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<document id="91EFFA7CF33CDB192349865BC3FAB816" ID-DOI="10.5281/zenodo.184828" ID-GBIF-Dataset="69cc2112-400c-4aa5-928c-5114601dd906" ID-ISSN="1175-5326" ID-Zenodo-Dep="184828" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460025117311" checkinUser="plazi" docAuthor="Mccranie, James R., Vieites, David R. &amp; Wake, David B." docDate="2008" docId="03B0937CFFB5FFFBFF4FF9B4C72A4974" docLanguage="en" docName="zt01930p017.pdf" docOrigin="Zootaxa 1930" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Oedipina leptopoda Mccranie, Vieites &amp; Wake, 2008, sp. nov." docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="FF89EB04FFB9FFF5FFD8FFE1C2704818" masterDocTitle="Description of a new divergent lineage and three new species of Honduran salamanders of the genus Oedipina (Caudata, Plethodontidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="13" updateTime="1698233873515" updateUser="plazi">
<mods:mods id="72C9DAC372A6963AE71A202A3136AA25" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C5AAD0B2F2E94448D927930FE2D85A7E">Description of a new divergent lineage and three new species of Honduran salamanders of the genus Oedipina (Caudata, Plethodontidae)</mods:title>
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<mods:name id="01C427EB1A0B03F47BE731D3C0AA8D7D" type="personal">
<mods:role id="C2ED4D7AB6E1A13A890FA0EA388EF441">
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</mods:role>
<mods:namePart id="D215CAD5C7683AFBDD85719D20D7AB62">Mccranie, James R.</mods:namePart>
</mods:name>
<mods:name id="488FAA9F0699312D9B10AC8790C47C50" type="personal">
<mods:role id="E68836CB16967587EE30517B2B32B8A1">
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<mods:namePart id="FA65FFC5DDDB4CF1EC77E015E4FB9375">Vieites, David R.</mods:namePart>
</mods:name>
<mods:name id="FF70EFC3C6EC88F1ECCD992568CE8B87" type="personal">
<mods:role id="16208EE8071FF960125CCD907C8CC825">
<mods:roleTerm id="2810B355872645E2E0C3A71CA3CF1353">Author</mods:roleTerm>
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<mods:namePart id="C1F649B045BFB5DF4473CF7A9F87BAFC">Wake, David B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C3CD36009035B91C137EA304B06B5226">text</mods:typeOfResource>
<mods:relatedItem id="FE3673D1C4942B1953E1B61DC95AE9A7" type="host">
<mods:titleInfo id="8CECD17261D72E5CFAC6B42C12B40DB1">
<mods:title id="1F638A5C8DDF3EFB0D7CCB6676F57BB3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A9F2B516C3F8E61AAA154CD1F01FB245">
<mods:date id="2B1EF12812334A3582F4E43EF4B63335">2008</mods:date>
<mods:detail id="49DB01F426F0B255C5B0D021B2648DE7" type="volume">
<mods:number id="92ED0B7C7347F4A03B19CD78BB830E8F">1930</mods:number>
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<mods:start id="69D6190277CB398C362CFD3292232C5C">1</mods:start>
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</mods:extent>
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<mods:classification id="71CA6FFB4D2454EDF350B1C9E7BDC282">journal article</mods:classification>
<mods:identifier id="01D7DE9F789EE176FDF1DED6EC0BBFB8" type="DOI">10.5281/zenodo.184828</mods:identifier>
<mods:identifier id="9DE5EDB77C259A6FE3FEE7471C963964" type="GBIF-Dataset">69cc2112-400c-4aa5-928c-5114601dd906</mods:identifier>
<mods:identifier id="27007B8848F87D71E5DA2EF1A8F771D7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E685B654B5B5776AFA9FCC5AAE02BC3C" type="Zenodo-Dep">184828</mods:identifier>
</mods:mods>
