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<document id="A2CDECAE3A872D0C06D6864806F0D9D3" ID-DOI="10.5281/zenodo.197589" ID-GBIF-Dataset="53fca97f-1dda-43cc-b95e-2eca97fde26c" ID-ISSN="1175-5326" ID-Zenodo-Dep="197589" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460273136847" checkinUser="plazi" docAuthor="Lavoué, Sébastien, Sullivan, John P. &amp; Arnegard, Matthew E." docDate="2010" docId="0394455B701BB453B8BFA228C3CCD06B" docLanguage="en" docName="zt02600p052.pdf" docOrigin="Zootaxa 2600" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Petrocephalus odzalaensis Lavoué, Sullivan &amp; Arnegard, 2010, n. sp." docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="FFAD3D23700EB44BB828A742C128D67C" masterDocTitle="African weakly electric fishes of the genus Petrocephalus (Osteoglossomorpha: Mormyridae) of Odzala National Park, Republic of the Congo (Lékoli River, Congo River basin) with description of five new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="22" updateTime="1698247377879" updateUser="plazi">
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<mods:title id="4C310481566D632C9F3DDCAB290C7ABB">African weakly electric fishes of the genus Petrocephalus (Osteoglossomorpha: Mormyridae) of Odzala National Park, Republic of the Congo (Lékoli River, Congo River basin) with description of five new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="C701D8043D3F248F041C2A21FD9D27B2">Lavoué, Sébastien</mods:namePart>
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<mods:namePart id="A457DBFB8C2AC0825332825E5388685C">Sullivan, John P.</mods:namePart>
</mods:name>
<mods:name id="5FB2FF0DC3C4E2417D1EC41E0F235A5A" type="personal">
<mods:role id="3B637997C5FE2EA2D74A8E91419158D0">
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<mods:namePart id="83C49814637578DCF8B17914F93B4507">Arnegard, Matthew E.</mods:namePart>
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<mods:title id="FD4F7DF112407D109E4FDA0BC2ED2AFF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2D7C99FF9C5C4737268B84B7C3412DC0">
<mods:date id="D602F51679B54948D7298B451B5D97E3">2010</mods:date>
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<mods:number id="EC1708B6B3B191154588EFD6906ADF91">2600</mods:number>
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<mods:identifier id="ACE53FB5D7384F7105D62B9458B371BF" type="DOI">10.5281/zenodo.197589</mods:identifier>
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<mods:identifier id="77D0BD917125F749F5CECD4A4D8D52C8" type="Zenodo-Dep">197589</mods:identifier>
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<treatment id="0394455B701BB453B8BFA228C3CCD06B" ID-DOI="http://doi.org/10.5281/zenodo.6202202" ID-GBIF-Taxon="119395498" ID-Zenodo-Dep="6202202" LSID="urn:lsid:zoobank.org:act:9D9893FC-28FA-4E99-82E7-3C6AB8E6926E" httpUri="http://treatment.plazi.org/id/0394455B701BB453B8BFA228C3CCD06B" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="C327A7C6701BB45EB8BFA228C1DAD04B" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8B82F44D701BB45EB8BFA228C336D3F8" blockId="21.[151,542,1386,1412]" box="[151,542,1386,1412]" pageId="21" pageNumber="22">
<heading id="D0CA4321701BB45EB8BFA228C336D3F8" bold="true" box="[151,542,1386,1412]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="B949285F701BB45EB8BFA228C336D3F8" bold="true" box="[151,542,1386,1412]" pageId="21" pageNumber="22">
<taxonomicName id="4C3D8FCE701BB45EB8BFA228C0FED3F8" ID-CoL="4FDXQ" box="[151,470,1386,1412]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="odzalaensis" status="sp. nov.">
<emphasis id="B949285F701BB45EB8BFA228C0FED3F8" bold="true" box="[151,470,1386,1412]" italics="true" pageId="21" pageNumber="22">Petrocephalus odzalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A9524701BB45EB9F4A228C336D3F8" box="[476,542,1386,1412]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B82F44D701BB45EB8BFA2F0C509D3B5" blockId="21.[151,1057,1458,1481]" box="[151,1057,1458,1481]" pageId="21" pageNumber="22">
[Odzala field identification and in
<bibRefCitation id="EFAC89BC701BB45EBA2FA2F0C3CCD3B5" box="[519,740,1458,1481]" pageId="21" pageNumber="22" refString="Lavoue, S., Arnegard, M. E., Sullivan, J. P. &amp; Hopkins, C. D. (2008) Petrocephalus of Odzala offer insights into evolutionary patterns of signal diversification in the Mormyridae, a family of weakly electrogenic fishes from Africa. Journal of Physiology - Paris, 102, 322 - 339." type="journal article">
Lavoué
<emphasis id="B949285F701BB45EBA75A2F0C3BBD3B5" box="[605,659,1458,1481]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2008)
</bibRefCitation>
:
<taxonomicName id="4C3D8FCE701BB45EBAC6A2F0C2AFD3B5" box="[750,903,1458,1481]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B949285F701BB45EBAC6A2F0C2AFD3B5" box="[750,903,1458,1481]" italics="true" pageId="21" pageNumber="22">Petrocephalus</emphasis>
</taxonomicName>
sp. 6, OTU 6]
</paragraph>
<paragraph id="8B82F44D701BB45EB8BFA2B5C1DAD04B" blockId="21.[151,1437,1527,2002]" pageId="21" pageNumber="22">
<emphasis id="B949285F701BB45EB8BFA2B5C1D2D06D" bold="true" box="[151,250,1527,1553]" pageId="21" pageNumber="22">Images.</emphasis>
<figureCitation id="1306E8C8701BB45EB92AA2BAC060D06E" box="[258,328,1528,1554]" captionStart="FIGURE 8" captionStartId="22.[151,258,1771,1795]" captionTargetBox="[188,1398,269,1750]" captionTargetId="figure@22.[188,1398,269,1751]" captionTargetPageId="22" captionText="FIGURE 8. Petrocephalus odzalaensis n. sp. of the Lékoli River system of Odzala National Park, Republic of the Congo. A. Photograph of a live specimen (no scale). B. Photograph of the preserved holotype (CU 88048; scale bar = 1.0 cm). C. EOD waveforms. One EOD was arbitrarily selected from each recorded individual and superimposed with the other recordings (N = number of individuals). All EODs are scaled to the same peak-to-peak voltage, and they are plotted (head-positive up) on the same time scale as overlays centered on the largest positive peak of each waveform (scale bar = 0.2 msec)." httpUri="https://zenodo.org/record/197597/files/figure.png" pageId="21" pageNumber="22">Fig. 8</figureCitation>
A, photo of a live specimen from Odzala and
<figureCitation id="1306E8C8701BB45EBB5CA2BAC293D06E" box="[884,955,1528,1554]" captionStart="FIGURE 8" captionStartId="22.[151,258,1771,1795]" captionTargetBox="[188,1398,269,1750]" captionTargetId="figure@22.[188,1398,269,1751]" captionTargetPageId="22" captionText="FIGURE 8. Petrocephalus odzalaensis n. sp. of the Lékoli River system of Odzala National Park, Republic of the Congo. A. Photograph of a live specimen (no scale). B. Photograph of the preserved holotype (CU 88048; scale bar = 1.0 cm). C. EOD waveforms. One EOD was arbitrarily selected from each recorded individual and superimposed with the other recordings (N = number of individuals). All EODs are scaled to the same peak-to-peak voltage, and they are plotted (head-positive up) on the same time scale as overlays centered on the largest positive peak of each waveform (scale bar = 0.2 msec)." httpUri="https://zenodo.org/record/197597/files/figure.png" pageId="21" pageNumber="22">Fig. 8</figureCitation>
B, photo of the preserved
<typeStatus id="54864AEF701BB45EBCDFA2BAC448D06E" box="[1271,1376,1528,1554]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
(CU 88048).
