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<mods:title id="C2FFDCFDCD098B3C2AB9DEA2B73ABDE6">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VIII. Pandinops Birula, 1913 (Scorpionidae) with description of two new species</mods:title>
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<paragraph id="8BA66419FFF0FFCFC221FB74FDF6FB45" blockId="3.[299,695,1134,1219]" box="[368,625,1164,1188]" pageId="3" pageNumber="2">
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<taxonomicName id="4C191F9AFFF0FFCFC221FB74FDF6FB45" ID-CoL="6D6C" authority="Birula, 1913" authorityName="Birula" authorityYear="1913" box="[368,625,1164,1188]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B96DB80BFFF0FFCFC221FB74FE62FB45" bold="true" box="[368,485,1164,1188]" italics="true" pageId="3" pageNumber="2">Pandinops</emphasis>
Birula, 1913
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3033792FFF0FFCFC2D1FB54FE0DFA83" pageId="3" pageNumber="2" type="description">
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(
<figureCitation id="1322789CFFF0FFCFC2D9FB54FE7BFB22" box="[392,508,1196,1219]" captionStart-0="Figures 18" captionStart-1="Figures 913" captionStart-10="Figures 5960" captionStart-11="Figures 6168" captionStart-12="Figure 69" captionStart-2="Figures 1421" captionStart-3="Figures 2225" captionStart-4="Figures 2631" captionStart-5="Figures 3244" captionStart-6="Figures 4548" captionStart-7="Figures 4954" captionStart-8="Figures 5556" captionStart-9="Figures 5758" captionStartId-0="4.[192,278,1409,1432]" captionStartId-1="5.[192,278,1708,1731]" captionStartId-10="16.[192,278,1752,1775]" captionStartId-11="18.[192,278,1718,1741]" captionStartId-12="19.[192,268,1374,1397]" captionStartId-2="6.[192,278,1593,1616]" captionStartId-3="8.[192,278,1500,1523]" captionStartId-4="9.[192,278,1527,1550]" captionStartId-5="10.[192,278,1608,1631]" captionStartId-6="11.[192,278,1001,1024]" captionStartId-7="12.[192,278,1393,1416]" captionStartId-8="13.[192,278,1683,1706]" captionStartId-9="15.[192,278,1692,1715]" captionTargetBox-0="[202,1433,176,1376]" captionTargetBox-1="[202,1433,170,1666]" captionTargetBox-10="[191,1440,167,1723]" captionTargetBox-11="[194,1427,215,1671]" captionTargetBox-12="[196,1436,149,1342]" captionTargetBox-2="[195,1433,175,1558]" captionTargetBox-3="[205,1428,148,1461]" captionTargetBox-4="[191,1437,167,1497]" captionTargetBox-5="[192,1439,177,1570]" captionTargetBox-6="[201,1430,149,964]" captionTargetBox-7="[199,1439,180,1350]" captionTargetBox-8="[191,1440,167,1654]" captionTargetBox-9="[191,1440,133,1663]" captionTargetId-0="figure-9@4.[192,1437,168,1379]" captionTargetId-1="figure-6@5.[192,1439,168,1678]" captionTargetId-10="figure-9@16.[192,1439,168,1722]" captionTargetId-11="figure-9@18.[192,1439,208,1680]" captionTargetId-12="figure-6@19.[192,1439,144,1345]" captionTargetId-2="figure-9@6.[192,1439,168,1563]" captionTargetId-3="figure-9@8.[192,1439,144,1470]" captionTargetId-4="figure-6@9.[192,1437,168,1497]" captionTargetId-5="figure-9@10.[192,1439,168,1579]" captionTargetId-6="figure-6@11.[192,1437,144,971]" captionTargetId-7="figure-9@12.[192,1439,168,1364]" captionTargetId-8="figure-6@13.[192,1439,168,1653]" captionTargetId-9="figure-6@15.[192,1439,144,1662]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="16" captionTargetPageId-11="18" captionTargetPageId-12="19" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="15" captionText-0="Figures 18: Figures 14. Pandinops eritreaensis, male holotype, pedipalp chela dorsal (1), external (2) and ventral (3) and patella external (4). Figures 57. Pandinops pococki, male holotype, pedipalp chela dorsal (5), external (6) and ventral (7). Figure 8. Pandinops bellicosus, male from &quot;Abyssinia or Egypt&quot;, possibly Eritrea, pedipalp chela dorsal." captionText-1="Figures 913: Pandinops friedrichi sp. n., male holotype. Figures 910. Dorsal (9) and ventral (10) views. Scale bar: 10 mm. Figures 1113. Metasoma and telson with