionStartId="5.[178,218,1523,1545]" captionTargetBox="[180,1058,229,1502]" captionTargetId="figure@5.[178,1085,225,1510]" captionTargetPageId="5" captionText="Figs 1-7: Myllaena graeca: (1) forebody; (2) male tergite VIII; (3) male sternite VIII; (4-6) median lobe of aedeagus in lateral and in ventral view; (7) spermatheca. Scale bars: 1: 1.0 mm; 2-3: 0.2 mm; 4-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776226" httpUri="https://zenodo.org/record/3776226/files/figure.png" pageId="3" pageNumber="1018">Fig. 2</figureCitation>
) oblong, posterior margin acutely pointed; sternite VIII (
<figureCitation id="13562A2B2909B836CDC8F9C6FB9E7196" box="[1013,1077,432,454]" captionStart="Figs 1-7" captionStartId="5.[178,218,1523,1545]" captionTargetBox="[180,1058,229,1502]" captionTargetId="figure@5.[178,1085,225,1510]" captionTargetPageId="5" captionText="Figs 1-7: Myllaena graeca: (1) forebody; (2) male tergite VIII; (3) male sternite VIII; (4-6) median lobe of aedeagus in lateral and in ventral view; (7) spermatheca. Scale bars: 1: 1.0 mm; 2-3: 0.2 mm; 4-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776226" httpUri="https://zenodo.org/record/3776226/files/figure.png" pageId="3" pageNumber="1018">Fig. 3</figureCitation>
) oblong and with convex posterior margin; median lobe of aedeagus (
<figureCitation id="13562A2B2909B836CD42F9BBFC7D71B3" box="[895,982,461,483]" captionStart="Figs 1-7" captionStartId="5.[178,218,1523,1545]" captionTargetBox="[180,1058,229,1502]" captionTargetId="figure@5.[178,1085,225,1510]" captionTargetPageId="5" captionText="Figs 1-7: Myllaena graeca: (1) forebody; (2) male tergite VIII; (3) male sternite VIII; (4-6) median lobe of aedeagus in lateral and in ventral view; (7) spermatheca. Scale bars: 1: 1.0 mm; 2-3: 0.2 mm; 4-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776226" httpUri="https://zenodo.org/record/3776226/files/figure.png" pageId="3" pageNumber="1018">Figs 4-6</figureCitation>
) approximately
<quantity id="4C959B4B2909B836CEC0F99DFEFC7250" box="[253,343,491,513]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="1018" unit="mm" value="0.45">0.45 mm</quantity>
long, with long and slender, bisinuate (lateral view) ventral process.
</paragraph>
<paragraph id="8BD236AE2909B836CE8FFA67FD167231" blockId="3.[178,1085,528,609]" pageId="3" pageNumber="1018">
<specimenCount id="9D6BFD272909B836CE8FFA67FF6C7276" box="[178,199,529,550]" pageId="3" pageNumber="1018" type="female"></specimenCount>
: tergite VIII oblong, of similar shape as that of male; posterior margin of sternite VIII truncate in the middle; spermatheca (
<figureCitation id="13562A2B2909B836CC16FA5BFDC37213" box="[555,616,557,579]" captionStart="Figs 1-7" captionStartId="5.[178,218,1523,1545]" captionTargetBox="[180,1058,229,1502]" captionTargetId="figure@5.[178,1085,225,1510]" captionTargetPageId="5" captionText="Figs 1-7: Myllaena graeca: (1) forebody; (2) male tergite VIII; (3) male sternite VIII; (4-6) median lobe of aedeagus in lateral and in ventral view; (7) spermatheca. Scale bars: 1: 1.0 mm; 2-3: 0.2 mm; 4-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776226" httpUri="https://zenodo.org/record/3776226/files/figure.png" pageId="3" pageNumber="1018">Fig. 7</figureCitation>
) of very conspicuous shape, proximal portion of capsule extremely long and with numerous coils.