</paragraph>
</subSubSection>
<subSubSection id="C327A7C6701BB45CB8EEA100C392D176" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8B82F44D701BB45EB8EEA100C3A3D0DB" blockId="21.[151,1437,1527,2002]" pageId="21" pageNumber="22">
<emphasis id="B949285F701BB45EB8EEA100C056D020" bold="true" box="[198,382,1602,1628]" pageId="21" pageNumber="22">
<typeStatus id="54864AEF701BB45EB8EEA100C029D020" box="[198,257,1602,1628]" pageId="21" pageNumber="22">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54864AEF701BB45EB9AFA100C0DFD020" box="[391,503,1602,1628]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
, CU 88048 (morpho,
<collectionCode id="ED2C6C88701BB45EBAD7A100C217D020" box="[767,831,1602,1628]" pageId="21" pageNumber="22">EOD</collectionCode>
), sex undet.,
<quantity id="4CC559A8701BB45EBBF6A100C561D020" box="[990,1097,1602,1628]" metricMagnitude="-2" metricUnit="m" metricValue="9.29" pageId="21" pageNumber="22" unit="mm" value="92.9">92.9 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBC7AA100C550D020" box="[1106,1144,1602,1628]" pageId="21" pageNumber="22">SL</collectionCode>
.
<collectingCountry id="F32AB4DD701BB45EBCACA100C4BCD020" box="[1156,1428,1602,1628]" name="Republic of the Congo" pageId="21" pageNumber="22">Republic of the Congo</collectingCountry>
, Lékénie River at Mboko débarcadère (
<collectingCountry id="F32AB4DD701BB45EBA75A12AC385D0FE" box="[605,685,1640,1666]" name="Republic of the Congo" pageId="21" pageNumber="22">Congo</collectingCountry>
basin), Odzala National Park, (
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0.62
<emphasis id="B949285F701BB45EBC7EA12AC54AD0FD" box="[1110,1122,1640,1665]" italics="true" pageId="21" pageNumber="22">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE09928A701BB45EBCA6A12AC5D1D0FE" box="[1166,1273,1640,1666]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="555" value="14.9">
14.90
<emphasis id="B949285F701BB45EBCF8A12AC5F4D0FD" box="[1232,1244,1640,1665]" italics="true" pageId="21" pageNumber="22">°</emphasis>
E
</geoCoordinate>
), J.P. Friel, S. Lavoué &amp; J.P. Sullivan coll.,
<date id="FF83D28D701BB45EB9C5A1CFC3AFD0DB" box="[493,647,1677,1703]" pageId="21" pageNumber="22" value="2002-08">August 2002</date>
.
</paragraph>
<paragraph id="8B82F44D701BB45DB8EEA1F0C0B4D6AB" blockId="21.[151,1437,1527,2002]" lastBlockId="22.[151,1436,152,215]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<typeStatus id="54864AEF701BB45EB8EEA1F0C014D0B0" box="[198,316,1714,1740]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
. CU 87850 (morpho,
<collectionCode id="ED2C6C88701BB45EBA11A1F0C351D0B0" box="[569,633,1714,1740]" pageId="21" pageNumber="22">EOD</collectionCode>
), male,
<quantity id="4CC559A8701BB45EBAFDA1F0C215D0B0" box="[725,829,1714,1740]" metricMagnitude="-2" metricUnit="m" metricValue="9.01" pageId="21" pageNumber="22" unit="mm" value="90.1">90.1 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBB6BA1F0C241D0B0" box="[835,873,1714,1740]" pageId="21" pageNumber="22">SL</collectionCode>
; CU 87857 (morpho,
<collectionCode id="ED2C6C88701BB45EBC4FA1F0C580D0B0" box="[1127,1192,1714,1740]" pageId="21" pageNumber="22">EOD</collectionCode>
), male,
<quantity id="4CC559A8701BB45EBD2BA1F0C443D0B0" box="[1283,1387,1714,1740]" metricMagnitude="-2" metricUnit="m" metricValue="9.26" pageId="21" pageNumber="22" unit="mm" value="92.6">92.6 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBD5AA1F0C4B0D0B0" box="[1394,1432,1714,1740]" pageId="21" pageNumber="22">SL</collectionCode>
; CU 88050 (morpho,
<collectionCode id="ED2C6C88701BB45EB9B8A19AC0F9D08E" box="[400,465,1752,1778]" pageId="21" pageNumber="22">EOD</collectionCode>
), male,
<quantity id="4CC559A8701BB45EBA07A19AC3B1D08E" box="[559,665,1752,1778]" metricMagnitude="-2" metricUnit="m" metricValue="9.04" pageId="21" pageNumber="22" unit="mm" value="90.4">90.4 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBA89A19AC3EFD08E" box="[673,711,1752,1778]" pageId="21" pageNumber="22">SL</collectionCode>
;
<collectionCode id="ED2C6C88701BB45EBAFCA19AC206D08E" LSID="urn:lsid:biocol.org:col:34925" box="[724,814,1752,1778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH</collectionCode>
251414 (ex CU 87843) (morpho,
<collectionCode id="ED2C6C88701BB45EBCE0A19AC421D08E" box="[1224,1289,1752,1778]" pageId="21" pageNumber="22">EOD</collectionCode>
), male, 87.0 mm
<collectionCode id="ED2C6C88701BB45EB8E4A1BFC1D9D16B" box="[204,241,1789,1815]" pageId="21" pageNumber="22">SL</collectionCode>
;
<collectionCode id="ED2C6C88701BB45EB8D6A1BFC070D16B" LSID="urn:lsid:biocol.org:col:34925" box="[254,344,1789,1815]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH</collectionCode>
251417 (ex CU 88056) (morpho,
<collectionCode id="ED2C6C88701BB45EBAC3A1BFC203D16B" box="[747,811,1789,1815]" pageId="21" pageNumber="22">EOD</collectionCode>
), male,
<quantity id="4CC559A8701BB45EBBA0A1BFC2D9D16B" box="[904,1009,1789,1815]" metricMagnitude="-2" metricUnit="m" metricValue="8.76" pageId="21" pageNumber="22" unit="mm" value="87.6">87.6 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBBD0A1BFC536D16B" box="[1016,1054,1789,1815]" pageId="21" pageNumber="22">SL</collectionCode>
;
<collectionCode id="ED2C6C88701BB45EBC02A1BFC5ACD16B" LSID="urn:lsid:biocol.org:col:34925" box="[1066,1156,1789,1815]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH</collectionCode>
251416 (ex CU 88054) (morpho,
<collectionCode id="ED2C6C88701BB45EB923A060C063D140" box="[267,331,1826,1852]" pageId="21" pageNumber="22">EOD</collectionCode>
), male,
<quantity id="4CC559A8701BB45EB981A060C33AD140" box="[425,530,1826,1852]" metricMagnitude="-2" metricUnit="m" metricValue="9.76" pageId="21" pageNumber="22" unit="mm" value="97.6">97.6 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBA32A060C368D140" box="[538,576,1826,1852]" pageId="21" pageNumber="22">SL</collectionCode>
; CU 88059 (morpho,
<collectionCode id="ED2C6C88701BB45EBB6BA060C2ACD140" box="[835,900,1826,1852]" pageId="21" pageNumber="22">EOD</collectionCode>
), male,
<quantity id="4CC559A8701BB45EBBC9A060C562D140" box="[993,1098,1826,1852]" metricMagnitude="-2" metricUnit="m" metricValue="9.93" pageId="21" pageNumber="22" unit="mm" value="99.3">99.3 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBC7AA060C550D140" box="[1106,1144,1826,1852]" pageId="21" pageNumber="22">SL</collectionCode>
;
<collectionCode id="ED2C6C88701BB45EBCACA060C5F6D140" LSID="urn:lsid:biocol.org:col:34925" box="[1156,1246,1826,1852]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH</collectionCode>
251415 (ex CU 87844) (morpho,
<collectionCode id="ED2C6C88701BB45EB944A00AC086D11E" box="[364,430,1864,1890]" pageId="21" pageNumber="22">EOD</collectionCode>
), male,
<quantity id="4CC559A8701BB45EBA26A00AC351D11E" box="[526,633,1864,1890]" metricMagnitude="-2" metricUnit="m" metricValue="8.73" pageId="21" pageNumber="22" unit="mm" value="87.3">87.3 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBAAAA00AC381D11E" box="[642,681,1864,1890]" pageId="21" pageNumber="22">SL</collectionCode>
.
