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<document id="7DAE5BE4CC79BB2D776B41F062C76546" ID-DOI="10.11646/zootaxa.3721.4.3" ID-GBIF-Dataset="2aad3a91-f5d8-4c11-b501-50065c1453fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="247749" ID-ZooBank="0A63253B-03A6-4332-903E-1A17D366C2D3" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460689205080" checkinUser="plazi" docAuthor="Schawaller, Wolfgang" docDate="2013" docId="603EBF35FFE9FF9859DDFF0BAA01E061" docLanguage="en" docName="zt03721p364.pdf" docOrigin="Zootaxa 3721 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cossyphodes mourgliai Ferrer 1993, sp. n." docType="treatment" docVersion="9" lastPageNumber="359" masterDocId="9C07C74DFFE1FF90594AFF9CAE7CE517" masterDocTitle="Cossyphodini (Coleoptera: Tenebrionidae: Pimeliinae) in South Africa, Namibia and adjacent regions: New species and records, key to genera, and Old World species catalogue" masterLastPageNumber="364" masterPageNumber="351" pageNumber="359" updateTime="1698351680976" updateUser="plazi">
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<mods:title id="8455790A4FC54B3153301A07C14C9316">Cossyphodini (Coleoptera: Tenebrionidae: Pimeliinae) in South Africa, Namibia and adjacent regions: New species and records, key to genera, and Old World species catalogue</mods:title>
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<mods:namePart id="602AB31087F93CD9C0160E4CB5562C77">Schawaller, Wolfgang</mods:namePart>
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<treatment id="603EBF35FFE9FF9859DDFF0BAA01E061" ID-DOI="http://doi.org/10.5281/zenodo.6154573" ID-GBIF-Taxon="119577661" ID-Zenodo-Dep="6154573" LSID="urn:lsid:plazi:treatment:603EBF35FFE9FF9859DDFF0BAA01E061" httpUri="http://treatment.plazi.org/id/603EBF35FFE9FF9859DDFF0BAA01E061" lastPageNumber="359" pageId="8" pageNumber="359">
<subSubSection id="A08D5DA8FFE9FF9859DDFF0BAE84E701" pageId="8" pageNumber="359" type="nomenclature">
<paragraph id="E8280E23FFE9FF9859DDFF0BAD9FE5A5" blockId="8.[151,995,151,211]" box="[151,995,151,178]" pageId="8" pageNumber="359">
<heading id="B360B94FFFE9FF9859DDFF0BAD9FE5A5" bold="true" box="[151,995,151,178]" fontSize="11" level="1" pageId="8" pageNumber="359" reason="1">
<emphasis id="DAE3D231FFE9FF9859DDFF0BAD9FE5A5" bold="true" box="[151,995,151,178]" pageId="8" pageNumber="359">
<taxonomicName id="2F9775A0FFE9FF9859DDFF0BAC2CE5A5" authority="Ferrer, 1993" authorityName="Ferrer" authorityYear="1993" box="[151,592,151,178]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="mourgliai">
<emphasis id="DAE3D231FFE9FF9859DDFF0BAFCCE5A6" bold="true" box="[151,432,151,177]" italics="true" pageId="8" pageNumber="359">Cossyphodes mourgliai</emphasis>
Ferrer, 1993
</taxonomicName>
(in Ferrer &amp; Collingwood 1993)
</emphasis>
</heading>
</paragraph>
<paragraph id="E8280E23FFE9FF9859DDFF26AE85E5C4" blockId="8.[151,995,151,211]" box="[151,249,186,211]" pageId="8" pageNumber="359">
(