</paragraph>
<paragraph id="8BD236AE2909B836CE8FFA06FCB073B6" blockId="3.[178,1085,624,998]" pageId="3" pageNumber="1018">
C o m p a r a t i v e n o t e s: This species is best identified based on the conspicuous shape of the spermatheca; no other West Palaearctic congener has a spermatheca of even remotely similar shape. In body size, habitus, and other characters,
<taxonomicName id="4C6D4D2D2909B836CD86FADAFB8F7291" authorityName="Kraatz" authorityYear="1858" box="[955,1060,684,705]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1018" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2909B836CD86FADAFB8F7291" box="[955,1060,684,705]" italics="true" pageId="3" pageNumber="1018">M. graeca</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C6D4D2D2909B836CF20FABFFE04728D" authorityName="Erichson" authorityYear="1837" box="[285,431,712,734]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1018" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B919EABC2909B836CF20FABFFE04728D" box="[285,431,712,734]" italics="true" pageId="3" pageNumber="1018">M. intermedia</emphasis>
</taxonomicName>
. Aside from the primary sexual characters, it differs from this species by (mostly) paler coloration, less slender antennae with less oblong antennomeres IV-X, and the shape of the female sternite VIII (
<taxonomicName id="4C6D4D2D2909B836CCCCFB72FC287348" authorityName="Erichson" authorityYear="1837" box="[753,899,771,793]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1018" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B919EABC2909B836CCCCFB72FC287348" box="[753,899,771,793]" italics="true" pageId="3" pageNumber="1018">M. intermedia</emphasis>
</taxonomicName>
: posterior margin convex). The median lobe of the aedeagus somewhat resembles that of
<emphasis id="B919EABC2909B836CD90FB57FBA77365" box="[941,1036,800,822]" italics="true" pageId="3" pageNumber="1018">M. dubia</emphasis>
, but differs by significantly smaller size (
<emphasis id="B919EABC2909B836CC1AFB48FD287303" box="[551,643,830,851]" italics="true" pageId="3" pageNumber="1018">M. dubia</emphasis>
: median lobe approximately
<quantity id="4C959B4B2909B836CD91FB4BFC517303" box="[940,1018,829,851]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="1018" unit="mm" value="0.6">0.6 mm</quantity>
long), a sinuate basal portion of the ventral process in lateral view (straight in
<emphasis id="B919EABC2909B836CD84FB2AFBB27320" box="[953,1049,859,881]" italics="true" pageId="3" pageNumber="1018">M. dubia</emphasis>
), a much smaller crista apicalis, and internal structures of different shape. For characters distinguishing
<taxonomicName id="4C6D4D2D2909B836CF74FBE0FE1973FB" authorityName="Kraatz" authorityYear="1858" box="[329,434,918,939]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1018" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2909B836CF74FBE0FE1973FB" box="[329,434,918,939]" italics="true" pageId="3" pageNumber="1018">M. graeca</emphasis>
</taxonomicName>
from the closely related
<taxonomicName id="4C6D4D2D2909B836CC8BFBE0FC9C73FB" authorityName="Assing" authorityYear="2018" box="[694,823,918,939]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1018" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="B919EABC2909B836CC8BFBE0FC9C73FB" box="[694,823,918,939]" italics="true" pageId="3" pageNumber="1018">M. mutabilis</emphasis>
</taxonomicName>
, with which it shares not only a similar habitus and other external characters, but also an aedeagus of highly similar shape, see the comparative notes in the following section.
</paragraph>
<paragraph id="8BD236AE2909B836CE8FFB80FD6574CD" blockId="3.[178,1085,1013,1182]" pageId="3" pageNumber="1018">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: Thisspecieshasreliablybeen recorded only from
<collectingCountry id="F37A763E2909B836CFBBFC65FE657478" box="[390,462,1043,1064]" name="Greece" pageId="3" pageNumber="1018">Greece</collectingCountry>
(mainland, Lesbos) and from North
<collectingCountry id="F37A763E2909B836CD64FC65FC087479" box="[857,931,1043,1065]" name="Turkey" pageId="3" pageNumber="1018">Turkey</collectingCountry>
(
<figureCitation id="13562A2B2909B836CD8EFC65FC527479" box="[947,1017,1043,1065]" captionStart="Map 1" captionStartId="4.[178,223,662,683]" captionTargetBox="[179,1086,212,653]" captionTargetId="figure@4.[179,1086,212,653]" captionTargetPageId="4" captionText="Map 1: Distributions of Myllaena graeca (based on revised records) (circles), M. mutabilis (triangles), and M. cretica (star)." figureDoi="http://doi.org/10.5281/zenodo.3776224" httpUri="https://zenodo.org/record/3776224/files/figure.png" pageId="3" pageNumber="1018">Map 1</figureCitation>
). Previous records from other regions should be regarded as erroneous for the time being. Based on external characters and on the morphology of the aedeagus, the two teneral males from Crete may belong to
<taxonomicName id="4C6D4D2D2909B836CC39FC1AFDC774D1" authorityName="Kraatz" authorityYear="1858" box="[516,620,1132,1153]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1018" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="B919EABC2909B836CC39FC1AFDC774D1" box="[516,620,1132,1153]" italics="true" pageId="3" pageNumber="1018">M. graeca</emphasis>
</taxonomicName>
, too. However, females would be required to reliably confirm the presence of this species in Crete.
</paragraph>
<paragraph id="8BD236AE2909B836CE8FFCDBFE6974AE" blockId="3.[178,1085,1197,1278]" pageId="3" pageNumber="1018">
The examined material was collected from wet moss, sifted from wet litter and debris near streams and waterfalls, or sifted from debris on a grassy roadside. The altitudes range from
<quantity id="4C959B4B2909B836CF15FC9EFE1474AE" box="[296,447,1256,1278]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="15.4" metricValueMin="1.6" pageId="3" pageNumber="1018" unit="m" value="850.0" valueMax="1540.0" valueMin="160.0">160 to 1540 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
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