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<document id="273FB36D190561894D2CF34DF1B4ADC9" ID-ISSN="1536-9307" ID-ZooBank="FAA7C294-622D-46A6-8A5C-D1541BF4B707" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638250336481" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2016" docId="03B0D50FFFF4FFC7C27AF8F9FE3CFB62" docLanguage="en" docName="Euscorpius.229.1-20.pdf" docOrigin="Euscorpius 229" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Pandinops friedrichi Kovarik 2016, sp. n." docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="FF89AD77FFF3FFCCC351FFF8FF87FFE1" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VIII. Pandinops Birula, 1913 (Scorpionidae) with description of two new species" masterLastPageNumber="20" masterPageNumber="1" pageNumber="6" updateTime="1699435275026" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="8BD733286BA418E03F6FBC3B540A232F">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VIII. Pandinops Birula, 1913 (Scorpionidae) with description of two new species</mods:title>
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<mods:date id="DAE3DBC64D53633C1B5DE4A3AFC64A4B">2016</mods:date>
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<taxonomicName id="4C191F9AFFF4FFCBC27AF8F9FDEBF8F8" ID-CoL="4CBT4" authority="Kovarik" authorityName="Kovarik" authorityYear="2016" box="[299,620,1793,1817]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="friedrichi" status="sp. nov.">
<emphasis id="B96DB80BFFF4FFCBC27AF8F9FD88F8F8" bold="true" box="[299,527,1793,1817]" italics="true" pageId="7" pageNumber="6">Pandinops friedrichi</emphasis>
Kovařík
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,
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(
<figureCitation id="1322789CFFF4FFCBC231F8D8FE52F8D6" box="[352,469,1824,1847]" captionStart-0="Figures 913" captionStart-1="Figures 1421" captionStartId-0="5.[192,278,1708,1731]" captionStartId-1="6.[192,278,1593,1616]" captionTargetBox-0="[202,1433,170,1666]" captionTargetBox-1="[195,1433,175,1558]" captionTargetId-0="figure-6@5.[192,1439,168,1678]" captionTargetId-1="figure-9@6.[192,1439,168,1563]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figures 913: Pandinops friedrichi sp. n., male holotype. Figures 910. Dorsal (9) and ventral (10) views. Scale bar: 10 mm. Figures 1113. Metasoma and telson with sternite VII in Fig. 11 and tergite VII in Fig.13, ventral (11) lateral (12) and dorsal (13) views. Scale bar: 5 mm." captionText-1="Figures 1421: Pandinops friedrichi sp. n., male holotype, pedipalp segments. Chela dorsal (14), external (15), and ventral (16). Patella dorsal (17), external (18) and ventral (19). Trochanter and femur dorsal (20) and ventral (21). Trichobothrial pattern is indicated in Figures 1420." pageId="7" pageNumber="6">Figs. 921</figureCitation>
,
<figureCitation id="1322789CFFF4FFCBC28EF8D8FE7BF8D6" box="[479,508,1824,1847]" captionStart="Figures 6168" captionStartId="18.[192,278,1718,1741]" captionTargetBox="[194,1427,215,1671]" captionTargetId="figure-9@18.[192,1439,208,1680]" captionTargetPageId="18" captionText="Figures 6168: Pandinops spp., metasoma and telson ventral. Figure 61. Pandinops bellicosus, male from &quot;Abyssinia or Egypt&quot;, possibly Eritrea. Figure 62. Pandinops colei, female holotype. Figure 63. Pandinops eritreaensis, male holotype. Figure 64. Pandinops friedrichi, sp. n., male holotype. Figure 65. Pandinops hawkeri, holotype of Pandinus pugilator. Figure 66. Pandinops peeli, male holotype. Figure 67. Pandinops pococki, male holotype. Figure 68. Pandinops turieli sp. n., female paratype. Scale bars: 10 mm." pageId="7" pageNumber="6">64</figureCitation>
,
