treatments-xml/data/03/91/02/0391026EFFAFFFA2FF0B7573FADAFAA1.xml

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<mods:title id="AFD7345F53E7A4313A5D68CE6BDDB8E7">Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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3.3 Key to 
<taxonomicName id="4C38C8FBFFAFFFADFE8C7573FE74FBC9" ID-CoL="8VTS6" authorityName="GANGLBAUER" authorityYear="1895" box="[356,448,1103,1135]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">species</taxonomicName>
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<key id="FFA9E449FFAFFFA2FF2B75A5FADAFAA1" lastPageId="9" lastPageNumber="210" pageId="6" pageNumber="207">
<keyStep id="30CCABDDFFAFFFADFF2B75A5FA22FB46" pageId="6" pageNumber="207">
<paragraph id="8B87B378FFAFFFADFF2B75A5FA22FB17" blockId="6.[195,1430,1177,1248]" box="[195,1430,1177,1201]" pageId="6" pageNumber="207">
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1. Species from 
<collectingCountry id="F32FF3E8FFAFFFADFE9575A5FE0AFB17" box="[381,446,1177,1201]" name="China" pageId="6" pageNumber="207">China</collectingCountry>
and 
<collectingCountry id="F32FF3E8FFAFFFADFE1B75A5FDF7FB17" box="[499,579,1177,1201]" name="Taiwan" pageId="6" pageNumber="207">Taiwan</collectingCountry>
. ....................................................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAFFFADFF2B75F4FA22FB46" blockId="6.[195,1430,1177,1248]" box="[195,1430,1224,1248]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B75F4FA22FB46" box="[195,1430,1224,1248]" pageId="6" pageNumber="207">– Distribution different ........................................................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAFFFADFF2B75CBFA22FA98" pageId="6" pageNumber="207">
<paragraph id="8B87B378FFAFFFADFF2B75CBFA22FAA9" blockId="6.[195,1430,1271,1342]" box="[195,1430,1271,1295]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B75CBFA22FAA9" box="[195,1430,1271,1295]" pageId="6" pageNumber="207">2. Pronotum and most of elytra blackish-brown to black. .................................................................................................. 3</keyLead>
</paragraph>
<paragraph id="8B87B378FFAFFFADFF2B741BFA22FA98" blockId="6.[195,1430,1271,1342]" box="[195,1430,1318,1342]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B741BFA22FA98" box="[195,1430,1318,1342]" pageId="6" pageNumber="207">
<heading id="D0CF0414FFAFFFADFF2B741BFF65FA98" box="[195,209,1319,1342]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="207" reason="4">–</heading>
Pronotum and most of elytra pale-reddish to pale-brown. ............................................................................................ 4
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</keyStep>
<keyStep id="30CCABDDFFAFFFADFF2B7469FA22F9BA" pageId="6" pageNumber="207">
<paragraph id="8B87B378FFAFFFADFF2B7469FA23FA0B" blockId="6.[195,1431,1365,1453]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B7469FA23FA0B" pageId="6" pageNumber="207">
3. Male pronotum with shallow impression. Antenna without sexual dimorphism. Tergite VIII sexually dimorphic, in male with broadly concave and in female with truncate posterior margin (
<figureCitation id="1303AFFDFFAFFFADFC187449FB34FA2B" box="[1008,1152,1397,1421]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="6" pageNumber="207">Figs 178, 182</figureCitation>
). Median lobe of aedeagus shaped as in 
<figureCitation id="1303AFFDFFAFFFADFE9974A9FDB4FA0B" box="[369,512,1429,1453]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="6" pageNumber="207">Figs 180–181</figureCitation>
. Distribution: 
<collectingCountry id="F32FF3E8FFAFFFADFD7174A9FD52FA0B" box="[665,742,1429,1453]" name="Taiwan" pageId="6" pageNumber="207">Taiwan</collectingCountry>
(
<figureCitation id="1303AFFDFFAFFFADFD1C74A9FC8FFA0B" box="[756,827,1429,1453]" captionStart="Map 7" captionStartId="46.[189,234,2053,2074]" captionTargetBox="[189,1436,1241,2044]" captionTargetId="figure-539@46.[189,1436,1241,2044]" captionTargetPageId="46" captionText="Map 7: Distributions of Anaulacaspis beesoni (black circles), A. pectinata (white star), A. schuelkei (black triangle), A. beijingensis (white circles), and A. taiwanica (black star) in the East Palaearctic region, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212643" httpUri="https://zenodo.org/record/5212643/files/figure.png" pageId="6" pageNumber="207">Map 7</figureCitation>
). ............................................................................ 
<taxonomicName id="4C38C8FBFFAFFFADFAC574A9FA23FA0B" ID-CoL="DJL3" authorityName="Pace" authorityYear="2008" box="[1325,1431,1429,1453]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="taiwanica">
<emphasis id="B94C6F6AFFAFFFADFAC574A9FA23FA0B" bold="true" box="[1325,1431,1429,1453]" italics="true" pageId="6" pageNumber="207">taiwanica</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8B87B378FFAFFFADFF2B74F9FA22F9BA" blockId="6.[195,1431,1476,1564]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B74F9FA22F9BA" pageId="6" pageNumber="207">
<heading id="D0CF0414FFAFFFADFF2B74F9FF65FA7A" box="[195,209,1477,1500]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="207" reason="4">–</heading>
Male pronotum with deep and extensive impression. Antenna sexually dimorphic, longer and more slender in male than in female. Tergite VIII (
<figureCitation id="1303AFFDFFAFFFADFDF974D8FDDCFA5A" box="[529,616,1508,1532]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="6" pageNumber="207">Fig. 173</figureCitation>
) strongly tranverse and with truncate posterior margin, without sexual dimorphism. Median lobe of aedeagus as in 
<figureCitation id="1303AFFDFFAFFFADFD927738FCBDF9BA" box="[634,777,1540,1564]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="6" pageNumber="207">Figs 175–176</figureCitation>
. Distribution: 
<collectingCountry id="F32FF3E8FFAFFFADFC4A7738FC52F9BA" box="[930,998,1540,1564]" name="China" pageId="6" pageNumber="207">China</collectingCountry>
: 
<collectingRegion id="49FC7D9AFFAFFFADFC077738FB89F9BA" box="[1007,1085,1540,1564]" country="China" name="Beijing" pageId="6" pageNumber="207">Beijing</collectingRegion>
, 
<collectingRegion id="49FC7D9AFFAFFFADFBAE7738FB2EF9BA" box="[1094,1178,1540,1564]" country="China" name="Shaanxi" pageId="6" pageNumber="207">Shaanxi</collectingRegion>
(
<figureCitation id="1303AFFDFFAFFFADFB407738FB5BF9BA" box="[1192,1263,1540,1564]" captionStart="Map 7" captionStartId="46.[189,234,2053,2074]" captionTargetBox="[189,1436,1241,2044]" captionTargetId="figure-539@46.[189,1436,1241,2044]" captionTargetPageId="46" captionText="Map 7: Distributions of Anaulacaspis beesoni (black circles), A. pectinata (white star), A. schuelkei (black triangle), A. beijingensis (white circles), and A. taiwanica (black star) in the East Palaearctic region, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212643" httpUri="https://zenodo.org/record/5212643/files/figure.png" pageId="6" pageNumber="207">Map 7</figureCitation>
). ... 
<taxonomicName id="4C38C8FBFFAFFFADFAF27738FA22F9BA" ID-CoL="DJHZ" baseAuthorityName="Pace" baseAuthorityYear="1998" box="[1306,1430,1540,1564]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="beijingensis">
<emphasis id="B94C6F6AFFAFFFADFAF27738FA22F9BA" bold="true" box="[1306,1430,1540,1564]" italics="true" pageId="6" pageNumber="207">beijingensis</emphasis>
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<paragraph id="8B87B378FFAFFFADFF2B770FFA22F90D" blockId="6.[195,1431,1587,1707]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B770FFA22F90D" pageId="6" pageNumber="207">
4. Antennomere XI pale-brown, paler than antennomeres V–X. Male head depressed dorsally. Posterior margin of tergite VIII concave and with unmodified thin setae (
<bibRefCitation id="EFA9CE89FFAFFFADFCF0776FFC15F9CD" author="ASSING, V." box="[792,929,1619,1643]" pageId="6" pageNumber="207" pagination="113 - 118" refId="ref48261" refString="ASSING, V. 2001: Two new species and a new name of Cordalia JACOBS, 1925 from Turkey and China (Insecta: Coleoptera: Staphylinidae: Aleocharinae). - Reichenbachia 34 (12): 113 - 118." type="journal article" year="2001">ASSING 2001</bibRefCitation>
: figure 12). Median lobe of aedeagus with broad (ventral view) and strongly curved (lateral view) ventral process (
<bibRefCitation id="EFA9CE89FFAFFFADFC65774FFBA2F92D" author="ASSING, V." box="[909,1046,1651,1675]" pageId="6" pageNumber="207" pagination="113 - 118" refId="ref48261" refString="ASSING, V. 2001: Two new species and a new name of Cordalia JACOBS, 1925 from Turkey and China (Insecta: Coleoptera: Staphylinidae: Aleocharinae). - Reichenbachia 34 (12): 113 - 118." type="journal article" year="2001">ASSING 2001</bibRefCitation>
: figures 10–11). Distribution: 
<collectingCountry id="F32FF3E8FFAFFFADFAA7774FFA27F92D" box="[1359,1427,1651,1675]" name="China" pageId="6" pageNumber="207">China</collectingCountry>
: 
<collectingRegion id="49FC7D9AFFAFFFADFF0177AFFE89F90D" box="[233,317,1683,1707]" country="China" name="Shaanxi" pageId="6" pageNumber="207">Shaanxi</collectingRegion>
(
<figureCitation id="1303AFFDFFAFFFADFEA277AFFE26F90D" box="[330,402,1683,1707]" captionStart="Map 7" captionStartId="46.[189,234,2053,2074]" captionTargetBox="[189,1436,1241,2044]" captionTargetId="figure-539@46.[189,1436,1241,2044]" captionTargetPageId="46" captionText="Map 7: Distributions of Anaulacaspis beesoni (black circles), A. pectinata (white star), A. schuelkei (black triangle), A. beijingensis (white circles), and A. taiwanica (black star) in the East Palaearctic region, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212643" httpUri="https://zenodo.org/record/5212643/files/figure.png" pageId="6" pageNumber="207">Map 7</figureCitation>
). .................................................................................................................................................... 
<taxonomicName id="4C38C8FBFFAFFFADFADD77AFFA22F90D" ID-CoL="DJKS" baseAuthorityName="ASSING" baseAuthorityYear="2001" box="[1333,1430,1683,1707]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="B94C6F6AFFAFFFADFADD77AFFA22F90D" bold="true" box="[1333,1430,1683,1707]" italics="true" pageId="6" pageNumber="207">schuelkei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAFFFADFF2B77FFFA22F89C" blockId="6.[195,1430,1730,1851]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B77FFFA22F89C" pageId="6" pageNumber="207">
<heading id="D0CF0414FFAFFFADFF2B77FFFF65F97C" box="[195,209,1731,1754]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="207" reason="4">–</heading>
Antennomere XI blackish-brown, not paler than antennomeres V–X. Male head unmodified, not depressed dorsally. Posterior margin of tergite VIII convex and with a comb-like row of modified, long and rather stout setae (
<figureCitation id="1303AFFDFFAFFFADFF18763EFE36F8BD" box="[240,386,1794,1819]" captionStart="Figs 187–210" captionStartId="19.[151,191,1941,1963]" captionTargetBox="[170,1381,238,1935]" captionTargetId="figure-145@19.[162,1387,232,1940]" captionText="Figs 187–210: Anaulacaspsis gilva (187–190), A. pectinata (191–195), and A. formosa (196–210; 199–200: Tunisia; 201–203: Morocco; 204–205: holotype of A. macra; 206: Murcia; 207–208: Andalucía): median lobe of aedeagus in lateral and in ventral view (187–188, 194–195, 199–208); female tergite VIII (189); female sternite VIII (190, 209); male tergite VIII (191, 196); posterior margin of male tergite VIII (192, 197); male sternite VIII (193, 198); spermatheca (210). Scale bars: 187–191, 193–196, 198–210: 0.1 mm; 192, 197: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884777" httpUri="https://zenodo.org/record/5884777/files/figure.png" pageId="6" pageNumber="207">Figs 191–192</figureCitation>
). Median lobe of aedeagus slender, with very slender and apically acute ventral process and with long dark structure in internal sac (
<figureCitation id="1303AFFDFFAFFFADFDB7761EFD46F89C" box="[607,754,1826,1850]" captionStart="Figs 187–210" captionStartId="19.[151,191,1941,1963]" captionTargetBox="[170,1381,238,1935]" captionTargetId="figure-145@19.[162,1387,232,1940]" captionText="Figs 187–210: Anaulacaspsis gilva (187–190), A. pectinata (191–195), and A. formosa (196–210; 199–200: Tunisia; 201–203: Morocco; 204–205: holotype of A. macra; 206: Murcia; 207–208: Andalucía): median lobe of aedeagus in lateral and in ventral view (187–188, 194–195, 199–208); female tergite VIII (189); female sternite VIII (190, 209); male tergite VIII (191, 196); posterior margin of male tergite VIII (192, 197); male sternite VIII (193, 198); spermatheca (210). Scale bars: 187–191, 193–196, 198–210: 0.1 mm; 192, 197: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884777" httpUri="https://zenodo.org/record/5884777/files/figure.png" pageId="6" pageNumber="207">Figs 194–195</figureCitation>
). Distribution: 
<collectingCountry id="F32FF3E8FFAFFFADFC7B761EFC63F89C" box="[915,983,1826,1850]" name="China" pageId="6" pageNumber="207">China</collectingCountry>
: 
<collectingRegion id="49FC7D9AFFAFFFADFC08761FFB80F89C" box="[992,1076,1827,1850]" country="China" name="Yunnan" pageId="6" pageNumber="207">Yunnan</collectingRegion>
(
<figureCitation id="1303AFFDFFAFFFADFBA9761FFB3DF89D" box="[1089,1161,1827,1851]" captionStart="Map 7" captionStartId="46.[189,234,2053,2074]" captionTargetBox="[189,1436,1241,2044]" captionTargetId="figure-539@46.[189,1436,1241,2044]" captionTargetPageId="46" captionText="Map 7: Distributions of Anaulacaspis beesoni (black circles), A. pectinata (white star), A. schuelkei (black triangle), A. beijingensis (white circles), and A. taiwanica (black star) in the East Palaearctic region, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212643" httpUri="https://zenodo.org/record/5212643/files/figure.png" pageId="6" pageNumber="207">Map 7</figureCitation>
). ....................... 