<figureCitation id="70AC12A6FFE9FF9859D5FF26AE8DE5C4" box="[159,241,186,211]" captionStart="FIGURES 21 26" captionStartId="9.[151,264,1722,1745]" captionTargetBox="[181,1404,195,1698]" captionTargetId="figure@9.[181,1405,193,1701]" captionTargetPageId="9" captionText="FIGURES 21 26. Dorsal view of African Cossyphodini. 21. Cossyphodes raffrayi Gestro, 1874, Tanzania, non-type HNHM. 22. Cossyphodes mourgliai Ferrer, 1993, Zambia, holotype ZSM. 23. Cossyphodes uviranus Basilewsky, 1951, Congo, nontype ZFMK. 24. Cossyphodes vandami Andreae, 1961, RSA / Pretoria, paratype MZLU. 25. Paramellops bewicki (Wollaston, 1861), RSA / Matjiesfontein, non-type TMSA. 26. Cossyphodites woodroofei (Péringuey, 1885), South Africa, non-type HNHM. Scale: 1 mm." httpUri="https://zenodo.org/record/247753/files/figure.png" pageId="8" pageNumber="359">Fig. 22</figureCitation>
)
</paragraph>
<paragraph id="E8280E23FFE9FF9859DDFE9EAFB8E490" pageId="8" pageNumber="359">
<table id="9A97FC83FFE9006F59DDFE9EABE0E490" box="[151,1436,258,391]" gridcols="10" gridrows="4" pageId="8" pageNumber="359">
<tr id="56A70C61FFE9006F59DDFE9EABE0E40C" box="[151,1436,258,283]" gridrow="0" pageId="8" pageNumber="359">
<th id="1576651DFFE9006F59DDFE9EACF1E40C" box="[151,653,258,283]" gridcol="0" gridrow="0" pageId="8" pageNumber="359">
<emphasis id="DAE3D231FFE9FF9859DDFE9EAFB8E40C" bold="true" box="[151,452,258,283]" pageId="8" pageNumber="359">Examined type material.</emphasis>
Zambia, Lusaka,
</th>
<th id="1576651DFFE9006F5BD2FE9EAC9FE40C" box="[664,739,258,283]" gridcol="1" gridrow="0" pageId="8" pageNumber="359">Kafue,</th>
<th id="1576651DFFE9006F5BA7FE9EAD4DE40C" box="[749,817,258,283]" gridcol="2" gridrow="0" pageId="8" pageNumber="359">Kafue</th>
<th id="1576651DFFE9006F5A76FE9EADFCE40C" box="[828,896,258,283]" gridcol="3" gridrow="0" pageId="8" pageNumber="359">River,</th>
<th id="1576651DFFE9006F5AC1FE9EADBFE40C" box="[907,963,258,283]" gridcol="4" gridrow="0" pageId="8" pageNumber="359">1200</th>
<th id="1576651DFFE9006F5A87FE9EAD95E40C" box="[973,1001,258,283]" gridcol="5" gridrow="0" pageId="8" pageNumber="359">m,</th>
<th id="1576651DFFE9006F5ABEFE9EAAB7E40C" box="[1012,1227,258,283]" gridcol="6" gridrow="0" pageId="8" pageNumber="359">22.XI.2.XII.1987,</th>
<th id="1576651DFFE9006F5D9CFE9EAA83E40C" box="[1238,1279,258,283]" gridcol="7" gridrow="0" pageId="8" pageNumber="359">leg.</th>
<th id="1576651DFFE9006F5C43FE9EAB5FE40C" box="[1289,1315,258,283]" gridcol="8" gridrow="0" pageId="8" pageNumber="359">R.</th>
<th id="1576651DFFE9006F5C67FE9EABE0E40C" box="[1325,1436,258,283]" gridcol="9" gridrow="0" pageId="8" pageNumber="359">Mourglia,</th>
</tr>
<tr id="56A70C61FFE9006F59DDFEBAABE0E428" box="[151,1436,294,319]" gridrow="1" pageId="8" pageNumber="359" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1576651DFFE9006F59DDFEBAACF1E428" box="[151,653,294,319]" gridcol="0" gridrow="1" pageId="8" pageNumber="359">
holotype ZSM (labelled as
<taxonomicName id="2F9775A0FFE9FF985882FEBAAC16E429" box="[456,618,294,318]" class="Insecta" family="Tenebrionidae" genus="Cossyphodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="DAE3D231FFE9FF985882FEBAAC16E429" box="[456,618,294,318]" italics="true" pageId="8" pageNumber="359">Cossyphodites</emphasis>
</taxonomicName>
).
</th>
</tr>
<tr id="56A70C61FFE9006F59DDFED6ABE0E474" box="[151,1436,330,355]" gridrow="2" pageId="8" pageNumber="359" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1576651DFFE9006F59DDFED6ACF1E474" box="[151,653,330,355]" gridcol="0" gridrow="2" pageId="8" pageNumber="359">
<emphasis id="DAE3D231FFE9FF98598DFED6AF19E474" bold="true" box="[199,357,330,355]" pageId="8" pageNumber="359">Type locality.</emphasis>
&quot;Kafue&quot;.
</th>
</tr>
<tr id="56A70C61FFE9006F59DDFEF2ABE0E490" box="[151,1436,366,391]" gridrow="3" pageId="8" pageNumber="359" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="1576651DFFE9006F59DDFEF2ACF1E490" box="[151,653,366,391]" gridcol="0" gridrow="3" pageId="8" pageNumber="359">
<emphasis id="DAE3D231FFE9FF98598DFEF2AF1CE490" bold="true" box="[199,352,366,391]" pageId="8" pageNumber="359">Distribution.</emphasis>
Zambia.
</th>
</tr>
</table>
</paragraph>
<paragraph id="E8280E23FFE9FF9859DDFE46AE84E701" blockId="8.[151,548,474,534]" pageId="8" pageNumber="359">
<taxonomicName id="2F9775A0FFE9FF9859DDFE46AFA1E4E3" box="[151,477,474,500]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="naukluftensis" status="sp. nov.">
<emphasis id="DAE3D231FFE9FF9859DDFE46AFA1E4E3" bold="true" box="[151,477,474,500]" italics="true" pageId="8" pageNumber="359">Cossyphodes naukluftensis</emphasis>
</taxonomicName>
<emphasis id="DAE3D231FFE9FF9858AEFE47AC58E4E3" bold="true" box="[484,548,475,500]" pageId="8" pageNumber="359">
<taxonomicNameLabel id="C1D06F4AFFE9FF9858AEFE47AC58E4E3" box="[484,548,475,500]" pageId="8" pageNumber="359" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="70AC12A6FFE9FF9859D5FE61AE8DE701" box="[159,241,509,534]" captionStart="FIGURES 7 13" captionStartId="4.[151,264,1864,1887]" captionTargetBox="[183,1401,306,1837]" captionTargetId="figure@4.[181,1405,303,1842]" captionTargetPageId="4" captionText="FIGURES 7 13. Dorsal view of African Cossyphodini. 7 8. Cossyphodes andreaei sp. n., RSA / Waterberg, paratype SMNS and aedeagus. 9. Cossyphodes arnoldi Brauns, 1925, Zimbabwe, lectotype TMSA. 10. Cossyphodes naukluftensis sp. n., Namibia / Naukluft, holotype TMSA. 11. Cossyphodes angolanus Basilewsky, 1952, holotype MRUC. 12. Cossyphodes wasmanni Reichensperger, 1915 (Cossyphodes brincki Andreae, 1961 syn. n., RSA / Hluhluwe, holotype MZLU). 13. Cossyphodes wasmanni Reichensperger, 1915, Zimbabwe, lectotype TMSA. Scale: 1 mm, for aedeagus: 0.5 mm." httpUri="https://zenodo.org/record/247751/files/figure.png" pageId="8" pageNumber="359">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE9FF9859DDFDD9ACF0E795" pageId="8" pageNumber="359" type="materials_examined">
<paragraph id="E8280E23FFE9FF9859DDFDD9ACF0E795" blockId="8.[151,1436,581,1398]" pageId="8" pageNumber="359">
<emphasis id="DAE3D231FFE9FF9859DDFDD9AFAAE749" bold="true" box="[151,470,581,606]" pageId="8" pageNumber="359">
<typeStatus id="372CB081FFE9FF9859DDFDD9AEAEE749" box="[151,210,581,606]" pageId="8" pageNumber="359">Type</typeStatus>
material.
<typeStatus id="372CB081FFE9FF98581FFDD9AFBEE749" box="[341,450,581,606]" pageId="8" pageNumber="359" type="holotype">Holotype</typeStatus>
(
</emphasis>
sex unknown
<emphasis id="DAE3D231FFE9FF985B3AFDD9AC89E749" bold="true" box="[624,757,581,606]" pageId="8" pageNumber="359">
):
<collectingCountry id="90804EB3FFE9FF985BC4FDD9AC89E749" box="[654,757,581,606]" name="Namibia" pageId="8" pageNumber="359">Namibia</collectingCountry>
</emphasis>
(labelled as S. W. Afr.), Naukluft Park,
<geoCoordinate id="8DA368E4FFE9FF985D96FDD9AB48E749" box="[1244,1332,581,606]" direction="south" orientation="latitude" pageId="8" pageNumber="359" precision="925" value="-24.266666">24°16'S</geoCoordinate>
/
<geoCoordinate id="8DA368E4FFE9FF985C74FDD9ABEBE749" box="[1342,1431,581,606]" direction="east" orientation="longitude" pageId="8" pageNumber="359" precision="925" value="16.25">16°15'E</geoCoordinate>
,
<date id="9C2928E3FFE9FF9859DDFDF5AF73E795" box="[151,271,617,642]" pageId="8" pageNumber="359" value="1974-10-26">26.X.1974</date>
, leg. S. Endrödy-Younga, TMSA.
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE9FF98598DFD11AADCE141" pageId="8" pageNumber="359" type="description">
<paragraph id="E8280E23FFE9FF98598DFD11AADCE141" blockId="8.[151,1436,581,1398]" pageId="8" pageNumber="359">
<emphasis id="DAE3D231FFE9FF98598DFD11AF25E7B1" bold="true" box="[199,345,653,678]" pageId="8" pageNumber="359">Description.</emphasis>
Body reddish brown without colour pattern. Body length
<quantity id="2F6FA3C6FFE9FF985AAAFD12AA44E7B2" box="[992,1080,654,678]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="359" unit="mm" value="2.3">2.3 mm</quantity>
. Head semicircular, clypeus not separated, clypeal lines indistinct, straight; anterior and lateral margins somewhat bent upwards, frons with a pair of distinct, complete external keels and a pair of a weak internal keels, anterior of internal keel with tubercle; head surface with regular microgranulation; eyes narrow sickle-shaped, composed by two rows of dark ocelli, each row with about five ocelli; antennae 11-segmented with two large apical antennomeres forming club. Pronotum 1.5 times broader than median length, posterior corner rectangular, acute, anterior corner rounded; all margins unbordered; surface with same microgranulation and microsetation as on head, disc on each side with three complete longitudinal keels, additionally with distinct medial keel, this medial keel shortly interrupted in anterior part, for direction, distances of these keels, see
<figureCitation id="70AC12A6FFE9FF985BECFC31AC84E6D1" box="[678,760,941,966]" captionStart="FIGURES 7 13" captionStartId="4.[151,264,1864,1887]" captionTargetBox="[183,1401,306,1837]" captionTargetId="figure@4.[181,1405,303,1842]" captionTargetPageId="4" captionText="FIGURES 7 13. Dorsal view of African Cossyphodini. 7 8. Cossyphodes andreaei sp. n., RSA / Waterberg, paratype SMNS and aedeagus. 9. Cossyphodes arnoldi Brauns, 1925, Zimbabwe, lectotype TMSA. 10. Cossyphodes naukluftensis sp. n., Namibia / Naukluft, holotype TMSA. 11. Cossyphodes angolanus Basilewsky, 1952, holotype MRUC. 12. Cossyphodes wasmanni Reichensperger, 1915 (Cossyphodes brincki Andreae, 1961 syn. n., RSA / Hluhluwe, holotype MZLU). 13. Cossyphodes wasmanni Reichensperger, 1915, Zimbabwe, lectotype TMSA. Scale: 1 mm, for aedeagus: 0.5 mm." httpUri="https://zenodo.org/record/247751/files/figure.png" pageId="8" pageNumber="359">Fig. 10</figureCitation>
; prosternal process elongate triangular. Elytra widest shortly before base, 1.4 times longer than broad; elytra with four primary and secondary keels, internal main keel only present near base of elytra, all main keels convergent from base towards tip, for direction, distances of these keels, see
<figureCitation id="70AC12A6FFE9FF98598EFB85AF66E125" box="[196,282,1049,1074]" captionStart="FIGURES 7 13" captionStartId="4.[151,264,1864,1887]" captionTargetBox="[183,1401,306,1837]" captionTargetId="figure@4.[181,1405,303,1842]" captionTargetPageId="4" captionText="FIGURES 7 13. Dorsal view of African Cossyphodini. 7 8. Cossyphodes andreaei sp. n., RSA / Waterberg, paratype SMNS and aedeagus. 9. Cossyphodes arnoldi Brauns, 1925, Zimbabwe, lectotype TMSA. 10. Cossyphodes naukluftensis sp. n., Namibia / Naukluft, holotype TMSA. 11. Cossyphodes angolanus Basilewsky, 1952, holotype MRUC. 12. Cossyphodes wasmanni Reichensperger, 1915 (Cossyphodes brincki Andreae, 1961 syn. n., RSA / Hluhluwe, holotype MZLU). 13. Cossyphodes wasmanni Reichensperger, 1915, Zimbabwe, lectotype TMSA. Scale: 1 mm, for aedeagus: 0.5 mm." httpUri="https://zenodo.org/record/247751/files/figure.png" pageId="8" pageNumber="359">Fig. 10</figureCitation>
; surface with the same microgranulation and microsetation as on head and pronotum. Legs without peculiarities, tarsal formula 5-4-4. Aedeagus unknown, sex not examined (see introduction).
