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<document id="D6E06DC28F1C52CF4110527C8C694CED" ID-DOI="10.37828/em.2023.61.9" ID-ISSN="2336-9744" ID-Zenodo-Dep="13246732" ID-ZooBank="urn:lsid:zoobank.org:pub:4EB022DC-076A-4C1C-BFBD-01F9B4342F0B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972049171" checkinUser="felipe" docAuthor="Dubatolov, Vladimir V., Korb, Stanislav K. &amp; Yakovlev, Roman V." docDate="2023" docId="601E954AFFB6FFF312A2269E8011D8C1" docLanguage="en" docName="EcolMontenegrina.61.88-123.pdf" docOrigin="Ecologica Montenegrina 61" docSource="http://dx.doi.org/10.37828/em.2023.61.9" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Triphysa nervosa subsp. arturi Dubatolov, Korb &amp; Yakovlev 2023, ssp. n." docType="treatment" docUuid="A9A360D0-F138-4D02-A80C-FAA7D298BF4C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="113" masterDocId="9C27ED32FFA4FFEA1302231E833CD90B" masterDocTitle="A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae)" masterLastPageNumber="123" masterPageNumber="88" pageNumber="106" updateTime="1722980517101" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FDE5A1571DF55FA4B91CED7FC59E0DBC">A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae)</mods:title>
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<mods:namePart id="259D2559C996D5B970321380BE1DF26C">Dubatolov, Vladimir V.</mods:namePart>
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<mods:affiliation id="921F911FD4B2D5DD71A4BE49D3C5EA3F">Institute of Systematics and Ecology of Animals, Siberian Branch of Russian Academy of Sciences, Frunze str. 11, Novosibirsk 630091 Russia. https: // orcid. org / 0000 - 0001 - 7687 - 2102</mods:affiliation>
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<mods:namePart id="97DE7B45C12825A0748D51220C88AEEB">Korb, Stanislav K.</mods:namePart>
<mods:affiliation id="F441E8770253269576E35255B2E3ADA9">Russian Entomological Society, Nizhny Novgorod Division P. O. Box 97, Nizhny Novgorod 603009 Russia.</mods:affiliation>
<mods:nameIdentifier id="5358512B94EBC3BB1EC7B5E3F597A771" type="email">stanislavkorb@list.ru</mods:nameIdentifier>
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<mods:namePart id="7ACB620BE9AB54CE65929AE02582B118">Yakovlev, Roman V.</mods:namePart>
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<mods:affiliation id="9A55AA33B92AFC376A23BEE88D935718">Altai State University, pr. Lenina 61, Barnaul, 656049, Russia. https: // orcid. org / 0000 - 0001 - 9512 - 8709 * Corresponding author. E-mail: stanislavkorb @ list. ru; https: // orcid. org / 0000 - 0002 - 1120 - 424 X</mods:affiliation>
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<mods:title id="949B1FA0EE8EB6A2808FAA99E5D98546">Ecologica Montenegrina</mods:title>
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<mods:date id="66DB478298CC9128404985CAF5142610">2023</mods:date>
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<treatment id="601E954AFFB6FFF312A2269E8011D8C1" ID-DOI="http://doi.org/10.5281/zenodo.13247354" ID-Zenodo-Dep="13247354" LSID="urn:lsid:zoobank.org:act:A9A360D0-F138-4D02-A80C-FAA7D298BF4C" httpUri="http://treatment.plazi.org/id/601E954AFFB6FFF312A2269E8011D8C1" lastPageId="25" lastPageNumber="113" pageId="18" pageNumber="106">
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<heading id="B3409330FFB6FFF812A2269E87AFDC91" box="[416,1171,1408,1434]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="106" reason="3">
<emphasis id="DAC3F84EFFB6FFF812A2269E87AFDC91" bold="true" box="[416,1171,1408,1434]" pageId="18" pageNumber="106">
<taxonomicName id="2FB75FDFFFB6FFF812A2269E8705DC91" authority="Dubatolov, Korb &amp; Yakovlev" authorityName="Dubatolov, Korb &amp; Yakovlev" authorityYear="2023" box="[416,1081,1408,1434]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="nervosa" status="subsp. nov." subSpecies="arturi">
<emphasis id="DAC3F84EFFB6FFF812A2269E8183DC91" bold="true" box="[416,703,1408,1434]" italics="true" pageId="18" pageNumber="106">Triphysa nervosa arturi</emphasis>