<taxonomicName id="4C38C8FBFFAFFFADFADA761EFA22F89C" ID-CoL="DJKG" authorityName="Assing" authorityYear="2016" box="[1330,1430,1826,1850]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="pectinata">
<emphasis id="B94C6F6AFFAFFFADFADA761EFA22F89C" bold="true" box="[1330,1430,1826,1850]" italics="true" pageId="6" pageNumber="207">pectinata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAFFFADFF2B766DFA23F85F" pageId="6" pageNumber="207">
<paragraph id="8B87B378FFAFFFADFF2B766DFA22F80C" blockId="6.[195,1431,1873,1962]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B766DFA22F80C" pageId="6" pageNumber="207">
5. Whole body dark-brown to black, sometimes with the posterior margin of the elytra more or less distinctly paler. Small species; length of forebody usually &lt;
<quantity id="4CC01E9DFFAFFFADFD5C764EFCBCF82F" box="[692,776,1906,1930]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="207" unit="mm" value="1.2">1.2 mm</quantity>
, only in one species from the Middle East up to 
<quantity id="4CC01E9DFFAFFFADFAE1764EFAEFF82F" box="[1289,1371,1906,1930]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="207" unit="mm" value="1.3">1.3 mm</quantity>
long. ................................................................................................................................................................................................. 6
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAFFFADFF2B76FDFA23F85F" blockId="6.[195,1431,1985,2041]" pageId="6" pageNumber="207">
<keyLead id="30C9104DFFAFFFADFF2B76FDFA23F85F" pageId="6" pageNumber="207">– Body partly paler; at least the humeral angles and the suture of the elytra, usually also the pronotum and parts of the abdomen, yellowish to reddish. ................................................................................................................................ 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF7771C5FADBFE46" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF7771C5FADBFED7" blockId="7.[159,1391,249,370]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF7771C5FADBFED7" pageId="7" pageNumber="208">
6. Scutellum coarsely rugosely sculptured. Anterior impressions of tergites III–V deep and with coarse punctures separated by longitudinal carinae. Tergite VIII without anterior median excision (
<figureCitation id="1303AFFDFFAEFFACFBF47025FBC2FE97" box="[1052,1142,281,305]" captionStart="Figs 233–236" captionStartId="21.[151,191,725,747]" captionTargetBox="[160,1374,246,704]" captionTargetId="figure-716@21.[151,1398,232,724]" captionTargetPageId="21" captionText="Figs 233–236: Anaulacaspsis inexpectata: male tergite VIII (233); posterior margin of male tergite VIII (234); male sternite VIII (235); female sternite VIII (236). Scale bars: 233, 235–236: 0.1 mm; 234: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5212626" httpUri="https://zenodo.org/record/5212626/files/figure.png" pageId="7" pageNumber="208">Fig. 233</figureCitation>
); posterior margin with a row of dense short modified setae (
<figureCitation id="1303AFFDFFAEFFACFDA97005FD2FFEF7" box="[577,667,313,337]" captionStart="Figs 233–236" captionStartId="21.[151,191,725,747]" captionTargetBox="[160,1374,246,704]" captionTargetId="figure-716@21.[151,1398,232,724]" captionTargetPageId="21" captionText="Figs 233–236: Anaulacaspsis inexpectata: male tergite VIII (233); posterior margin of male tergite VIII (234); male sternite VIII (235); female sternite VIII (236). Scale bars: 233, 235–236: 0.1 mm; 234: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5212626" httpUri="https://zenodo.org/record/5212626/files/figure.png" pageId="7" pageNumber="208">Fig. 234</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="1303AFFDFFAEFFACFC0E7005FBC2FEF7" box="[998,1142,313,337]" captionStart="Figs 211–232" captionStartId="20.[189,229,1941,1963]" captionTargetBox="[201,1414,238,1920]" captionTargetId="figure-41@20.[200,1426,232,1940]" captionText="Figs 211–232: Anaulacaspsis formosa (211), A. elegans (212–220), A. beesoni (221–228; 224–225: lectotype), and A. inexpectata (229–232; 231–232: holotype): habitus (211; photo by Marc Tronquet); male tergite VIII (212, 221); male sternite VIII (213, 222); median lobe of aedeagus in lateral and in ventral view (214–217, 224–227, 229–232); female tergite VIII (218); posterior margin of female tergite VIII (219); female sternite VIII (220, 228); postero-lateral portion of sternite VIII (223). Scale bars: 211: 1.0 mm; 212–218, 220–222, 224–232: 0.1 mm; 219, 223: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884779" httpUri="https://zenodo.org/record/5884779/files/figure.png" pageId="7" pageNumber="208">Figs 229–232</figureCitation>
. Spermatheca with long capsule, not comma-shaped. Distribution: 
<collectingCountry id="F32FF3E8FFAEFFACFD617066FD0CFED7" box="[649,696,346,369]" name="Iran" pageId="7" pageNumber="208">Iran</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFAEFFACFD2A7066FC8CFED4" box="[706,824,346,370]" name="Kyrgyzstan" pageId="7" pageNumber="208">Kyrgyzstan</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFCAE7065FC39FED7" box="[838,909,345,369]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="7" pageNumber="208">Map 5</figureCitation>
). ...................................................... 
<taxonomicName id="4C38C8FBFFAEFFACFB1B7065FADBFED7" ID-CoL="8VTS6" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[1267,1391,345,369]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="inexpectata">
<emphasis id="B94C6F6AFFAEFFACFB1B7065FADBFED7" bold="true" box="[1267,1391,345,369]" italics="true" pageId="7" pageNumber="208">inexpectata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF7770B5FADBFE46" blockId="7.[159,1392,392,480]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF7770B5FADBFE46" pageId="7" pageNumber="208">– Scutellum neither coarsely nor rugosely sculptured. Anterior impressions with fine punctation and without carinae. Tergite VIII with anterior median excision; posterior margin with unmodified setae. Spermatheca comma-shaped. ................................................................................................................................................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF7770C4FADBFD99" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF7770C4FADBFDB6" blockId="7.[159,1391,503,575]" box="[159,1391,503,528]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF7770C4FADBFDB6" box="[159,1391,503,528]" pageId="7" pageNumber="208">7. Posterior margin of elytra sharply yellow. ........................................................................................................................ 8</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF77731BFADBFD99" blockId="7.[159,1391,503,575]" box="[159,1391,551,575]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF77731BFADBFD99" box="[159,1391,551,575]" pageId="7" pageNumber="208">– Elytra either of uniformly dark coloration or with diffusely paler posterior margin. ............................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF77736AFADBFCBA" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF77736AFADBFD0B" blockId="7.[159,1391,598,686]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF77736AFADBFD0B" pageId="7" pageNumber="208">
8. Posterior portion of male head depressed or shallowly impressed. Male pronotum with extensive and rather deep impression. Median lobe of aedeagus larger, approximately 
<quantity id="4CC01E9DFFAEFFACFCC4734AFC3EFD2B" box="[812,906,630,654]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="7" pageNumber="208" unit="mm" value="0.38">0.38 mm</quantity>
long, and shaped as in 
<figureCitation id="1303AFFDFFAEFFACFB9E734AFAB1FD28" box="[1142,1285,630,654]" captionStart="Figs 136–163" captionStartId="17.[151,191,1971,1993]" captionTargetBox="[153,1394,237,1965]" captionTargetId="figure-144@17.[152,1399,232,1970]" captionText="Figs 136–163: Anaulacaspsis excisa (136–141), A. pamphylica (142–145), A. flavomarginata (146–150), A. laevigata (151–158; 153–154: lectotype; 155: Pelopónnisos; 156: Lesbos),and A. iberica (159–163): maletergite VIII(136, 142,146, 151); male sternite VIII (137, 143, 147, 152); median lobe of aedeagus in lateral and in ventral view (138–139, 144–145, 148–149, 153–156, 159–162); female sternite VIII (141, 150, 158);female tergite VIII (140); apex of ventral process of aedeagus in lateral view (157, 163). Scalebars:0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884773" httpUri="https://zenodo.org/record/5884773/files/figure.png" pageId="7" pageNumber="208">Figs 144–145</figureCitation>
. Distribution: South 
<collectingCountry id="F32FF3E8FFAEFFACFEA973AAFE38FD08" box="[321,396,662,686]" name="Turkey" pageId="7" pageNumber="208">Turkey</collectingCountry>
(
<collectingRegion id="49FC7D9AFFAEFFACFE7173AAFE44FD08" box="[409,496,662,686]" country="Turkey" name="Antalya" pageId="7" pageNumber="208">Antalya</collectingRegion>
) (
<figureCitation id="1303AFFDFFAEFFACFDEC73AAFDF8FD08" box="[516,588,662,686]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="7" pageNumber="208">Map 5</figureCitation>
). ........................................................................................................... 
<taxonomicName id="4C38C8FBFFAEFFACFB1A73A9FADBFD0B" ID-CoL="66MD5" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[1266,1391,661,685]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="pamphylica">
<emphasis id="B94C6F6AFFAEFFACFB1A73A9FADBFD0B" bold="true" box="[1266,1391,661,685]" italics="true" pageId="7" pageNumber="208">pamphylica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF7773F9FADBFCBA" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF7773F9FADBFCBA" pageId="7" pageNumber="208">
– Male head without appreciable depression or impression posteriorly. Male pronotum without distinct impression. Median lobe of aedeagus smaller, approximately 
<quantity id="4CC01E9DFFAEFFACFD1473D9FCEFFD5A" box="[764,859,741,765]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="208" unit="mm" value="0.3">0.30 mm</quantity>
long, and shaped as in 
<figureCitation id="1303AFFDFFAEFFACFBA573D9FB68FD5B" box="[1101,1244,741,765]" captionStart="Figs 136–163" captionStartId="17.[151,191,1971,1993]" captionTargetBox="[153,1394,237,1965]" captionTargetId="figure-144@17.[152,1399,232,1970]" captionText="Figs 136–163: Anaulacaspsis excisa (136–141), A. pamphylica (142–145), A. flavomarginata (146–150), A. laevigata (151–158; 153–154: lectotype; 155: Pelopónnisos; 156: Lesbos),and A. iberica (159–163): maletergite VIII(136, 142,146, 151); male sternite VIII (137, 143, 147, 152); median lobe of aedeagus in lateral and in ventral view (138–139, 144–145, 148–149, 153–156, 159–162); female sternite VIII (141, 150, 158);female tergite VIII (140); apex of ventral process of aedeagus in lateral view (157, 163). Scalebars:0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884773" httpUri="https://zenodo.org/record/5884773/files/figure.png" pageId="7" pageNumber="208">Figs 148–149</figureCitation>
. Distribution: 
<collectingCountry id="F32FF3E8FFAEFFACFF2C7239FEAEFCBB" box="[196,282,773,797]" name="Albania" pageId="7" pageNumber="208">Albania</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFAEFFACFECC7239FEDAFCBB" box="[292,366,773,797]" name="Greece" pageId="7" pageNumber="208">Greece</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFE947239FE77FCBB" box="[380,451,773,797]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="7" pageNumber="208">Map 5</figureCitation>
). .......................................................................................................................... 