<collectingCountry id="F32AB4DD701BB45EBA9DA00AC2E1D11E" box="[693,969,1864,1890]" name="Republic of the Congo" pageId="21" pageNumber="22">Republic of the Congo</collectingCountry>
, Lékénie River at Mboko débarcadère (
<collectingCountry id="F32AB4DD701BB45EB8B7A02FC1C7D1FB" box="[159,239,1901,1927]" name="Republic of the Congo" pageId="21" pageNumber="22">Congo</collectingCountry>
basin), Odzala National Park, (
<geoCoordinate id="EE09928A701BB45EBA4DA02FC3EED1FB" box="[613,710,1901,1927]" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="555" value="0.62">
0.62
<emphasis id="B949285F701BB45EBAB2A02FC38ED1FA" box="[666,678,1901,1926]" italics="true" pageId="21" pageNumber="22">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE09928A701BB45EBAFBA02FC216D1FB" box="[723,830,1901,1927]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="555" value="14.9">
14.90
<emphasis id="B949285F701BB45EBB3DA02FC209D1FA" box="[789,801,1901,1926]" italics="true" pageId="21" pageNumber="22">°</emphasis>
E
</geoCoordinate>
), J.P. Friel, S. Lavoué &amp; J.P. Sullivan coll.,
<date id="FF83D28D701BB45EBD6FA02FC1FED1D0" pageId="21" pageNumber="22" value="2002-08">August 2002</date>
. CU 88049 (morpho,
<collectionCode id="ED2C6C88701BB45EB9F1A0D0C331D1D0" box="[473,537,1938,1964]" pageId="21" pageNumber="22">EOD</collectionCode>
; caudal peduncle dissected by
<bibRefCitation id="EFAC89BC701BB45EBBAEA0D0C550D1D0" box="[902,1144,1938,1964]" pageId="21" pageNumber="22" refString="Lavoue, S., Arnegard, M. E., Sullivan, J. P. &amp; Hopkins, C. D. (2008) Petrocephalus of Odzala offer insights into evolutionary patterns of signal diversification in the Mormyridae, a family of weakly electrogenic fishes from Africa. Journal of Physiology - Paris, 102, 322 - 339." type="journal article">
Lavoué
<emphasis id="B949285F701BB45EBBCEA0D1C50AD1D0" box="[998,1058,1939,1964]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2008)
</bibRefCitation>
to sample electric organ for histological examination), male,
<quantity id="4CC559A8701BB45EBA5CA0FAC3CED1AE" box="[628,742,1976,2002]" metricMagnitude="-2" metricUnit="m" metricValue="9.32" pageId="21" pageNumber="22" unit="mm" value="93.2">93.2 mm</quantity>
<collectionCode id="ED2C6C88701BB45EBAC7A0FAC230D1AE" box="[751,792,1976,2002]" pageId="21" pageNumber="22">SL</collectionCode>
.
<collectingCountry id="F32AB4DD701BB45EBB0EA0FAC565D1AE" box="[806,1101,1976,2002]" name="Republic of the Congo" pageId="21" pageNumber="22">Republic of the Congo</collectingCountry>
, Lékénie River at Mboko débarcadère (
<collectingCountry id="F32AB4DD7018B45DB91EA7DAC0AFD6CE" box="[310,391,152,178]" name="Republic of the Congo" pageId="22" pageNumber="23">Congo</collectingCountry>
basin), Odzala National Park, (
<geoCoordinate id="EE09928A7018B45DBB28A7DAC24AD6CE" box="[768,866,152,178]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="0.62">
0.62
<emphasis id="B949285F7018B45DBB1EA7DAC26AD6CD" box="[822,834,152,177]" italics="true" pageId="22" pageNumber="23">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE09928A7018B45DBB46A7DAC2F2D6CE" box="[878,986,152,178]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="14.9">
14.90
<emphasis id="B949285F7018B45DBB99A7DAC295D6CD" box="[945,957,152,177]" italics="true" pageId="22" pageNumber="23">°</emphasis>
E
</geoCoordinate>
), J.P. Friel, S. Lavoué &amp; J.P. Sullivan coll.,
<date id="FF83D28D7018B45DB8F1A7FFC0BFD6AB" box="[217,407,189,215]" pageId="22" pageNumber="23" value="2002-08-12">12 August 2002</date>
.
</paragraph>
<caption id="DF42A4C57018B45DB8BFA1A9C02ED1DF" httpUri="https://zenodo.org/record/197597/files/figure.png" pageId="22" pageNumber="23" targetBox="[188,1398,269,1750]" targetPageId="22">
<paragraph id="8B82F44D7018B45DB8BFA1A9C02ED1DF" blockId="22.[151,1436,1771,1955]" pageId="22" pageNumber="23">
<emphasis id="B949285F7018B45DB8BFA1A9C036D17F" bold="true" box="[151,286,1771,1795]" pageId="22" pageNumber="23">FIGURE 8.</emphasis>
<taxonomicName id="4C3D8FCE7018B45DB90EA1AEC366D17F" box="[294,590,1772,1795]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="odzalaensis" status="sp. nov.">
<emphasis id="B949285F7018B45DB90EA1AEC366D17F" box="[294,590,1772,1795]" italics="true" pageId="22" pageNumber="23">Petrocephalus odzalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A95247018B45DBA7EA1A9C3BED17F" box="[598,662,1771,1795]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B949285F7018B45DBA7EA1A9C3BED17F" bold="true" box="[598,662,1771,1795]" pageId="22" pageNumber="23">n. sp.</emphasis>
</taxonomicNameLabel>
of the Lékoli River system of Odzala National Park, Republic of the Congo. A. Photograph of a live specimen (no scale). B. Photograph of the preserved holotype (CU 88048; scale bar = 1.0 cm). C. EOD waveforms. One EOD was arbitrarily selected from each recorded individual and superimposed with the other recordings (
<emphasis id="B949285F7018B45DB97BA00EC04DD11F" box="[339,357,1868,1891]" italics="true" pageId="22" pageNumber="23">N</emphasis>
= number of individuals). All EODs are scaled to the same peak-to-peak voltage, and they are plotted (head-positive up) on the same time scale as overlays centered on the largest positive peak of each waveform (scale bar = 0.2 msec).
</paragraph>
</caption>
<caption id="DF42A4C57019B45CB8BFA7D5C5ACD773" ID-Table-UUID="DF42A4C57019B45CB8BFA7D5C5ACD773" httpUri="http://table.plazi.org/id/DF42A4C57019B45CB8BFA7D5C5ACD773" pageId="23" pageNumber="24" targetBox="[151,1436,378,1279]" targetIsTable="true" targetPageId="23">
<paragraph id="8B82F44D7019B45CB8BFA7D5C5ACD773" blockId="23.[151,1436,151,359]" pageId="23" pageNumber="24">
<emphasis id="B949285F7019B45CB8BFA7D5C026D6D3" bold="true" box="[151,270,151,175]" pageId="23" pageNumber="24">TABLE 6.</emphasis>
Principal morphometric ratios and meristic counts for the holotype (CU 88048) and nine paratypes (CU 87857, CU 88049, CU 87850, CU 88050, CU 88059, AMNH 250928, AMNH 250929, AMNH 250930, AMNH 250931) of
<taxonomicName id="4C3D8FCE7019B45CB89CA79AC0E7D693" box="[180,463,216,239]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="odzalaensis" status="sp. nov.">
<emphasis id="B949285F7019B45CB89CA79AC0E7D693" box="[180,463,216,239]" italics="true" pageId="23" pageNumber="24">Petrocephalus odzalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A95247019B45CB9FEA795C33AD693" box="[470,530,215,239]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B949285F7019B45CB9FEA795C33AD693" bold="true" box="[470,530,215,239]" pageId="23" pageNumber="24">n. sp.</emphasis>
</taxonomicNameLabel>
(Abbreviations: o= sex category &quot;other&quot; [i.e., not an obvious mature male, but rather a female or immature male]; StdDev= standard deviation; MinMax= minimummaximum).