sternite VII in Fig. 11 and tergite VII in Fig.13, ventral (11) lateral (12) and dorsal (13) views. Scale bar: 5 mm." captionText-10="Figures 5960: Pandinops turieli sp. n., burrows at the type locality." captionText-11="Figures 6168: Pandinops spp., metasoma and telson ventral. Figure 61. Pandinops bellicosus, male from &quot;Abyssinia or Egypt&quot;, possibly Eritrea. Figure 62. Pandinops colei, female holotype. Figure 63. Pandinops eritreaensis, male holotype. Figure 64. Pandinops friedrichi, sp. n., male holotype. Figure 65. Pandinops hawkeri, holotype of Pandinus pugilator. Figure 66. Pandinops peeli, male holotype. Figure 67. Pandinops pococki, male holotype. Figure 68. Pandinops turieli sp. n., female paratype. Scale bars: 10 mm." captionText-12="Figure 69: Map showing confirmed distribution of Pandinops spp. There are not included locality datas for all old specimens deposited mainly in Italian museums (for example Borelli, 1919: 374; Borelli, 1925: 16; and Kovařík &amp; Whitman, 2005: 114) that include large collections from this region, because taxonomical position of these old specimens and validity of their locality datas were not checked. Scorpions in photos inside map are holotype of Pandinops bellicosus (above) and holotype of Pandinops hawkeri (bottom)." captionText-2="Figures 1421: Pandinops friedrichi sp. n., male holotype, pedipalp segments. Chela dorsal (14), external (15), and ventral (16). Patella dorsal (17), external (18) and ventral (19). Trochanter and femur dorsal (20) and ventral (21). Trichobothrial pattern is indicated in Figures 1420." captionText-3="Figures 2225: Pandinops turieli sp. n. Figures 2223. Male holotype in dorsal (22) and ventral (23) views. Figures 2425. Female paratype in dorsal (24) and ventral (25) views. Scale bar: 10 mm." captionText-4="Figures 2631: Pandinops turieli sp. n. Figures 26, 28. Male holotype, chelicerae, carapace and tergites IIII (26) and coxosternal area with sternites IIIIV (28). Figures 27, 2931. Female paratype, chelicerae, carapace and tergites III (27), coxosternal area and sternites IIIIV (29), and left chelicera dorsal (30) and ventral (31) views." captionText-5="Figures 3244: Pandinops turieli sp. n., pedipalp segments. Figures 3235. Male holotype. Chela dorsal (32), external (33), and ventral (34). Patella external (35). Figures 3644. Female paratype. Chela dorsal (36), external (37), and ventral (38). Patella dorsal (39), external (40) and ventral (41). Trochanter and femur dorsal (42). Fixed (43) and movable finger (44) dentitions. Trichobothrial pattern is indicated in Figures 35, 3642." captionText-6="Figures 4548: Pandinops turieli sp. n., female paratype, right legs IIV, retrolateral aspect. Abbreviations: vst, ventral subterminal; vt, ventral terminal; pd, prolateral distal; pst, prolateral subterminal; rm, retrolateral medial; rt, retrolateral terminal." captionText-7="Figures 4954: Pandinops turieli sp. n., metasoma and telson. Figures 4951. Male holotype, lateral (49), dorsal (50), and ventral (51) views. Figures 5254. Female paratype, lateral (52), dorsal (53), and ventral (54) views. Scale bar: 10 mm." captionText-8="Figures 5556: Pandinops turieli sp. n., in vivo habitus. Female (55) and male (56) paratypes." captionText-9="Figures 5758: Pandinops turieli sp. n., female paratype eating cricket in laboratory (57) and the type locality (58)." pageId="3" pageNumber="2">Figs. 169</figureCitation>
,
<tableCitation id="C69B51A2FFF0FFCFC156FB54FDD0FB22" box="[519,599,1196,1219]" captionStart="Table 1" captionStartId="17.[192,257,798,821]" captionText="Table 1: Comparative measurements of adults of Pandinops friedrichi sp. n., and P. turieli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="3" pageNumber="2">Table 1</tableCitation>
)
</paragraph>
<paragraph id="8BA66419FFF0FFCFC391FB17FE0DFA83" blockId="3.[192,803,1263,1378]" pageId="3" pageNumber="2">
