133 lines
16 KiB
XML
133 lines
16 KiB
XML
<document id="DF19FCF816DDB4F395DAD118924229F3" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE36BFFFDFF75FC15B8B1F95A" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Hydraenopsis) dilutipes" docType="treatment" docVersion="9" lastPageNumber="109" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="109" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:mods id="D878AE8C9A458609FF7CAD0D5886B8D0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="52EA920ACAD687CA0829EE3853773407">
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<mods:title id="BB87CC1815C209F84034378909E60299">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="A79A142C9955CA3BAB95130DA5CDF3DF" type="personal">
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<mods:namePart id="227AC47FE678D3A42D37F95B4733102E">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="2EF7E749FA463B87015033F6B71D8607">text</mods:typeOfResource>
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<mods:titleInfo id="1778BFA94A799F5471AC8F15905C776A">
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<mods:title id="67519330677E722C9E3AC42152FA73BB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="482D140F0EB94C64E76C5F8C12A0BC6D">
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<mods:date id="7687A3126EDD649C1F818867A6417DC6">2017</mods:date>
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<mods:number id="42CB3CFD5478D505AC85FDD8039A0992">2017-11-03</mods:number>
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<mods:detail id="FE79BBD2A5669D827A3BE73DB2B34916" type="volume">
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<mods:number id="A0BF28B5BE59DD0E972E694F8FB1E9E5">4342</mods:number>
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<mods:detail id="25243C33EBC1790E8BFB4B1A6BDE23F3" type="issue">
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<mods:classification id="8220176D466E592E95306BF654934559">journal article</mods:classification>
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<mods:identifier id="973C286D7E71FB3BC29893A88827A93D" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="A3533E5A03D765579EBCD6F6F1A9BC5E" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="82A83DC3ECDB7B957DAC57C22BBAC291" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="491C381FC3DF8FF55DCADB56C84EBE9B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="671072AC00B281CA5362CB14B819C93F" type="Zenodo-Dep">1041066</mods:identifier>
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<treatment id="038587BBE36BFFFDFF75FC15B8B1F95A" ID-DOI="http://doi.org/10.5281/zenodo.5323606" ID-GBIF-Taxon="137233439" ID-Zenodo-Dep="5323606" LSID="urn:lsid:plazi:treatment:038587BBE36BFFFDFF75FC15B8B1F95A" httpUri="http://treatment.plazi.org/id/038587BBE36BFFFDFF75FC15B8B1F95A" lastPageNumber="109" pageId="108" pageNumber="109">
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<subSubSection id="C3366526E36BFFFDFF75FC15B8FFFBB7" pageId="108" pageNumber="109" type="nomenclature">
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<paragraph id="8B9336ADE36BFFFDFF75FC15BBD2FC4F" blockId="108.[151,750,944,970]" box="[151,750,944,970]" pageId="108" pageNumber="109">
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<heading id="D0DB81C1E36BFFFDFF75FC15BBD2FC4F" bold="true" box="[151,750,944,970]" fontSize="11" level="1" pageId="108" pageNumber="109" reason="1">
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<emphasis id="B958EABFE36BFFFDFF75FC15BBD2FC4F" bold="true" box="[151,750,944,970]" pageId="108" pageNumber="109">
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<taxonomicName id="4C2C4D2EE36BFFFDFF75FC15BB01FC4F" ID-CoL="3N394" box="[151,573,944,970]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="dilutipes" subGenus="Hydraenopsis">
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<emphasis id="B958EABFE36BFFFDFF75FC15B82CFC4F" bold="true" box="[151,272,944,970]" italics="true" pageId="108" pageNumber="109">Hydraena</emphasis>
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(
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<emphasis id="B958EABFE36BFFFDFEC2FC15B8FAFC4F" bold="true" box="[288,454,944,970]" italics="true" pageId="108" pageNumber="109">Hydraenopsis</emphasis>
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)
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<emphasis id="B958EABFE36BFFFDFE35FC15BB01FC4F" bold="true" box="[471,573,944,970]" italics="true" pageId="108" pageNumber="109">dilutipes</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE36BFFFDFF75FC52B8FFFBB7" blockId="108.[151,1437,1014,1759]" pageId="108" pageNumber="109">
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Seven species are included in this group:
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<emphasis id="B958EABFE36BFFFDFD9BFC52B835FBB6" italics="true" pageId="108" pageNumber="109">