<figureCitation id="1322789CFFF4FFCBC156F8D8FDA3F8D6" box="[519,548,1824,1847]" captionStart="Figure 69" captionStartId="19.[192,268,1374,1397]" captionTargetBox="[196,1436,149,1342]" captionTargetId="figure-6@19.[192,1439,144,1345]" captionTargetPageId="19" captionText="Figure 69: Map showing confirmed distribution of Pandinops spp. There are not included locality datas for all old specimens deposited mainly in Italian museums (for example Borelli, 1919: 374; Borelli, 1925: 16; and Kovařík &amp; Whitman, 2005: 114) that include large collections from this region, because taxonomical position of these old specimens and validity of their locality datas were not checked. Scorpions in photos inside map are holotype of Pandinops bellicosus (above) and holotype of Pandinops hawkeri (bottom)." pageId="7" pageNumber="6">69</figureCitation>
,
<tableCitation id="C69B51A2FFF4FFCBC17EF8D8FDF8F8D6" box="[559,639,1824,1847]" captionStart="Table 1" captionStartId="17.[192,257,798,821]" captionText="Table 1: Comparative measurements of adults of Pandinops friedrichi sp. n., and P. turieli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="7" pageNumber="6">Table 1</tableCitation>
)
<emphasis id="B96DB80BFFF4FFCBC393F8C7FC98F8B7" bold="true" box="[194,799,1855,1878]" pageId="7" pageNumber="6">http://zoobank.org/urn:lsid:zoobank.org:act:6093783</emphasis>
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<emphasis id="B96DB80BFFF4FFCBC249F8A6FD4EF894" bold="true" box="[280,713,1886,1909]" pageId="7" pageNumber="6">E-A723-4CB7-8C9C-BC3F066EF6AB</emphasis>
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<taxonomicName id="4C191F9AFFF4FFCBC06FFF3FFAF4FF3F" authority=": Kovarik, 2003: 150" authorityName=": Kovarik" authorityPageNumber="150" authorityYear="2003" box="[830,1395,199,222]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="peeli" subGenus="Pandinops">
<emphasis id="B96DB80BFFF4FFCBC06FFF3FFC24FF3F" box="[830,931,199,222]" italics="true" pageId="7" pageNumber="6">Pandinus</emphasis>
(
<emphasis id="B96DB80BFFF4FFCBC0EDFF3FFBA9FF3F" box="[956,1070,199,222]" italics="true" pageId="7" pageNumber="6">Pandinops</emphasis>
)
<emphasis id="B96DB80BFFF4FFCBC717FF3FFBFDFF3F" box="[1094,1146,199,222]" italics="true" pageId="7" pageNumber="6">peeli</emphasis>
: Kovařík, 2003: 150
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(in part); Kovařík &amp; Whitman, 2005: 114; Kovařík, 2009: 53, 118, figs. 323324 (in part).
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,
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.1989, leg.
<collectorName id="26EC01CFFFF4FFCBC0BEFE44FBC2FE32" box="[1007,1093,444,467]" pageId="7" pageNumber="6">Politzar</collectorName>
,
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,
<figureCitation id="1322789CFFF4FFCBC7A2FE44FAEFFE32" box="[1267,1384,444,467]" captionStart-0="Figures 913" captionStart-1="Figures 1421" captionStartId-0="5.[192,278,1708,1731]" captionStartId-1="6.[192,278,1593,1616]" captionTargetBox-0="[202,1433,170,1666]" captionTargetBox-1="[195,1433,175,1558]" captionTargetId-0="figure-6@5.[192,1439,168,1678]" captionTargetId-1="figure-9@6.[192,1439,168,1563]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figures 913: Pandinops friedrichi sp. n., male holotype. Figures 910. Dorsal (9) and ventral (10) views. Scale bar: 10 mm. Figures 1113. Metasoma and telson with sternite VII in Fig. 11 and tergite VII in Fig.13, ventral (11) lateral (12) and dorsal (13) views. Scale bar: 5 mm." captionText-1="Figures 1421: Pandinops friedrichi sp. n., male holotype, pedipalp segments. Chela dorsal (14), external (15), and ventral (16). Patella dorsal (17), external (18) and ventral (19). Trochanter and femur dorsal (20) and ventral (21). Trichobothrial pattern is indicated in Figures 1420." pageId="7" pageNumber="6">Figs. 921</figureCitation>
,
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from
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(figs.
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, 2009: 118),
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.