<treatment id="03B0937CFFB5FFFBFF4FF9B4C72A4974" ID-DOI="http://doi.org/10.5281/zenodo.5624603" ID-GBIF-Taxon="119382042" ID-Zenodo-Dep="5624603" LSID="urn:lsid:plazi:treatment:03B0937CFFB5FFFBFF4FF9B4C72A4974" httpUri="http://treatment.plazi.org/id/03B0937CFFB5FFFBFF4FF9B4C72A4974" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="C30371E1FFB5FFF9FF4FF9B4C0564E8A" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BA6226AFFB5FFF9FF4FF9B4C3974E77" blockId="12.[151,550,1621,1682]" box="[151,487,1621,1647]" pageId="12" pageNumber="13">
<heading id="D0EE9506FFB5FFF9FF4FF9B4C3974E77" bold="true" box="[151,487,1621,1647]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B96DFE78FFB5FFF9FF4FF9B4C3974E77" bold="true" box="[151,487,1621,1647]" pageId="12" pageNumber="13">
<taxonomicName id="4C1959E9FFB5FFF9FF4FF9B4C3F24E77" ID-CoL="6SJLJ" box="[151,386,1621,1647]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="leptopoda" status="sp. nov.">
<emphasis id="B96DFE78FFB5FFF9FF4FF9B4C3F24E77" bold="true" box="[151,386,1621,1647]" italics="true" pageId="12" pageNumber="13">Oedipina leptopoda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4303FFB5FFF9FE51F9B4C3974E77" box="[393,487,1621,1647]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6226AFFB5FFF9FF4FF999C0564E8A" blockId="12.[151,550,1621,1682]" box="[151,550,1656,1682]" pageId="12" pageNumber="13">Narrow-footed Worm Salamander</paragraph>
</subSubSection>
<subSubSection id="C30371E1FFB5FFF9FF4FF929C0744EE7" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8BA6226AFFB5FFF9FF4FF929C0744EE7" blockId="12.[151,1436,1736,1791]" pageId="12" pageNumber="13">
<taxonomicName id="4C1959E9FFB5FFF9FF4FF929C30B4EC7" ID-CoL="48W9V" box="[151,379,1736,1759]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="cyclocauda">
<emphasis id="B96DFE78FFB5FFF9FF4FF929C30B4EC7" box="[151,379,1736,1759]" italics="true" pageId="12" pageNumber="13">Oedipina cyclocauda</emphasis>
</taxonomicName>
(part), Brame 1968,
<bibRefCitation id="EF885F9BFFB5FFF9FDB9F929C14E4EC7" author="Good" box="[609,830,1736,1759]" pageId="12" pageNumber="13" refString="Good, D. A. &amp; Wake, D. B. (1998; dated 1997) Phylogenetic and taxonomic implications of protein variation in the Mesoamerican salamander genus Oedipina (Caudata: Plethodontidae). Revista de Biologia Tropical, 48 (3), 1185 - 1208." type="journal article" year="1998">
Good &amp;
<collectingCountry id="F30E62FAFFB5FFF9FD19F929C08D4EC7" box="[705,765,1736,1759]" name="Wake Island" pageId="12" pageNumber="13">Wake</collectingCountry>
1998
</bibRefCitation>
,
<bibRefCitation id="EF885F9BFFB5FFF9FC92F929C6004EC7" author="Garcia-Paris" box="[842,1136,1736,1759]" pageId="12" pageNumber="13" refString="Garcia-Paris, M. &amp; Wake, D. B. (2000) Molecular phylogenetic analysis of relationships of the tropical salamander genera Oedipina and Nototriton, with descriptions of a new genus and three new species. Copeia, 2000 (1), 42 - 70." type="journal article" year="2000">
García-París &amp;
<collectingCountry id="F30E62FAFFB5FFF9FC2CF929C6404EC7" box="[1012,1072,1736,1759]" name="Wake Island" pageId="12" pageNumber="13">Wake</collectingCountry>
2000
</bibRefCitation>
,
<bibRefCitation id="EF885F9BFFB5FFF9FBA4F929C7E84EC7" author="McCranie" box="[1148,1432,1736,1759]" pageId="12" pageNumber="13" refString="McCranie, J. R. &amp; Wilson, L. D. (2002) The Amphibians of Honduras. Society for the Study of Amphibians and Reptiles, Ithaca, New York, 625 pp." type="book" year="2002">McCranie &amp; Wilson 2002</bibRefCitation>
,