Dubatolov, Korb &amp; Yakovlev
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,
<taxonomicNameLabel id="C1F04535FFB6FFF81744269E87AFDC91" box="[1094,1171,1408,1434]" pageId="18" pageNumber="106" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="A0AD77D7FFB6FFF8119826A180A1DF88" pageId="18" pageNumber="106" type="description">
<paragraph id="E808245CFFB6FFF8119826A180A6DCD1" blockId="18.[289,1298,1408,1498]" box="[666,922,1471,1498]" pageId="18" pageNumber="106">
<figureCitation id="708C38D9FFB6FFF8119826A1802EDCD1" box="[666,786,1471,1498]" captionStart="Figure 3" captionStartId="7.[151,226,1794,1818]" captionTargetBox="[227,1360,201,1775]" captionTargetId="figure-22@7.[227,1360,189,1775]" captionTargetPageId="7" captionText="Figure 3. Triphysa sp., imago: a – T. nervosa tuvinica, holotype (ISEAN), upperside, b – underside; c – T. nervosa tuvinica, paratype, male (RYB), upperside, d – underside; e – T. nervosa tuvinica, paratype, female (RYB), upperside, f – underside; g – T. nervosa tuvinica, paratype, female (RYB), upperside, h – underside; i – T. nervosa arturi, holotype (ISEAN), upperside, j – underside; k – T. nervosa arturi, paratype, male (RYB), upperside, l – underside; m – T. nervosa arturi, paratype, female (RYB), upperside, n – underside; o – T. nervosa arturi, paratype, female (ISEAN), upperside, p – underside; q – T. nervosa kobdoensis, holotype (ZISP), upperside, r – underside; s – T. nervosa kobdoensis, paratype, male (RYB), upperside, t – underside; u – T. nervosa kobdoensis, paratype, female (RYB), upperside, v – underside." figureDoi="http://doi.org/10.5281/zenodo.13246742" httpUri="https://zenodo.org/record/13246742/files/figure.png" pageId="18" pageNumber="106">Figs. 3i–p</figureCitation>
,
<figureCitation id="708C38D9FFB6FFF8101C26A18001DCD1" box="[798,829,1471,1498]" captionStart="Figure 6" captionStartId="12.[151,226,1968,1992]" captionTargetBox="[211,1376,217,1925]" captionTargetId="figure-8@12.[210,1376,189,1249]" captionTargetPageId="12" captionText="Figure 6. Distribution map of Triphysa species: a (above) – general map; b (below) – northern-eastern part of area of T. nervosa nervosa." figureDoi="http://doi.org/10.5281/zenodo.13246750" httpUri="https://zenodo.org/record/13246750/files/figure.png" pageId="18" pageNumber="106">6a</figureCitation>
,
<figureCitation id="708C38D9FFB6FFF8104B26A18054DCD1" box="[841,872,1471,1498]" captionStart="Figures 12−13" captionStartId="21.[151,237,1978,2002]" captionTargetBox="[171,1415,190,1951]" captionTargetId="figure-22@21.[171,1415,189,1953]" captionTargetPageId="21" captionText="Figures 12−13. Habitats of Triphysa: 12 – T. nervosa tuvinica (type locality, photo by P. Kosachev); 13 – T. nervosa arturi (type locality, photo by P. Kosachev)." figureDoi="http://doi.org/10.5281/zenodo.13246762" httpUri="https://zenodo.org/record/13246762/files/figure.png" pageId="18" pageNumber="106">13</figureCitation>
,
<figureCitation id="708C38D9FFB6FFF8107726A180AADCD1" box="[885,918,1471,1498]" captionStart="Figures 25–30" captionStartId="27.[151,237,1970,1994]" captionTargetBox="[231,1373,193,1939]" captionTargetId="figure-22@27.[209,1378,189,1952]" captionTargetPageId="27" captionText="Figures 25–30. Male genitalia (lateral projection) of Triphysa: 25 – T. nervosa glacialis, Altai, Ukok, Mai-Pak, h–2300 m; 26 – T. nervosa tuvinica, paratype; 27 – T. nervosa arturi, paratype; 28 – T. nervosa kobdoensis, paratype; 29 – T. nervosa mongolaltaica, holotype; 30 – T. nervosa biocellata, lectotype." figureDoi="http://doi.org/10.5281/zenodo.13246776" httpUri="https://zenodo.org/record/13246776/files/figure.png" pageId="18" pageNumber="106">27</figureCitation>
.
</paragraph>
<paragraph id="E808245CFFB6FFF81395251D80A1DF88" blockId="18.[151,1436,1538,1970]" pageId="18" pageNumber="106">
<emphasis id="DAC3F84EFFB6FFF81395251D821FDF16" bold="true" box="[151,291,1539,1565]" italics="true" pageId="18" pageNumber="106">Description</emphasis>
.