</paragraph>
</subSubSection>
<subSubSection id="C3033792FFF4FFCBC06FFD4AFB49FD06" pageId="7" pageNumber="6" type="etymology">
<paragraph id="8BA66419FFF4FFCBC06FFD4AFB49FD06" blockId="7.[830,1440,690,743]" pageId="7" pageNumber="6">
ETYMOLOGY. It is a pleasure to name this species after Stefan Friedrich (Munich,
<collectingCountry id="F30E2489FFF4FFCBC70AFD28FB44FD06" box="[1115,1219,720,743]" name="Germany" pageId="7" pageNumber="6">Germany</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3033792FFF4FFCBC06FFCF6FACBFA8A" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8BA66419FFF4FFCBC06FFCF6FACBFA8A" blockId="7.[830,1441,782,1387]" pageId="7" pageNumber="6">
DIAGNOSIS. Total length
<quantity id="4CE1C9FCFFF4FFCBC731FCF6FB59FCC4" box="[1120,1246,782,805]" metricMagnitude="-2" metricUnit="m" metricValue="7.05" metricValueMax="8.0" metricValueMin="6.1" pageId="7" pageNumber="6" unit="mm" value="70.5" valueMax="80.0" valueMin="61.0">6180 mm</quantity>
. Base color uniformly reddish brown to black, legs and telson yellow, pedipalp chela orange to reddish brown. Carapace smooth in middle, covered by granules along margins. External trichobothria on patella number 1415 (5
<emphasis id="B96DB80BFFF4FFCBC63BFC70FA04FC7E" box="[1386,1411,904,927]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
, 3
<emphasis id="B96DB80BFFF4FFCBC06FFC5FFCE5FC5F" box="[830,866,935,958]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
, 2
<emphasis id="B96DB80BFFF4FFCBC0D7FC5FFC22FC5F" box="[902,933,935,958]" italics="true" pageId="7" pageNumber="6">em</emphasis>
, 12
<emphasis id="B96DB80BFFF4FFCBC0B5FC5FFB85FC5F" box="[996,1026,935,958]" italics="true" pageId="7" pageNumber="6">est</emphasis>
, 3
<emphasis id="B96DB80BFFF4FFCBC777FC5FFBBEFC5F" box="[1062,1081,935,958]" italics="true" pageId="7" pageNumber="6">et</emphasis>
); ventral trichobothria on patella number 2228; internal trichobothria on chela number 67, ventral trichobothria on chela number 912. Pedipalp chela hirsute rather sparsely. Pedipalp chela dorsally smooth to bumpy, without pointed granules, lobe smooth. Chela internal smooth sparsely granulated mainly in anterior part, with two longitudinal carinae indicated by 46 granules. Chela of male length/ width ratio is 1.621.69. Pectinal teeth number
<quantity id="4CE1C9FCFFF4FFCBC7B6FB64FACAFB52" box="[1255,1357,1180,1203]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="7" pageNumber="6" unit="in" value="15.5" valueMax="16.0" valueMin="15.0">1516 in</quantity>
males. Sternite VII densely granulated. Metasomal segments I V ventrally densely granulated; metasomal segments I IV with ventral carinae absent. Spiniform formula of tarsomere II = 3/4: 3/4: 3/5: 3/5. Tarsomere II with 2 spines on inclined anteroventral surface. Length to width ratio of male metasomal segment V is 2.142.17.
</paragraph>
</subSubSection>
<subSubSection id="C3033792FFF4FFC7C06FFA6AFE3CFB62" lastPageId="11" lastPageNumber="10" pageId="7" pageNumber="6" type="description">
<paragraph id="8BA66419FFF4FFCBC06FFA6AFB95F99E" blockId="7.[830,1441,1426,1909]" pageId="7" pageNumber="6">
DESCRIPTION. The adult males are
<quantity id="4CE1C9FCFFF4FFCBC792FA6AFABFFA48" box="[1219,1336,1426,1449]" metricMagnitude="-2" metricUnit="m" metricValue="7.05" metricValueMax="8.0" metricValueMin="6.1" pageId="7" pageNumber="6" unit="mm" value="70.5" valueMax="80.0" valueMin="61.0">6180 mm</quantity>
long, female unknown. The habitus is shown in
<figureCitation id="1322789CFFF4FFCBC7ABFA48FAE8FA26" box="[1274,1391,1456,1479]" captionStart="Figures 913" captionStartId="5.[192,278,1708,1731]" captionTargetBox="[202,1433,170,1666]" captionTargetId="figure-6@5.[192,1439,168,1678]" captionTargetPageId="5" captionText="Figures 913: Pandinops friedrichi sp. n., male holotype. Figures 910. Dorsal (9) and ventral (10) views. Scale bar: 10 mm. Figures 1113. Metasoma and telson with sternite VII in Fig. 11 and tergite VII in Fig.13, ventral (11) lateral (12) and dorsal (13) views. Scale bar: 5 mm." pageId="7" pageNumber="6">Figs. 910</figureCitation>
. For position and distribution of the trichobothria of pedipalps see
<figureCitation id="1322789CFFF4FFCBC0FDFA16FBB6F9E4" box="[940,1073,1518,1541]" captionStart="Figures 1421" captionStartId="6.[192,278,1593,1616]" captionTargetBox="[195,1433,175,1558]" captionTargetId="figure-9@6.[192,1439,168,1563]" captionTargetPageId="6" captionText="Figures 1421: Pandinops friedrichi sp. n., male holotype, pedipalp segments. Chela dorsal (14), external (15), and ventral (16). Patella dorsal (17), external (18) and ventral (19). Trochanter and femur dorsal (20) and ventral (21). Trichobothrial pattern is indicated in Figures 1420." pageId="7" pageNumber="6">Figs. 1420</figureCitation>
. External trichobothria on patella number 1415 (5
<emphasis id="B96DB80BFFF4FFCBC753F9F4FB9CF9C2" box="[1026,1051,1548,1571]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
, 3
<emphasis id="B96DB80BFFF4FFCBC711F9F4FBE4F9C2" box="[1088,1123,1548,1571]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
, 2
<emphasis id="B96DB80BFFF4FFCBC7D8F9F4FB2FF9C2" box="[1161,1192,1548,1571]" italics="true" pageId="7" pageNumber="6">em</emphasis>
, 12
<emphasis id="B96DB80BFFF4FFCBC7B9F9F4FA81F9C2" box="[1256,1286,1548,1571]" italics="true" pageId="7" pageNumber="6">est</emphasis>
, 3
<emphasis id="B96DB80BFFF4FFCBC67BF9F4FABAF9C2" box="[1322,1341,1548,1571]" italics="true" pageId="7" pageNumber="6">et</emphasis>
); ventral trichobothria on patella number 2228; internal trichobothria on chela number 67, ventral trichobothria on chela number 912.