<bibRefCitation id="EF885F9BFFB5FFF9FF1FF909C0704EE7" author="McCranie" box="[199,512,1768,1791]" pageId="12" pageNumber="13" refString="McCranie, J. R. &amp; Castaneda, F. E. (2007) Guia de Campo de los Anfibios de Honduras. Bibliomania!, Salt Lake City, 304 pp." type="book" year="2007">McCranie &amp; Castañeda 2007</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30371E1FFB5FFF9FF4FF8CEC6AE4FF2" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BA6226AFFB5FFF9FF4FF8CEC05E4FDA" blockId="12.[151,1436,1839,2026]" pageId="12" pageNumber="13">
<emphasis id="B96DFE78FFB5FFF9FF4FF8CEC3604F51" bold="true" box="[151,272,1839,1865]" pageId="12" pageNumber="13">
<typeStatus id="54A29CC8FFB5FFF9FF4FF8CEC37B4F51" box="[151,267,1839,1865]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED08BAAFFFB5FFF9FEC0F8D1C3294F52" LSID="urn:lsid:biocol.org:col:34777" box="[280,345,1840,1866]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="12" pageNumber="13">MVZ</collectionCode>
167772, a subadult female from
<quantity id="4CE18F8FFFB5FFF9FD07F8D1C15A4F52" box="[735,810,1840,1866]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="13" unit="km" value="32.0">32 km</quantity>
(road) W of Yoro on road to Morazán,
<geoCoordinate id="EE2D44ADFFB5FFF9FB23F8D1C7E74F52" box="[1275,1431,1840,1866]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="15.26748">15.267480 N</geoCoordinate>
,
<geoCoordinate id="EE2D44ADFFB5FFF9FF4FF8B9C3474F6A" box="[151,311,1880,1906]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="-87.43482">87.434820 W</geoCoordinate>
, Dept. Yoro,
<collectingCountry id="F30E62FAFFB5FFF9FE0AF8B9C0374F6A" box="[466,583,1880,1906]" name="Honduras" pageId="12" pageNumber="13">Honduras</collectingCountry>
, collected
<date id="FFA704AAFFB5FFF9FD1BF8B9C1074F6A" box="[707,887,1880,1906]" pageId="12" pageNumber="13" value="1979-01-08">8 January 1979</date>
by E. J. Koford and J. F. Lynch.
<emphasis id="B96DFE78FFB5FFF9FF1DF89EC34E4F81" bold="true" box="[197,318,1919,1945]" pageId="12" pageNumber="13">
<typeStatus id="54A29CC8FFB5FFF9FF1DF89EC3494F81" box="[197,313,1919,1945]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectionCode id="ED08BAAFFFB5FFF9FE92F861C3FB4F82" LSID="urn:lsid:biocol.org:col:34777" box="[330,395,1920,1946]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="12" pageNumber="13">MVZ</collectionCode>
<date id="FFA704AAFFB5FFF9FE4FF861C3824F82" box="[407,498,1920,1946]" pageId="12" pageNumber="13" value="78-10-17">171078</date>
, adult male from Montaña de Yoro,
<quantity id="4CE18F8FFFB5FFF9FC6FF861C67E4F82" box="[951,1038,1920,1946]" metricMagnitude="3" metricUnit="m" metricValue="6.6" pageId="12" pageNumber="13" unit="km" value="6.6">6.6 km</quantity>
(road) S of Yoro,
<geoCoordinate id="EE2D44ADFFB5FFF9FB2FF861C7E74F82" box="[1271,1431,1920,1946]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="15.07365">15.073650 N</geoCoordinate>
,
<geoCoordinate id="EE2D44ADFFB5FFF9FF4FF849C3694FDA" box="[151,281,1960,1986]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="-87.1333">87.1333 W</geoCoordinate>
, Dept. Yoro,
<collectingCountry id="F30E62FAFFB5FFF9FE6CF849C05A4FDA" box="[436,554,1960,1986]" name="Honduras" pageId="12" pageNumber="13">Honduras</collectingCountry>
.
</paragraph>
<paragraph id="8BA6226AFFB5FFF9FF1DF82EC6AE4FF2" blockId="12.[151,1436,1839,2026]" box="[197,1246,1999,2026]" pageId="12" pageNumber="13">
<emphasis id="B96DFE78FFB5FFF9FF1DF82EC3D64FF1" bold="true" box="[197,422,1999,2025]" pageId="12" pageNumber="13">Referred material</emphasis>
(1).