<emphasis id="DAC3F84EFFB6FFF81236251C824CDF16" box="[308,368,1538,1565]" italics="true" pageId="18" pageNumber="106">Male</emphasis>
. Forewing length
<quantity id="2F4F89B9FFB6FFF8114E251D81F3DF16" box="[588,719,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="18" pageNumber="106" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
. Wings dark with white fringes and wide light border along the marginal and costal (fore wing) borders. Central cell with white veins. Underside with greatly reduced postdiscal spots.
<emphasis id="DAC3F84EFFB6FFF8126525588283DF6A" box="[359,447,1606,1633]" italics="true" pageId="18" pageNumber="106">Female</emphasis>
. Forewing length
<quantity id="2F4F89B9FFB6FFF811A125588015DF6A" box="[675,809,1606,1633]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="18" pageNumber="106" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
. Upperside white. Underside with strongly reduced eyespots. Forewing underside in its angle with expressed lighting.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFB6FFF813F42594822BDE64" pageId="18" pageNumber="106" type="diagnosis">
<paragraph id="E808245CFFB6FFF813F42594822BDE64" blockId="18.[151,1436,1538,1970]" pageId="18" pageNumber="106">
<emphasis id="DAC3F84EFFB6FFF813F425948253DFAF" bold="true" box="[246,367,1674,1700]" italics="true" pageId="18" pageNumber="106">Diagnosis</emphasis>
. This new subspecies (as far as
<emphasis id="DAC3F84EFFB6FFF811F425948044DFAE" box="[758,888,1674,1701]" italics="true" pageId="18" pageNumber="106">kobdoensis</emphasis>
) is an intermediate taxon between the groups
<taxonomicName id="2FB75FDFFFB6FFF8139525B583C0DFCD" authorityName="A. Bang-Haas" authorityYear="1912" box="[151,252,1707,1734]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="106" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="DAC3F84EFFB6FFF8139525B583C0DFCD" box="[151,252,1707,1734]" italics="true" pageId="18" pageNumber="106">glacialis</emphasis>
</taxonomicName>
and
<taxonomicName id="2FB75FDFFFB6FFF8123925B5828CDFCD" authorityName="Staudinger" authorityYear="1901" box="[315,432,1707,1734]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="106" phylum="Arthropoda" rank="species" species="biocellata">
<emphasis id="DAC3F84EFFB6FFF8123925B5828CDFCD" box="[315,432,1707,1734]" italics="true" pageId="18" pageNumber="106">biocellata</emphasis>
</taxonomicName>
. Specimen are very small, males with wide white bordering on the wing upperside (the bordering often extends to the costal area), in the central cell white veins are visible (as in the previous subspecies, which is larger and with larger postdiscal eyespots). Postdiscal eyespots on the underside are reduced, but the quite large cubital spots on the forewings can remain; they are always without white centers. Underside always with contrasting bright veins. Female has brightening on the forewing underside in the anal angle.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFB6FFF813F424688139DEB9" pageId="18" pageNumber="106" type="etymology">
<paragraph id="E808245CFFB6FFF813F424688139DEB9" blockId="18.[151,1436,1538,1970]" pageId="18" pageNumber="106">
<emphasis id="DAC3F84EFFB6FFF813F42468824BDE9B" bold="true" box="[246,375,1910,1936]" italics="true" pageId="18" pageNumber="106">Etymology</emphasis>
. New subspecies named after Artur Yakovlev, son of the third author, who helped to collect a part of the
<typeStatus id="370C9AFEFFB6FFF812832486828FDEB9" box="[385,435,1944,1970]" pageId="18" pageNumber="106">type</typeStatus>
series.