</paragraph>
<paragraph id="8BA66419FFF4FFCBC06FF97FFB18F8F8" blockId="7.[830,1441,1426,1909]" pageId="7" pageNumber="6">
<emphasis id="B96DB80BFFF4FFCBC06FF97FFC3EF97F" bold="true" box="[830,953,1671,1694]" pageId="7" pageNumber="6">Coloration</emphasis>
(
<figureCitation id="1322789CFFF4FFCBC09EF97FFBCDF97F" box="[975,1098,1671,1694]" captionStart="Figures 913" captionStartId="5.[192,278,1708,1731]" captionTargetBox="[202,1433,170,1666]" captionTargetId="figure-6@5.[192,1439,168,1678]" captionTargetPageId="5" captionText="Figures 913: Pandinops friedrichi sp. n., male holotype. Figures 910. Dorsal (9) and ventral (10) views. Scale bar: 10 mm. Figures 1113. Metasoma and telson with sternite VII in Fig. 11 and tergite VII in Fig.13, ventral (11) lateral (12) and dorsal (13) views. Scale bar: 5 mm." pageId="7" pageNumber="6">Figs. 910</figureCitation>
). The base color is uniformly reddish brown to black, legs and telson are yellow, pedipalp chela is orange to reddish brown, chelicerae are yellow to orange, and carapace is reddish brown to black, lighter in the anterior part.
</paragraph>
<paragraph id="8BA66419FFF4FFC4C06FF8D8FD77F8CF" blockId="7.[830,1441,1426,1909]" lastBlockId="8.[192,802,1600,1899]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<emphasis id="B96DB80BFFF4FFCBC06FF8D8FBDCF8D6" bold="true" box="[830,1115,1824,1847]" pageId="7" pageNumber="6">Carapace and mesosoma</emphasis>
(
<figureCitation id="1322789CFFF4FFCBC73AF8D8FB65F8D6" box="[1131,1250,1824,1847]" captionStart="Figures 913" captionStartId="5.[192,278,1708,1731]" captionTargetBox="[202,1433,170,1666]" captionTargetId="figure-6@5.[192,1439,168,1678]" captionTargetPageId="5" captionText="Figures 913: Pandinops friedrichi sp. n., male holotype. Figures 910. Dorsal (9) and ventral (10) views. Scale bar: 10 mm. Figures 1113. Metasoma and telson with sternite VII in Fig. 11 and tergite VII in Fig.13, ventral (11) lateral (12) and dorsal (13) views. Scale bar: 5 mm." pageId="7" pageNumber="6">Figs. 910</figureCitation>
). The entire carapace is smooth in the middle, covered by granules along margins. The anterior margin of the carapace is symmetrically concave, medially strongly convex, and it bears several macrosetae. The tergites are smooth and can be very finely granulated, mainly tergite VI; tergite VII is strongly granulated. The pectinal tooth count is
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males, female unknown. The pectine marginal tips extend to quarter of the fourth sternite in the male. The sternites are smooth, without carinae, but with two longitudinal furrows; sternite VII is densely granulated.