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<taxonomicName id="4C2C4D2EE36BFFFDFD9BFC52BA3DFB8A" box="[633,769,1014,1039]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="dilutipes">H. dilutipes</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE36BFFFDFCEDFC52BA87FB8A" authorityName="Perkins" authorityYear="2017" box="[783,955,1014,1039]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="dilutipoides">H. dilutipoides</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE36BFFFDFC2BFC52BD56FB8A" authorityName="Perkins" authorityYear="2017" box="[969,1130,1014,1039]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="pilotumida">H. pilotumida</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE36BFFFDFB9AFC52BDC7FB8A" authorityName="Perkins" authorityYear="2017" box="[1144,1275,1014,1039]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="pilobova">H pilobova</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE36BFFFDFAEBFC52BC4FFB8A" authorityName="Perkins" authorityYear="2017" box="[1289,1395,1014,1039]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="divisa">H. divisa</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE36BFFFDFA63FC52B839FBB6" authorityName="Perkins" authorityYear="2017" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="longiloba">H. longiloba</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="4C2C4D2EE36BFFFDFEA2FBB9B881FBB6" authorityName="Perkins" authorityYear="2017" box="[320,445,1050,1075]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="renalisa">
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<emphasis id="B958EABFE36BFFFDFEA2FBB9B881FBB6" box="[320,445,1050,1075]" italics="true" pageId="108" pageNumber="109">H. renalisa</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E36BFFFDFF25FB9BB8B1F95A" pageId="108" pageNumber="109" type="description">
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<paragraph id="8B9336ADE36BFFFDFF25FB9BB9D3FB1A" blockId="108.[151,1437,1014,1759]" pageId="108" pageNumber="109">
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Members of this group are moderately large to large (ca.
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<quantity id="4CD49B48E36BFFFDFCB6FB9BBAC0FBD2" box="[852,1020,1086,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.815" metricValueMax="2.1" metricValueMin="1.53" pageId="108" pageNumber="109" unit="mm" value="1.815" valueMax="2.1" valueMin="1.53">1.53–2.10 mm</quantity>
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), dark brown, with coarse and dense dorsal punctation. Males have distinctive metatibial brushes. The plaques vary in size and position according to the species.
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</paragraph>
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<paragraph id="8B9336ADE36BFFFDFF25FB0FB843F9CA" blockId="108.[151,1437,1014,1759]" pageId="108" pageNumber="109">
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The aedeagi have a similar general plan: the distal piece is very elongate and lobate, at the base of which is a complex multilobed structure that bears the gonopore; the basal orifice has a characteristic shape and orientation. All species have a large right paramere, but there is variation in the size of the left paramere, which is sometimes absent (only a pore in that location). In the holotype of
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<taxonomicName id="4C2C4D2EE36BFFFDFCEBFAB2BAB0FAAA" box="[777,908,1302,1327]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="dilutipes">
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<emphasis id="B958EABFE36BFFFDFCEBFAB2BAB0FAAA" box="[777,908,1302,1327]" italics="true" pageId="108" pageNumber="109">H. dilutipes</emphasis>
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</taxonomicName>
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the left paramere is long, but exceedingly thin and appears to be flexible and slightly twisted (
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<figureCitation id="13172A28E36BFFFDFD22FA9FBA1AFAD6" box="[704,806,1338,1363]" captionStart="FIGURE 100" captionStartId="113.[151,250,1877,1899]" captionText="FIGURE 100. Aedeagus of Hydraena dilutipes, lectotype." pageId="108" pageNumber="109">Fig. 100</figureCitation>
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); in other specimens of this species it appears that the paramere has been broken, and only the basal "stub" is present (
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<figureCitation id="13172A28E36BFFFDFC49FAFBBD2FFAF2" box="[939,1043,1374,1399]" captionStart="FIGURES 101 – 102" captionStartId="114.[151,264,1648,1671]" captionTargetBox="[280,1304,194,1597]" captionTargetId="figure@114.[264,1323,179,1628]" captionTargetPageId="114" captionText="FIGURES 101 – 102. Aedeagi of Hydraena dilutipes, non-type, and Hydraena dilutipoides, holotype." httpUri="https://zenodo.org/record/1041110/files/figure.png" pageId="108" pageNumber="109">Fig. 101</figureCitation>
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). Males in the group have thick musculature surrounding the aedeagus, and it is likely that a very thin and fragile paramere, with an especially weak point near the base, could be broken during dissection. In two other species of this group the remnant of the left paramere tapers to a very fine point, perhaps indicating that it has not been broken during dissection (e.g.,
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<figureCitation id="13172A28E36BFFFDFA92FA6FB9F9F982" captionStart="FIGURES 101 – 102" captionStartId="114.[151,264,1648,1671]" captionTargetBox="[280,1304,194,1597]" captionTargetId="figure@114.[264,1323,179,1628]" captionTargetPageId="114" captionText="FIGURES 101 – 102. Aedeagi of Hydraena dilutipes, non-type, and Hydraena dilutipoides, holotype." httpUri="https://zenodo.org/record/1041110/files/figure.png" pageId="108" pageNumber="109">Fig. 102</figureCitation>
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).