<collectionCode id="ED08BAAFFFB5FFF9FE06F831C0434FF2" LSID="urn:lsid:biocol.org:col:34795" box="[478,563,2000,2026]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="13">FMNH</collectionCode>
34683 (subadult), Portillo Grande, Dept. Yoro,
<collectingCountry id="F30E62FAFFB5FFF9FBBCF831C6AA4FF2" box="[1124,1242,2000,2026]" name="Honduras" pageId="12" pageNumber="13">Honduras</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30371E1FFB4FFF8FF1DFF76C3E04A6A" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BA6226AFFB4FFF8FF1DFF76C3E04A6A" blockId="13.[151,1437,151,2007]" pageId="13" pageNumber="14">
<emphasis id="B96DFE78FFB4FFF8FF1DFF76C33548A9" bold="true" box="[197,325,151,177]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
A small to moderate sized (
<typeStatus id="54A29CC8FFB4FFF8FD4BFF79C08B48AA" box="[659,763,152,178]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
is a subadult at
<quantity id="4CE18F8FFFB4FFF8FC66FF79C65848AA" box="[958,1064,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="13" pageNumber="14" unit="mm" value="29.6">29.6 mm</quantity>
SL) member of
<taxonomicName id="4C1959E9FFB4FFF8FB35FF79C72A48A9" box="[1261,1370,152,177]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B96DFE78FFB4FFF8FB35FF79C72A48A9" box="[1261,1370,152,177]" italics="true" pageId="13" pageNumber="14">Oedipina</emphasis>
</taxonomicName>
. Distinguished from
<taxonomicName id="4C1959E9FFB4FFF8FE84FF5CC02748CE" box="[348,599,189,214]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="cyclocauda">
<emphasis id="B96DFE78FFB4FFF8FE84FF5CC02748CE" box="[348,599,189,214]" italics="true" pageId="13" pageNumber="14">Oedipina cyclocauda</emphasis>
</taxonomicName>
of
<collectingCountry id="F30E62FAFFB4FFF8FD5AFF5CC17348CF" box="[642,771,189,215]" name="Costa Rica" pageId="13" pageNumber="14">Costa Rica</collectingCountry>
(the species with which this population was previously identified by
<bibRefCitation id="EF885F9BFFB4FFF8FEE1FF03C05D48E4" author="Good" box="[313,557,226,252]" pageId="13" pageNumber="14" refString="Good, D. A. &amp; Wake, D. B. (1998; dated 1997) Phylogenetic and taxonomic implications of protein variation in the Mesoamerican salamander genus Oedipina (Caudata: Plethodontidae). Revista de Biologia Tropical, 48 (3), 1185 - 1208." type="journal article" year="1998">
Good &amp;
<collectingCountry id="F30E62FAFFB4FFF8FE7DFF03C39748E4" box="[421,487,226,252]" name="Wake Island" pageId="13" pageNumber="14">Wake</collectingCountry>
1998
</bibRefCitation>
and
<bibRefCitation id="EF885F9BFFB4FFF8FDB3FF03C1C148E4" author="Garcia-Paris" box="[619,945,226,252]" pageId="13" pageNumber="14" refString="Garcia-Paris, M. &amp; Wake, D. B. (2000) Molecular phylogenetic analysis of relationships of the tropical salamander genera Oedipina and Nototriton, with descriptions of a new genus and three new species. Copeia, 2000 (1), 42 - 70." type="journal article" year="2000">
García-París &amp;
<collectingCountry id="F30E62FAFFB4FFF8FCF1FF03C11B48E4" box="[809,875,226,252]" name="Wake Island" pageId="13" pageNumber="14">Wake</collectingCountry>
2000
</bibRefCitation>
) by its inferred larger size and narrower hands and feet with less discrete digits. It differs from other described
<collectingCountry id="F30E62FAFFB4FFF8FC25FEE9C602493A" box="[1021,1138,264,290]" name="Honduras" pageId="13" pageNumber="14">Honduran</collectingCountry>
species of the subgenus
<taxonomicName id="4C1959E9FFB4FFF8FF4FFECCC374495E" box="[151,260,301,326]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B96DFE78FFB4FFF8FF4FFECCC374495E" box="[151,260,301,326]" italics="true" pageId="13" pageNumber="14">Oedipina</emphasis>
</taxonomicName>
(
<emphasis id="B96DFE78FFB4FFF8FECFFECCC078495E" box="[279,520,301,326]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C1959E9FFB4FFF8FECFFECCC3F1495E" box="[279,385,301,326]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ignea">O. ignea</taxonomicName>
,
<taxonomicName id="4C1959E9FFB4FFF8FE56FECCC073495E" box="[398,515,301,326]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="stuarti">O. stuarti</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C1959E9FFB4FFF8FD9CFECCC0C8495E" box="[580,696,301,326]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B96DFE78FFB4FFF8FD9CFECCC0C8495E" box="[580,696,301,326]" italics="true" pageId="13" pageNumber="14">O. taylori</emphasis>