</paragraph>
<caption id="DF663491FFFBFFC4C391FA24FCF8F9EA" pageId="8" pageNumber="7" startId="8.[192,278,1500,1523]" targetBox="[205,1428,148,1461]" targetPageId="8">
<paragraph id="8BA66419FFFBFFC4C391FA24FCF8F9EA" blockId="8.[192,1440,1500,1547]" pageId="8" pageNumber="7">
<emphasis id="B96DB80BFFFBFFC4C391FA24FEEDFA12" bold="true" box="[192,362,1500,1523]" pageId="8" pageNumber="7">Figures 2225:</emphasis>
<taxonomicName id="4C191F9AFFFBFFC4C220FA26FD9CFA12" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" box="[369,539,1502,1523]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
<emphasis id="B96DB80BFFFBFFC4C220FA26FD9CFA12" box="[369,539,1502,1523]" italics="true" pageId="8" pageNumber="7">Pandinops turieli</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFFBFFC4C173FA26FD75FA12" bold="true" box="[546,754,1502,1523]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="A25E0570FFFBFFC4C173FA26FDDDFA12" box="[546,602,1502,1523]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
Figures 2223
</emphasis>
. Male holotype in dorsal (22) and ventral (23) views.
<emphasis id="B96DB80BFFFBFFC4C658FA26FA1DFA12" bold="true" box="[1289,1434,1502,1523]" pageId="8" pageNumber="7">Figures 2425</emphasis>
. Female paratype in dorsal (24) and ventral (25) views. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="8BA66419FFFBFFC4C391F8CEFAE2F8CF" blockId="8.[192,802,1600,1899]" lastBlockId="8.[830,1440,1600,1899]" pageId="8" pageNumber="7">
<emphasis id="B96DB80BFFFBFFC4C391F8CEFE38F8AC" bold="true" box="[192,447,1846,1869]" pageId="8" pageNumber="7">Metasoma and telson</emphasis>
(
<figureCitation id="1322789CFFFBFFC4C284F8CEFDDAF8AC" box="[469,605,1846,1869]" captionStart="Figures 913" captionStartId="5.[192,278,1708,1731]" captionTargetBox="[202,1433,170,1666]" captionTargetId="figure-6@5.[192,1439,168,1678]" captionTargetPageId="5" captionText="Figures 913: Pandinops friedrichi sp. n., male holotype. Figures 910. Dorsal (9) and ventral (10) views. Scale bar: 10 mm. Figures 1113. Metasoma and telson with sternite VII in Fig. 11 and tergite VII in Fig.13, ventral (11) lateral (12) and dorsal (13) views. Scale bar: 5 mm." pageId="8" pageNumber="7">Figs. 1113</figureCitation>
). The metasomal segments IIV bear a total of 6 sparsely granulated carinae. The ventral carinae are absent. The fifth segment has five variously developed and granulated carinae. The dorsal and lateral surfaces of the segments are smooth with several solitary granules. The metasomal segments IV ventrally densely granulated with the same intensity on all segments. The entire metasoma and telson are sparsely hirsute. The telson is smooth, elongate, with the aculeus approximately as long as vesicle or shorter.
</paragraph>
<paragraph id="8BA66419FFFBFFC5C06FF8CEFB35F921" blockId="8.[830,1440,1600,1899]" lastBlockId="9.[830,1441,1643,1912]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="B96DB80BFFFBFFC4C06FF8CEFC2AF8AC" bold="true" box="[830,941,1846,1869]" pageId="8" pageNumber="7">Pedipalps</emphasis>
(
<figureCitation id="1322789CFFFBFFC4C0EFF8CEFBC5F8AC" box="[958,1090,1846,1869]" captionStart="Figures 1421" captionStartId="6.[192,278,1593,1616]" captionTargetBox="[195,1433,175,1558]" captionTargetId="figure-9@6.[192,1439,168,1563]" captionTargetPageId="6" captionText="Figures 1421: Pandinops friedrichi sp. n., male holotype, pedipalp segments. Chela dorsal (14), external (15), and ventral (16). Patella dorsal (17), external (18) and ventral (19). Trochanter and femur dorsal (20) and ventral (21). Trichobothrial pattern is indicated in Figures 1420." pageId="8" pageNumber="7">Figs. 1421</figureCitation>
). The pedipalps are hirsute. The femur is granulated dorsally and bears four carinae composed of several strong granules. The patella is smooth, finely granulated on internal surface only, there are two externo lateral smooth carinae and one internal carina is indicated by 57 big granules. Pedipalp chela is dorsally smooth to bumpy, without carinae and pointed granules, lobe smooth. Several well defined granules are present on external suface. The chela internal is smooth, sparsely granulated mainly in anterior part, with two short characteristic smooth longitudinal carinae indicated by 46 granules. The movable and fixed fingers of the pedipalp with distinct granules in a row divided into 57 rows by big concave granules.
</paragraph>
<caption id="DF663491FFFAFFC5C391FA0FFBD6F9DE" pageId="9" pageNumber="8" startId="9.[192,278,1527,1550]" targetBox="[191,1437,167,1497]" targetPageId="9">
<paragraph id="8BA66419FFFAFFC5C391FA0FFBD6F9DE" blockId="9.[192,1440,1527,1599]" pageId="9" pageNumber="8">
<emphasis id="B96DB80BFFFAFFC5C391FA0FFEEAF9EF" bold="true" box="[192,365,1527,1550]" pageId="9" pageNumber="8">Figures 2631:</emphasis>
<taxonomicName id="4C191F9AFFFAFFC5C228FA01FDA1F9EF" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" box="[377,550,1529,1550]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
<emphasis id="B96DB80BFFFAFFC5C228FA01FDA1F9EF" box="[377,550,1529,1550]" italics="true" pageId="9" pageNumber="8">Pandinops turieli</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFFAFFC5C160FA01FD60F9EF" bold="true" box="[561,743,1529,1550]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A25E0570FFFAFFC5C160FA01FDEBF9EF" box="[561,620,1529,1550]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
Figures 26
</emphasis>
,
<emphasis id="B96DB80BFFFAFFC5C1A9FA01FC97F9EF" bold="true" box="[760,784,1529,1550]" pageId="9" pageNumber="8">28</emphasis>
. Male holotype, chelicerae, carapace and tergites IIII (26) and coxosternal area with sternites IIIIV (28).
<emphasis id="B96DB80BFFFAFFC5C128F9EAFD6FF9C6" bold="true" box="[633,744,1554,1575]" pageId="9" pageNumber="8">Figures 27</emphasis>
,
<emphasis id="B96DB80BFFFAFFC5C1A9F9EAFCB3F9C6" bold="true" box="[760,820,1554,1575]" pageId="9" pageNumber="8">2931</emphasis>
. Female paratype, chelicerae, carapace and tergites III (27), coxosternal area and sternites IIIIV (29), and left chelicera dorsal (30) and ventral (31) views.
</paragraph>
</caption>
<paragraph id="8BA66419FFFAFFC5C06FF93FFC15F8B8" blockId="9.[830,1441,1643,1912]" pageId="9" pageNumber="8">
<emphasis id="B96DB80BFFFAFFC5C06FF93FFCF4F93F" bold="true" box="[830,883,1735,1758]" pageId="9" pageNumber="8">Legs</emphasis>
. All legs are without distinct carinae and smooth. The tarsomeres are hirsute with setae and macrosetae more densely on legs III. For spiniform formula of tarsomeres of legs see paragraph under
<taxonomicName id="4C191F9AFFFAFFC5C67FF8DBFA27F8DB" authorityName="Birula" authorityYear="1913" box="[1326,1440,1827,1850]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B96DB80BFFFAFFC5C67FF8DBFA27F8DB" box="[1326,1440,1827,1850]" italics="true" pageId="9" pageNumber="8">Pandinops</emphasis>
</taxonomicName>
section.
</paragraph>
<paragraph id="8BA66419FFFAFFC5C06FF899FBFFF899" blockId="9.[830,1441,1643,1912]" box="[830,1144,1889,1912]" pageId="9" pageNumber="8">
<emphasis id="B96DB80BFFFAFFC5C06FF899FC62F899" bold="true" box="[830,997,1889,1912]" pageId="9" pageNumber="8">Measurements</emphasis>
. See
<tableCitation id="C69B51A2FFFAFFC5C771F899FBFFF899" box="[1056,1144,1889,1912]" captionStart="Table 1" captionStartId="17.[192,257,798,821]" captionText="Table 1: Comparative measurements of adults of Pandinops friedrichi sp. n., and P. turieli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="9" pageNumber="8">Table 1.</tableCitation>
</paragraph>
<caption id="DF663491FFF9FFC6C391F9B0FD5CF94B" pageId="10" pageNumber="9" startId="10.[192,278,1608,1631]" targetBox="[192,1439,177,1570]" targetPageId="10">
<paragraph id="8BA66419FFF9FFC6C391F9B0FD5CF94B" blockId="10.[192,1441,1608,1706]" pageId="10" pageNumber="9">
<emphasis id="B96DB80BFFF9FFC6C391F9B0FEEDF9BE" bold="true" box="[192,362,1608,1631]" pageId="10" pageNumber="9">Figures 3244:</emphasis>
<taxonomicName id="4C191F9AFFF9FFC6C223F9B2FD9BF9BE" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" box="[370,540,1610,1631]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
<emphasis id="B96DB80BFFF9FFC6C223F9B2FD9BF9BE" box="[370,540,1610,1631]" italics="true" pageId="10" pageNumber="9">Pandinops turieli</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFF9FFC6C172F9B2FDDCF9BE" bold="true" box="[547,603,1610,1631]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A25E0570FFF9FFC6C172F9B2FDDCF9BE" box="[547,603,1610,1631]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, pedipalp segments.
<emphasis id="B96DB80BFFF9FFC6C07AF9B2FC3BF9BE" bold="true" box="[811,956,1610,1631]" pageId="10" pageNumber="9">Figures 3235</emphasis>
. Male holotype. Chela dorsal (32), external (33), and ventral (34). Patella external (35).
<emphasis id="B96DB80BFFF9FFC6C167F99BFD42F999" bold="true" box="[566,709,1635,1656]" pageId="10" pageNumber="9">Figures 3644</emphasis>
. Female paratype. Chela dorsal (36), external (37), and ventral (38). Patella dorsal (39), external (40) and ventral (41). Trochanter and femur dorsal (42). Fixed (43) and movable finger (44) dentitions. Trichobothrial pattern is indicated in Figures 35, 3642.
</paragraph>
</caption>
<paragraph id="8BA66419FFF9FFC6C391F927FA1DF891" blockId="10.[192,801,1759,1904]" lastBlockId="10.[830,1441,1759,1904]" pageId="10" pageNumber="9">
AFFINITIES. The described features distinguish
<taxonomicName id="4C191F9AFFF9FFC6C197F927FF75F8F5" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="friedrichi" status="sp. nov.">
<emphasis id="B96DB80BFFF9FFC6C197F927FF75F8F5" italics="true" pageId="10" pageNumber="9">P. friedrichi</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFF9FFC6C3ABF905FEB0F8F5" bold="true" box="[250,311,1789,1812]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A25E0570FFF9FFC6C3ABF905FEB0F8F5" box="[250,311,1789,1812]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from all other species of the genus. They are recounted in the key below. Kovařík (2009: 53, 118, figs. 323324) misidentified the specimen from MZUF collection as
<taxonomicName id="4C191F9AFFF9FFC6C205F8A1FE2DF891" authorityName=": Kovarik" authorityYear="2003" box="[340,426,1881,1904]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="peeli">
<emphasis id="B96DB80BFFF9FFC6C205F8A1FE2DF891" box="[340,426,1881,1904]" italics="true" pageId="10" pageNumber="9">P. peeli</emphasis>
</taxonomicName>
, which has also granulated metasomal segments III ventrally but differently than
<taxonomicName id="4C191F9AFFF9FFC6C6D8F927FC24F8F5" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="friedrichi" status="sp. nov.">
<emphasis id="B96DB80BFFF9FFC6C6D8F927FC24F8F5" italics="true" pageId="10" pageNumber="9">P. friedrichi</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFF9FFC6C0FDF905FC6DF8F5" bold="true" box="[940,1002,1789,1812]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A25E0570FFF9FFC6C0FDF905FC6DF8F5" box="[940,1002,1789,1812]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(see the key below and
<figureCitation id="1322789CFFF9FFC6C7AAF905FAD4F8F5" box="[1275,1363,1789,1812]" captionStart="Figures 6168" captionStartId="18.[192,278,1718,1741]" captionTargetBox="[194,1427,215,1671]" captionTargetId="figure-9@18.[192,1439,208,1680]" captionTargetPageId="18" captionText="Figures 6168: Pandinops spp., metasoma and telson ventral. Figure 61. Pandinops bellicosus, male from &quot;Abyssinia or Egypt&quot;, possibly Eritrea. Figure 62. Pandinops colei, female holotype. Figure 63. Pandinops eritreaensis, male holotype. Figure 64. Pandinops friedrichi, sp. n., male holotype. Figure 65. Pandinops hawkeri, holotype of Pandinus pugilator. Figure 66. Pandinops peeli, male holotype. Figure 67. Pandinops pococki, male holotype. Figure 68. Pandinops turieli sp. n., female paratype. Scale bars: 10 mm." pageId="10" pageNumber="9">Figs. 64</figureCitation>
versus 66). These two species occur in different areas of distribution.