<taxonomicName id="4C38C8FBFFAEFFACFB207238FADBFCBA" ID-CoL="DJJF" authorityName="Assing" authorityYear="2016" box="[1224,1391,772,796]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="flavomarginata">
<emphasis id="B94C6F6AFFAEFFACFB207238FADBFCBA" bold="true" box="[1224,1391,772,796]" italics="true" pageId="7" pageNumber="208">flavomarginata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF777208FADBFC5D" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF777208FADBFC2A" blockId="7.[159,1391,820,908]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF777208FADBFC2A" pageId="7" pageNumber="208">
9. Pronotum strongly transverse, approximately 1.2 times as broad as long. Forebody with fine punctation. Posterolateral portions of tergite and sternite VIII each with a cluster of conspicuously modified, apically dilated setae (
<figureCitation id="1303AFFDFFAEFFACFF247248FE93FC2A" box="[204,295,884,908]" captionStart="Figs 211–232" captionStartId="20.[189,229,1941,1963]" captionTargetBox="[201,1414,238,1920]" captionTargetId="figure-41@20.[200,1426,232,1940]" captionText="Figs 211–232: Anaulacaspsis formosa (211), A. elegans (212–220), A. beesoni (221–228; 224–225: lectotype), and A. inexpectata (229–232; 231–232: holotype): habitus (211; photo by Marc Tronquet); male tergite VIII (212, 221); male sternite VIII (213, 222); median lobe of aedeagus in lateral and in ventral view (214–217, 224–227, 229–232); female tergite VIII (218); posterior margin of female tergite VIII (219); female sternite VIII (220, 228); postero-lateral portion of sternite VIII (223). Scale bars: 211: 1.0 mm; 212–218, 220–222, 224–232: 0.1 mm; 219, 223: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884779" httpUri="https://zenodo.org/record/5884779/files/figure.png" pageId="7" pageNumber="208">Fig. 223</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="1303AFFDFFAEFFACFD947248FCB8FC2A" box="[636,780,884,908]" captionStart="Figs 211–232" captionStartId="20.[189,229,1941,1963]" captionTargetBox="[201,1414,238,1920]" captionTargetId="figure-41@20.[200,1426,232,1940]" captionText="Figs 211–232: Anaulacaspsis formosa (211), A. elegans (212–220), A. beesoni (221–228; 224–225: lectotype), and A. inexpectata (229–232; 231–232: holotype): habitus (211; photo by Marc Tronquet); male tergite VIII (212, 221); male sternite VIII (213, 222); median lobe of aedeagus in lateral and in ventral view (214–217, 224–227, 229–232); female tergite VIII (218); posterior margin of female tergite VIII (219); female sternite VIII (220, 228); postero-lateral portion of sternite VIII (223). Scale bars: 211: 1.0 mm; 212–218, 220–222, 224–232: 0.1 mm; 219, 223: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884779" httpUri="https://zenodo.org/record/5884779/files/figure.png" pageId="7" pageNumber="208">Figs 224–227</figureCitation>
. Distribution: North 
<collectingCountry id="F32FF3E8FFAEFFACFC047248FB90FC2A" box="[1004,1060,884,908]" name="India" pageId="7" pageNumber="208">India</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFBD97248FBCDFC2A" box="[1073,1145,884,908]" captionStart="Map 7" captionStartId="46.[189,234,2053,2074]" captionTargetBox="[189,1436,1241,2044]" captionTargetId="figure-539@46.[189,1436,1241,2044]" captionTargetPageId="46" captionText="Map 7: Distributions of Anaulacaspis beesoni (black circles), A. pectinata (white star), A. schuelkei (black triangle), A. beijingensis (white circles), and A. taiwanica (black star) in the East Palaearctic region, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212643" httpUri="https://zenodo.org/record/5212643/files/figure.png" pageId="7" pageNumber="208">Map 7</figureCitation>
). ...................... 
<taxonomicName id="4C38C8FBFFAEFFACFAC87248FADBFC2A" ID-CoL="6HNZP" baseAuthorityName="CAMERON" baseAuthorityYear="1939" box="[1312,1391,884,908]" class="Insecta" family="Staphylinidae" genus="Falagria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="beesoni">
<emphasis id="B94C6F6AFFAEFFACFAC87248FADBFC2A" bold="true" box="[1312,1391,884,908]" italics="true" pageId="7" pageNumber="208">beesoni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF777298FADBFC5D" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF777298FADBFC5D" pageId="7" pageNumber="208">
– Pronotum less transverse, usually 1.10–1.15 times as broad as long. Postero-lateral portions of sternite VIII without clusters of setae; postero-lateral portions of tergite VIII often with clusters of setae, but these setae long and thin, not dilated apically. Unknown from 
<collectingCountry id="F32FF3E8FFAEFFACFDAC72DFFDCBFC5D" box="[580,639,995,1019]" name="India" pageId="7" pageNumber="208">India</collectingCountry>
. .................................................................................................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF77752EFADBFB7F" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF77752EFADBFBCC" blockId="7.[159,1391,1042,1130]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF77752EFADBFBCC" pageId="7" pageNumber="208">
10. Head and pronotum with extremely fine punctation (slightly more distinct in male 
<taxonomicName id="4C38C8FBFFAEFFACFBF2752FFB36FB8C" ID-CoL="DJKB" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[1050,1154,1043,1066]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="nigrina">
<emphasis id="B94C6F6AFFAEFFACFBF2752FFB36FB8C" box="[1050,1154,1043,1066]" italics="true" pageId="7" pageNumber="208">A. nigrina</emphasis>
</taxonomicName>
). Tergite VIII posterolaterally with or without cluster of long thin setae on either side. Male pronotum with indistinct impression at most. .................................................................................................................................................................................... 11
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF7775BEFADBFB7F" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF7775BEFADBFB7F" pageId="7" pageNumber="208">– Head with fine, but distinct punctation. Tergite VIII postero-laterally always with cluster of long thin setae on either side. Male pronotum with dense and distinct punctation and often with median impression posteriorly. ....................................................................................................................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF7675CDFADBFA1E" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF7675CDFADBFAEF" blockId="7.[158,1391,1264,1353]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF7675CDFADBFAEF" pageId="7" pageNumber="208">11. Tergite VIII postero-laterally with a cluster of long thin setae on either side, anterior margin with narrower and deeper median excision. Male sternite VIII strongly produced posteriorly. Posterior margin of female sternite VIII convex, not concave in the middle. ................................................................................................................................. 12</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF76745CFADBFA1E" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF76745CFADBFA1E" pageId="7" pageNumber="208">– Tergite VIII postero-laterally without a cluster of long thin setae on either side, anterior margin with broader and shallower median excision. Male sternite VIII less produced posteriorly, broadly convex to obtusely pointed. Posterior margin of female sternite VIII concave in the middle. ............................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF7674F3FADAF930" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF7674F3FADBF981" blockId="7.[158,1391,1487,1575]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF7674F3FADBF981" pageId="7" pageNumber="208">
12. Posterior margin of tergite VIII truncate (
<figureCitation id="1303AFFDFFAEFFACFD9E74F3FD77FA41" box="[630,707,1487,1511]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="7" pageNumber="208">Fig. 52</figureCitation>
). Male sternite VIII more strongly produced posteriorly (
<figureCitation id="1303AFFDFFAEFFACFAFE74F3FAD7FA41" box="[1302,1379,1487,1511]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="7" pageNumber="208">Fig. 53</figureCitation>
). Median lobe of aedeagus 
<quantity id="4CC01E9DFFAEFFACFE3974D3FDDEF9A0" box="[465,618,1519,1543]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="7" pageNumber="208" unit="mm" value="0.22" valueMax="0.23" valueMin="0.21">0.21–0.23 mm</quantity>
long and shaped as in 
<figureCitation id="1303AFFDFFAEFFACFCB374D3FC64F9A1" box="[859,976,1519,1543]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="7" pageNumber="208">Figs 54–55</figureCitation>
. Distribution: West 
<collectingCountry id="F32FF3E8FFAEFFACFB4D74D3FB66F9A0" box="[1189,1234,1519,1542]" name="Iran" pageId="7" pageNumber="208">Iran</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFB3774D3FA92F9A1" box="[1247,1318,1519,1543]" captionStart="Map 2" captionStartId="27.[151,196,2023,2044]" captionTargetBox="[151,1398,1241,2014]" captionTargetId="figure-455@27.[151,1398,1241,2014]" captionTargetPageId="27" captionText="Map 2: Distributions of Anaulacaspis nigra (black circles), A. pseudonigra (white circle), A. reticulata (white triangle), A. libanotica (black triangles), A. caucasica (white diamonds), A. truncata (black diamonds), and A. sinuata (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212628" httpUri="https://zenodo.org/record/5212628/files/figure.png" pageId="7" pageNumber="208">Map 2</figureCitation>
). ........ ..................................................................................................................................................................................... 
<taxonomicName id="4C38C8FBFFAEFFACFAF77733FADBF981" ID-CoL="DJKW" authorityName="Assing" authorityYear="2016" box="[1311,1391,1551,1575]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="sinuata">
<emphasis id="B94C6F6AFFAEFFACFAF77733FADBF981" bold="true" box="[1311,1391,1551,1575]" italics="true" pageId="7" pageNumber="208">sinuata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF767702FADAF930" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF767702FADAF930" pageId="7" pageNumber="208">
– Posterior margin of tergite VIII concave (
<figureCitation id="1303AFFDFFAEFFACFD917702FD71F9F0" box="[633,709,1598,1622]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="7" pageNumber="208">Fig. 57</figureCitation>
). Male sternite VIII obtusely pointed posteriorly. Median lobe of aedeagus larger, 
<quantity id="4CC01E9DFFAEFFACFE9B7762FDB8F9D3" box="[371,524,1630,1654]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="7" pageNumber="208" unit="mm" value="0.285" valueMax="0.3" valueMin="0.27">0.27–0.30 mm</quantity>
long, and shaped as in 
<figureCitation id="1303AFFDFFAEFFACFCEA7762FCC3F9D0" box="[770,887,1630,1654]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="7" pageNumber="208">Figs 59–61</figureCitation>
. Distribution: Middle East from Northeast 
<collectingCountry id="F32FF3E8FFAEFFACFAAA7762FADBF9D3" box="[1346,1391,1630,1653]" name="Iran" pageId="7" pageNumber="208">Iran</collectingCountry>
eastwards to 
<collectingCountry id="F32FF3E8FFAEFFACFEA67742FE70F930" box="[334,452,1662,1686]" name="Kyrgyzstan" pageId="7" pageNumber="208">Kyrgyzstan</collectingCountry>
and 
<collectingCountry id="F32FF3E8FFAEFFACFE107742FDCEF930" box="[504,634,1662,1686]" name="Afghanistan" pageId="7" pageNumber="208">Afghanistan</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFD607742FD7AF930" box="[648,718,1662,1686]" captionStart="Map 3" captionStartId="28.[189,234,2028,2049]" captionTargetBox="[189,1436,1371,2017]" captionTargetId="figure-589@28.[189,1436,1371,2017]" captionTargetPageId="28" captionText="Map 3: Distributions of Anaulacaspis formosa (black circles), A. elegans (white circles), A. nigrina (black diamonds), A. eminens (white diamonds), and A. nigerrima (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212634" httpUri="https://zenodo.org/record/5212634/files/figure.png" pageId="7" pageNumber="208">Map 3</figureCitation>
). ........................................................................................ 
<taxonomicName id="4C38C8FBFFAEFFACFAEB7742FADAF930" ID-CoL="3DSZM" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[1283,1390,1662,1686]" class="Insecta" family="Staphylinidae" genus="Falagria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="B94C6F6AFFAEFFACFAEB7742FADAF930" bold="true" box="[1283,1390,1662,1686]" italics="true" pageId="7" pageNumber="208">nigerrima</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF767791FADBF892" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF767791FADAF8A3" blockId="7.[158,1391,1709,1797]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF767791FADAF8A3" pageId="7" pageNumber="208">
13. Male head with or without indistinct depression in postero-median portion. Male pronotum shallowly and usually extensively depressed or impressed in the middle). Posterior margin of tergite VIII shallowly concave (
<figureCitation id="1303AFFDFFAEFFACFAE177F1FAD7F943" box="[1289,1379,1741,1765]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="7" pageNumber="208">Fig. 167</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="1303AFFDFFAEFFACFDE177D1FD23F8A3" box="[521,663,1773,1797]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="7" pageNumber="208">Figs 169–171</figureCitation>
. Distribution: Greek island Lesbos, 
<collectingCountry id="F32FF3E8FFAEFFACFBFA77D1FBE9F8A3" box="[1042,1117,1773,1797]" name="Turkey" pageId="7" pageNumber="208">Turkey</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFAEFFACFB8F77D1FB20F8A2" box="[1127,1172,1773,1796]" name="Iran" pageId="7" pageNumber="208">Iran</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFB4977D1FB53F8A3" box="[1185,1255,1773,1797]" captionStart="Map 3" captionStartId="28.[189,234,2028,2049]" captionTargetBox="[189,1436,1371,2017]" captionTargetId="figure-589@28.[189,1436,1371,2017]" captionTargetPageId="28" captionText="Map 3: Distributions of Anaulacaspis formosa (black circles), A. elegans (white circles), A. nigrina (black diamonds), A. eminens (white diamonds), and A. nigerrima (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212634" httpUri="https://zenodo.org/record/5212634/files/figure.png" pageId="7" pageNumber="208">Map 3</figureCitation>
). .... 