</taxonomicName>
) in the following ways: from
<taxonomicName id="4C1959E9FFB4FFF8FBFBFECCC614495E" box="[1059,1124,301,326]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ignea">
<emphasis id="B96DFE78FFB4FFF8FBFBFECCC614495E" box="[1059,1124,301,326]" italics="true" pageId="13" pageNumber="14">ignea</emphasis>
</taxonomicName>
in having more maxillary teeth and narrower hands and feet; from
<taxonomicName id="4C1959E9FFB4FFF8FDA5FEB3C0B94973" box="[637,713,338,363]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="stuarti">
<emphasis id="B96DFE78FFB4FFF8FDA5FEB3C0B94973" box="[637,713,338,363]" italics="true" pageId="13" pageNumber="14">stuarti</emphasis>
</taxonomicName>
in being smaller, having much narrower hands and feet, and lacking glandular spots on its head and body; from
<taxonomicName id="4C1959E9FFB4FFF8FD2AFE99C1304989" box="[754,832,376,401]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B96DFE78FFB4FFF8FD2AFE99C1304989" box="[754,832,376,401]" italics="true" pageId="13" pageNumber="14">taylori</emphasis>
</taxonomicName>
in having a broader head, narrower hands and feet, shorter body and tail, and 19-20 costal grooves; from members of the subgenus
<emphasis id="B96DFE78FFB4FFF8FB9AFE7CC6CC49AE" box="[1090,1212,413,438]" italics="true" pageId="13" pageNumber="14">Oeditriton</emphasis>
(
<taxonomicName id="4C1959E9FFB4FFF8FB15FE7CC73649AE" box="[1229,1350,413,438]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="quadra">
<emphasis id="B96DFE78FFB4FFF8FB15FE7CC73649AE" box="[1229,1350,413,438]" italics="true" pageId="13" pageNumber="14">O. quadra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1959E9FFB4FFF8FAA7FE7CC29049C3" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="kasios">
<emphasis id="B96DFE78FFB4FFF8FAA7FE7CC29049C3" italics="true" pageId="13" pageNumber="14">O. kasios</emphasis>
</taxonomicName>
) in being larger (based on the size of the
<typeStatus id="54A29CC8FFB4FFF8FD10FE23C14149C4" box="[712,817,450,476]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
), having relatively smaller limbs and hands and feet, and a rounded body and tail; from all described
<collectingCountry id="F30E62FAFFB4FFF8FD09FE09C1364A1A" box="[721,838,488,514]" name="Honduras" pageId="13" pageNumber="14">Honduran</collectingCountry>
species of the subgenus
<emphasis id="B96DFE78FFB4FFF8FBB5FE09C6874A19" box="[1133,1271,488,513]" italics="true" pageId="13" pageNumber="14">Oedopinola</emphasis>
(
<emphasis id="B96DFE78FFB4FFF8FAD1FE09C36A4A3E" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C1959E9FFB4FFF8FAD1FE09C7E74A19" box="[1289,1431,488,513]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="elongata">O. elongata</taxonomicName>
,
<taxonomicName id="4C1959E9FFB4FFF8FF4FFDECC36A4A3E" box="[151,282,525,550]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="gephyra">O. gephyra</taxonomicName>
</emphasis>
and
<taxonomicName id="4C1959E9FFB4FFF8FE8AFDECC3B34A3E" box="[338,451,525,550]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="tomasi">
<emphasis id="B96DFE78FFB4FFF8FE8AFDECC3B34A3E" box="[338,451,525,550]" italics="true" pageId="13" pageNumber="14">O. tomasi</emphasis>
</taxonomicName>
) by having 19–20 costal grooves. In addition to these features, this species differs in allozymic (
<bibRefCitation id="EF885F9BFFB4FFF8FEC1FDD3C0754A54" author="Good" box="[281,517,562,588]" pageId="13" pageNumber="14" refString="Good, D. A. &amp; Wake, D. B. (1998; dated 1997) Phylogenetic and taxonomic implications of protein variation in the Mesoamerican salamander genus Oedipina (Caudata: Plethodontidae). Revista de Biologia Tropical, 48 (3), 1185 - 1208." type="journal article" year="1998">
Good &amp;
<collectingCountry id="F30E62FAFFB4FFF8FE59FDD3C3B34A54" box="[385,451,562,588]" name="Wake Island" pageId="13" pageNumber="14">Wake</collectingCountry>
1998
</bibRefCitation>
) and mtDNA characters (