</paragraph>
</subSubSection>
<caption id="BCC874D4FFB7FFF9139524B88006DED7" ID-DOI="http://doi.org/10.5281/zenodo.13246756" ID-Zenodo-Dep="13246756" httpUri="https://zenodo.org/record/13246756/files/figure.png" pageId="19" pageNumber="107" startId="19.[151,237,1958,1982]" targetBox="[162,1425,193,1926]" targetPageId="19" targetType="figure">
<paragraph id="E808245CFFB7FFF9139524B88006DED7" blockId="19.[151,1437,1957,2013]" pageId="19" pageNumber="107">
<emphasis id="DAC3F84EFFB7FFF9139524B88215DEB5" bold="true" box="[151,297,1958,1982]" pageId="19" pageNumber="107">Figures 8–9.</emphasis>
Habitats of
<taxonomicName id="2FB75FDFFFB7FFF912BF24BB8120DEB6" authorityName="Zeller" authorityYear="1850" box="[445,540,1957,1981]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="DAC3F84EFFB7FFF912BF24BB8120DEB6" box="[445,540,1957,1981]" italics="true" pageId="19" pageNumber="107">Triphysa</emphasis>
</taxonomicName>
: 8 –
<taxonomicName id="2FB75FDFFFB7FFF9116124B8801ADEB6" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[611,806,1957,1981]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="107" phylum="Arthropoda" rank="subSpecies" species="phryne" subSpecies="phryne">
<emphasis id="DAC3F84EFFB7FFF9116124B8801ADEB6" box="[611,806,1957,1981]" italics="true" pageId="19" pageNumber="107">T. phryne phryne</emphasis>
</taxonomicName>
, Russia, Volgograd reg. (photo by A. Samus’); 9 –
<taxonomicName id="2FB75FDFFFB7FFF9168524B88204DED7" authority=", SE" authorityName="SE" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="107" phylum="Arthropoda" rank="species" species="issykkulica">
<emphasis id="DAC3F84EFFB7FFF9168524B88230DED7" italics="true" pageId="19" pageNumber="107">T. issykkulica</emphasis>
, SE
</taxonomicName>
Kazakhstan, Alma-Ata reg. (photo by S. Korb).
</paragraph>
</caption>
<caption id="BCC874D4FFB0FFFE13952485817FDEFB" ID-DOI="http://doi.org/10.5281/zenodo.13246758" ID-Zenodo-Dep="13246758" httpUri="https://zenodo.org/record/13246758/files/figure.png" pageId="20" pageNumber="108" startId="20.[151,237,1947,1971]" targetBox="[233,1356,191,1928]" targetPageId="20" targetType="figure">
<paragraph id="E808245CFFB0FFFE13952485817FDEFB" blockId="20.[151,1436,1947,2032]" pageId="20" pageNumber="108">
<emphasis id="DAC3F84EFFB0FFFE13952485827CDEB8" bold="true" box="[151,320,1947,1971]" pageId="20" pageNumber="108">Figures 10−11.</emphasis>
Habitats of
<taxonomicName id="2FB75FDFFFB0FFFE12C52485811ADEB8" authorityName="Zeller" authorityYear="1850" box="[455,550,1947,1971]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="DAC3F84EFFB0FFFE12C52485811ADEB8" box="[455,550,1947,1971]" italics="true" pageId="20" pageNumber="108">Triphysa</emphasis>
</taxonomicName>
, and female of
<taxonomicName id="2FB75FDFFFB0FFFE11D32482809BDEB8" authorityName="A. Bang-Haas (Altai-Sayan Mts., Tannu-Ola Mts., Minusinsk Valley" authorityYear="1912" box="[721,935,1947,1971]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="108" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="glacialis">
<emphasis id="DAC3F84EFFB0FFFE11D32482809BDEB8" box="[721,935,1947,1971]" italics="true" pageId="20" pageNumber="108">T. nervosa glacialis</emphasis>
</taxonomicName>
in nature: 10 –
<taxonomicName id="2FB75FDFFFB0FFFE175724828665DEB8" authority=", SE" authorityName="A. Bang-Haas (Altai-Sayan Mts., Tannu-Ola Mts., Minusinsk Valley" authorityYear="1912" box="[1109,1369,1947,1971]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="108" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="glacialis">
<emphasis id="DAC3F84EFFB0FFFE175724828617DEB8" box="[1109,1323,1947,1971]" italics="true" pageId="20" pageNumber="108">T. nervosa glacialis</emphasis>
, SE
</taxonomicName>
Altai, Ukok plateau, near Bertek, h–2400 m (photo by R. Yakovlev); 11 −
<taxonomicName id="2FB75FDFFFB0FFFE107B24A48772DEDA" authorityName="A. Bang-Haas (Altai-Sayan Mts., Tannu-Ola Mts., Minusinsk Valley" authorityYear="1912" box="[889,1102,1977,2001]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="108" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="glacialis">
<emphasis id="DAC3F84EFFB0FFFE107B24A48772DEDA" box="[889,1102,1977,2001]" italics="true" pageId="20" pageNumber="108">T. nervosa glacialis</emphasis>
</taxonomicName>
– SE Altai, near Chagan-Uzun vill., h–2200 m (photo by A. Volynkin).