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE9FF98598DFBFDAFA1E045" pageId="8" pageNumber="359" type="diagnosis">
<paragraph id="E8280E23FFE9FF98598DFBFDAFA1E045" blockId="8.[151,1436,581,1398]" pageId="8" pageNumber="359">
<emphasis id="DAE3D231FFE9FF98598DFBFDAF3DE16D" bold="true" box="[199,321,1121,1146]" pageId="8" pageNumber="359">Diagnosis.</emphasis>
<taxonomicName id="2F9775A0FFE9FF985802FBFEAF81E16E" box="[328,509,1121,1145]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="naukluftensis" status="sp. nov.">
<emphasis id="DAE3D231FFE9FF985802FBFEAF81E16E" box="[328,509,1121,1145]" italics="true" pageId="8" pageNumber="359">C. naukluftensis</emphasis>
</taxonomicName>
<emphasis id="DAE3D231FFE9FF985B4EFBFDAC3FE16D" bold="true" box="[516,579,1121,1146]" pageId="8" pageNumber="359">
<taxonomicNameLabel id="C1D06F4AFFE9FF985B4EFBFDAC3FE16D" box="[516,579,1121,1146]" pageId="8" pageNumber="359" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="2F9775A0FFE9FF985B9FFBFEAD94E16D" authority="Brauns, 1925" authorityName="Brauns" authorityYear="1925" box="[725,1000,1121,1146]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="arnoldi">
<emphasis id="DAE3D231FFE9FF985B9FFBFEAD34E16E" box="[725,840,1121,1145]" italics="true" pageId="8" pageNumber="359">C. arnoldi</emphasis>
Brauns, 1925
</taxonomicName>
, known from
<collectingCountry id="90804EB3FFE9FF985DCEFBFDAA80E16D" box="[1156,1276,1121,1146]" name="Zimbabwe" pageId="8" pageNumber="359">Zimbabwe</collectingCountry>
(
<figureCitation id="70AC12A6FFE9FF985C46FBFDAB2CE16D" box="[1292,1360,1121,1146]" captionStart="FIGURES 7 13" captionStartId="4.[151,264,1864,1887]" captionTargetBox="[183,1401,306,1837]" captionTargetId="figure@4.[181,1405,303,1842]" captionTargetPageId="4" captionText="FIGURES 7 13. Dorsal view of African Cossyphodini. 7 8. Cossyphodes andreaei sp. n., RSA / Waterberg, paratype SMNS and aedeagus. 9. Cossyphodes arnoldi Brauns, 1925, Zimbabwe, lectotype TMSA. 10. Cossyphodes naukluftensis sp. n., Namibia / Naukluft, holotype TMSA. 11. Cossyphodes angolanus Basilewsky, 1952, holotype MRUC. 12. Cossyphodes wasmanni Reichensperger, 1915 (Cossyphodes brincki Andreae, 1961 syn. n., RSA / Hluhluwe, holotype MZLU). 13. Cossyphodes wasmanni Reichensperger, 1915, Zimbabwe, lectotype TMSA. Scale: 1 mm, for aedeagus: 0.5 mm." httpUri="https://zenodo.org/record/247751/files/figure.png" pageId="8" pageNumber="359">Fig. 9</figureCitation>
). Both share the narrow body shape, the head with a pair of distinct keels and with a tubercle before the internal keel, and the distinct pronotal keels with interrupted medial keel. Both species can be separated by a completely different pattern of main elytral keels (