<taxonomicName id="4C38C8FBFFAEFFACFAF777D1FADAF8A3" ID-CoL="DJKB" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[1311,1390,1773,1797]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="nigrina">
<emphasis id="B94C6F6AFFAEFFACFAF777D1FADAF8A3" bold="true" box="[1311,1390,1773,1797]" italics="true" pageId="7" pageNumber="208">nigrina</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF767621FADBF892" box="[158,1391,1820,1844]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF767621FADBF892" box="[158,1391,1820,1844]" pageId="7" pageNumber="208">– Head and pronotum without sexual dimorphism. Posterior margin of tergite VIII truncate to weakly convex. ... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFAEFFACFF767677FADEF7B4" pageId="7" pageNumber="208">
<paragraph id="8B87B378FFAEFFACFF767677FADBF805" blockId="7.[158,1391,1867,1955]" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF767677FADBF805" pageId="7" pageNumber="208">
14. Median lobe of aedeagus (
<figureCitation id="1303AFFDFFAEFFACFE307677FDDDF8C5" box="[472,617,1867,1891]" captionStart="Figs 136–163" captionStartId="17.[151,191,1971,1993]" captionTargetBox="[153,1394,237,1965]" captionTargetId="figure-144@17.[152,1399,232,1970]" captionText="Figs 136–163: Anaulacaspsis excisa (136–141), A. pamphylica (142–145), A. flavomarginata (146–150), A. laevigata (151–158; 153–154: lectotype; 155: Pelopónnisos; 156: Lesbos),and A. iberica (159–163): maletergite VIII(136, 142,146, 151); male sternite VIII (137, 143, 147, 152); median lobe of aedeagus in lateral and in ventral view (138–139, 144–145, 148–149, 153–156, 159–162); female sternite VIII (141, 150, 158);female tergite VIII (140); apex of ventral process of aedeagus in lateral view (157, 163). Scalebars:0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884773" httpUri="https://zenodo.org/record/5884773/files/figure.png" pageId="7" pageNumber="208">Figs 159–163</figureCitation>
) slightly smaller; ventral process longer in relation basal capsule, apically more slender in lateral view; crista apicalis more prominent. Posterior margin of female sternite VIII more distinctly concave in the middle (
<figureCitation id="1303AFFDFFAEFFACFE5476B7FDA1F805" box="[444,533,1931,1955]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="7" pageNumber="208">Fig. 166</figureCitation>
). Distribution: 
<collectingCountry id="F32FF3E8FFAEFFACFD5F76B7FD47F805" box="[695,755,1931,1955]" name="Spain" pageId="7" pageNumber="208">Spain</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFCE876B7FCFDF805" box="[768,841,1931,1955]" captionStart="Map 4" captionStartId="33.[151,196,2022,2043]" captionTargetBox="[151,1398,1203,2012]" captionTargetId="figure-561@33.[151,1398,1203,2012]" captionTargetPageId="33" captionText="Map 4: Distributions of Anaulacaspis desertorum (black circles), A. naevula (white circles), A. iberica (white stars), A. laevigata (white diamonds), A. gratilla (black diamonds), and A. excisa (black stars) in the West Palaearctic and Middle Asia, based on revised records. The question mark indicates the records without specified locality of A. gratilla from Iraq." figureDoi="http://doi.org/10.5281/zenodo.5212637" httpUri="https://zenodo.org/record/5212637/files/figure.png" pageId="7" pageNumber="208">Map 4</figureCitation>
). ......................................................................... 
<taxonomicName id="4C38C8FBFFAEFFACFACD76B7FADBF805" ID-CoL="DJJQ" authorityName="Assing" authorityYear="2016" box="[1317,1391,1931,1955]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="iberica">
<emphasis id="B94C6F6AFFAEFFACFACD76B7FADBF805" bold="true" box="[1317,1391,1931,1955]" italics="true" pageId="7" pageNumber="208">iberica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFAEFFACFF767687FADEF7B4" pageId="7" pageNumber="208">
<keyLead id="30C9104DFFAEFFACFF767687FADEF7B4" pageId="7" pageNumber="208">
<heading id="D0CF0414FFAEFFACFF767687FF18F874" box="[158,172,1979,2002]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="208" reason="4">–</heading>
Median lobe of aedeagus (
<figureCitation id="1303AFFDFFAEFFACFE3B7686FDD7F874" box="[467,611,1978,2002]" captionStart="Figs 136–163" captionStartId="17.[151,191,1971,1993]" captionTargetBox="[153,1394,237,1965]" captionTargetId="figure-144@17.[152,1399,232,1970]" captionText="Figs 136–163: Anaulacaspsis excisa (136–141), A. pamphylica (142–145), A. flavomarginata (146–150), A. laevigata (151–158; 153–154: lectotype; 155: Pelopónnisos; 156: Lesbos),and A. iberica (159–163): maletergite VIII(136, 142,146, 151); male sternite VIII (137, 143, 147, 152); median lobe of aedeagus in lateral and in ventral view (138–139, 144–145, 148–149, 153–156, 159–162); female sternite VIII (141, 150, 158);female tergite VIII (140); apex of ventral process of aedeagus in lateral view (157, 163). Scalebars:0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884773" httpUri="https://zenodo.org/record/5884773/files/figure.png" pageId="7" pageNumber="208">Figs 153–157</figureCitation>
) slightly larger; ventral process shorter in relation to basal capsule, apically less slender in lateral view; crista apicalis less prominent. Posterior margin of female sternite VIII very shallowly concave in the middle (
<figureCitation id="1303AFFDFFAEFFACFE5476C6FDA1F7B4" box="[444,533,2042,2066]" captionStart="Figs 136–163" captionStartId="17.[151,191,1971,1993]" captionTargetBox="[153,1394,237,1965]" captionTargetId="figure-144@17.[152,1399,232,1970]" captionText="Figs 136–163: Anaulacaspsis excisa (136–141), A. pamphylica (142–145), A. flavomarginata (146–150), A. laevigata (151–158; 153–154: lectotype; 155: Pelopónnisos; 156: Lesbos),and A. iberica (159–163): maletergite VIII(136, 142,146, 151); male sternite VIII (137, 143, 147, 152); median lobe of aedeagus in lateral and in ventral view (138–139, 144–145, 148–149, 153–156, 159–162); female sternite VIII (141, 150, 158);female tergite VIII (140); apex of ventral process of aedeagus in lateral view (157, 163). Scalebars:0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884773" httpUri="https://zenodo.org/record/5884773/files/figure.png" pageId="7" pageNumber="208">Fig. 158</figureCitation>
). Distribution Ponto-Mediterranean: 
<collectingCountry id="F32FF3E8FFAEFFACFC4C76C6FC5BF7B4" box="[932,1007,2042,2066]" name="Turkey" pageId="7" pageNumber="208">Turkey</collectingCountry>
, Balkans, 
<collectingCountry id="F32FF3E8FFAEFFACFBB176C6FB3CF7B4" box="[1113,1160,2042,2066]" name="Italy" pageId="7" pageNumber="208">Italy</collectingCountry>
(
<figureCitation id="1303AFFDFFAEFFACFB7D76C6FB69F7B4" box="[1173,1245,2042,2066]" captionStart="Map 4" captionStartId="33.[151,196,2022,2043]" captionTargetBox="[151,1398,1203,2012]" captionTargetId="figure-561@33.[151,1398,1203,2012]" captionTargetPageId="33" captionText="Map 4: Distributions of Anaulacaspis desertorum (black circles), A. naevula (white circles), A. iberica (white stars), A. laevigata (white diamonds), A. gratilla (black diamonds), and A. excisa (black stars) in the West Palaearctic and Middle Asia, based on revised records. The question mark indicates the records without specified locality of A. gratilla from Iraq." figureDoi="http://doi.org/10.5281/zenodo.5212637" httpUri="https://zenodo.org/record/5212637/files/figure.png" pageId="7" pageNumber="208">Map 4</figureCitation>
). ... laevigata
</keyLead>
</paragraph>
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15. Ventral process of aedeagus in ventral view with a pronounced projection on either side (
<figureCitation id="1303AFFDFFA1FFA3FB7271C5FA8DFEB7" box="[1178,1337,249,273]" captionStart="Figs 31–48" captionStartId="12.[189,229,2000,2022]" captionTargetBox="[193,1432,245,1963]" captionTargetId="figure-41@12.[189,1436,232,1971]" captionText="Figs 31–48: Anaulacaspsis caucasica (31–34), A. pseudonigra (35–41), A. reticulata (42–46), and A. libanotica (47–48): male tergite VIII (31, 35, 43); male sternite VIII (32, 36, 44); median lobe of aedeagus in lateral and in ventral view (33–34, 37–39, 45–48); apical portion of median lobe in ventral view (40); female sternite VIII (41); sutural portion of elytra (42). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884769" httpUri="https://zenodo.org/record/5884769/files/figure.png" pageId="8" pageNumber="209">Figs 34, 39–40</figureCitation>
). ...... 16
</keyLead>
</paragraph>
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<heading id="D0CF0414FFA1FFA3FF2D7015FF67FEE6" box="[197,211,297,320]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="209" reason="4">–</heading>
Ventral process of aedeagus in ventral view sometimes angled laterally, but without pronounced projection. ............................................................................................................................................................................... 17
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16. Median lobe of aedeagus 
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long and shaped as in 
<figureCitation id="1303AFFDFFA1FFA3FC65704BFBB7FE36" box="[909,1027,375,400]" captionStart="Figs 31–48" captionStartId="12.[189,229,2000,2022]" captionTargetBox="[193,1432,245,1963]" captionTargetId="figure-41@12.[189,1436,232,1971]" captionText="Figs 31–48: Anaulacaspsis caucasica (31–34), A. pseudonigra (35–41), A. reticulata (42–46), and A. libanotica (47–48): male tergite VIII (31, 35, 43); male sternite VIII (32, 36, 44); median lobe of aedeagus in lateral and in ventral view (33–34, 37–39, 45–48); apical portion of median lobe in ventral view (40); female sternite VIII (41); sutural portion of elytra (42). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884769" httpUri="https://zenodo.org/record/5884769/files/figure.png" pageId="8" pageNumber="209">Figs 37–40</figureCitation>
. Distribution: North 
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: 
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(
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). ............................................................................................................................................................ 
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<emphasis id="B94C6F6AFFA1FFA3FAF870ABFA20FE09" bold="true" box="[1296,1428,407,431]" italics="true" pageId="8" pageNumber="209">pseudonigra</emphasis>
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</paragraph>
<paragraph id="8B87B378FFA1FFA3FF2C70FBFA21FE59" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
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- Median lobe of aedeagus larger, approximately 
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long, and shaped as in 
<figureCitation id="1303AFFDFFA1FFA3FBFD70FBFB3DFE79" box="[1045,1161,455,479]" captionStart="Figs 31–48" captionStartId="12.[189,229,2000,2022]" captionTargetBox="[193,1432,245,1963]" captionTargetId="figure-41@12.[189,1436,232,1971]" captionText="Figs 31–48: Anaulacaspsis caucasica (31–34), A. pseudonigra (35–41), A. reticulata (42–46), and A. libanotica (47–48): male tergite VIII (31, 35, 43); male sternite VIII (32, 36, 44); median lobe of aedeagus in lateral and in ventral view (33–34, 37–39, 45–48); apical portion of median lobe in ventral view (40); female sternite VIII (41); sutural portion of elytra (42). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884769" httpUri="https://zenodo.org/record/5884769/files/figure.png" pageId="8" pageNumber="209">Figs 33–34</figureCitation>
. Distribution: 
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, North 
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(
<figureCitation id="1303AFFDFFA1FFA3FE8370DBFE07FE59" box="[363,435,487,511]" captionStart="Map 2" captionStartId="27.[151,196,2023,2044]" captionTargetBox="[151,1398,1241,2014]" captionTargetId="figure-455@27.[151,1398,1241,2014]" captionTargetPageId="27" captionText="Map 2: Distributions of Anaulacaspis nigra (black circles), A. pseudonigra (white circle), A. reticulata (white triangle), A. libanotica (black triangles), A. caucasica (white diamonds), A. truncata (black diamonds), and A. sinuata (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212628" httpUri="https://zenodo.org/record/5212628/files/figure.png" pageId="8" pageNumber="209">Map 2</figureCitation>
). ............................................................................................................................................. 
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<emphasis id="B94C6F6AFFA1FFA3FAC670DBFA21FE59" bold="true" box="[1326,1429,487,511]" italics="true" pageId="8" pageNumber="209">caucasica</emphasis>
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17. Distribution: Middle East (
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, 
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) (
<figureCitation id="1303AFFDFFA1FFA3FD53732AFCB6FD88" box="[699,770,534,558]" captionStart="Map 2" captionStartId="27.[151,196,2023,2044]" captionTargetBox="[151,1398,1241,2014]" captionTargetId="figure-455@27.[151,1398,1241,2014]" captionTargetPageId="27" captionText="Map 2: Distributions of Anaulacaspis nigra (black circles), A. pseudonigra (white circle), A. reticulata (white triangle), A. libanotica (black triangles), A. caucasica (white diamonds), A. truncata (black diamonds), and A. sinuata (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212628" httpUri="https://zenodo.org/record/5212628/files/figure.png" pageId="8" pageNumber="209">Map 2</figureCitation>
). Median lobe of aedeagus larger, approximately 
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long, and shaped as in 
<figureCitation id="1303AFFDFFA1FFA3FE48730AFDA2FDE8" box="[416,534,566,590]" captionStart="Figs 31–48" captionStartId="12.[189,229,2000,2022]" captionTargetBox="[193,1432,245,1963]" captionTargetId="figure-41@12.[189,1436,232,1971]" captionText="Figs 31–48: Anaulacaspsis caucasica (31–34), A. pseudonigra (35–41), A. reticulata (42–46), and A. libanotica (47–48): male tergite VIII (31, 35, 43); male sternite VIII (32, 36, 44); median lobe of aedeagus in lateral and in ventral view (33–34, 37–39, 45–48); apical portion of median lobe in ventral view (40); female sternite VIII (41); sutural portion of elytra (42). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884769" httpUri="https://zenodo.org/record/5884769/files/figure.png" pageId="8" pageNumber="209">Figs 47–48</figureCitation>
. .............................................................................................................................. 
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<emphasis id="B94C6F6AFFA1FFA3FAC17309FA21FDEB" bold="true" box="[1321,1429,565,589]" italics="true" pageId="8" pageNumber="209">libanotica</emphasis>
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<keyLead id="30C9104DFFA1FFA3FF2C7359FA21FDDB" box="[196,1429,613,637]" pageId="8" pageNumber="209">– Absent from the Middle East. Median lobe of aedeagus smaller. ............................................................................... 18</keyLead>
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</keyStep>
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18. Antennomere IV distinctly transverse. Pronotum, elytra, and abdomen with distinct microreticulation (
<figureCitation id="1303AFFDFFA1FFA3FAD473A8FA3EFD0A" box="[1340,1418,660,684]" captionStart="Figs 31–48" captionStartId="12.[189,229,2000,2022]" captionTargetBox="[193,1432,245,1963]" captionTargetId="figure-41@12.[189,1436,232,1971]" captionText="Figs 31–48: Anaulacaspsis caucasica (31–34), A. pseudonigra (35–41), A. reticulata (42–46), and A. libanotica (47–48): male tergite VIII (31, 35, 43); male sternite VIII (32, 36, 44); median lobe of aedeagus in lateral and in ventral view (33–34, 37–39, 45–48); apical portion of median lobe in ventral view (40); female sternite VIII (41); sutural portion of elytra (42). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884769" httpUri="https://zenodo.org/record/5884769/files/figure.png" pageId="8" pageNumber="209">Fig. 42</figureCitation>
). Posterior margin of tergite VIII convex (
<figureCitation id="1303AFFDFFA1FFA3FD6C7388FD7BFD6A" box="[644,719,692,716]" captionStart="Figs 31–48" captionStartId="12.[189,229,2000,2022]" captionTargetBox="[193,1432,245,1963]" captionTargetId="figure-41@12.[189,1436,232,1971]" captionText="Figs 31–48: Anaulacaspsis caucasica (31–34), A. pseudonigra (35–41), A. reticulata (42–46), and A. libanotica (47–48): male tergite VIII (31, 35, 43); male sternite VIII (32, 36, 44); median lobe of aedeagus in lateral and in ventral view (33–34, 37–39, 45–48); apical portion of median lobe in ventral view (40); female sternite VIII (41); sutural portion of elytra (42). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884769" httpUri="https://zenodo.org/record/5884769/files/figure.png" pageId="8" pageNumber="209">Fig. 43</figureCitation>
). Median lobe of aedeagus 
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long and shaped as in 
<figureCitation id="1303AFFDFFA1FFA3FAF67388FA25FD6A" box="[1310,1425,692,716]" captionStart="Figs 31–48" captionStartId="12.[189,229,2000,2022]" captionTargetBox="[193,1432,245,1963]" captionTargetId="figure-41@12.[189,1436,232,1971]" captionText="Figs 31–48: Anaulacaspsis caucasica (31–34), A. pseudonigra (35–41), A. reticulata (42–46), and A. libanotica (47–48): male tergite VIII (31, 35, 43); male sternite VIII (32, 36, 44); median lobe of aedeagus in lateral and in ventral view (33–34, 37–39, 45–48); apical portion of median lobe in ventral view (40); female sternite VIII (41); sutural portion of elytra (42). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884769" httpUri="https://zenodo.org/record/5884769/files/figure.png" pageId="8" pageNumber="209">Figs 45–46</figureCitation>
; ventral process broad in ventral view and without membranous projection at base (lateral view). Distribution: 
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(
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) (
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). ................................................................................................................................... 
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<emphasis id="B94C6F6AFFA1FFA3FAC573C8FA21FCAA" bold="true" box="[1325,1429,756,780]" italics="true" pageId="8" pageNumber="209">reticulata</emphasis>
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<paragraph id="8B87B378FFA1FFA3FF2C7218FA20FC1D" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C7218FA20FC1D" pageId="8" pageNumber="209">
<heading id="D0CF0414FFA1FFA3FF2C7218FF66FC9D" box="[196,210,804,827]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="209" reason="4">–</heading>
Antennomere IV approximately as long as broad. Pronotum and elytra without appreciable microreticulation. Posterior margin of tergite VIII weakly to strongly concave (
<figureCitation id="1303AFFDFFA1FFA3FC69727FFC57FCFD" box="[897,995,835,859]" captionStart="Figs 1–12" captionStartId="10.[189,229,2029,2051]" captionTargetBox="[202,1418,263,1965]" captionTargetId="figure-41@10.[189,1436,251,1990]" captionTargetPageId="10" captionText="Figs 1–12: Anaulacaspsis nigra. Maxilla (1); labium (2); labrum (3); male tergite VIII (4–9); male sternite VIII (10–12). Scale bars: 1–2, 4–12: 0.1 mm; 3: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5212610" httpUri="https://zenodo.org/record/5212610/files/figure.png" pageId="8" pageNumber="209">Figs 4–9</figureCitation>
). Median lobe of aedeagus larger, 
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long; ventral process of different shape, at base usually with, rarely without, membranous projection in lateral view (
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). Widespread from northeastern 
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and 
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eastwards to 
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(
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). ................................................................................................................................................................................ 
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<emphasis id="B94C6F6AFFA1FFA3FAB2729FFA20FC1D" bold="true" box="[1370,1428,931,955]" italics="true" pageId="8" pageNumber="209">nigra</emphasis>
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19. Tergite VIII postero-laterally with a cluster of long thin setae on either side. Small species; length of forebody 
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at most. Posterior margin of tergite VIII with conspicuous median projection or strongly convex, except in 
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<emphasis id="B94C6F6AFFA1FFA3FEEE752FFEAAFB8C" box="[262,286,1043,1066]" italics="true" pageId="8" pageNumber="209">A.</emphasis>
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. ....................................................................................................................................................................... 20
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA1FFA3FF2C757EFA21FBDF" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C757EFA21FBDF" pageId="8" pageNumber="209">
<heading id="D0CF0414FFA1FFA3FF2C757EFF66FBFF" box="[196,210,1090,1113]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="209" reason="4">–</heading>
Tergite VIII postero-laterally without cluster of long thin setae on either side. Mostly larger species. Posterior margin of tergite VIII always without median process. ............................................................................................... 24
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA1FFA3FF2C75ADFA21FB5E" pageId="8" pageNumber="209">
<paragraph id="8B87B378FFA1FFA3FF2C75ADFA21FB0F" blockId="8.[196,1429,249,2048]" box="[196,1429,1168,1193]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C75ADFA21FB0F" box="[196,1429,1168,1193]" pageId="8" pageNumber="209">20. Posterior margin of tergite VIII truncate to convex, without a conspicuous median process. ............................... 21</keyLead>
</paragraph>
<paragraph id="8B87B378FFA1FFA3FF2C75FCFA21FB5E" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C75FCFA21FB5E" pageId="8" pageNumber="209">
<heading id="D0CF0414FFA1FFA3FF2C75FCFF66FB71" box="[196,210,1216,1239]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="209" reason="4">–</heading>
Posterior margin of tergite VIII with a conspicuous median process (
<figureCitation id="1303AFFDFFA1FFA3FC5F75FCFB9AFB71" box="[951,1070,1215,1240]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="8" pageNumber="209">Figs 62–63</figureCitation>
, 
<figureCitation id="1303AFFDFFA1FFA3FBD075FCFB72FB7E" box="[1080,1222,1215,1240]" captionStart="Figs 70–90" captionStartId="14.[189,229,2000,2022]" captionTargetBox="[196,1434,240,1957]" captionTargetId="figure-41@14.[189,1436,232,1971]" captionTargetPageId="14" captionText="Figs 70–90: Anaulacaspsis truncata (70–75), A. convexa (76–83), and A. eminens (84–90): male tergite VIII (70, 76, 84); male sternite VIII (71, 77, 85); median lobe of aedeagus in lateral and in ventral view (72–73, 78–80, 86–88); female tergite VIII (74, 82, 89); female sternite VIII (75, 83, 90); apical portion of aedeagus in ventral view (81). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212618" httpUri="https://zenodo.org/record/5212618/files/figure.png" pageId="8" pageNumber="209">70, 74, 76, 82</figureCitation>
). Male sternite VIII more strongly produced posteriorly. .............................................................................................................................. 22
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA1FFA3FF2C7433FA21FA70" pageId="8" pageNumber="209">
<paragraph id="8B87B378FFA1FFA3FF2C7433FA21FAC0" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C7433FA21FAC0" pageId="8" pageNumber="209">
21. Posterior margin of tergite VIII truncate to weakly convex (
<figureCitation id="1303AFFDFFA1FFA3FC977433FC6DFA81" box="[895,985,1295,1319]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="8" pageNumber="209">Fig. 121</figureCitation>
). Median lobe of aedeagus shaped as in 
<figureCitation id="1303AFFDFFA1FFA3FF027413FE54FAE1" box="[234,480,1327,1351]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="8" pageNumber="209">Figs 123–126, 130–131</figureCitation>
. Posterior margin of female sternite VIII weakly concave in the middle (
<figureCitation id="1303AFFDFFA1FFA3FB0D7413FA8BFAE1" box="[1253,1343,1327,1351]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="8" pageNumber="209">Fig. 127</figureCitation>
). Widespread from 
<collectingCountry id="F32FF3E8FFA1FFA3FE997473FE7CFAC1" box="[369,456,1359,1383]" name="Ukraine" pageId="8" pageNumber="209">Ukraine</collectingCountry>
in the west to Middle Asia in the east (
<figureCitation id="1303AFFDFFA1FFA3FC8C7473FC19FAC1" box="[868,941,1359,1383]" captionStart="Map 4" captionStartId="33.[151,196,2022,2043]" captionTargetBox="[151,1398,1203,2012]" captionTargetId="figure-561@33.[151,1398,1203,2012]" captionTargetPageId="33" captionText="Map 4: Distributions of Anaulacaspis desertorum (black circles), A. naevula (white circles), A. iberica (white stars), A. laevigata (white diamonds), A. gratilla (black diamonds), and A. excisa (black stars) in the West Palaearctic and Middle Asia, based on revised records. The question mark indicates the records without specified locality of A. gratilla from Iraq." figureDoi="http://doi.org/10.5281/zenodo.5212637" httpUri="https://zenodo.org/record/5212637/files/figure.png" pageId="8" pageNumber="209">Map 4</figureCitation>
). .............................................................. 
<taxonomicName id="4C38C8FBFFA1FFA3FAAE7472FA21FAC0" ID-CoL="DJJN" baseAuthorityName="Erichson" baseAuthorityYear="1839" box="[1350,1429,1358,1382]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="gratilla">
<emphasis id="B94C6F6AFFA1FFA3FAAE7472FA21FAC0" bold="true" box="[1350,1429,1358,1382]" italics="true" pageId="8" pageNumber="209">gratilla</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA1FFA3FF2C7442FA21FA70" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C7442FA21FA70" pageId="8" pageNumber="209">
– Posterior margin of tergite VIII strongly convex (
<figureCitation id="1303AFFDFFA1FFA3FD1E7442FCD9FA30" box="[758,877,1406,1430]" captionStart="Figs 70–90" captionStartId="14.[189,229,2000,2022]" captionTargetBox="[196,1434,240,1957]" captionTargetId="figure-41@14.[189,1436,232,1971]" captionTargetPageId="14" captionText="Figs 70–90: Anaulacaspsis truncata (70–75), A. convexa (76–83), and A. eminens (84–90): male tergite VIII (70, 76, 84); male sternite VIII (71, 77, 85); median lobe of aedeagus in lateral and in ventral view (72–73, 78–80, 86–88); female tergite VIII (74, 82, 89); female sternite VIII (75, 83, 90); apical portion of aedeagus in ventral view (81). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212618" httpUri="https://zenodo.org/record/5212618/files/figure.png" pageId="8" pageNumber="209">Figs 84, 89</figureCitation>
). Median lobe of aedeagus with strongly prominent crista apicalis and with ventral process of completely different shape (
<figureCitation id="1303AFFDFFA1FFA3FC0B74A2FBE8FA10" box="[995,1116,1438,1462]" captionStart="Figs 70–90" captionStartId="14.[189,229,2000,2022]" captionTargetBox="[196,1434,240,1957]" captionTargetId="figure-41@14.[189,1436,232,1971]" captionTargetPageId="14" captionText="Figs 70–90: Anaulacaspsis truncata (70–75), A. convexa (76–83), and A. eminens (84–90): male tergite VIII (70, 76, 84); male sternite VIII (71, 77, 85); median lobe of aedeagus in lateral and in ventral view (72–73, 78–80, 86–88); female tergite VIII (74, 82, 89); female sternite VIII (75, 83, 90); apical portion of aedeagus in ventral view (81). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212618" httpUri="https://zenodo.org/record/5212618/files/figure.png" pageId="8" pageNumber="209">Figs 86–88</figureCitation>
). Posterior margin of female sternite VIII convex, without median concavity. Distribution: South 
<collectingCountry id="F32FF3E8FFA1FFA3FC577482FC5AFA73" box="[959,1006,1470,1493]" name="Iran" pageId="8" pageNumber="209">Iran</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA1FFA3FC1F7482FBCDFA70" box="[1015,1145,1470,1494]" name="Afghanistan" pageId="8" pageNumber="209">Afghanistan</collectingCountry>
(
<figureCitation id="1303AFFDFFA1FFA3FB6F7482FB79FA70" box="[1159,1229,1470,1494]" captionStart="Map 3" captionStartId="28.[189,234,2028,2049]" captionTargetBox="[189,1436,1371,2017]" captionTargetId="figure-589@28.[189,1436,1371,2017]" captionTargetPageId="28" captionText="Map 3: Distributions of Anaulacaspis formosa (black circles), A. elegans (white circles), A. nigrina (black diamonds), A. eminens (white diamonds), and A. nigerrima (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212634" httpUri="https://zenodo.org/record/5212634/files/figure.png" pageId="8" pageNumber="209">Map 3</figureCitation>
). ............. 
<taxonomicName id="4C38C8FBFFA1FFA3FAD47482FA21FA70" ID-CoL="DJJD" authorityName="Assing" authorityYear="2016" box="[1340,1429,1470,1494]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="eminens">
<emphasis id="B94C6F6AFFA1FFA3FAD47482FA21FA70" bold="true" box="[1340,1429,1470,1494]" italics="true" pageId="8" pageNumber="209">eminens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA1FFA3FF2C74D1FA20F9D2" pageId="8" pageNumber="209">
<paragraph id="8B87B378FFA1FFA3FF2C74D1FA20F983" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C74D1FA20F983" pageId="8" pageNumber="209">
22. Median projection of posterior margin of tergite VIII narrower (
<figureCitation id="1303AFFDFFA1FFA3FC7D74D1FBBBF9A3" box="[917,1039,1517,1541]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="8" pageNumber="209">Figs 62–63</figureCitation>
). Median lobe of aedeagus shaped as in 
<figureCitation id="1303AFFDFFA1FFA3FEEE7731FECFF983" box="[262,379,1549,1573]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="8" pageNumber="209">Figs 65–69</figureCitation>
. Distribution: 
<collectingCountry id="F32FF3E8FFA1FFA3FDFD7731FDF0F982" box="[533,580,1549,1572]" name="Iran" pageId="8" pageNumber="209">Iran</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA1FFA3FDA57731FD77F983" box="[589,707,1549,1573]" name="Kyrgyzstan" pageId="8" pageNumber="209">Kyrgyzstan</collectingCountry>
(
<figureCitation id="1303AFFDFFA1FFA3FD397731FCADF983" box="[721,793,1549,1573]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="8" pageNumber="209">Map 5</figureCitation>
). ....................................................................................... 
<taxonomicName id="4C38C8FBFFA1FFA3FAA17731FA20F983" ID-CoL="DJKJ" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[1353,1428,1549,1573]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="persica">
<emphasis id="B94C6F6AFFA1FFA3FAA17731FA20F983" bold="true" box="[1353,1428,1549,1573]" italics="true" pageId="8" pageNumber="209">persica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA1FFA3FF2C7701FA20F9D2" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C7701FA20F9D2" pageId="8" pageNumber="209">
<heading id="D0CF0414FFA1FFA3FF2C7701FF79F9F2" box="[196,205,1597,1620]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="209" reason="4">-</heading>
Median projection of posterior margin of tergite VIII broader. Median lobe of aedeagus of different shape. ..... ....................................................................................................................................................................................... 23
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA1FFA3FF2C77B7FA20F894" pageId="8" pageNumber="209">
<paragraph id="8B87B378FFA1FFA3FF2C77B7FA21F965" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C77B7FA21F965" pageId="8" pageNumber="209">
23. Tergite VIII with broader and apically distinctly truncate median projection posteriorly (
<figureCitation id="1303AFFDFFA1FFA3FB6177B7FAB5F905" box="[1161,1281,1675,1699]" captionStart="Figs 70–90" captionStartId="14.[189,229,2000,2022]" captionTargetBox="[196,1434,240,1957]" captionTargetId="figure-41@14.[189,1436,232,1971]" captionTargetPageId="14" captionText="Figs 70–90: Anaulacaspsis truncata (70–75), A. convexa (76–83), and A. eminens (84–90): male tergite VIII (70, 76, 84); male sternite VIII (71, 77, 85); median lobe of aedeagus in lateral and in ventral view (72–73, 78–80, 86–88); female tergite VIII (74, 82, 89); female sternite VIII (75, 83, 90); apical portion of aedeagus in ventral view (81). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212618" httpUri="https://zenodo.org/record/5212618/files/figure.png" pageId="8" pageNumber="209">Figs 70, 74</figureCitation>
). Median lobe of aedeagus shaped as in 
<figureCitation id="1303AFFDFFA1FFA3FE1C7797FDDDF965" box="[500,617,1707,1731]" captionStart="Figs 70–90" captionStartId="14.[189,229,2000,2022]" captionTargetBox="[196,1434,240,1957]" captionTargetId="figure-41@14.[189,1436,232,1971]" captionTargetPageId="14" captionText="Figs 70–90: Anaulacaspsis truncata (70–75), A. convexa (76–83), and A. eminens (84–90): male tergite VIII (70, 76, 84); male sternite VIII (71, 77, 85); median lobe of aedeagus in lateral and in ventral view (72–73, 78–80, 86–88); female tergite VIII (74, 82, 89); female sternite VIII (75, 83, 90); apical portion of aedeagus in ventral view (81). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212618" httpUri="https://zenodo.org/record/5212618/files/figure.png" pageId="8" pageNumber="209">Figs 72–73</figureCitation>
. Distribution: Northwest 
<collectingCountry id="F32FF3E8FFA1FFA3FC917790FC12F965" box="[889,934,1708,1731]" name="Iran" pageId="8" pageNumber="209">Iran</collectingCountry>
(
<figureCitation id="1303AFFDFFA1FFA3FC5B7797FC4EF965" box="[947,1018,1707,1731]" captionStart="Map 2" captionStartId="27.[151,196,2023,2044]" captionTargetBox="[151,1398,1241,2014]" captionTargetId="figure-455@27.[151,1398,1241,2014]" captionTargetPageId="27" captionText="Map 2: Distributions of Anaulacaspis nigra (black circles), A. pseudonigra (white circle), A. reticulata (white triangle), A. libanotica (black triangles), A. caucasica (white diamonds), A. truncata (black diamonds), and A. sinuata (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212628" httpUri="https://zenodo.org/record/5212628/files/figure.png" pageId="8" pageNumber="209">Map 2</figureCitation>
). ............................................... 
<taxonomicName id="4C38C8FBFFA1FFA3FAD07790FA21F965" ID-CoL="DJL5" authorityName="Assing" authorityYear="2016" box="[1336,1429,1708,1731]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="B94C6F6AFFA1FFA3FAD07790FA21F965" bold="true" box="[1336,1429,1708,1731]" italics="true" pageId="8" pageNumber="209">truncata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA1FFA3FF2C77E7FA20F894" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C77E7FA20F894" pageId="8" pageNumber="209">
<heading id="D0CF0414FFA1FFA3FF2C77E7FF66F954" box="[196,210,1755,1778]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="209" reason="4">–</heading>
Tergite VIII with less broad and apically convex median projection posteriorly (
<figureCitation id="1303AFFDFFA1FFA3FBB677E6FB6CF954" box="[1118,1240,1754,1778]" captionStart="Figs 70–90" captionStartId="14.[189,229,2000,2022]" captionTargetBox="[196,1434,240,1957]" captionTargetId="figure-41@14.[189,1436,232,1971]" captionTargetPageId="14" captionText="Figs 70–90: Anaulacaspsis truncata (70–75), A. convexa (76–83), and A. eminens (84–90): male tergite VIII (70, 76, 84); male sternite VIII (71, 77, 85); median lobe of aedeagus in lateral and in ventral view (72–73, 78–80, 86–88); female tergite VIII (74, 82, 89); female sternite VIII (75, 83, 90); apical portion of aedeagus in ventral view (81). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212618" httpUri="https://zenodo.org/record/5212618/files/figure.png" pageId="8" pageNumber="209">Figs 76, 82</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="1303AFFDFFA1FFA3FE6077C6FE4BF8B4" box="[392,511,1786,1810]" captionStart="Figs 70–90" captionStartId="14.[189,229,2000,2022]" captionTargetBox="[196,1434,240,1957]" captionTargetId="figure-41@14.[189,1436,232,1971]" captionTargetPageId="14" captionText="Figs 70–90: Anaulacaspsis truncata (70–75), A. convexa (76–83), and A. eminens (84–90): male tergite VIII (70, 76, 84); male sternite VIII (71, 77, 85); median lobe of aedeagus in lateral and in ventral view (72–73, 78–80, 86–88); female tergite VIII (74, 82, 89); female sternite VIII (75, 83, 90); apical portion of aedeagus in ventral view (81). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212618" httpUri="https://zenodo.org/record/5212618/files/figure.png" pageId="8" pageNumber="209">Figs 78–81</figureCitation>
, with distinctly broader ventral process in ventral view. Distribution: 
<collectingCountry id="F32FF3E8FFA1FFA3FB0D77C6FA85F8B4" box="[1253,1329,1786,1810]" name="Turkey" pageId="8" pageNumber="209">Turkey</collectingCountry>
(
<figureCitation id="1303AFFDFFA1FFA3FAD677C6FA33F8B4" box="[1342,1415,1786,1810]" captionStart="Map 6" captionStartId="45.[151,196,2053,2074]" captionTargetBox="[151,1398,1396,2042]" captionTargetId="figure-556@45.[151,1398,1396,2042]" captionTargetPageId="45" captionText="Map 6: Distributions of Anaulacaspis convexa (white circles), A. seclusa (black circles), and A. gilva (white diamonds) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212641" httpUri="https://zenodo.org/record/5212641/files/figure.png" pageId="8" pageNumber="209">Map 6</figureCitation>
). .................................................................................................................................................................................... 
<taxonomicName id="4C38C8FBFFA1FFA3FAD47627FA20F894" ID-CoL="DJJ5" authorityName="Assing" authorityYear="2016" box="[1340,1428,1819,1842]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="B94C6F6AFFA1FFA3FAD47627FA20F894" bold="true" box="[1340,1428,1819,1842]" italics="true" pageId="8" pageNumber="209">convexa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA1FFA3FF2C7676FA21F836" pageId="8" pageNumber="209">
<paragraph id="8B87B378FFA1FFA3FF2C7676FA21F8C7" blockId="8.[196,1429,249,2048]" box="[196,1429,1865,1890]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C7676FA21F8C7" box="[196,1429,1865,1890]" pageId="8" pageNumber="209">
24. Posterior margin of tergite VIII with a row of strongly modified, stout and very short setae (
<figureCitation id="1303AFFDFFA1FFA3FB5E7675FAA6F8C4" box="[1206,1298,1865,1890]" captionStart="Figs 187–210" captionStartId="19.[151,191,1941,1963]" captionTargetBox="[170,1381,238,1935]" captionTargetId="figure-145@19.[162,1387,232,1940]" captionText="Figs 187–210: Anaulacaspsis gilva (187–190), A. pectinata (191–195), and A. formosa (196–210; 199–200: Tunisia; 201–203: Morocco; 204–205: holotype of A. macra; 206: Murcia; 207–208: Andalucía): median lobe of aedeagus in lateral and in ventral view (187–188, 194–195, 199–208); female tergite VIII (189); female sternite VIII (190, 209); male tergite VIII (191, 196); posterior margin of male tergite VIII (192, 197); male sternite VIII (193, 198); spermatheca (210). Scale bars: 187–191, 193–196, 198–210: 0.1 mm; 192, 197: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884777" httpUri="https://zenodo.org/record/5884777/files/figure.png" pageId="8" pageNumber="209">Figs 197</figureCitation>
, 
<figureCitation id="1303AFFDFFA1FFA3FAF47676FAF3F8C4" box="[1308,1351,1866,1890]" captionStart="Figs 211–232" captionStartId="20.[189,229,1941,1963]" captionTargetBox="[201,1414,238,1920]" captionTargetId="figure-41@20.[200,1426,232,1940]" captionText="Figs 211–232: Anaulacaspsis formosa (211), A. elegans (212–220), A. beesoni (221–228; 224–225: lectotype), and A. inexpectata (229–232; 231–232: holotype): habitus (211; photo by Marc Tronquet); male tergite VIII (212, 221); male sternite VIII (213, 222); median lobe of aedeagus in lateral and in ventral view (214–217, 224–227, 229–232); female tergite VIII (218); posterior margin of female tergite VIII (219); female sternite VIII (220, 228); postero-lateral portion of sternite VIII (223). Scale bars: 211: 1.0 mm; 212–218, 220–222, 224–232: 0.1 mm; 219, 223: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884779" httpUri="https://zenodo.org/record/5884779/files/figure.png" pageId="8" pageNumber="209">219</figureCitation>
). .... 25
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA1FFA3FF2C7645FA21F836" blockId="8.[196,1429,249,2048]" box="[196,1429,1912,1936]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C7645FA21F836" box="[196,1429,1912,1936]" pageId="8" pageNumber="209">
<heading id="D0CF0414FFA1FFA3FF2C7645FF66F836" box="[196,210,1913,1936]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="209" reason="4">–</heading>
Posterior margin of tergite VIII mostly with unmodified, in one species with weakly modified long setae. ...... 26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA1FFA2FF2C769BFADBFEF6" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
<paragraph id="8B87B378FFA1FFA3FF2C769BFA21F859" blockId="8.[196,1429,249,2048]" pageId="8" pageNumber="209">
<keyLead id="30C9104DFFA1FFA3FF2C769BFA21F859" pageId="8" pageNumber="209">
25. Pronotum with weakly pronounced sexual dimorphim; male pronotum with or without depression or very shallow median impression posteriorly. Median lobe of aedeagus (
<figureCitation id="1303AFFDFFA1FFA3FCB276F4FC5EF846" box="[858,1002,1992,2016]" captionStart="Figs 187–210" captionStartId="19.[151,191,1941,1963]" captionTargetBox="[170,1381,238,1935]" captionTargetId="figure-145@19.[162,1387,232,1940]" captionText="Figs 187–210: Anaulacaspsis gilva (187–190), A. pectinata (191–195), and A. formosa (196–210; 199–200: Tunisia; 201–203: Morocco; 204–205: holotype of A. macra; 206: Murcia; 207–208: Andalucía): median lobe of aedeagus in lateral and in ventral view (187–188, 194–195, 199–208); female tergite VIII (189); female sternite VIII (190, 209); male tergite VIII (191, 196); posterior margin of male tergite VIII (192, 197); male sternite VIII (193, 198); spermatheca (210). Scale bars: 187–191, 193–196, 198–210: 0.1 mm; 192, 197: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884777" httpUri="https://zenodo.org/record/5884777/files/figure.png" pageId="8" pageNumber="209">Figs 199–208</figureCitation>
) with relatively shorter ventral process. Distribution: South Spain, Northwest Africa eastwards to 
<collectingCountry id="F32FF3E8FFA1FFA3FCA776D4FC3EF7A6" box="[847,906,2024,2048]" name="Libya" pageId="8" pageNumber="209">Libya</collectingCountry>
(
<figureCitation id="1303AFFDFFA1FFA3FC7F76D4FC69F7A6" box="[919,989,2024,2048]" captionStart="Map 3" captionStartId="28.[189,234,2028,2049]" captionTargetBox="[189,1436,1371,2017]" captionTargetId="figure-589@28.[189,1436,1371,2017]" captionTargetPageId="28" captionText="Map 3: Distributions of Anaulacaspis formosa (black circles), A. elegans (white circles), A. nigrina (black diamonds), A. eminens (white diamonds), and A. nigerrima (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212634" httpUri="https://zenodo.org/record/5212634/files/figure.png" pageId="8" pageNumber="209">Map 3</figureCitation>
). ..................................................... formosa
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA0FFA2FF7771C5FADBFEF6" blockId="9.[159,1392,248,337]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF7771C5FADBFEF6" pageId="9" pageNumber="210">
<heading id="D0CF0414FFA0FFA2FF7771C5FF19FEB6" box="[159,173,249,272]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="210" reason="4">–</heading>
Pronotum with pronounced sexual dimorphism; male pronotum with distinct and extensive median impression. Median lobe of aedeagus with relatively longer ventral process (
<figureCitation id="1303AFFDFFA0FFA2FCB47024FC5BFE97" box="[860,1007,280,305]" captionStart="Figs 211–232" captionStartId="20.[189,229,1941,1963]" captionTargetBox="[201,1414,238,1920]" captionTargetId="figure-41@20.[200,1426,232,1940]" captionText="Figs 211–232: Anaulacaspsis formosa (211), A. elegans (212–220), A. beesoni (221–228; 224–225: lectotype), and A. inexpectata (229–232; 231–232: holotype): habitus (211; photo by Marc Tronquet); male tergite VIII (212, 221); male sternite VIII (213, 222); median lobe of aedeagus in lateral and in ventral view (214–217, 224–227, 229–232); female tergite VIII (218); posterior margin of female tergite VIII (219); female sternite VIII (220, 228); postero-lateral portion of sternite VIII (223). Scale bars: 211: 1.0 mm; 212–218, 220–222, 224–232: 0.1 mm; 219, 223: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884779" httpUri="https://zenodo.org/record/5884779/files/figure.png" pageId="9" pageNumber="210">Figs 214–217</figureCitation>
). Distribution: Middle East (
<collectingCountry id="F32FF3E8FFA0FFA2FAF27025FADFFE97" box="[1306,1387,281,305]" name="Cyprus" pageId="9" pageNumber="210">Cyprus</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FF2D7004FE91FEF6" box="[197,293,312,336]" name="Lebanon" pageId="9" pageNumber="210">Lebanon</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FEC67004FEDEFEF6" box="[302,362,312,336]" name="Israel" pageId="9" pageNumber="210">Israel</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FE9C7004FE09FEF6" box="[372,445,312,336]" name="Jordan" pageId="9" pageNumber="210">Jordan</collectingCountry>
, Sinai Peninsula, 
<collectingCountry id="F32FF3E8FFA0FFA2FD9E7004FCB6FEF6" box="[630,770,312,336]" name="Saudi Arabia" pageId="9" pageNumber="210">Saudi Arabia</collectingCountry>
) (
<figureCitation id="1303AFFDFFA0FFA2FCFE7005FCE8FEF7" box="[790,860,313,337]" captionStart="Map 3" captionStartId="28.[189,234,2028,2049]" captionTargetBox="[189,1436,1371,2017]" captionTargetId="figure-589@28.[189,1436,1371,2017]" captionTargetPageId="28" captionText="Map 3: Distributions of Anaulacaspis formosa (black circles), A. elegans (white circles), A. nigrina (black diamonds), A. eminens (white diamonds), and A. nigerrima (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212634" httpUri="https://zenodo.org/record/5212634/files/figure.png" pageId="9" pageNumber="210">Map 3</figureCitation>
). ..................................................................... 
<taxonomicName id="4C38C8FBFFA0FFA2FAC97004FADBFEF6" ID-CoL="66MCQ" baseAuthorityName="Cameron" baseAuthorityYear="1944" box="[1313,1391,312,336]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B94C6F6AFFA0FFA2FAC97004FADBFEF6" bold="true" box="[1313,1391,312,336]" italics="true" pageId="9" pageNumber="210">elegans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA0FFA2FF77705BFADBFE69" pageId="9" pageNumber="210">
<paragraph id="8B87B378FFA0FFA2FF77705BFADBFE39" blockId="9.[159,1391,359,416]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF77705BFADBFE39" pageId="9" pageNumber="210">
26. Median lobe of aedeagus with crista apicalis of very conspicuous shape (
<figureCitation id="1303AFFDFFA0FFA2FC4C705BFB81FED9" box="[932,1077,359,383]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="9" pageNumber="210">Figs 134–135</figureCitation>
). Distribution: North 
<collectingCountry id="F32FF3E8FFA0FFA2FAFD705BFADBFED9" box="[1301,1391,359,383]" name="Pakistan" pageId="9" pageNumber="210">Pakistan</collectingCountry>
(
<figureCitation id="1303AFFDFFA0FFA2FF2470B4FEA7FE39" box="[204,275,391,416]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="9" pageNumber="210">Map 5</figureCitation>
). ..................................................................................................................................................................... 
<taxonomicName id="4C38C8FBFFA0FFA2FAF670BBFADBFE39" ID-CoL="DJJ6" authorityName="Assing" authorityYear="2016" box="[1310,1391,391,415]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="cristata">
<emphasis id="B94C6F6AFFA0FFA2FAF670BBFADBFE39" bold="true" box="[1310,1391,391,415]" italics="true" pageId="9" pageNumber="210">cristata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA0FFA2FF77708BFADBFE69" blockId="9.[159,1391,439,463]" box="[159,1391,439,463]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF77708BFADBFE69" box="[159,1391,439,463]" pageId="9" pageNumber="210">
<heading id="D0CF0414FFA0FFA2FF77708BFF19FE68" box="[159,173,439,462]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="210" reason="4">–</heading>
Median lobe of aedeagus of completely different shape. .............................................................................................. 27
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA0FFA2FF7770DAFADBFD2B" pageId="9" pageNumber="210">
<paragraph id="8B87B378FFA0FFA2FF7770DAFADBFD9B" blockId="9.[159,1391,486,574]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF7770DAFADBFD9B" pageId="9" pageNumber="210">
27. Posterior margin of tergite VIII with a comb-like row of rather long and slightly modified setae (
<figureCitation id="1303AFFDFFA0FFA2FB3B70DAFA9BFE58" box="[1235,1327,486,510]" captionStart="Figs 164–186" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[195,1433,238,1952]" captionTargetId="figure-129@18.[189,1436,232,1970]" captionText="Figs 164–186: Anaulacaspsis iberica (164–166), A. nigrina (167–172), A. beijingensis (173–177), A. taiwanica (178–183), and A. gilva (184–186): male tergite VIII (164, 167, 173, 178, 184); male sternite VIII (165, 168, 174, 179, 185); female sternite VIII (166, 172, 177, 183); median lobe of aedeagus in lateral and in ventral view (169–171, 175–176, 180–181); female tergite VIII (182); spermatheca (186). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884775" httpUri="https://zenodo.org/record/5884775/files/figure.png" pageId="9" pageNumber="210">Figs 184</figureCitation>
, 
<figureCitation id="1303AFFDFFA0FFA2FAD170DAFAD7FE58" box="[1337,1379,486,510]" captionStart="Figs 187–210" captionStartId="19.[151,191,1941,1963]" captionTargetBox="[170,1381,238,1935]" captionTargetId="figure-145@19.[162,1387,232,1940]" captionText="Figs 187–210: Anaulacaspsis gilva (187–190), A. pectinata (191–195), and A. formosa (196–210; 199–200: Tunisia; 201–203: Morocco; 204–205: holotype of A. macra; 206: Murcia; 207–208: Andalucía): median lobe of aedeagus in lateral and in ventral view (187–188, 194–195, 199–208); female tergite VIII (189); female sternite VIII (190, 209); male tergite VIII (191, 196); posterior margin of male tergite VIII (192, 197); male sternite VIII (193, 198); spermatheca (210). Scale bars: 187–191, 193–196, 198–210: 0.1 mm; 192, 197: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884777" httpUri="https://zenodo.org/record/5884777/files/figure.png" pageId="9" pageNumber="210">189</figureCitation>
). Median lobe of aedeagus very slender, with long, slender, and apically acute ventral process, and with long dark structure in internal sac (
<figureCitation id="1303AFFDFFA0FFA2FE27731AFDD5FD98" box="[463,609,550,574]" captionStart="Figs 187–210" captionStartId="19.[151,191,1941,1963]" captionTargetBox="[170,1381,238,1935]" captionTargetId="figure-145@19.[162,1387,232,1940]" captionText="Figs 187–210: Anaulacaspsis gilva (187–190), A. pectinata (191–195), and A. formosa (196–210; 199–200: Tunisia; 201–203: Morocco; 204–205: holotype of A. macra; 206: Murcia; 207–208: Andalucía): median lobe of aedeagus in lateral and in ventral view (187–188, 194–195, 199–208); female tergite VIII (189); female sternite VIII (190, 209); male tergite VIII (191, 196); posterior margin of male tergite VIII (192, 197); male sternite VIII (193, 198); spermatheca (210). Scale bars: 187–191, 193–196, 198–210: 0.1 mm; 192, 197: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5884777" httpUri="https://zenodo.org/record/5884777/files/figure.png" pageId="9" pageNumber="210">Figs 187–188</figureCitation>
). Distribution: North 
<collectingCountry id="F32FF3E8FFA0FFA2FCA2731AFC11FD98" box="[842,933,550,574]" name="Pakistan" pageId="9" pageNumber="210">Pakistan</collectingCountry>
(
<figureCitation id="1303AFFDFFA0FFA2FC5A731AFC4DFD98" box="[946,1017,550,574]" captionStart="Map 7" captionStartId="46.[189,234,2053,2074]" captionTargetBox="[189,1436,1241,2044]" captionTargetId="figure-539@46.[189,1436,1241,2044]" captionTargetPageId="46" captionText="Map 7: Distributions of Anaulacaspis beesoni (black circles), A. pectinata (white star), A. schuelkei (black triangle), A. beijingensis (white circles), and A. taiwanica (black star) in the East Palaearctic region, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212643" httpUri="https://zenodo.org/record/5212643/files/figure.png" pageId="9" pageNumber="210">Map 7</figureCitation>
). ................................................ 