<taxonomicName id="4C191F9AFFF0FFCFC391FB17FD4FFAE7" authority="Birula, 1913: 419 - 422" authorityName="Birula" authorityPageNumber="419 - 422" authorityYear="1913" box="[192,712,1263,1286]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Pandinops">
<emphasis id="B96DB80BFFF0FFCFC391FB17FEA2FAE7" box="[192,293,1263,1286]" italics="true" pageId="3" pageNumber="2">Pandinus</emphasis>
(
<emphasis id="B96DB80BFFF0FFCFC26DFB17FE29FAE7" box="[316,430,1263,1286]" italics="true" pageId="3" pageNumber="2">Pandinops</emphasis>
) Birula, 1913: 419422
</taxonomicName>
, fig. b;
<bibRefCitation id="EF8819E8FFF0FFCFC3A1FAF6FE43FAC4" author="VACHON" box="[240,452,1294,1317]" pageId="3" pageNumber="2" pagination="857 - 958" refId="ref9842" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e serie, 140 (Zoologie, 104): 857 - 958." type="journal article" year="1974">Vachon, 1974: 921</bibRefCitation>
, 953, figs. 116118; Fet, 2000: 469; Kovařík, 2009: 5153, 115118, figs. 294 298, 304327.
</paragraph>
</subSubSection>
<subSubSection id="C3033792FFF0FFCFC391FA73FD12FA43" box="[192,661,1419,1442]" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8BA66419FFF0FFCFC391FA73FD12FA43" blockId="3.[192,661,1419,1442]" box="[192,661,1419,1442]" pageId="3" pageNumber="2">
<materialsCitation id="3B716E44FFF0FFCFC391FA73FD12FA43" box="[192,661,1419,1442]" collectionCode="SPECIES" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A2DABBFFF0FFCFC391FA73FF7FFA43" box="[192,248,1419,1442]" pageId="3" pageNumber="2">TYPE</typeStatus>
<collectionCode id="ED08FCDCFFF0FFCFC3AFFA77FED6FA43" box="[254,337,1423,1442]" pageId="3" pageNumber="2">SPECIES</collectionCode>
.
<taxonomicName id="4C191F9AFFF0FFCFC20FFA73FD12FA43" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[350,661,1419,1442]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="peeli">
<emphasis id="B96DB80BFFF0FFCFC20FFA73FE79FA43" box="[350,510,1419,1442]" italics="true" pageId="3" pageNumber="2">Pandinus peeli</emphasis>
Pocock, 1900
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C3033792FFF0FFCFC391FA34FC0DFCE7" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8BA66419FFF0FFCFC391FA34FB8FFEDB" blockId="3.[192,803,1484,1906]" lastBlockId="3.[830,1440,199,314]" pageId="3" pageNumber="2">
DIAGNOSIS. Total length
<quantity id="4CE1C9FCFFF0FFCFC2B2FA34FDE6FA02" box="[483,609,1484,1507]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.5" metricValueMin="5.5" pageId="3" pageNumber="2" unit="mm" value="75.0" valueMax="95.0" valueMin="55.0">5595 mm</quantity>
. External trichobothria on patella number 1316 (56
<emphasis id="B96DB80BFFF0FFCFC10EFA12FDFFF9E0" box="[607,632,1514,1537]" italics="true" pageId="3" pageNumber="2">eb</emphasis>
, 24
<emphasis id="B96DB80BFFF0FFCFC1E4FA12FD5EF9E0" box="[693,729,1514,1537]" italics="true" pageId="3" pageNumber="2">esb</emphasis>
, 2
<emphasis id="B96DB80BFFF0FFCFC1ADFA12FC9CF9E0" box="[764,795,1514,1537]" italics="true" pageId="3" pageNumber="2">em</emphasis>
, 12
<emphasis id="B96DB80BFFF0FFCFC3A5F9F1FE95F9C1" box="[244,274,1545,1568]" italics="true" pageId="3" pageNumber="2">est</emphasis>
, 3
<emphasis id="B96DB80BFFF0FFCFC26CF9F1FED7F9C1" box="[317,336,1545,1568]" italics="true" pageId="3" pageNumber="2">et</emphasis>
); ventral trichobothria on patella number 2235; internal trichobothria on chela number 68; ventral trichobothria on chela number 913. Pedipalp chela manus lobiform. Movable fingers of pedipalp, length of segments of pedipalps, and telson without noticeable sexual dimorphism. Pectines with fulcra. Pectinal teeth number 1121. Sternum subpentagonal, longer than wide. Carapace without distinct carinae. Dentate margin of pedipalp chela movable finger with distinct granules divided into 57 rows. Tergites IVI of mesosoma bear one carina. Stridulation organ located on pedipalp coxae and first pair of legs, but can be reduced. Metasomal segments IIV with paired parallel ventral median carinae or without carinae. Telson without subaculear tubercle. Legs with one pedal spur, retrolateral spur absent.