<figureCitation id="70AC12A6FFE9FF9858A2FB51AC1FE1F1" box="[488,611,1229,1254]" captionStart="FIGURES 7 13" captionStartId="4.[151,264,1864,1887]" captionTargetBox="[183,1401,306,1837]" captionTargetId="figure@4.[181,1405,303,1842]" captionTargetPageId="4" captionText="FIGURES 7 13. Dorsal view of African Cossyphodini. 7 8. Cossyphodes andreaei sp. n., RSA / Waterberg, paratype SMNS and aedeagus. 9. Cossyphodes arnoldi Brauns, 1925, Zimbabwe, lectotype TMSA. 10. Cossyphodes naukluftensis sp. n., Namibia / Naukluft, holotype TMSA. 11. Cossyphodes angolanus Basilewsky, 1952, holotype MRUC. 12. Cossyphodes wasmanni Reichensperger, 1915 (Cossyphodes brincki Andreae, 1961 syn. n., RSA / Hluhluwe, holotype MZLU). 13. Cossyphodes wasmanni Reichensperger, 1915, Zimbabwe, lectotype TMSA. Scale: 1 mm, for aedeagus: 0.5 mm." httpUri="https://zenodo.org/record/247751/files/figure.png" pageId="8" pageNumber="359">Figs. 910</figureCitation>
). In
<taxonomicName id="2F9775A0FFE9FF985BD0FB52AD2CE1F2" box="[666,848,1229,1253]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="naukluftensis" status="sp. nov.">
<emphasis id="DAE3D231FFE9FF985BD0FB52AD2CE1F2" box="[666,848,1229,1253]" italics="true" pageId="8" pageNumber="359">C. naukluftensis</emphasis>
</taxonomicName>
<emphasis id="DAE3D231FFE9FF985A12FB51ADE5E1F1" bold="true" box="[856,921,1229,1254]" pageId="8" pageNumber="359">
<taxonomicNameLabel id="C1D06F4AFFE9FF985A12FB51ADE5E1F1" box="[856,921,1229,1254]" pageId="8" pageNumber="359" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the elytral keels are bent outwards near base and convergent from base towards tip, in
<taxonomicName id="2F9775A0FFE9FF985BCCFB6EAC81E01E" box="[646,765,1265,1289]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="arnoldi">
<emphasis id="DAE3D231FFE9FF985BCCFB6EAC81E01E" box="[646,765,1265,1289]" italics="true" pageId="8" pageNumber="359">C. arnoldi</emphasis>
</taxonomicName>
the main keels are parallel in anterior half of elytra and convergent only in posterior half; in
<taxonomicName id="2F9775A0FFE9FF985B72FA8AAC92E03A" box="[568,750,1301,1325]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="naukluftensis" status="sp. nov.">
<emphasis id="DAE3D231FFE9FF985B72FA8AAC92E03A" box="[568,750,1301,1325]" italics="true" pageId="8" pageNumber="359">C. naukluftensis</emphasis>
</taxonomicName>
<emphasis id="DAE3D231FFE9FF985BBCFA89AD4BE039" bold="true" box="[758,823,1301,1326]" pageId="8" pageNumber="359">
<taxonomicNameLabel id="C1D06F4AFFE9FF985BBCFA89AD4BE039" box="[758,823,1301,1326]" pageId="8" pageNumber="359" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the internal main keel is present only near base, in
<taxonomicName id="2F9775A0FFE9FF985CC8FA8AAE96E046" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="arnoldi">
<emphasis id="DAE3D231FFE9FF985CC8FA8AAE96E046" italics="true" pageId="8" pageNumber="359">C. arnoldi</emphasis>
</taxonomicName>
this keel is complete.
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE9FF98598DFAC1AA01E061" box="[199,1149,1373,1398]" pageId="8" pageNumber="359" type="etymology">
<paragraph id="E8280E23FFE9FF98598DFAC1AA01E061" blockId="8.[151,1436,581,1398]" box="[199,1149,1373,1398]" pageId="8" pageNumber="359">
<emphasis id="DAE3D231FFE9FF98598DFAC1AF32E061" bold="true" box="[199,334,1373,1398]" pageId="8" pageNumber="359">Etymology.</emphasis>
Named after the Naukluft Mountains, where the
<typeStatus id="372CB081FFE9FF985A3EFAC1ADAAE061" box="[884,982,1373,1398]" pageId="8" pageNumber="359" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>