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<taxonomicName id="4C2C4D2EE36BFFFDFF39FA4AB867F982" authorityName="Perkins" authorityYear="2017" box="[219,347,1518,1543]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="renalisa">
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<emphasis id="B958EABFE36BFFFDFF39FA4AB867F982" box="[219,347,1518,1543]" italics="true" pageId="108" pageNumber="109">H. renalisa</emphasis>
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</taxonomicName>
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has a long left paramere, whereas
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<taxonomicName id="4C2C4D2EE36BFFFDFD10FA4ABAA0F982" authorityName="Perkins" authorityYear="2017" box="[754,924,1518,1543]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="dilutipoides">
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<emphasis id="B958EABFE36BFFFDFD10FA4ABAA0F982" box="[754,924,1518,1543]" italics="true" pageId="108" pageNumber="109">H. dilutipoides</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2C4D2EE36BFFFDFC35FA4ABD5BF982" authorityName="Perkins" authorityYear="2017" box="[983,1127,1518,1543]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="longiloba">
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<emphasis id="B958EABFE36BFFFDFC35FA4ABD5BF982" box="[983,1127,1518,1543]" italics="true" pageId="108" pageNumber="109">H. longiloba</emphasis>
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</taxonomicName>
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have a very short, sharply pointed "paramere", and in
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<taxonomicName id="4C2C4D2EE36BFFFDFE3BF9B1BB7EF9AE" authorityName="Perkins" authorityYear="2017" box="[473,578,1554,1579]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="divisa">
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<emphasis id="B958EABFE36BFFFDFE3BF9B1BB7EF9AE" box="[473,578,1554,1579]" italics="true" pageId="108" pageNumber="109">H. divisa</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE36BFFFDFDB1F9B1BBCEF9AE" authorityName="Perkins" authorityYear="2017" box="[595,754,1554,1579]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="pilotumida">
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<emphasis id="B958EABFE36BFFFDFDB1F9B1BBCEF9AE" box="[595,754,1554,1579]" italics="true" pageId="108" pageNumber="109">H. pilotumida</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C2C4D2EE36BFFFDFCD5F9B1BA82F9AE" authorityName="Perkins" authorityYear="2017" box="[823,958,1554,1579]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="pilobova">
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<emphasis id="B958EABFE36BFFFDFCD5F9B1BA82F9AE" box="[823,958,1554,1579]" italics="true" pageId="108" pageNumber="109">H. pilobova</emphasis>
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</taxonomicName>
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only a pore appears to be present in the appropriate location.
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</paragraph>
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<paragraph id="8B9336ADE36BFFFDFF25F9FEB8B1F95A" blockId="108.[151,1437,1014,1759]" pageId="108" pageNumber="109">
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Even though some of these species have only one functional paramere (the right), they are placed in the subgenus
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<taxonomicName id="4C2C4D2EE36BFFFDFEEBF9DBB899F912" box="[265,421,1662,1687]" class="Insecta" family="Hydraenidae" genus="Hydraenopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE36BFFFDFEEBF9DBB899F912" box="[265,421,1662,1687]" italics="true" pageId="108" pageNumber="109">Hydraenopsis</emphasis>
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</taxonomicName>
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because of the following characters: 1) the genae do not have a posterior ridge; 2) the metaventrite does not have an anteromedian carina; and 3) the mesotibiae of males do not have prominent spines on the medial margin.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |