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<document id="9F9109AB34214C98CFAC78B3F79246DB" ID-DOI="10.11646/zootaxa.3721.4.3" ID-GBIF-Dataset="2aad3a91-f5d8-4c11-b501-50065c1453fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="247749" ID-ZooBank="0A63253B-03A6-4332-903E-1A17D366C2D3" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460689205080" checkinUser="plazi" docAuthor="Schawaller, Wolfgang" docDate="2013" docId="603EBF35FFE4FF9659DDFCF0AA57E472" docLanguage="en" docName="zt03721p364.pdf" docOrigin="Zootaxa 3721 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cossyphodes braunsi Andreae 1961" docType="treatment" docVersion="9" lastPageNumber="357" masterDocId="9C07C74DFFE1FF90594AFF9CAE7CE517" masterDocTitle="Cossyphodini (Coleoptera: Tenebrionidae: Pimeliinae) in South Africa, Namibia and adjacent regions: New species and records, key to genera, and Old World species catalogue" masterLastPageNumber="364" masterPageNumber="351" pageNumber="356" updateTime="1698351680976" updateUser="plazi">
<mods:mods id="923301ADC0BEBEC989E8561C2BEFB07E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AF1F0B318696FD703AF4C023ECAD4B8E">
<mods:title id="EBD652E32D9FEF0AED22F55C674B0DD1">Cossyphodini (Coleoptera: Tenebrionidae: Pimeliinae) in South Africa, Namibia and adjacent regions: New species and records, key to genera, and Old World species catalogue</mods:title>
</mods:titleInfo>
<mods:name id="92586F1A093D894C52F186ADE48E00C5" type="personal">
<mods:role id="2D55A93FA72C2D272926C867913571DD">
<mods:roleTerm id="EA27CBFCD9374EAAA8E1421066662FBE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EBEB70BAAF093C19AFB8FD9C62E94AFA">Schawaller, Wolfgang</mods:namePart>
</mods:name>
<mods:typeOfResource id="6A3FD52766BB2FCF56ED48E9AB606CE6">text</mods:typeOfResource>
<mods:relatedItem id="7D70D7CDD3F4F8B5EE33369CF4F5BB9C" type="host">
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<mods:title id="8D9AE37304338E5A1DD6A2391F40A8D6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="27B0AECDF53FB37C784FE06BB2FBF415">
<mods:date id="AD5CC34787486257FD85B4CF21A4A7D9">2013</mods:date>
<mods:detail id="65C6458FADD5718A29621537EAA3EDBD" type="volume">
<mods:number id="D18EB6D31C03EF19B9F9DFFCE214937F">3721</mods:number>
</mods:detail>
<mods:detail id="182043FAFE7DD7065FB6B56F2F14C227" type="issue">
<mods:number id="F5B914F5F7E508208968EABC4D62D2D5">4</mods:number>
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<mods:start id="58204678A4D53FB6BFA3CAEAA78E99AB">351</mods:start>
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<mods:classification id="03F8C4BEFA2E67675FA32125D8B1AC26">journal article</mods:classification>
<mods:identifier id="66392C37F70D7E43EF0DB9F2E7556741" type="DOI">10.11646/zootaxa.3721.4.3</mods:identifier>
<mods:identifier id="9EDC6DA0AE19643F6F8BB81B03776E5C" type="GBIF-Dataset">2aad3a91-f5d8-4c11-b501-50065c1453fc</mods:identifier>
<mods:identifier id="C4DD806BE54A00F85A4198B851436FEF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AE68BD663C45CFD39423A834F22F48EA" type="Zenodo-Dep">247749</mods:identifier>
<mods:identifier id="2C551BBC97CF13676591B6A404CD7C4E" type="ZooBank">0A63253B-03A6-4332-903E-1A17D366C2D3</mods:identifier>
</mods:mods>