</paragraph>
</caption>
<paragraph id="8B82F44D7019B45CBA8CA668C518D743" blockId="23.[151,1436,151,359]" box="[676,1072,298,319]" pageId="23" pageNumber="24">
<typeStatus id="54864AEF7019B45CBA8CA668C3D5D743" box="[676,765,298,319]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
(o)
<typeStatus id="54864AEF7019B45CBBB0A668C2DDD743" box="[920,1013,298,319]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
(n=9)
</paragraph>
<paragraph id="8B82F44D7019B45CBBB0A610C5E7D333" blockId="23.[151,1436,151,359]" lastBlockId="23.[159,1231,1338,1668]" pageId="23" pageNumber="24">MinMax Mean StdDev MinMax Median</paragraph>
<paragraph id="8B82F44D7019B45CB8B7A638C406D283" pageId="23" pageNumber="24">
<table id="F93D06ED70194BB4B8BFA638C4B4D283" box="[151,1436,378,1279]" gridcols="4" gridrows="20" pageId="23" pageNumber="24">
<tr id="350DF60F70194BB4B8BFA638C4B4D7F3" box="[151,1436,378,399]" gridrow="0" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA638C331D7F3" box="[151,537,378,399]" gridcol="0" gridrow="0" pageId="23" pageNumber="24">Standard length (mm)</th>
<th id="76DC9F7370194BB4BA8CA638C3E6D7F3" box="[676,718,378,399]" gridcol="1" gridrow="0" pageId="23" pageNumber="24">92.9</th>
<th id="76DC9F7370194BB4BBB0A638C2D0D7F3" box="[920,1016,378,399]" gridcol="2" gridrow="0" pageId="23" pageNumber="24">87.099.3</th>
<th id="76DC9F7370194BB4BCADA638C4B4D7F3" box="[1157,1436,378,399]" gridcol="3" gridrow="0" pageId="23" pageNumber="24">91.7 4.5</th>
</tr>
<tr id="350DF60F70194BB4B8BFA6E0C4B4D7CB" box="[151,1436,418,439]" gridrow="1" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA6E0C331D7CB" box="[151,537,418,439]" gridcol="0" gridrow="1" pageId="23" pageNumber="24">Head length (mm)</th>
<td id="76DC9F7370194BB4BA8CA6E0C3E6D7CB" box="[676,718,418,439]" gridcol="1" gridrow="1" pageId="23" pageNumber="24">23.3</td>
<td id="76DC9F7370194BB4BBB0A6E0C2D0D7CB" box="[920,1016,418,439]" gridcol="2" gridrow="1" pageId="23" pageNumber="24">22.726.6</td>
<td id="76DC9F7370194BB4BCADA6E0C4B4D7CB" box="[1157,1436,418,439]" gridcol="3" gridrow="1" pageId="23" pageNumber="24">23.7 1.6</td>
</tr>
<tr id="350DF60F70194BB4B8BFA6B0C4B4D47B" box="[151,1436,498,519]" gridrow="2" pageId="23" pageNumber="24" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76DC9F7370194BB4B8BFA6B0C331D47B" box="[151,537,498,519]" gridcol="0" gridrow="2" pageId="23" pageNumber="24">Ratio of standard length (SL):</th>
</tr>
<tr id="350DF60F70194BB4B8BFA558C4B4D453" box="[151,1436,538,559]" gridrow="3" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA558C331D453" box="[151,537,538,559]" gridcol="0" gridrow="3" pageId="23" pageNumber="24">SL/body height (H)</th>
<td id="76DC9F7370194BB4BA8CA558C3E6D453" box="[676,718,538,559]" gridcol="1" gridrow="3" pageId="23" pageNumber="24">2.6</td>
<td id="76DC9F7370194BB4BBB0A558C2D0D453" box="[920,1016,538,559]" gridcol="2" gridrow="3" pageId="23" pageNumber="24">2.72.9</td>
<td id="76DC9F7370194BB4BCADA558C4B4D453" box="[1157,1436,538,559]" gridcol="3" gridrow="3" pageId="23" pageNumber="24">2.8 0.0</td>
</tr>
<tr id="350DF60F70194BB4B8BFA500C4B4D42B" box="[151,1436,578,599]" gridrow="4" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA500C331D42B" box="[151,537,578,599]" gridcol="0" gridrow="4" pageId="23" pageNumber="24">SL/head length (HL)</th>
<td id="76DC9F7370194BB4BA8CA500C3E6D42B" box="[676,718,578,599]" gridcol="1" gridrow="4" pageId="23" pageNumber="24">4.0</td>
<td id="76DC9F7370194BB4BBB0A500C2D0D42B" box="[920,1016,578,599]" gridcol="2" gridrow="4" pageId="23" pageNumber="24">3.74.0</td>
<td id="76DC9F7370194BB4BCADA500C4B4D42B" box="[1157,1436,578,599]" gridcol="3" gridrow="4" pageId="23" pageNumber="24">3.9 0.1</td>
</tr>
<tr id="350DF60F70194BB4B8BFA528C4B4D403" box="[151,1436,618,639]" gridrow="5" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA528C331D403" box="[151,537,618,639]" gridcol="0" gridrow="5" pageId="23" pageNumber="24">SL/pre-dorsal distance (PDD)</th>
<td id="76DC9F7370194BB4BA8CA528C3E6D403" box="[676,718,618,639]" gridcol="1" gridrow="5" pageId="23" pageNumber="24">1.6</td>
<td id="76DC9F7370194BB4BBB0A528C2D0D403" box="[920,1016,618,639]" gridcol="2" gridrow="5" pageId="23" pageNumber="24">1.51.6</td>
<td id="76DC9F7370194BB4BCADA528C4B4D403" box="[1157,1436,618,639]" gridcol="3" gridrow="5" pageId="23" pageNumber="24">1.6 0.0</td>
</tr>
<tr id="350DF60F70194BB4B8BFA5D0C4B4D4DB" box="[151,1436,658,679]" gridrow="6" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA5D0C331D4DB" box="[151,537,658,679]" gridcol="0" gridrow="6" pageId="23" pageNumber="24">SL/pre-anal distance (PAD)</th>
<td id="76DC9F7370194BB4BA8CA5D0C3E6D4DB" box="[676,718,658,679]" gridcol="1" gridrow="6" pageId="23" pageNumber="24">1.7</td>
<td id="76DC9F7370194BB4BBB0A5D0C2D0D4DB" box="[920,1016,658,679]" gridcol="2" gridrow="6" pageId="23" pageNumber="24">1.61.7</td>
<td id="76DC9F7370194BB4BCADA5D0C4B4D4DB" box="[1157,1436,658,679]" gridcol="3" gridrow="6" pageId="23" pageNumber="24">1.6 0.0</td>
</tr>
<tr id="350DF60F70194BB4B8BFA5F8C4B4D4B3" box="[151,1436,698,719]" gridrow="7" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA5F8C331D4B3" box="[151,537,698,719]" gridcol="0" gridrow="7" pageId="23" pageNumber="24">SL/dorsal fin length (DFL)</th>
<td id="76DC9F7370194BB4BA8CA5F8C3E6D4B3" box="[676,718,698,719]" gridcol="1" gridrow="7" pageId="23" pageNumber="24">4.5</td>
<td id="76DC9F7370194BB4BBB0A5F8C2D0D4B3" box="[920,1016,698,719]" gridcol="2" gridrow="7" pageId="23" pageNumber="24">4.44.7</td>
<td id="76DC9F7370194BB4BCADA5F8C4B4D4B3" box="[1157,1436,698,719]" gridcol="3" gridrow="7" pageId="23" pageNumber="24">4.6 0.2</td>
</tr>
<tr id="350DF60F70194BB4B8BFA5A0C4B4D48B" box="[151,1436,738,759]" gridrow="8" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA5A0C331D48B" box="[151,537,738,759]" gridcol="0" gridrow="8" pageId="23" pageNumber="24">SL/anal fin length (AFL)</th>
<td id="76DC9F7370194BB4BA8CA5A0C3E6D48B" box="[676,718,738,759]" gridcol="1" gridrow="8" pageId="23" pageNumber="24">3.3</td>