</paragraph>
<paragraph id="8BA66419FFF0FFCFC06FFE98FBD1FE77" blockId="3.[830,1441,352,774]" pageId="3" pageNumber="2">
SPINIFORM FORMULA OF TARSOMERES OF LEGS OF
<taxonomicName id="4C191F9AFFF0FFCFC06FFE87FC31FE77" authorityName="Birula" authorityYear="1913" box="[830,950,383,406]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B96DB80BFFF0FFCFC06FFE87FC31FE77" box="[830,950,383,406]" italics="true" pageId="3" pageNumber="2">PANDINOPS</emphasis>
</taxonomicName>
(
<figureCitation id="1322789CFFF0FFCFC092FE87FBCDFE77" box="[963,1098,383,406]" captionStart="Figures 4548" captionStartId="11.[192,278,1001,1024]" captionTargetBox="[201,1430,149,964]" captionTargetId="figure-6@11.[192,1437,144,971]" captionTargetPageId="11" captionText="Figures 4548: Pandinops turieli sp. n., female paratype, right legs IIV, retrolateral aspect. Abbreviations: vst, ventral subterminal; vt, ventral terminal; pd, prolateral distal; pst, prolateral subterminal; rm, retrolateral medial; rt, retrolateral terminal." pageId="3" pageNumber="2">FIGS. 4548</figureCitation>
).
</paragraph>
<paragraph id="8BA66419FFF0FFCFC06FFE66FBC5FDAF" blockId="3.[830,1441,352,774]" pageId="3" pageNumber="2">
<emphasis id="B96DB80BFFF0FFCFC06FFE66FC4DFE54" box="[830,970,414,437]" italics="true" pageId="3" pageNumber="2">Tarsomere I</emphasis>
. Spiniform macroseta
<emphasis id="B96DB80BFFF0FFCFC785FE66FB68FE54" box="[1236,1263,414,437]" italics="true" pageId="3" pageNumber="2">pd</emphasis>
,
<emphasis id="B96DB80BFFF0FFCFC655FE66FA90FE54" box="[1284,1303,414,437]" italics="true" pageId="3" pageNumber="2">vt</emphasis>
,
<emphasis id="B96DB80BFFF0FFCFC67DFE66FAB9FE54" box="[1324,1342,414,437]" italics="true" pageId="3" pageNumber="2">rt</emphasis>
,
<emphasis id="B96DB80BFFF0FFCFC602FE66FAF6FE54" box="[1363,1393,414,437]" italics="true" pageId="3" pageNumber="2">vst</emphasis>
are present on legs IIV, but
<emphasis id="B96DB80BFFF0FFCFC71FFE44FBEEFE32" box="[1102,1129,444,467]" italics="true" pageId="3" pageNumber="2">pd</emphasis>
can be replaced by seta in
<taxonomicName id="4C191F9AFFF0FFCFC6D8FE44FC2EFE13" baseAuthorityName="L. Koch" baseAuthorityYear="1875" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bellicosus">
<emphasis id="B96DB80BFFF0FFCFC6D8FE44FC2EFE13" italics="true" pageId="3" pageNumber="2">P. bellicosus</emphasis>
</taxonomicName>
as intraspecific variability;
<emphasis id="B96DB80BFFF0FFCFC7B4FE23FA83FE13" box="[1253,1284,475,498]" italics="true" pageId="3" pageNumber="2">pst</emphasis>
is present on legs IIIIV;
<emphasis id="B96DB80BFFF0FFCFC081FE02FC62FDF0" box="[976,997,506,529]" italics="true" pageId="3" pageNumber="2">pt</emphasis>
and
<emphasis id="B96DB80BFFF0FFCFC777FE02FBC2FDF0" box="[1062,1093,506,529]" italics="true" pageId="3" pageNumber="2">vm</emphasis>
are absent on all legs;
<emphasis id="B96DB80BFFF0FFCFC632FE02FA06FDF0" box="[1379,1409,506,529]" italics="true" pageId="3" pageNumber="2">rm</emphasis>
is present on legs IIV, but is often replaced by seta or spiniform seta (
<figureCitation id="1322789CFFF0FFCFC0B4FDCFFBB2FDAF" box="[997,1077,567,590]" captionStart="Figures 4548" captionStartId="11.[192,278,1001,1024]" captionTargetBox="[201,1430,149,964]" captionTargetId="figure-6@11.[192,1437,144,971]" captionTargetPageId="11" captionText="Figures 4548: Pandinops turieli sp. n., female paratype, right legs IIV, retrolateral aspect. Abbreviations: vst, ventral subterminal; vt, ventral terminal; pd, prolateral distal; pst, prolateral subterminal; rm, retrolateral medial; rt, retrolateral terminal." pageId="3" pageNumber="2">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="8BA66419FFF0FFCFC06FFDAEFC0DFCE7" blockId="3.[830,1441,352,774]" pageId="3" pageNumber="2">
<emphasis id="B96DB80BFFF0FFCFC06FFDAEFC4AFD8C" box="[830,973,598,621]" italics="true" pageId="3" pageNumber="2">Tarsomere II</emphasis>
. Spiniform formula is 3/4: 3/4: 3/4-5: 3/4- 5. Tarsomere II with 2 spines on inclined anteroventral surface, but
<taxonomicName id="4C191F9AFFF0FFCFC096FD6BFBAEFD4B" authority="Kovařík, 2016" authorityName="Kovařík" authorityYear="2016" box="[967,1065,659,682]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="turieli" status="sp. nov.">
<emphasis id="B96DB80BFFF0FFCFC096FD6BFBAEFD4B" box="[967,1065,659,682]" italics="true" pageId="3" pageNumber="2">P. turieli</emphasis>
</taxonomicName>
<emphasis id="B96DB80BFFF0FFCFC762FD6BFBF6FD4B" bold="true" box="[1075,1137,659,682]" pageId="3" pageNumber="2">
<taxonomicNameLabel id="A25E0570FFF0FFCFC762FD6BFBF6FD4B" box="[1075,1137,659,682]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a seta on leg III transformed to &quot;spiniform seta&quot; which indicates not well developed third spine (
<figureCitation id="1322789CFFF0FFCFC71BFD28FB27FD06" box="[1098,1184,720,743]" captionStart="Figures 4548" captionStartId="11.[192,278,1001,1024]" captionTargetBox="[201,1430,149,964]" captionTargetId="figure-6@11.[192,1437,144,971]" captionTargetPageId="11" captionText="Figures 4548: Pandinops turieli sp. n., female paratype, right legs IIV, retrolateral aspect. Abbreviations: vst, ventral subterminal; vt, ventral terminal; pd, prolateral distal; pst, prolateral subterminal; rm, retrolateral medial; rt, retrolateral terminal." pageId="3" pageNumber="2">Fig. 47</figureCitation>
) as intraspecific variability.
</paragraph>
</subSubSection>
<subSubSection id="C3033792FFF0FFCFC06FFCD4FBB6FC1A" pageId="3" pageNumber="2" type="discussion">
<paragraph id="8BA66419FFF0FFCFC06FFCD4FBB6FC1A" blockId="3.[830,1440,812,1019]" pageId="3" pageNumber="2">
COMMENTS. Since no data about macrosetae of tarsomeres and other characters (for example number of ventral trichobothria on pedipalp patella) have been published yet, and only the limited number of specimens of the rare
<taxonomicName id="4C191F9AFFF0FFCFC09CFC5FFBB8FC5F" authorityName="Birula" authorityYear="1913" box="[973,1087,935,958]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B96DB80BFFF0FFCFC09CFC5FFBB8FC5F" box="[973,1087,935,958]" italics="true" pageId="3" pageNumber="2">Pandinops</emphasis>
</taxonomicName>
genus were examined in this study, I cannot define intraspecific variability exactly for most of the characters.
</paragraph>
</subSubSection>
</treatment>
</document>