<treatment id="603EBF35FFE4FF9659DDFCF0AA57E472" ID-DOI="http://doi.org/10.5281/zenodo.6154567" ID-GBIF-Taxon="119577668" ID-Zenodo-Dep="6154567" LSID="urn:lsid:plazi:treatment:603EBF35FFE4FF9659DDFCF0AA57E472" httpUri="http://treatment.plazi.org/id/603EBF35FFE4FF9659DDFCF0AA57E472" lastPageId="6" lastPageNumber="357" pageId="5" pageNumber="356">
<subSubSection id="A08D5DA8FFE4FF9559DDFCF0AC86E6FB" pageId="5" pageNumber="356" type="nomenclature">
<paragraph id="E8280E23FFE4FF9559DDFCF0AC2DE691" blockId="5.[151,593,876,935]" box="[151,593,876,902]" pageId="5" pageNumber="356">
<heading id="B360B94FFFE4FF9559DDFCF0AC2DE691" bold="true" box="[151,593,876,902]" fontSize="11" level="1" pageId="5" pageNumber="356" reason="1">
<taxonomicName id="2F9775A0FFE4FF9559DDFCF0AC2DE691" authority="Andreae, 1961" authorityName="Andreae" authorityYear="1961" box="[151,593,876,902]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="braunsi">
<emphasis id="DAE3D231FFE4FF9559DDFCF0AC2DE691" bold="true" box="[151,593,876,902]" pageId="5" pageNumber="356">
<emphasis id="DAE3D231FFE4FF9559DDFCF0AFEAE691" bold="true" box="[151,406,876,902]" italics="true" pageId="5" pageNumber="356">Cossyphodes braunsi</emphasis>
Andreae, 1961
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="E8280E23FFE4FF9559DDFC12AE85E6B0" blockId="5.[151,593,876,935]" box="[151,249,910,935]" pageId="5" pageNumber="356">
(
<figureCitation id="70AC12A6FFE4FF9559D5FC12AE8DE6B0" box="[159,241,910,935]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[185,1404,195,1721]" captionTargetId="figure@7.[181,1405,193,1725]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Dorsal view of African Cossyphodini. 14. Cossyphodes braunsi Andreae, 1961, RSA / Port Elizabeth, nontype HNHM. 15 – 16. Cossyphodes caecus sp. n., RSA / Richtersveld, paratype SMNS and aedeagus. 17. Cossyphodes kundelunguensis Basilewsky, 1950, Congo, paratype ZSM. 18. Cossyphodes freudei Basilewsky, 1957, Namibia, holotype ZSM. 19. Cossyphodes freudei Basilewsky, 1957 (Cossyphodes uhligi Ferrer, 2004 syn. n., Namibia, holotype MNB). 16. Cossyphodes machadoi Basilewsky, 1952, holotype MRUC. Scale: 1 mm." httpUri="https://zenodo.org/record/247752/files/figure.png" pageId="5" pageNumber="356">Fig. 14</figureCitation>
)
</paragraph>
<paragraph id="E8280E23FFE4FF9559DDFC4AAC86E6FB" blockId="5.[151,762,982,1004]" box="[151,762,982,1004]" pageId="5" pageNumber="356">
<taxonomicName id="2F9775A0FFE4FF9559DDFC4AAC64E6FB" authority="Wasmann, 1899" authorityName="Wasmann" authorityYear="1899" box="[151,536,982,1004]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="bewicki">
<emphasis id="DAE3D231FFE4FF9559DDFC4AAF12E6FB" box="[151,366,982,1004]" italics="true" pageId="5" pageNumber="356">Cossyphodes bewicki</emphasis>
Wasmann, 1899
</taxonomicName>
(not Wollaston, 1861)
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE4FF9559DDFB85AC56E1D5" pageId="5" pageNumber="356" type="materials_examined">
<paragraph id="E8280E23FFE4FF9559DDFB85AC75E126" blockId="5.[151,1436,1049,1254]" box="[151,521,1049,1074]" pageId="5" pageNumber="356">
<emphasis id="DAE3D231FFE4FF9559DDFB85AFC2E125" bold="true" box="[151,446,1049,1074]" pageId="5" pageNumber="356">
Examined
<typeStatus id="372CB081FFE4FF95585DFB85AF37E125" box="[279,331,1049,1074]" pageId="5" pageNumber="356">type</typeStatus>
material.
</emphasis>
None.