<td id="76DC9F7370194BB4BBB0A5A0C2D0D48B" box="[920,1016,738,759]" gridcol="2" gridrow="8" pageId="23" pageNumber="24">3.33.6</td>
<td id="76DC9F7370194BB4BCADA5A0C4B4D48B" box="[1157,1436,738,759]" gridcol="3" gridrow="8" pageId="23" pageNumber="24">3.4 0.1</td>
</tr>
<tr id="350DF60F70194BB4B8BFA448C4B4D563" box="[151,1436,778,799]" gridrow="9" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA448C331D563" box="[151,537,778,799]" gridcol="0" gridrow="9" pageId="23" pageNumber="24">SL/caudal peduncle length (CPL)</th>
<td id="76DC9F7370194BB4BA8CA448C3E6D563" box="[676,718,778,799]" gridcol="1" gridrow="9" pageId="23" pageNumber="24">6.9</td>
<td id="76DC9F7370194BB4BBB0A448C2D0D563" box="[920,1016,778,799]" gridcol="2" gridrow="9" pageId="23" pageNumber="24">6.17.1</td>
<td id="76DC9F7370194BB4BCADA448C4B4D563" box="[1157,1436,778,799]" gridcol="3" gridrow="9" pageId="23" pageNumber="24">6.8 0.3</td>
</tr>
<tr id="350DF60F70194BB4B8BFA470C4B4D53B" box="[151,1436,818,839]" gridrow="10" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA470C331D53B" box="[151,537,818,839]" gridcol="0" gridrow="10" pageId="23" pageNumber="24">SL/mouth width (MW)</th>
<td id="76DC9F7370194BB4BA8CA470C3E6D53B" box="[676,718,818,839]" gridcol="1" gridrow="10" pageId="23" pageNumber="24">17.4</td>
<td id="76DC9F7370194BB4BBB0A470C2D0D53B" box="[920,1016,818,839]" gridcol="2" gridrow="10" pageId="23" pageNumber="24">16.018.3</td>
<td id="76DC9F7370194BB4BCADA470C4B4D53B" box="[1157,1436,818,839]" gridcol="3" gridrow="10" pageId="23" pageNumber="24">16.8 0.8</td>
</tr>
<tr id="350DF60F70194BB4B8BFA4C0C4B4D5EB" box="[151,1436,898,919]" gridrow="11" pageId="23" pageNumber="24" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76DC9F7370194BB4B8BFA4C0C331D5EB" box="[151,537,898,919]" gridcol="0" gridrow="11" pageId="23" pageNumber="24">Ratio of head length (HL):</th>
</tr>
<tr id="350DF60F70194BB4B8BFA4E8C4B4D5C3" box="[151,1436,938,959]" gridrow="12" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA4E8C331D5C3" box="[151,537,938,959]" gridcol="0" gridrow="12" pageId="23" pageNumber="24">HL/snout length (SNL)</th>
<td id="76DC9F7370194BB4BA8CA4E8C3E6D5C3" box="[676,718,938,959]" gridcol="1" gridrow="12" pageId="23" pageNumber="24">6.6</td>
<td id="76DC9F7370194BB4BBB0A4E8C2D0D5C3" box="[920,1016,938,959]" gridcol="2" gridrow="12" pageId="23" pageNumber="24">6.28.3</td>
<td id="76DC9F7370194BB4BCADA4E8C4B4D5C3" box="[1157,1436,938,959]" gridcol="3" gridrow="12" pageId="23" pageNumber="24">7.3 0.7</td>
</tr>
<tr id="350DF60F70194BB4B8BFA490C4B4D59B" box="[151,1436,978,999]" gridrow="13" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA490C331D59B" box="[151,537,978,999]" gridcol="0" gridrow="13" pageId="23" pageNumber="24">HL/mouth width (MW)</th>
<td id="76DC9F7370194BB4BA8CA490C3E6D59B" box="[676,718,978,999]" gridcol="1" gridrow="13" pageId="23" pageNumber="24">4.3</td>
<td id="76DC9F7370194BB4BBB0A490C2D0D59B" box="[920,1016,978,999]" gridcol="2" gridrow="13" pageId="23" pageNumber="24">4.04.8</td>
<td id="76DC9F7370194BB4BCADA490C4B4D59B" box="[1157,1436,978,999]" gridcol="3" gridrow="13" pageId="23" pageNumber="24">4.3 0.2</td>
</tr>
<tr id="350DF60F70194BB4B8BFA4B8C4B4D273" box="[151,1436,1018,1039]" gridrow="14" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA4B8C331D273" box="[151,537,1018,1039]" gridcol="0" gridrow="14" pageId="23" pageNumber="24">HL/eye diameter (ED)</th>
<td id="76DC9F7370194BB4BA8CA4B8C3E6D273" box="[676,718,1018,1039]" gridcol="1" gridrow="14" pageId="23" pageNumber="24">3.7</td>
<td id="76DC9F7370194BB4BBB0A4B8C2D0D273" box="[920,1016,1018,1039]" gridcol="2" gridrow="14" pageId="23" pageNumber="24">3.84.2</td>
<td id="76DC9F7370194BB4BCADA4B8C4B4D273" box="[1157,1436,1018,1039]" gridcol="3" gridrow="14" pageId="23" pageNumber="24">4.0 0.1</td>
</tr>
<tr id="350DF60F70194BB4B8BFA360C4B4D24B" box="[151,1436,1058,1079]" gridrow="15" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA360C331D24B" box="[151,537,1058,1079]" gridcol="0" gridrow="15" pageId="23" pageNumber="24">HL/interorbital width (IOW)</th>
<td id="76DC9F7370194BB4BA8CA360C3E6D24B" box="[676,718,1058,1079]" gridcol="1" gridrow="15" pageId="23" pageNumber="24">2.7</td>
<td id="76DC9F7370194BB4BBB0A360C2D0D24B" box="[920,1016,1058,1079]" gridcol="2" gridrow="15" pageId="23" pageNumber="24">2.53.0</td>
<td id="76DC9F7370194BB4BCADA360C4B4D24B" box="[1157,1436,1058,1079]" gridcol="3" gridrow="15" pageId="23" pageNumber="24">2.8 0.2</td>
</tr>
<tr id="350DF60F70194BB4B8BFA308C4B4D223" box="[151,1436,1098,1119]" gridrow="16" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA308C331D223" box="[151,537,1098,1119]" gridcol="0" gridrow="16" pageId="23" pageNumber="24">HL/head width (HW)</th>
<td id="76DC9F7370194BB4BA8CA308C3E6D223" box="[676,718,1098,1119]" gridcol="1" gridrow="16" pageId="23" pageNumber="24">1.7</td>
<td id="76DC9F7370194BB4BBB0A308C2D0D223" box="[920,1016,1098,1119]" gridcol="2" gridrow="16" pageId="23" pageNumber="24">1.71.8</td>
<td id="76DC9F7370194BB4BCADA308C4B4D223" box="[1157,1436,1098,1119]" gridcol="3" gridrow="16" pageId="23" pageNumber="24">1.7 0.1</td>
</tr>
<tr id="350DF60F70194BB4B8BFA330C4B4D2FB" box="[151,1436,1138,1159]" gridrow="17" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA330C331D2FB" box="[151,537,1138,1159]" gridcol="0" gridrow="17" pageId="23" pageNumber="24">HL/mouth position (MP)</th>
<td id="76DC9F7370194BB4BA8CA330C3E6D2FB" box="[676,718,1138,1159]" gridcol="1" gridrow="17" pageId="23" pageNumber="24">4.7</td>
<td id="76DC9F7370194BB4BBB0A330C2D0D2FB" box="[920,1016,1138,1159]" gridcol="2" gridrow="17" pageId="23" pageNumber="24">4.25.0</td>
<td id="76DC9F7370194BB4BCADA330C4B4D2FB" box="[1157,1436,1138,1159]" gridcol="3" gridrow="17" pageId="23" pageNumber="24">4.5 0.3</td>
</tr>
<tr id="350DF60F70194BB4B8BFA380C4B4D2AB" box="[151,1436,1218,1239]" gridrow="18" pageId="23" pageNumber="24" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76DC9F7370194BB4B8BFA380C331D2AB" box="[151,537,1218,1239]" gridcol="0" gridrow="18" pageId="23" pageNumber="24">Ratio of caudal peduncle length (CPL):</th>