<taxonomicName id="4C191F9AFFF9FFC6C0FDF8C3FB85F8B3" authorityName=": Kovarik" authorityYear="2003" box="[940,1026,1851,1874]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="peeli">
<emphasis id="B96DB80BFFF9FFC6C0FDF8C3FB85F8B3" box="[940,1026,1851,1874]" italics="true" pageId="10" pageNumber="9">P. peeli</emphasis>
</taxonomicName>
is known from the northern Somaliland, and
<taxonomicName id="4C191F9AFFF9FFC6C0E5F8A1FBB0F891" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" box="[948,1079,1881,1904]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="friedrichi" status="sp. nov.">
<emphasis id="B96DB80BFFF9FFC6C0E5F8A1FBB0F891" box="[948,1079,1881,1904]" italics="true" pageId="10" pageNumber="9">P. friedrichi</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFF9FFC6C76CF8A1FBFEF891" bold="true" box="[1085,1145,1881,1904]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A25E0570FFF9FFC6C76CF8A1FBFEF891" box="[1085,1145,1881,1904]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from south
<collectingCountry id="F30E2489FFF9FFC6C659F8A1FAD8F891" box="[1288,1375,1881,1904]" name="Somalia" pageId="10" pageNumber="9">Somalia</collectingCountry>
(
<figureCitation id="1322789CFFF9FFC6C63FF8A1FA1DF891" box="[1390,1434,1881,1904]" captionStart="Figure 69" captionStartId="19.[192,268,1374,1397]" captionTargetBox="[196,1436,149,1342]" captionTargetId="figure-6@19.[192,1439,144,1345]" captionTargetPageId="19" captionText="Figure 69: Map showing confirmed distribution of Pandinops spp. There are not included locality datas for all old specimens deposited mainly in Italian museums (for example Borelli, 1919: 374; Borelli, 1925: 16; and Kovařík &amp; Whitman, 2005: 114) that include large collections from this region, because taxonomical position of these old specimens and validity of their locality datas were not checked. Scorpions in photos inside map are holotype of Pandinops bellicosus (above) and holotype of Pandinops hawkeri (bottom)." pageId="10" pageNumber="9">Fig.</figureCitation>
</paragraph>
<caption id="DF663491FFF8FFC7C391FC11FA11FBF9" pageId="11" pageNumber="10" startId="11.[192,278,1001,1024]" targetBox="[201,1430,149,964]" targetPageId="11">
<paragraph id="8BA66419FFF8FFC7C391FC11FA11FBF9" blockId="11.[192,1440,1001,1048]" pageId="11" pageNumber="10">
<emphasis id="B96DB80BFFF8FFC7C391FC11FEEAFBE1" bold="true" box="[192,365,1001,1024]" pageId="11" pageNumber="10">Figures 4548:</emphasis>
<taxonomicName id="4C191F9AFFF8FFC7C228FC12FDA2FC1E" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" box="[377,549,1002,1023]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
<emphasis id="B96DB80BFFF8FFC7C228FC12FDA2FC1E" box="[377,549,1002,1023]" italics="true" pageId="11" pageNumber="10">Pandinops turieli</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFF8FFC7C161FC12FDEBFC1E" bold="true" box="[560,620,1002,1023]" pageId="11" pageNumber="10">
<taxonomicNameLabel id="A25E0570FFF8FFC7C161FC12FDEBFC1E" box="[560,620,1002,1023]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female paratype, right legs IIV, retrolateral aspect. Abbreviations:
<emphasis id="B96DB80BFFF8FFC7C660FC12FACBFC1E" box="[1329,1356,1002,1023]" italics="true" pageId="11" pageNumber="10">vst</emphasis>
, ventral subterminal;
<emphasis id="B96DB80BFFF8FFC7C26FFBFBFEC8FBF9" box="[318,335,1027,1048]" italics="true" pageId="11" pageNumber="10">vt</emphasis>
, ventral terminal;
<emphasis id="B96DB80BFFF8FFC7C150FBFBFD9EFBF9" box="[513,537,1027,1048]" italics="true" pageId="11" pageNumber="10">pd</emphasis>
, prolateral distal;
<emphasis id="B96DB80BFFF8FFC7C196FBFBFD64FBF9" box="[711,739,1027,1048]" italics="true" pageId="11" pageNumber="10">pst</emphasis>
, prolateral subterminal;
<emphasis id="B96DB80BFFF8FFC7C09EFBFBFC6EFBF9" box="[975,1001,1027,1048]" italics="true" pageId="11" pageNumber="10">rm</emphasis>
, retrolateral medial;
<emphasis id="B96DB80BFFF8FFC7C7E2FBFBFB44FBF9" box="[1203,1219,1027,1048]" italics="true" pageId="11" pageNumber="10">rt</emphasis>
, retrolateral terminal.
</paragraph>
</caption>
<paragraph id="8BA66419FFF8FFC7C391FBB5FE3CFB62" blockId="11.[192,802,1101,1155]" pageId="11" pageNumber="10">
69) where it represents the southern limits of the genus
<taxonomicName id="4C191F9AFFF8FFC7C391FB94FEB5FB62" authorityName="Birula" authorityYear="1913" box="[192,306,1132,1155]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B96DB80BFFF8FFC7C391FB94FEB5FB62" box="[192,306,1132,1155]" italics="true" pageId="11" pageNumber="10">Pandinops</emphasis>
</taxonomicName>
distribution.
</paragraph>
</subSubSection>
</treatment>
</document>