<figureCitation id="13223EEFFFB4FFF8FCE9FDD3C1044A54" box="[817,884,562,588]" captionStart="FIGURE 1" captionStartId="4.[151,255,1802,1826]" captionTargetBox="[175,1422,674,1766]" captionTargetId="figure@4.[151,1436,635,1796]" captionTargetPageId="4" captionText="FIGURE 1. Maximum likelihood phylogram. Upper values represent ML bootstrap supports, while lower numbers represent Bayesian posterior probabilities (times 100). Values lower than 50 % were not included. Species described as new are highlighted in gray. Asterisks for species of Oedipina indicate specimens from Honduras." httpUri="https://zenodo.org/record/184829/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
) from all members of the genus in which those characters are known
</paragraph>
</subSubSection>
<subSubSection id="C30371E1FFB4FFF8FF1DFD9CC7534F94" pageId="13" pageNumber="14" type="description">
<paragraph id="8BA6226AFFB4FFF8FF1DFD9CC1084D0A" blockId="13.[151,1437,151,2007]" pageId="13" pageNumber="14">
<emphasis id="B96DFE78FFB4FFF8FF1DFD9CC32D4A8F" bold="true" box="[197,349,637,663]" pageId="13" pageNumber="14">Description.</emphasis>
This is a slender species of moderate size, judging from the fact that the
<typeStatus id="54A29CC8FFB4FFF8FB6FFD9CC76C4A8F" box="[1207,1308,637,663]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
is an adult male, the smaller sex in this genus, at a size larger than that of several other species in the subgenus. The head is relatively broad (SL 7.4-9.8 times head width), rounded rather than flattened, and with a narrowly rounded snout (expanded and broadly rounded in the
<typeStatus id="54A29CC8FFB4FFF8FD7AFD0CC17A4B1F" box="[674,778,749,775]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
, an adult male 47.9 SL). Nostrils are small and conspicuous. Nasolabial protuberances are moderate. Eyes are of moderate size and barely extend beyond the lateral margins of the head. The suborbital groove does not intercept the lip line. There are 5 small premaxillary teeth in the
<typeStatus id="54A29CC8FFB4FFF8FF3BFCBCC3394B6F" box="[227,329,861,887]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and a single slightly larger tooth in the adult male
<typeStatus id="54A29CC8FFB4FFF8FC69FCBCC66A4B6F" box="[945,1050,861,887]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
. The teeth are near the front but inside of the mouth, well separated from the maxillary teeth. There are 29 maxillary teeth in the
<typeStatus id="54A29CC8FFB4FFF8FADBFC63C7194B84" box="[1283,1385,898,924]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and
<quantity id="4CE18F8FFFB4FFF8FF4FFC49C2A24BDA" box="[151,210,936,962]" metricMagnitude="0" metricUnit="m" metricValue="1.016" pageId="13" pageNumber="14" unit="in" value="40.0">40 in</quantity>
the
<typeStatus id="54A29CC8FFB4FFF8FEDCFC49C31D4BDA" box="[260,365,936,962]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
. The
<typeStatus id="54A29CC8FFB4FFF8FE75FC49C0634BDA" box="[429,531,936,962]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
has 10 vomerine teeth and the
<typeStatus id="54A29CC8FFB4FFF8FC58FC49C1954BDA" box="[896,997,936,962]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
has 17. The vomerine teeth are small and in an arched series. The
<typeStatus id="54A29CC8FFB4FFF8FE33FC2CC0214BFF" box="[491,593,973,999]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and
<typeStatus id="54A29CC8FFB4FFF8FD54FC2CC0814BFF" box="[652,753,973,999]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
have 19 costal grooves between the slender limbs, with a limb interval of
<quantity id="4CE18F8FFFB4FFF8FE8DFC13C3E14C14" box="[341,401,1010,1036]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="13" pageNumber="14" unit="in" value="12.0">12 in</quantity>
both specimens. Hands and feet are extraordinarily narrow, small and flattened (
<figureCitation id="13223EEFFFB4FFF8FA9DFC13C7FC4C14" box="[1349,1420,1010,1036]" captionStart="FIGURE 3" captionStartId="9.[151,255,834,858]" captionTargetBox="[178,1429,193,799]" captionTargetId="figure@9.[151,1436,181,808]" captionTargetPageId="9" captionText="FIGURE 3. Schematic drawings of dorsal views of the limbs of Oedipina. From left to right, right hand and right foot of holotype of O. leptopoda (MVZ 167772), right foot of holotype of O. quadra (MVZ 257761), right foot of holotype of O. kasios (MVZ 232825). All drawings are at the same scale." httpUri="https://zenodo.org/record/184831/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
). In the
<typeStatus id="54A29CC8FFB4FFF8FF3CFBF9C33B4C2A" box="[228,331,1048,1074]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
digits can be discerned only because there are ill-defined grooves on the dorsal surfaces that suggest where digital borders lie. All of the digits are tightly fused, all the way to their tips in the
<typeStatus id="54A29CC8FFB4FFF8FAD1FBDCC7004C4F" box="[1289,1392,1085,1111]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
but there are small, rounded, independent tips of the three longest digits of both forelimbs and hind limbs of the
<typeStatus id="54A29CC8FFB4FFF8FF4FFB69C3704CBA" box="[151,256,1160,1186]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
. In the
<typeStatus id="54A29CC8FFB4FFF8FE84FB69C3B34CBA" box="[348,451,1160,1186]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
the longest digit is pointed, and it extends less then
<quantity id="4CE18F8FFFB4FFF8FB99FB69C6ED4CBA" box="[1089,1181,1160,1186]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="0.2">0.2 mm</quantity>
from the ends of the neighboring digits. The
<typeStatus id="54A29CC8FFB4FFF8FE6FFB4CC06E4CDF" box="[439,542,1197,1223]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
lacks a tail and has been almost completely eviscerated (for tissue sample used in allozyme and DNA studies). The
<typeStatus id="54A29CC8FFB4FFF8FDA0FB33C0AD4CF4" box="[632,733,1234,1260]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
has a rather stout, round tail that is broken
<quantity id="4CE18F8FFFB4FFF8FB0BFB33C74B4CF4" box="[1235,1339,1234,1260]" metricMagnitude="-2" metricUnit="m" metricValue="4.4399999999999995" pageId="13" pageNumber="14" unit="mm" value="44.4">44.4 mm</quantity>
from its base, and it doubtless greatly exceeded SL in the intact animal.
</paragraph>
<paragraph id="8BA6226AFFB4FFF8FF1DFAFCC0894E7A" blockId="13.[151,1437,151,2007]" pageId="13" pageNumber="14">
<emphasis id="B96DFE78FFB4FFF8FF1DFAFCC7784D2F" bold="true" box="[197,1288,1309,1335]" pageId="13" pageNumber="14">
Measurements (in millimeters), limb interval and tooth counts of the female
<typeStatus id="54A29CC8FFB4FFF8FB4DFAFCC7734D2F" box="[1173,1283,1309,1335]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
.
</emphasis>
Head width 4.0; snout to gular fold (head length) 5.4; head depth at posterior angle of jaw 2.1; eyelid width 0.5; eyelid length 1.3; eye to nostril 1.2; anterior rim of eye to snout 1.3; horizontal orbital diameter 1.0; interorbital distance 1.7; distance separating eyelids 1.1; nostril diameter 0.2; snout projecting beyond mandible 0.2; distance from eye to distal end of postorbital groove 1.5; snout to posterior angle of vent (SL) 29.6; snout to anterior angle of vent 28.3; snout to forelimb 7.5; axilla to groin 18.2; limb interval 12; shoulder width 2.7; tail length 33.2; tail width at base 1.5; tail depth at base 1.8; forelimb length (to tip of longest digit) 3.9; hind limb length 4.2; forelimb foot width 0.6; hind limb foot width 0.8; free length of longest digit on hind limb 0.2. Numbers of teeth: premaxillary 5; maxillary 29; vomerine 10.
</paragraph>
<paragraph id="8BA6226AFFB4FFF8FF1DF98CC0024F04" blockId="13.[151,1437,151,2007]" pageId="13" pageNumber="14">
<emphasis id="B96DFE78FFB4FFF8FF1DF98CC1334E9E" box="[197,835,1645,1670]" italics="true" pageId="13" pageNumber="14">
Coloration of the
<typeStatus id="54A29CC8FFB4FFF8FE7FF98CC07C4E9E" box="[423,524,1645,1670]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and
<typeStatus id="54A29CC8FFB4FFF8FD8AF98CC0CA4E9E" box="[594,698,1645,1670]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
in alcohol
</emphasis>
. The
<typeStatus id="54A29CC8FFB4FFF8FC4AF98CC18B4E9F" box="[914,1019,1645,1671]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, while apparently well preserved, appears to be faded and the overall coloration is now moderate brown. The head, especially the snout, is significantly paler than the rest of the specimen. The
<typeStatus id="54A29CC8FFB4FFF8FD3BF959C1384ECA" box="[739,840,1720,1746]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
is all black, darker dorsally and laterally than ventrally. The swollen snout and the nasolabial protuberances are much lighter, pale gray. The small and inconspicuous mental gland is pale gray.
</paragraph>
<paragraph id="8BA6226AFFB4FFF8FF1DF8C6C7534F94" blockId="13.[151,1437,151,2007]" pageId="13" pageNumber="14">
<emphasis id="B96DFE78FFB4FFF8FF1DF8C6C0744F59" bold="true" box="[197,516,1831,1857]" pageId="13" pageNumber="14">Habitat and distribution.</emphasis>
The species is known only from three specimens collected many years ago. Field notes are precise with respect to locality, two from west and one from south of Yoro (
<figureCitation id="13223EEFFFB4FFF8FB12F8ACC7624F7F" box="[1226,1298,1869,1895]" captionStart="FIGURE 4" captionStartId="10.[151,255,1003,1027]" captionTargetBox="[151,1435,197,977]" captionTargetId="figure@10.[151,1436,197,977]" captionTargetPageId="10" captionText="FIGURE 4. Map showing the collecting localities of Oedipina quadra, O. kasios, and O. leptopoda. The open circle represents a literature record (see McCranie 2006 b)." httpUri="https://zenodo.org/record/184832/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
), but vague with respect to habitat. The estimated elevation for these specimens is from ca.
<quantity id="4CE18F8FFFB4FFF8FBE3F893C6954F94" box="[1083,1253,1906,1932]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="13" pageNumber="14" unit="m" value="1000.0" valueMax="1300.0" valueMin="700.0">700 to 1300 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C30371E1FFB4FFFBFF1DF876C04C493A" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BA6226AFFB4FFFBFF1DF876C04C493A" blockId="13.[151,1437,151,2007]" lastBlockId="14.[151,1437,152,364]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B96DFE78FFB4FFF8FF1DF876C3234FA9" bold="true" box="[197,339,1943,1969]" pageId="13" pageNumber="14">Comments.</emphasis>
The specimen referred to this species is a poorly preserved subadult with sufficient morphological detail to make it a likely member of this species, and its collecting site is also appropriate for this assignment. The
<typeStatus id="54A29CC8FFB7FFFBFE86FF79C3B348AA" box="[350,451,152,178]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
is an intact specimen with good preservation of color, whereas the
<typeStatus id="54A29CC8FFB7FFFBFB02FF79C73048AA" box="[1242,1344,152,178]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
is eviscerated and faded but otherwise well preserved. However, because the two specimens are not syntopic and because we have molecular data only from the
<typeStatus id="54A29CC8FFB7FFFBFD61FF03C15348E4" box="[697,803,226,252]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
, we elected to use the specimen for which we have the best documentation as the
<typeStatus id="54A29CC8FFB7FFFBFE16FEE9C048493A" box="[462,568,264,290]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30371E1FFB7FFFBFF1DFECCC72A4974" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BA6226AFFB7FFFBFF1DFECCC72A4974" blockId="14.[151,1437,152,364]" pageId="14" pageNumber="15">
<emphasis id="B96DFE78FFB7FFFBFF1DFECCC322495F" bold="true" box="[197,338,301,327]" pageId="14" pageNumber="15">Etymology.</emphasis>
The specific name
<taxonomicName id="4C1959E9FFB7FFFBFDE0FECCC0DD495E" box="[568,685,301,326]" class="Amphibia" family="Plethodontidae" genus="Oedipina" kingdom="Animalia" order="Caudata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leptopoda">
<emphasis id="B96DFE78FFB7FFFBFDE0FECCC0DD495E" box="[568,685,301,326]" italics="true" pageId="14" pageNumber="15">leptopoda</emphasis>
</taxonomicName>
is derived from the Greek
<emphasis id="B96DFE78FFB7FFFBFC32FECCC65E495E" box="[1002,1070,301,326]" italics="true" pageId="14" pageNumber="15">lepto-</emphasis>
for small or fine and
<emphasis id="B96DFE78FFB7FFFBFAF4FECCC702495E" box="[1324,1394,301,326]" italics="true" pageId="14" pageNumber="15">podos</emphasis>
for foot, and is used as an adjective in reference to the very small and narrow hands and feet of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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