<taxonomicName id="4C38C8FBFFA0FFA2FAD37319FADBFD9B" ID-CoL="DJJJ" authorityName="Assing" authorityYear="2016" box="[1339,1391,549,573]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="gilva">
<emphasis id="B94C6F6AFFA0FFA2FAD37319FADBFD9B" bold="true" box="[1339,1391,549,573]" italics="true" pageId="9" pageNumber="210">gilva</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA0FFA2FF777369FADBFD2B" blockId="9.[159,1391,597,653]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF777369FADBFD2B" pageId="9" pageNumber="210">
<heading id="D0CF0414FFA0FFA2FF777369FF19FDCA" box="[159,173,597,620]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="210" reason="4">–</heading>
Posterior margin of tergite VIII with unmodified setae. Median lobe of aedeagus much less slender and without long dark structure in internal sac. ................................................................................................................................. 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA0FFA2FF777398FADBFCCD" pageId="9" pageNumber="210">
<paragraph id="8B87B378FFA0FFA2FF777398FADAFCBA" blockId="9.[159,1391,676,796]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF777398FADAFCBA" pageId="9" pageNumber="210">
28. Antenna very slender, 
<quantity id="4CC01E9DFFA0FFA2FE477398FD98FD1A" box="[431,556,676,700]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="9" pageNumber="210" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
long; antennomeres IV–VII distinctly oblong, VIII–IX approximately as long as broad, and X weakly transverse. Posterior margin of tergite VIII (
<figureCitation id="1303AFFDFFA0FFA2FCB073F8FC57FD7A" box="[856,995,708,732]" captionStart="Figs 136–163" captionStartId="17.[151,191,1971,1993]" captionTargetBox="[153,1394,237,1965]" captionTargetId="figure-144@17.[152,1399,232,1970]" captionText="Figs 136–163: Anaulacaspsis excisa (136–141), A. pamphylica (142–145), A. flavomarginata (146–150), A. laevigata (151–158; 153–154: lectotype; 155: Pelopónnisos; 156: Lesbos),and A. iberica (159–163): maletergite VIII(136, 142,146, 151); male sternite VIII (137, 143, 147, 152); median lobe of aedeagus in lateral and in ventral view (138–139, 144–145, 148–149, 153–156, 159–162); female sternite VIII (141, 150, 158);female tergite VIII (140); apex of ventral process of aedeagus in lateral view (157, 163). Scalebars:0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884773" httpUri="https://zenodo.org/record/5884773/files/figure.png" pageId="9" pageNumber="210">Figs 136, 140</figureCitation>
) concavely excised in the middle, setae at posterior margin short, middle of posterior margin without marginal setae. Median lobe of aedeagus shaped as in 
<figureCitation id="1303AFFDFFA0FFA2FF2C7238FEE5FCBA" box="[196,337,772,796]" captionStart="Figs 136–163" captionStartId="17.[151,191,1971,1993]" captionTargetBox="[153,1394,237,1965]" captionTargetId="figure-144@17.[152,1399,232,1970]" captionText="Figs 136–163: Anaulacaspsis excisa (136–141), A. pamphylica (142–145), A. flavomarginata (146–150), A. laevigata (151–158; 153–154: lectotype; 155: Pelopónnisos; 156: Lesbos),and A. iberica (159–163): maletergite VIII(136, 142,146, 151); male sternite VIII (137, 143, 147, 152); median lobe of aedeagus in lateral and in ventral view (138–139, 144–145, 148–149, 153–156, 159–162); female sternite VIII (141, 150, 158);female tergite VIII (140); apex of ventral process of aedeagus in lateral view (157, 163). Scalebars:0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884773" httpUri="https://zenodo.org/record/5884773/files/figure.png" pageId="9" pageNumber="210">Figs 138–139</figureCitation>
; ventral process subapically obtusely angled in lateral view. Distribution: 
<collectingCountry id="F32FF3E8FFA0FFA2FBAF7239FBC7FCBA" box="[1095,1139,773,796]" name="Iran" pageId="9" pageNumber="210">Iran</collectingCountry>
(
<figureCitation id="1303AFFDFFA0FFA2FB977238FB72FCBA" box="[1151,1222,772,796]" captionStart="Map 4" captionStartId="33.[151,196,2022,2043]" captionTargetBox="[151,1398,1203,2012]" captionTargetId="figure-561@33.[151,1398,1203,2012]" captionTargetPageId="33" captionText="Map 4: Distributions of Anaulacaspis desertorum (black circles), A. naevula (white circles), A. iberica (white stars), A. laevigata (white diamonds), A. gratilla (black diamonds), and A. excisa (black stars) in the West Palaearctic and Middle Asia, based on revised records. The question mark indicates the records without specified locality of A. gratilla from Iraq." figureDoi="http://doi.org/10.5281/zenodo.5212637" httpUri="https://zenodo.org/record/5212637/files/figure.png" pageId="9" pageNumber="210">Map 4</figureCitation>
). ............. 
<taxonomicName id="4C38C8FBFFA0FFA2FAC67238FADAFCBA" ID-CoL="66MD2" authorityName="Assing" authorityYear="2016" box="[1326,1390,772,796]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="excisa">
<emphasis id="B94C6F6AFFA0FFA2FAC67238FADAFCBA" bold="true" box="[1326,1390,772,796]" italics="true" pageId="9" pageNumber="210">excisa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA0FFA2FF777208FADBFCCD" blockId="9.[159,1391,819,875]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF777208FADBFCCD" pageId="9" pageNumber="210">
<heading id="D0CF0414FFA0FFA2FF777208FF19FCED" box="[159,173,820,843]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="210" reason="4">–</heading>
Antenna less slender. Posterior margin of tergite VIII without median concavity and with uninterrupted row of longer setae. Median lobe of aedeagus of different shape. ........................................................................................... 29
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA0FFA2FF7772BEFADBFB8F" pageId="9" pageNumber="210">
<paragraph id="8B87B378FFA0FFA2FF7772BEFADBFC7C" blockId="9.[159,1391,898,986]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF7772BEFADBFC7C" pageId="9" pageNumber="210">
29. Coloration generally paler; head usually reddish to reddish-brown; medio-lateral infuscate spot on elytra usually brown to dark-brown; abdominal segments V–VII usually brown to dark-brown. Median lobe of aedeagus shaped as in 
<figureCitation id="1303AFFDFFA0FFA2FF1472FEFECAFC7C" box="[252,382,962,986]" captionStart="Figs 91–112" captionStartId="15.[151,191,1998,2020]" captionTargetBox="[157,1394,238,1964]" captionTargetId="figure-41@15.[151,1398,232,1971]" captionTargetPageId="15" captionText="Figs 91–112:Anaulacaspsisnaevula(91–101;93–94:SaudiArabia;95–96:paralectotype;97–98: Iran;99–100:holotypeofA. elegansides), A. desertorum (102–107), and A. seclusa (108–112): male tergite VIII (91, 102, 108); male sternite VIII (92, 103, 109); median lobe of aedeagus in lateral and in ventral view (93–100, 104–107, 110–111); female sternite VIII (101, 112). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212624" httpUri="https://zenodo.org/record/5212624/files/figure.png" pageId="9" pageNumber="210">Figs 93–100</figureCitation>
. Distribution: North 
<collectingCountry id="F32FF3E8FFA0FFA2FDB772FEFD10FC7C" box="[607,676,962,986]" name="Sudan" pageId="9" pageNumber="210">Sudan</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FD4572FEFC8DFC7C" box="[685,825,962,986]" name="Saudi Arabia" pageId="9" pageNumber="210">Saudi Arabia</collectingCountry>
, South 
<collectingCountry id="F32FF3E8FFA0FFA2FC6F72FFFC02FC7C" box="[903,950,963,986]" name="Iran" pageId="9" pageNumber="210">Iran</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FC2872FEFBAFFC7C" box="[960,1051,962,986]" name="Pakistan" pageId="9" pageNumber="210">Pakistan</collectingCountry>
(
<figureCitation id="1303AFFDFFA0FFA2FBC072FEFBC5FC7C" box="[1064,1137,962,986]" captionStart="Map 4" captionStartId="33.[151,196,2022,2043]" captionTargetBox="[151,1398,1203,2012]" captionTargetId="figure-561@33.[151,1398,1203,2012]" captionTargetPageId="33" captionText="Map 4: Distributions of Anaulacaspis desertorum (black circles), A. naevula (white circles), A. iberica (white stars), A. laevigata (white diamonds), A. gratilla (black diamonds), and A. excisa (black stars) in the West Palaearctic and Middle Asia, based on revised records. The question mark indicates the records without specified locality of A. gratilla from Iraq." figureDoi="http://doi.org/10.5281/zenodo.5212637" httpUri="https://zenodo.org/record/5212637/files/figure.png" pageId="9" pageNumber="210">Map 4</figureCitation>
). ....................... 
<taxonomicName id="4C38C8FBFFA0FFA2FAF072FEFADBFC7C" ID-CoL="DJK7" baseAuthorityName="Erichson" baseAuthorityYear="1839" box="[1304,1391,962,986]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="naevula">
<emphasis id="B94C6F6AFFA0FFA2FAF072FEFADBFC7C" bold="true" box="[1304,1391,962,986]" italics="true" pageId="9" pageNumber="210">naevula</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA0FFA2FF7772CEFADBFB8F" blockId="9.[159,1391,1009,1066]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF7772CEFADBFB8F" pageId="9" pageNumber="210">
<heading id="D0CF0414FFA0FFA2FF7772CEFF19FBAF" box="[159,173,1010,1033]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="210" reason="4">–</heading>
Coloration generally darker; head usually dark-brown to blackish; medio-lateral infuscate spot on elytra darkbrown to black, usually larger and more defined; abdominal segments V–VII blackish-brown to black. ........... 30
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCABDDFFA0FFA2FF77757DFADAFAA1" pageId="9" pageNumber="210">
<paragraph id="8B87B378FFA0FFA2FF77757DFADBFB3E" blockId="9.[159,1391,1088,1177]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF77757DFADBFB3E" pageId="9" pageNumber="210">
30. Pronotum with weakly pronounced sexual dimorphism, in male with shallow and small impression. Median lobe of aedeagus with apically longer and more slender ventral process in lateral view (
<figureCitation id="1303AFFDFFA0FFA2FBF6755CFB1AFBDF" box="[1054,1198,1120,1145]" captionStart="Figs 91–112" captionStartId="15.[151,191,1998,2020]" captionTargetBox="[157,1394,238,1964]" captionTargetId="figure-41@15.[151,1398,232,1971]" captionTargetPageId="15" captionText="Figs 91–112:Anaulacaspsisnaevula(91–101;93–94:SaudiArabia;95–96:paralectotype;97–98: Iran;99–100:holotypeofA. elegansides), A. desertorum (102–107), and A. seclusa (108–112): male tergite VIII (91, 102, 108); male sternite VIII (92, 103, 109); median lobe of aedeagus in lateral and in ventral view (93–100, 104–107, 110–111); female sternite VIII (101, 112). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212624" httpUri="https://zenodo.org/record/5212624/files/figure.png" pageId="9" pageNumber="210">Figs 110–111</figureCitation>
, 
<figureCitation id="1303AFFDFFA0FFA2FB50755CFAA3FBDE" box="[1208,1303,1120,1144]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="9" pageNumber="210">115–118</figureCitation>
). Distribution: 
<collectingCountry id="F32FF3E8FFA0FFA2FEFF75BDFEF2FB3E" box="[279,326,1153,1176]" name="Iran" pageId="9" pageNumber="210">Iran</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FEB875BCFE5CFB3E" box="[336,488,1152,1176]" name="Turkmenistan" pageId="9" pageNumber="210">Turkmenistan</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FE1A75BCFDC0FB3E" box="[498,628,1152,1176]" name="Afghanistan" pageId="9" pageNumber="210">Afghanistan</collectingCountry>
(
<figureCitation id="1303AFFDFFA0FFA2FD6975BDFD7DFB3E" box="[641,713,1152,1177]" captionStart="Map 6" captionStartId="45.[151,196,2053,2074]" captionTargetBox="[151,1398,1396,2042]" captionTargetId="figure-556@45.[151,1398,1396,2042]" captionTargetPageId="45" captionText="Map 6: Distributions of Anaulacaspis convexa (white circles), A. seclusa (black circles), and A. gilva (white diamonds) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212641" httpUri="https://zenodo.org/record/5212641/files/figure.png" pageId="9" pageNumber="210">Map 6</figureCitation>
). .............................................................................................. 
<taxonomicName id="4C38C8FBFFA0FFA2FACC75BCFADBFB3E" ID-CoL="DJKT" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[1316,1391,1152,1176]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="seclusa">
<emphasis id="B94C6F6AFFA0FFA2FACC75BCFADBFB3E" bold="true" box="[1316,1391,1152,1176]" italics="true" pageId="9" pageNumber="210">seclusa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87B378FFA0FFA2FF77758CFADAFAA1" blockId="9.[159,1390,1200,1288]" pageId="9" pageNumber="210">
<keyLead id="30C9104DFFA0FFA2FF77758CFADAFAA1" pageId="9" pageNumber="210">
<heading id="D0CF0414FFA0FFA2FF77758CFF19FB61" box="[159,173,1200,1223]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="210" reason="4">–</heading>
Pronotum with pronounced sexual dimorphism, in male with deep and extensive impression. Median lobe of aedeagus with apically shorter and less slender ventral process in lateral view (
<figureCitation id="1303AFFDFFA0FFA2FC1275ECFB39FB4E" box="[1018,1165,1232,1256]" captionStart="Figs 91–112" captionStartId="15.[151,191,1998,2020]" captionTargetBox="[157,1394,238,1964]" captionTargetId="figure-41@15.[151,1398,232,1971]" captionTargetPageId="15" captionText="Figs 91–112:Anaulacaspsisnaevula(91–101;93–94:SaudiArabia;95–96:paralectotype;97–98: Iran;99–100:holotypeofA. elegansides), A. desertorum (102–107), and A. seclusa (108–112): male tergite VIII (91, 102, 108); male sternite VIII (92, 103, 109); median lobe of aedeagus in lateral and in ventral view (93–100, 104–107, 110–111); female sternite VIII (101, 112). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212624" httpUri="https://zenodo.org/record/5212624/files/figure.png" pageId="9" pageNumber="210">Figs 104–107</figureCitation>
). Distribution: North Africa and Middle East (
<collectingCountry id="F32FF3E8FFA0FFA2FE2175CCFDA1FAAE" box="[457,533,1264,1288]" name="Jordan" pageId="9" pageNumber="210">Jordan</collectingCountry>
, 
<collectingCountry id="F32FF3E8FFA0FFA2FDF675CCFD1EFAAE" box="[542,682,1264,1288]" name="Saudi Arabia" pageId="9" pageNumber="210">Saudi Arabia</collectingCountry>
) (
<figureCitation id="1303AFFDFFA0FFA2FD5675CCFCB3FAAE" box="[702,775,1264,1288]" captionStart="Map 4" captionStartId="33.[151,196,2022,2043]" captionTargetBox="[151,1398,1203,2012]" captionTargetId="figure-561@33.[151,1398,1203,2012]" captionTargetPageId="33" captionText="Map 4: Distributions of Anaulacaspis desertorum (black circles), A. naevula (white circles), A. iberica (white stars), A. laevigata (white diamonds), A. gratilla (black diamonds), and A. excisa (black stars) in the West Palaearctic and Middle Asia, based on revised records. The question mark indicates the records without specified locality of A. gratilla from Iraq." figureDoi="http://doi.org/10.5281/zenodo.5212637" httpUri="https://zenodo.org/record/5212637/files/figure.png" pageId="9" pageNumber="210">Map 4</figureCitation>
). ............................................................................ desertorum
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>