</paragraph>
<paragraph id="E8280E23FFE4FF95598DFBA1ACA5E189" blockId="5.[151,1436,1049,1254]" pageId="5" pageNumber="356">
<emphasis id="DAE3D231FFE4FF95598DFBA1AF0EE141" bold="true" box="[199,370,1085,1110]" pageId="5" pageNumber="356">New material.</emphasis>
<collectingCountry id="90804EB3FFE4FF955837FBA1AC6FE141" box="[381,531,1085,1110]" name="South Africa" pageId="5" pageNumber="356">South Africa</collectingCountry>
, Eastern Cape, Port Elizabeth, Algoa Bay, leg. H. Brauns, 2 ex. HNHM (det. Basilewsky as
<taxonomicName id="2F9775A0FFE4FF955874FBFEAFCFE16E" box="[318,435,1121,1145]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="bewicki">
<emphasis id="DAE3D231FFE4FF955874FBFEAFCFE16E" box="[318,435,1121,1145]" italics="true" pageId="5" pageNumber="356">C. bewicki</emphasis>
</taxonomicName>
), 1 ex. BMNH (det. as
<taxonomicName id="2F9775A0FFE4FF955BF7FBFEAD4FE16E" box="[701,819,1121,1145]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="bewicki">
<emphasis id="DAE3D231FFE4FF955BF7FBFEAD4FE16E" box="[701,819,1121,1145]" italics="true" pageId="5" pageNumber="356">C. bewicki</emphasis>
</taxonomicName>
).—
<collectingCountry id="90804EB3FFE4FF955A17FBFDAD8CE16D" box="[861,1008,1121,1146]" name="South Africa" pageId="5" pageNumber="356">South Africa</collectingCountry>
(labelled as Kapland), Port Elizabeth, leg. H. Brauns, 2 ex. ZFMK (coll. Reichensperger).
</paragraph>
<paragraph id="E8280E23FFE4FF95598DFB35AC56E1D5" blockId="5.[151,1436,1049,1254]" box="[199,554,1193,1218]" pageId="5" pageNumber="356">
<emphasis id="DAE3D231FFE4FF95598DFB35AF19E1D5" bold="true" box="[199,357,1193,1218]" pageId="5" pageNumber="356">
<typeStatus id="372CB081FFE4FF95598DFB35AF7EE1D5" box="[199,258,1193,1218]" pageId="5" pageNumber="356">Type</typeStatus>
locality.
</emphasis>
“Port Elisabeth”.
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE4FF95598DFB51AF51E07A" pageId="5" pageNumber="356" type="distribution">
<paragraph id="E8280E23FFE4FF95598DFB51ACD1E1F1" blockId="5.[151,1436,1049,1254]" box="[199,685,1229,1254]" pageId="5" pageNumber="356">
<emphasis id="DAE3D231FFE4FF95598DFB51AF1CE1F1" bold="true" box="[199,352,1229,1254]" pageId="5" pageNumber="356">Distribution.</emphasis>
<collectingCountry id="90804EB3FFE4FF95582DFB51AF8AE1F1" box="[359,502,1229,1254]" name="South Africa" pageId="5" pageNumber="356">South Africa</collectingCountry>
(Eastern Cape).
</paragraph>
<paragraph id="E8280E23FFE4FF9559DDFAADAF51E07A" blockId="5.[151,465,1329,1389]" pageId="5" pageNumber="356">
<taxonomicName id="2F9775A0FFE4FF9559DDFAADAFF6E05C" box="[151,394,1329,1355]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="caecus" status="sp. nov.">
<emphasis id="DAE3D231FFE4FF9559DDFAADAFF6E05C" bold="true" box="[151,394,1329,1355]" italics="true" pageId="5" pageNumber="356">Cossyphodes caecus</emphasis>
</taxonomicName>
<emphasis id="DAE3D231FFE4FF9558DBFAAEAFADE05C" bold="true" box="[401,465,1330,1355]" pageId="5" pageNumber="356">
<taxonomicNameLabel id="C1D06F4AFFE4FF9558DBFAAEAFADE05C" box="[401,465,1330,1355]" pageId="5" pageNumber="356" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="70AC12A6FFE4FF9559D5FAC8AF5AE07A" box="[159,294,1364,1389]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[185,1404,195,1721]" captionTargetId="figure@7.[181,1405,193,1725]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Dorsal view of African Cossyphodini. 14. Cossyphodes braunsi Andreae, 1961, RSA / Port Elizabeth, nontype HNHM. 15 – 16. Cossyphodes caecus sp. n., RSA / Richtersveld, paratype SMNS and aedeagus. 17. Cossyphodes kundelunguensis Basilewsky, 1950, Congo, paratype ZSM. 18. Cossyphodes freudei Basilewsky, 1957, Namibia, holotype ZSM. 19. Cossyphodes freudei Basilewsky, 1957 (Cossyphodes uhligi Ferrer, 2004 syn. n., Namibia, holotype MNB). 16. Cossyphodes machadoi Basilewsky, 1952, holotype MRUC. Scale: 1 mm." httpUri="https://zenodo.org/record/247752/files/figure.png" pageId="5" pageNumber="356">Figs. 15–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE4FF9559DDFA00AE90E353" pageId="5" pageNumber="356" type="materials_examined">
<paragraph id="E8280E23FFE4FF9559DDFA00AE90E353" blockId="5.[151,1436,1436,2037]" pageId="5" pageNumber="356">
<emphasis id="DAE3D231FFE4FF9559DDFA00ACD0E0A2" bold="true" box="[151,684,1436,1461]" pageId="5" pageNumber="356">
<typeStatus id="372CB081FFE4FF9559DDFA00AEAEE0A2" box="[151,210,1436,1461]" pageId="5" pageNumber="356">Type</typeStatus>
material.
<typeStatus id="372CB081FFE4FF95581EFA00AFBDE0A2" box="[340,449,1436,1461]" pageId="5" pageNumber="356" type="holotype">Holotype</typeStatus>
(♂):
<collectingCountry id="90804EB3FFE4FF955B42FA00ACDBE0A2" box="[520,679,1436,1461]" name="South Africa" pageId="5" pageNumber="356">South Africa</collectingCountry>
,
</emphasis>
Northern Cape, Richtersveld NP, Doringpoort,
<geoCoordinate id="8DA368E4FFE4FF955D96FA00AB48E0A2" box="[1244,1332,1436,1461]" direction="south" orientation="latitude" pageId="5" pageNumber="356" precision="925" value="-28.566668">28°34'S</geoCoordinate>
/
<geoCoordinate id="8DA368E4FFE4FF955C77FA00ABEBE0A2" box="[1341,1431,1436,1461]" direction="east" orientation="longitude" pageId="5" pageNumber="356" precision="925" value="16.933332">16°56'E</geoCoordinate>
,
<date id="9C2928E3FFE4FF9559DDFA23AF77E0CF" box="[151,267,1471,1496]" pageId="5" pageNumber="356" value="1976-09-07">7.IX.1976</date>
, leg. S. Endrödy-Younga, TMSA.—
<emphasis id="DAE3D231FFE4FF955BE6FA23AD51E0CF" bold="true" box="[684,813,1471,1496]" pageId="5" pageNumber="356">
<typeStatus id="372CB081FFE4FF955BE6FA23AD54E0CF" box="[684,808,1471,1496]" pageId="5" pageNumber="356" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as
<typeStatus id="372CB081FFE4FF955A9DFA23AA40E0CF" box="[983,1084,1471,1496]" pageId="5" pageNumber="356" type="holotype">holotype</typeStatus>
, 1 ex. TMSA, 1 ex. SMNS.— Western Cape, Elands Bay,
<date id="9C2928E3FFE4FF9558A9FA78AC3EE0EB" box="[483,578,1508,1532]" pageId="5" pageNumber="356" value="1948-11">XI.1948</date>
, leg. C. Koch, 3 ex. TMSA, 1 ex. SMNS.—
<collectingCountry id="90804EB3FFE4FF955D14FA78AA84E0EA" box="[1118,1272,1508,1533]" name="South Africa" pageId="5" pageNumber="356">South Africa</collectingCountry>
, Mpumalanga (labelled as N Transvaal), Waterberg, Haakdoringboom Farm,
<geoCoordinate id="8DA368E4FFE4FF955A12F99BADD0E337" box="[856,940,1543,1568]" direction="south" orientation="latitude" pageId="5" pageNumber="356" precision="925" value="-24.183332">24°11'S</geoCoordinate>
/
<geoCoordinate id="8DA368E4FFE4FF955AFCF99BAA6EE337" box="[950,1042,1543,1568]" direction="east" orientation="longitude" pageId="5" pageNumber="356" precision="925" value="27.833334">27°50'E</geoCoordinate>
,
<date id="9C2928E3FFE4FF955D55F99BAAEAE337" box="[1055,1174,1543,1568]" pageId="5" pageNumber="356" value="1976-02-13">13.II.1976</date>
, leg. A. Strydom, 1 ex. TMSA.
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE4FF95598DF9D3ABF1E2E2" pageId="5" pageNumber="356" type="description">
<paragraph id="E8280E23FFE4FF95598DF9D3ABF1E2E2" blockId="5.[151,1436,1436,2037]" pageId="5" pageNumber="356">
<emphasis id="DAE3D231FFE4FF95598DF9D3AF25E37F" bold="true" box="[199,345,1615,1640]" pageId="5" pageNumber="356">Description.</emphasis>
Body dark brown without colour pattern. Body length
<quantity id="2F6FA3C6FFE4FF955A9BF9D3AA21E370" box="[977,1117,1615,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="5" pageNumber="356" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 mm</quantity>
. Head semicircular, clypeus not separated, clypeal lines indistinct, straight; anterior and lateral margins somewhat bent upwards, frons with two pairs of weak keels, internal keels reduced to two tubercles each; head surface with regular microgranulation; eyes completely absent, a short longitudinal keel instead of ocelli; antennae 11-segmented with two large apical antennomeres forming club. Pronotum 1.7 times broader than median length, posterior corner rectangular, acute, anterior corner rounded; all margins unbordered; surface with same microgranulation and microsetation as on head, disc on each side with three distinct, complete longitudinal keels, additionally with a distinct medial keel, for direction, distances of these keels, see
<figureCitation id="70AC12A6FFE4FF955B24F8D0ACBAE272" box="[622,710,1868,1893]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[185,1404,195,1721]" captionTargetId="figure@7.[181,1405,193,1725]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Dorsal view of African Cossyphodini. 14. Cossyphodes braunsi Andreae, 1961, RSA / Port Elizabeth, nontype HNHM. 15 – 16. Cossyphodes caecus sp. n., RSA / Richtersveld, paratype SMNS and aedeagus. 17. Cossyphodes kundelunguensis Basilewsky, 1950, Congo, paratype ZSM. 18. Cossyphodes freudei Basilewsky, 1957, Namibia, holotype ZSM. 19. Cossyphodes freudei Basilewsky, 1957 (Cossyphodes uhligi Ferrer, 2004 syn. n., Namibia, holotype MNB). 16. Cossyphodes machadoi Basilewsky, 1952, holotype MRUC. Scale: 1 mm." httpUri="https://zenodo.org/record/247752/files/figure.png" pageId="5" pageNumber="356">Fig. 15</figureCitation>
; prosternal process elongate triangular with protruding acute posterior angle. Elytra widest shortly before base, 1.4 times longer than broad; elytra with four primary keels and distinct secondary keels, internal main keel present only on anterior third of elytra, for direction, distances of these keels, see
<figureCitation id="70AC12A6FFE4FF955858F824AF14E2C6" box="[274,360,1976,2001]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[185,1404,195,1721]" captionTargetId="figure@7.[181,1405,193,1725]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Dorsal view of African Cossyphodini. 14. Cossyphodes braunsi Andreae, 1961, RSA / Port Elizabeth, nontype HNHM. 15 – 16. Cossyphodes caecus sp. n., RSA / Richtersveld, paratype SMNS and aedeagus. 17. Cossyphodes kundelunguensis Basilewsky, 1950, Congo, paratype ZSM. 18. Cossyphodes freudei Basilewsky, 1957, Namibia, holotype ZSM. 19. Cossyphodes freudei Basilewsky, 1957 (Cossyphodes uhligi Ferrer, 2004 syn. n., Namibia, holotype MNB). 16. Cossyphodes machadoi Basilewsky, 1952, holotype MRUC. Scale: 1 mm." httpUri="https://zenodo.org/record/247752/files/figure.png" pageId="5" pageNumber="356">Fig. 15</figureCitation>
; surface with the same microgranulation and microsetation as on head and pronotum. Legs without peculiarities, tarsal formula 5-4-4 as usual for the genus. Aedeagus with trianguar acute apicale (
<figureCitation id="70AC12A6FFE4FF955C67F840ABFCE2E2" box="[1325,1408,2012,2037]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[185,1404,195,1721]" captionTargetId="figure@7.[181,1405,193,1725]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Dorsal view of African Cossyphodini. 14. Cossyphodes braunsi Andreae, 1961, RSA / Port Elizabeth, nontype HNHM. 15 – 16. Cossyphodes caecus sp. n., RSA / Richtersveld, paratype SMNS and aedeagus. 17. Cossyphodes kundelunguensis Basilewsky, 1950, Congo, paratype ZSM. 18. Cossyphodes freudei Basilewsky, 1957, Namibia, holotype ZSM. 19. Cossyphodes freudei Basilewsky, 1957 (Cossyphodes uhligi Ferrer, 2004 syn. n., Namibia, holotype MNB). 16. Cossyphodes machadoi Basilewsky, 1952, holotype MRUC. Scale: 1 mm." httpUri="https://zenodo.org/record/247752/files/figure.png" pageId="5" pageNumber="356">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE7FF96598DFF0BAEA4E457" pageId="6" pageNumber="357" type="diagnosis">
<paragraph id="E8280E23FFE7FF96598DFF0BAEA4E457" blockId="6.[151,1436,151,357]" pageId="6" pageNumber="357">
<emphasis id="DAE3D231FFE7FF96598DFF0BAF3DE5A7" bold="true" box="[199,321,151,176]" pageId="6" pageNumber="357">Diagnosis.</emphasis>
<taxonomicName id="2F9775A0FFE7FF965802FF04AC51E5A7" box="[328,557,152,176]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="caecus" status="sp. nov.">
<emphasis id="DAE3D231FFE7FF965802FF04AC51E5A7" box="[328,557,152,176]" italics="true" pageId="6" pageNumber="357">Cossyphodes caecus</emphasis>
</taxonomicName>
<emphasis id="DAE3D231FFE7FF965B7EFF0BAC0FE5A7" bold="true" box="[564,627,151,176]" pageId="6" pageNumber="357">
<taxonomicNameLabel id="C1D06F4AFFE7FF965B7EFF0BAC0FE5A7" box="[564,627,151,176]" pageId="6" pageNumber="357" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be recognised by the lack of eyes without any ocelli present, by two pairs of weak keels on the head, by distinct complete keels on the pronotum, and by distinct main keels as well as present secondary keels on the elytra.
<taxonomicName id="2F9775A0FFE7FF965B03FF7CAD45E5EF" box="[585,825,224,248]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="braunsi">
<emphasis id="DAE3D231FFE7FF965B03FF7CAD45E5EF" box="[585,825,224,248]" italics="true" pageId="6" pageNumber="357">Cossyphodes braunsi</emphasis>
</taxonomicName>
from Eastern Cape (
<figureCitation id="70AC12A6FFE7FF965D6CFF43AA05E5EF" box="[1062,1145,223,248]" captionStart="FIGURES 14 – 20" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[185,1404,195,1721]" captionTargetId="figure@7.[181,1405,193,1725]" captionTargetPageId="7" captionText="FIGURES 14 – 20. Dorsal view of African Cossyphodini. 14. Cossyphodes braunsi Andreae, 1961, RSA / Port Elizabeth, nontype HNHM. 15 – 16. Cossyphodes caecus sp. n., RSA / Richtersveld, paratype SMNS and aedeagus. 17. Cossyphodes kundelunguensis Basilewsky, 1950, Congo, paratype ZSM. 18. Cossyphodes freudei Basilewsky, 1957, Namibia, holotype ZSM. 19. Cossyphodes freudei Basilewsky, 1957 (Cossyphodes uhligi Ferrer, 2004 syn. n., Namibia, holotype MNB). 16. Cossyphodes machadoi Basilewsky, 1952, holotype MRUC. Scale: 1 mm." httpUri="https://zenodo.org/record/247752/files/figure.png" pageId="6" pageNumber="357">Fig. 14</figureCitation>
) also lacks ocelli, but the head is without any keels or tubercles, the pronotum with only weak external keels, and the elytra with extinct inner keels.
</paragraph>
</subSubSection>
<subSubSection id="A08D5DA8FFE7FF96598DFED0AA57E472" box="[199,1067,332,357]" pageId="6" pageNumber="357" type="etymology">
<paragraph id="E8280E23FFE7FF96598DFED0AA57E472" blockId="6.[151,1436,151,357]" box="[199,1067,332,357]" pageId="6" pageNumber="357">
<emphasis id="DAE3D231FFE7FF96598DFED0AF32E472" bold="true" box="[199,334,332,357]" pageId="6" pageNumber="357">Etymology.</emphasis>
The name refers to the lack of eyes,
<taxonomicName id="2F9775A0FFE7FF965BA1FED1AD4AE473" box="[747,822,333,356]" class="Insecta" family="Tenebrionidae" genus="Cossyphodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="caecus">caecus</taxonomicName>
in Latin means blind.
</paragraph>
</subSubSection>
</treatment>
</document>
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