</tr>
<tr id="350DF60F70194BB4B8BFA3A8C4B4D283" box="[151,1436,1258,1279]" gridrow="19" pageId="23" pageNumber="24">
<th id="76DC9F7370194BB4B8BFA3A8C331D283" box="[151,537,1258,1279]" gridcol="0" gridrow="19" pageId="23" pageNumber="24">CPL/caudal peduncle depth (CPD)</th>
<td id="76DC9F7370194BB4BA8CA3A8C3E6D283" box="[676,718,1258,1279]" gridcol="1" gridrow="19" pageId="23" pageNumber="24">2.1</td>
<td id="76DC9F7370194BB4BBB0A3A8C2D0D283" box="[920,1016,1258,1279]" gridcol="2" gridrow="19" pageId="23" pageNumber="24">1.92.3</td>
<td id="76DC9F7370194BB4BCADA3A8C4B4D283" box="[1157,1436,1258,1279]" gridcol="3" gridrow="19" pageId="23" pageNumber="24">2.1 0.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8B82F44D7019B45CB8B7A220C013D30B" blockId="23.[159,1231,1338,1668]" box="[159,315,1378,1399]" pageId="23" pageNumber="24">Meristic counts:</paragraph>
<paragraph id="8B82F44D7019B45CB8B7A2C8C32ED048" blockId="23.[159,1231,1338,1668]" pageId="23" pageNumber="24">
Dorsal fin branched rays (DR) 22 2022 21 Anal fin branched rays (AR) 29 2729 28 Number of scales in the lateral line (SLL) 38 3638 37 Number of scale rows between the anterior base of
<date id="FF83D28D7019B45CBA8CA140C5B5D06B" box="[676,1181,1538,1559]" pageId="23" pageNumber="24" value="1911-12-10" valueMax="1911-12-14" valueMin="1911-12-10">11 1014 12</date>
the anal fin and the lateral line (SDL)
</paragraph>
<paragraph id="8B82F44D7019B45CB8B7A105C392D176" blockId="23.[159,1231,1338,1668]" lastBlockId="23.[151,1437,1738,2026]" pageId="23" pageNumber="24">
Number of teeth in the upper jaw (TUJ)
<date id="FF83D28D7019B45CBA8CA105C5B3D020" box="[676,1179,1607,1628]" pageId="23" pageNumber="24" value="1911-08-11" valueMax="1911-12-11" valueMin="1911-08-11">11 812 11</date>
Number of teeth in the lower jaw (TLJ) 18 1823 20
<emphasis id="B949285F7019B45CB8EEA188C088D098" bold="true" box="[198,416,1738,1764]" pageId="23" pageNumber="24">Other specimens.</emphasis>
We examined 19 other specimens from Odzala National Park (specimen list provided in the section &quot;additional material examined&quot;).
</paragraph>
</subSubSection>
<subSubSection id="C327A7C67019B453B8EEA057C592D75E" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8B82F44D7019B453B8EEA057C592D75E" blockId="23.[151,1437,1738,2026]" lastBlockId="24.[151,1437,152,1559]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="B949285F7019B45CB8EEA057C06ED153" bold="true" box="[198,326,1813,1839]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
<taxonomicName id="4C3D8FCE7019B45CB97AA057C3B8D152" box="[338,656,1813,1838]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="odzalaensis" status="sp. nov.">
<emphasis id="B949285F7019B45CB97AA057C3B8D152" box="[338,656,1813,1838]" italics="true" pageId="23" pageNumber="24">Petrocephalus odzalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A95247019B45CBAB4A057C3CBD153" box="[668,739,1813,1839]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B949285F7019B45CBAB4A057C3CBD153" bold="true" box="[668,739,1813,1839]" pageId="23" pageNumber="24">n. sp.</emphasis>
</taxonomicNameLabel>
is distinguished from all other
<taxonomicName id="4C3D8FCE7019B45CBC47A057C43FD152" box="[1135,1303,1813,1838]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B949285F7019B45CBC47A057C43FD152" box="[1135,1303,1813,1838]" italics="true" pageId="23" pageNumber="24">Petrocephalus</emphasis>
</taxonomicName>
species in Central Africa by the following combination of characteristics. Dorsal fin shorter than anal fin. Dorsal fin with a maximum of 22 branched rays (range = 2022). Anal fin with a minimum of 27 branched rays (range = 2729). Mouth sub-terminal; ratio between head length and mouth position is between 4.2 and 5.0. Eye small (3.7 ≤ HL/ED ≤ 4.2). Body pinkish-gray, darker dorsally, with the presence of three distinct pigmentation patches: (1) a distinct ovoid black mark situated below the anterior base of the dorsal fin on each side of the body; (2) a small black mark at the base of each pectoral fin; (3) an ovoid black mark on each side that is centered at the base of the caudal fin, not extending onto the upper and lower lobes of this fin. EOD of normal polarity, appearing triphasic at low gain, with two main phases and a small third phase. A final, fourth phase may be present, but it is always extremely small (&lt;1.5% of total peak-to-peak amplitude).
</paragraph>
</subSubSection>
<subSubSection id="C327A7C67016B453B8EEA66FC5C1D24E" pageId="24" pageNumber="25" type="description">
<paragraph id="8B82F44D7016B453B8EEA66FC06DD5E0" blockId="24.[151,1437,152,1559]" pageId="24" pageNumber="25">
<emphasis id="B949285F7016B453B8EEA66FC077D73B" bold="true" box="[198,351,301,327]" pageId="24" pageNumber="25">Description.</emphasis>
Morphometric ratios and meristic data for the
<typeStatus id="54864AEF7016B453BBABA66FC2C1D73B" box="[899,1001,301,327]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
and
<typeStatus id="54864AEF7016B453BC0AA66FC5BAD73B" box="[1058,1170,301,327]" pageId="24" pageNumber="25" type="paratype">paratypes</typeStatus>
are presented in
<tableCitation id="C6BFC1F67016B453BD73A66FC181D710" captionStart="TABLE 6" captionStartId="23.[151,242,151,175]" captionTargetBox="[151,1436,378,1279]" captionTargetPageId="23" captionText="TABLE 6. Principal morphometric ratios and meristic counts for the holotype (CU 88048) and nine paratypes (CU 87857, CU 88049, CU 87850, CU 88050, CU 88059, AMNH 250928, AMNH 250929, AMNH 250930, AMNH 250931) of Petrocephalus odzalaensis n. sp. (Abbreviations: o = sex category &quot; other &quot; [i. e., not an obvious mature male, but rather a female or immature male]; Std Dev = standard deviation; Min Max = minimum maximum)." httpUri="http://table.plazi.org/id/DF42A4C57019B45CB8BFA7D5C5ACD773" pageId="24" pageNumber="25" tableUuid="DF42A4C57019B45CB8BFA7D5C5ACD773">Table 6</tableCitation>
.
<taxonomicName id="4C3D8FCE7016B453B89EA610C0DFD717" box="[182,503,338,363]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="odzalaensis" status="sp. nov.">
<emphasis id="B949285F7016B453B89EA610C0DFD717" box="[182,503,338,363]" italics="true" pageId="24" pageNumber="25">Petrocephalus odzalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A95247016B453B9D7A610C36CD710" box="[511,580,338,364]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B949285F7016B453B9D7A610C36CD710" bold="true" box="[511,580,338,364]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
is a medium sized species within the genus (maximum SL observed =
<quantity id="4CC559A87016B453B8BFA63AC02FD7EE" box="[151,263,376,402]" metricMagnitude="-2" metricUnit="m" metricValue="9.93" pageId="24" pageNumber="25" unit="mm" value="99.3">99.3 mm</quantity>
;
<typeStatus id="54864AEF7016B453B93CA63AC056D7EE" box="[276,382,376,402]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
=
<quantity id="4CC559A87016B453B9B7A63AC339D7EE" box="[415,529,376,402]" metricMagnitude="-2" metricUnit="m" metricValue="9.29" pageId="24" pageNumber="25" unit="mm" value="92.9">92.9 mm</quantity>
). Body ovoid, longer than high (2.6 ≤ SL/H ≤ 2.9,
<typeStatus id="54864AEF7016B453BC57A63AC5CFD7EE" box="[1151,1255,376,402]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
average = 2.8,
<typeStatus id="54864AEF7016B453B8BFA6DFC1D6D7CB" box="[151,254,413,439]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 2.6) and laterally compressed. Head length between 3.7 and 4.0 times in standard length (
<typeStatus id="54864AEF7016B453BD1DA6DFC4B4D7CB" box="[1333,1436,413,439]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
average = 3.9,
<typeStatus id="54864AEF7016B453B96CA680C083D7A0" box="[324,427,450,476]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 4.0). Snout short (6.2 ≤ HL/SNL ≤ 8.3,
<typeStatus id="54864AEF7016B453BBB4A680C529D7A0" box="[924,1025,450,476]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
average = 7.3,
<typeStatus id="54864AEF7016B453BC9EA680C435D7A0" box="[1206,1309,450,476]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 6.6) and round. Mouth small (4.0 ≤ HL/MW ≤ 4.8,
<typeStatus id="54864AEF7016B453BAA3A6AAC3D8D47E" box="[651,752,488,514]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
average = 4.3,
<typeStatus id="54864AEF7016B453BB8DA6AAC524D47E" box="[933,1036,488,514]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 4.3), sub-terminal, opening just under the anterior half of the eye. Teeth small and bicuspid,
<quantity id="4CC559A87016B453BB4EA54FC2E8D45B" box="[870,960,525,551]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="3.048" metricValueMin="2.032" pageId="24" pageNumber="25" unit="in" value="10.0" valueMax="12.0" valueMin="8.0">812 in</quantity>
a single row in the upper jaw (
<typeStatus id="54864AEF7016B453BD1CA54FC4B3D45B" box="[1332,1435,525,551]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
median = 11,
<typeStatus id="54864AEF7016B453B912A570C089D430" box="[314,417,562,588]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 11),
<quantity id="4CC559A87016B453B9DFA570C348D430" box="[503,608,562,588]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.842" metricValueMin="4.572" pageId="24" pageNumber="25" unit="in" value="20.5" valueMax="23.0" valueMin="18.0">1823 in</quantity>
a single row in the lower jaw (
<typeStatus id="54864AEF7016B453BBFAA570C511D430" box="[978,1081,562,588]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
median = 20,
<typeStatus id="54864AEF7016B453BCCDA570C464D430" box="[1253,1356,562,588]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 18). Dorsal and anal fins originate in the posterior half of the body (1.5 ≤ SL/PDD ≤ 1.6 and 1.6 ≤ SL/PAD ≤ 1.7, respectively). Pre-dorsal distance slightly greater than the preanal distance (1.0 ≤ PDD/PAD ≤ 1.1). Dorsal fin with 2022 branched rays (
<typeStatus id="54864AEF7016B453BA27A5E0C352D4C0" box="[527,634,674,700]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
median = 21,
<typeStatus id="54864AEF7016B453BB04A5E0C2BED4C0" box="[812,918,674,700]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 22). Anal fin with 2729 branched rays (
<typeStatus id="54864AEF7016B453B8B7A58AC020D49E" box="[159,264,712,738]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
median = 28,
<typeStatus id="54864AEF7016B453B99FA58AC308D49E" box="[439,544,712,738]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 29). Scales cover the body, except for the head. Lateral line visible and complete with 3638 (
<typeStatus id="54864AEF7016B453B989A5AFC322D57B" box="[417,522,749,775]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 38) pored scales along its length. Ten to 14 scales (
<typeStatus id="54864AEF7016B453BCA3A5AFC5DAD57B" box="[1163,1266,749,775]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
average = 12,
<typeStatus id="54864AEF7016B453B8BFA450C1D6D550" box="[151,254,786,812]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 11) between the anterior base of the anal fin and the lateral line. Caudal peduncle thin (1.9 ≤ CPL/ CPD ≤ 2.3,
<typeStatus id="54864AEF7016B453B90CA47AC0A1D52E" box="[292,393,824,850]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
average = 2.1,
<typeStatus id="54864AEF7016B453BA68A47AC38FD52E" box="[576,679,824,850]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
= 2.1). Twelve scales around the caudal peduncle. Skin on head thick, turning opaque with formalin fixation. Knollenorgans clustered into the three distinct &quot;rosettes&quot; of
<bibRefCitation id="EFAC89BC7016B453B8BFA4C0C069D5E0" author="Harder" box="[151,321,898,924]" pageId="24" pageNumber="25" refString="Harder, W. (1968) Zum Aufbau der epidermalen Sinnesorgane der Mormyridae (Mormyriformes, Teleostei). Zeitschrift fur Zellforschung, 89, 212 - 224." type="journal article" year="1968">Harder (1968)</bibRefCitation>
.
</paragraph>
<paragraph id="8B82F44D7016B453B8EEA4E5C5C1D24E" blockId="24.[151,1437,152,1559]" pageId="24" pageNumber="25">
<emphasis id="B949285F7016B453B8EEA4E5C0AFD5BD" bold="true" box="[198,391,935,961]" pageId="24" pageNumber="25">Live coloration</emphasis>
(
<figureCitation id="1306E8C87016B453B9B3A4EAC0CBD5BE" box="[411,483,936,962]" captionStart="FIGURE 8" captionStartId="22.[151,258,1771,1795]" captionTargetBox="[188,1398,269,1750]" captionTargetId="figure@22.[188,1398,269,1751]" captionTargetPageId="22" captionText="FIGURE 8. Petrocephalus odzalaensis n. sp. of the Lékoli River system of Odzala National Park, Republic of the Congo. A. Photograph of a live specimen (no scale). B. Photograph of the preserved holotype (CU 88048; scale bar = 1.0 cm). C. EOD waveforms. One EOD was arbitrarily selected from each recorded individual and superimposed with the other recordings (N = number of individuals). All EODs are scaled to the same peak-to-peak voltage, and they are plotted (head-positive up) on the same time scale as overlays centered on the largest positive peak of each waveform (scale bar = 0.2 msec)." httpUri="https://zenodo.org/record/197597/files/figure.png" pageId="24" pageNumber="25">Fig. 8</figureCitation>
A). Body background color pinkish-gray, darker dorsally. Pigmentation pattern consisting of three characteristic black patches: (1) a distinct ovoid black mark below the anterior base of the dorsal fin; (2) a small black mark at the base of the pectoral fin; and (3) an ovoid black mark centered at the base of the caudal fin, which does not extend onto the upper and lower lobes. Fins translucent.
</paragraph>
</subSubSection>
<subSubSection id="C327A7C67016B453B8EEA37FC0BDD3FE" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8B82F44D7016B453B8EEA37FC06CD2DE" blockId="24.[151,1437,152,1559]" pageId="24" pageNumber="25">
<emphasis id="B949285F7016B453B8EEA37FC077D22B" bold="true" box="[198,351,1085,1111]" pageId="24" pageNumber="25">Distribution</emphasis>
(
<figureCitation id="1306E8C87016B453B959A37FC09FD22B" box="[369,439,1085,1111]" captionStart="FIGURE 1" captionStartId="2.[151,255,1561,1585]" captionTargetBox="[169,1408,388,1531]" captionTargetId="figure@2.[151,1435,383,1536]" captionTargetPageId="2" captionText="FIGURE 1. Map of Africa (right) showing the entire Congo River and its main tributaries together with the neighboring Lower Guinea province (first inset at center). The boundary between the Congo and Lower Guinea provinces is shown by a dashed line. Filled black circles indicate type localities for all Petrocephalus species that were described from either province and were considered valid just prior to the present study. Exact type localities are unknown for P. b a l a y i and P. sauvagii. Based on our understanding of the chronology of Central African exploration at the time of the descriptions, we estimate these type localities to be the following: the vicinity of Pool Malebo on the Lower Congo River near the modern cities of Brazzaville and Kinshasa [4.25 ° S, 15.29 ° E] for P. sauvagii; and the vicinity of the present-day city Lambaréné on the Lower Ogooué River [0.7 ° S, 10.22 ° E] for P. b a l a y i. Second inset at the far left: detailed map of Odzala National Park, located in the Congo River basin. Within the boundaries of the park, forest is shown in green and savannah in yellow. The red oval indicates our collection area (a segment of the Lékoli drainage system, including the lower parts of two of its tributaries, the Pandaka and Lékénie Rivers)." httpUri="https://zenodo.org/record/197590/files/figure.png" pageId="24" pageNumber="25">Fig. 1</figureCitation>
). Endemic to the
<collectingCountry id="F32AB4DD7016B453BAA7A37FC3F5D22B" box="[655,733,1085,1111]" name="Republic of the Congo" pageId="24" pageNumber="25">Congo</collectingCountry>
basin. Abundant in Odzala. We collected
<taxonomicName id="4C3D8FCE7016B453BCFCA37FC454D22A" box="[1236,1404,1085,1110]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="odzalaensis" status="sp. nov.">
<emphasis id="B949285F7016B453BCFCA37FC454D22A" box="[1236,1404,1085,1110]" italics="true" pageId="24" pageNumber="25">P. odzalaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A95247016B453BDACA37FC192D200" pageId="24" pageNumber="25" rank="species">
<emphasis id="B949285F7016B453BDACA37FC192D200" bold="true" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
at several localities along the main course of the Lékoli River and in some small tributary creeks flowing through forest.
</paragraph>
<paragraph id="8B82F44D7016B453B8EEA3EFC0BDD3FE" blockId="24.[151,1437,152,1559]" pageId="24" pageNumber="25">
<emphasis id="B949285F7016B453B8EEA3EFC32BD2BB" bold="true" box="[198,515,1197,1223]" pageId="24" pageNumber="25">Electric organ discharge</emphasis>
(
<figureCitation id="1306E8C87016B453BA31A3EFC34CD2BB" box="[537,612,1197,1223]" captionStart="FIGURE 8" captionStartId="22.[151,258,1771,1795]" captionTargetBox="[188,1398,269,1750]" captionTargetId="figure@22.[188,1398,269,1751]" captionTargetPageId="22" captionText="FIGURE 8. Petrocephalus odzalaensis n. sp. of the Lékoli River system of Odzala National Park, Republic of the Congo. A. Photograph of a live specimen (no scale). B. Photograph of the preserved holotype (CU 88048; scale bar = 1.0 cm). C. EOD waveforms. One EOD was arbitrarily selected from each recorded individual and superimposed with the other recordings (N = number of individuals). All EODs are scaled to the same peak-to-peak voltage, and they are plotted (head-positive up) on the same time scale as overlays centered on the largest positive peak of each waveform (scale bar = 0.2 msec)." httpUri="https://zenodo.org/record/197597/files/figure.png" pageId="24" pageNumber="25">Fig. 8</figureCitation>
C). The EOD waveform is typical for the genus, similar to EODs produced by many other
<taxonomicName id="4C3D8FCE7016B453B997A391C34FD290" box="[447,615,1235,1260]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B949285F7016B453B997A391C34FD290" box="[447,615,1235,1260]" italics="true" pageId="24" pageNumber="25">Petrocephalus</emphasis>
</taxonomicName>
species. Total EOD duration ranges from 0.231 to 0.339 msec, based on 1.5% voltage deviations from baseline relative to peak-peak amplitude. No EOD sex differences are apparent in the specimens recorded thus far.
<bibRefCitation id="EFAC89BC7016B453BA85A25FC289D34B" box="[685,929,1309,1335]" pageId="24" pageNumber="25" refString="Lavoue, S., Arnegard, M. E., Sullivan, J. P. &amp; Hopkins, C. D. (2008) Petrocephalus of Odzala offer insights into evolutionary patterns of signal diversification in the Mormyridae, a family of weakly electrogenic fishes from Africa. Journal of Physiology - Paris, 102, 322 - 339." type="journal article">
Lavoué
<emphasis id="B949285F7016B453BB25A25FC261D34A" box="[781,841,1309,1334]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
(2008)
</bibRefCitation>
provide additional statistics for waveform landmarks and other EOD measurements. Histological examination confirms that electrocytes are
<typeStatus id="54864AEF7016B453BD04A200C477D320" box="[1324,1375,1346,1372]" pageId="24" pageNumber="25">type</typeStatus>
NPp (
<bibRefCitation id="EFAC89BC7016B453B8B7A22AC0A1D3FE" author="Lavoue" box="[159,393,1384,1410]" pageId="24" pageNumber="25" refString="Lavoue, S., Arnegard, M. E., Sullivan, J. P. &amp; Hopkins, C. D. (2008) Petrocephalus of Odzala offer insights into evolutionary patterns of signal diversification in the Mormyridae, a family of weakly electrogenic fishes from Africa. Journal of Physiology - Paris, 102, 322 - 339." type="journal article" year="2008">
Lavoué
<emphasis id="B949285F7016B453B928A22AC012D3FD" box="[256,314,1384,1409]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
, 2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327A7C67016B453B8EEA2CFC377D38E" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8B82F44D7016B453B8EEA2CFC377D38E" blockId="24.[151,1437,152,1559]" pageId="24" pageNumber="25">
<emphasis id="B949285F7016B453B8EEA2CFC068D3DB" bold="true" box="[198,320,1421,1447]" pageId="24" pageNumber="25">Remarks.</emphasis>
Based on museum records from the
<collectingCountry id="F32AB4DD7016B453BAD4A2CFC262D3DB" box="[764,842,1421,1447]" name="Republic of the Congo" pageId="24" pageNumber="25">Congo</collectingCountry>
basin (
<emphasis id="B949285F7016B453BB8CA2CFC53FD3DA" box="[932,1047,1421,1446]" italics="true" pageId="24" pageNumber="25">pers. obs.</emphasis>
),
<taxonomicName id="4C3D8FCE7016B453BC19A2CFC444D3DA" box="[1073,1388,1421,1446]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="odzalaensis">
<emphasis id="B949285F7016B453BC19A2CFC444D3DA" box="[1073,1388,1421,1446]" italics="true" pageId="24" pageNumber="25">Petrocephalus odzalaensis</emphasis>
</taxonomicName>
has been misidentified as
<taxonomicName id="4C3D8FCE7016B453B988A2F1C3B1D3B0" box="[416,665,1459,1484]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="simus">
<emphasis id="B949285F7016B453B988A2F1C3B1D3B0" box="[416,665,1459,1484]" italics="true" pageId="24" pageNumber="25">Petrocephalus simus</emphasis>
</taxonomicName>
in several instances. The latter species is endemic to the Lower
<collectingCountry id="F32AB4DD7016B453B8BFA29AC1C4D38E" box="[151,236,1496,1522]" name="Guinea-Bissau" pageId="24" pageNumber="25">Guinea</collectingCountry>
province (
<bibRefCitation id="EFAC89BC7016B453B940A29AC37BD38E" author="Lavoue" box="[360,595,1496,1522]" pageId="24" pageNumber="25" refString="Lavoue, S., Hopkins, C. D. &amp; Kamdem Toham, A. (2004) The Petrocephalus (Pisces, Osteoglossomorpha, Mormyridae) of Gabon, Central Africa, with the description of a new species. Zoosystema, 26, 511 - 535." type="journal article" year="2004">
Lavoué
<emphasis id="B949285F7016B453B9E2A29AC32CD38D" box="[458,516,1496,1521]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
, 2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327A7C67016B453B8EEA2BFC3CCD06B" box="[198,740,1533,1559]" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8B82F44D7016B453B8EEA2BFC3CCD06B" blockId="24.[151,1437,152,1559]" box="[198,740,1533,1559]" pageId="24" pageNumber="25">
<emphasis id="B949285F7016B453B8EEA2BFC07BD06B" bold="true" box="[198,339,1533,1559]" pageId="24" pageNumber="25">Etymology.</emphasis>
Named for Odzala National Park.
</paragraph>
</subSubSection>
</treatment>
</document>