</paragraph>
</caption>
<caption id="BCC874D4FFB1FFFF139524A4814DDEFB" ID-DOI="http://doi.org/10.5281/zenodo.13246762" ID-Zenodo-Dep="13246762" httpUri="https://zenodo.org/record/13246762/files/figure.png" pageId="21" pageNumber="109" startId="21.[151,237,1978,2002]" targetBox="[171,1415,190,1951]" targetPageId="21" targetType="figure">
<paragraph id="E808245CFFB1FFFF139524A4814DDEFB" blockId="21.[151,1436,1978,2032]" pageId="21" pageNumber="109">
<emphasis id="DAC3F84EFFB1FFFF139524A4827DDED9" bold="true" box="[151,321,1978,2002]" pageId="21" pageNumber="109">Figures 12−13.</emphasis>
Habitats of
<taxonomicName id="2FB75FDFFFB1FFFF12CB24A48114DED9" authorityName="Zeller" authorityYear="1850" box="[457,552,1978,2002]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="DAC3F84EFFB1FFFF12CB24A48114DED9" box="[457,552,1978,2002]" italics="true" pageId="21" pageNumber="109">Triphysa</emphasis>
</taxonomicName>
: 12 –
<taxonomicName id="2FB75FDFFFB1FFFF117324A5807EDED9" authorityName="Dubatolov, Korb et Yakovlev" authorityYear="2023" box="[625,834,1979,2002]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="tuvinica">
<emphasis id="DAC3F84EFFB1FFFF117324A5807EDED9" box="[625,834,1979,2002]" italics="true" pageId="21" pageNumber="109">T. nervosa tuvinica</emphasis>
</taxonomicName>
(type locality, photo by P. Kosachev); 13
<emphasis id="DAC3F84EFFB1FFFF161624A583E9DEFB" italics="true" pageId="21" pageNumber="109">
–
<taxonomicName id="2FB75FDFFFB1FFFF162924A583E9DEFB" authorityName="Dubatolov, Korb &amp; Yakovlev" authorityYear="2023" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="arturi">T. nervosa arturi</taxonomicName>
</emphasis>
(type locality, photo by P. Kosachev).
</paragraph>
</caption>
<caption id="BCC874D4FFB2FFFC139524DC81E9DEF3" ID-DOI="http://doi.org/10.5281/zenodo.13246764" ID-Zenodo-Dep="13246764" httpUri="https://zenodo.org/record/13246764/files/figure.png" pageId="22" pageNumber="110" startId="22.[151,237,1986,2010]" targetBox="[229,1356,189,1960]" targetPageId="22" targetType="figure">
<paragraph id="E808245CFFB2FFFC139524DC81E9DEF3" blockId="22.[151,1437,1986,2040]" pageId="22" pageNumber="110">
<emphasis id="DAC3F84EFFB2FFFC139524DC8201DED1" bold="true" box="[151,317,1986,2010]" pageId="22" pageNumber="110">Figures 14–15.</emphasis>
Habitats of
<taxonomicName id="2FB75FDFFFB2FFFC12C024DC811DDED1" authorityName="Zeller" authorityYear="1850" box="[450,545,1986,2010]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="DAC3F84EFFB2FFFC12C024DC811DDED1" box="[450,545,1986,2010]" italics="true" pageId="22" pageNumber="110">Triphysa</emphasis>
</taxonomicName>
: 14 –
<taxonomicName id="2FB75FDFFFB2FFFC116424DD806FDED1" authorityName="Dubatolov &amp; Korb &amp; Yakovlev" authorityYear="2023" box="[614,851,1986,2010]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="110" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="kobdoensis">
<emphasis id="DAC3F84EFFB2FFFC116424DD806FDED1" box="[614,851,1986,2010]" italics="true" pageId="22" pageNumber="110">T. nervosa kobdoensis</emphasis>
</taxonomicName>
(type locality, photo by D. Ryzhkov); 15 –
<taxonomicName id="2FB75FDFFFB2FFFC162924DD820DDEF3" authorityName="Dubatolov &amp; Korb &amp; Yakovlev" authorityYear="2023" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="110" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="mongolaltaica">
<emphasis id="DAC3F84EFFB2FFFC162924DD820DDEF3" italics="true" pageId="22" pageNumber="110">T. nervosa mongolaltaica</emphasis>
</taxonomicName>
(type locality, photo by A. Kechaikin).
</paragraph>
</caption>
<caption id="BCC874D4FFB3FFFD139524AA801ADEE0" ID-DOI="http://doi.org/10.5281/zenodo.13246766" ID-Zenodo-Dep="13246766" httpUri="https://zenodo.org/record/13246766/files/figure.png" pageId="23" pageNumber="111" startId="23.[151,237,1972,1996]" targetBox="[191,1394,190,1950]" targetPageId="23" targetType="figure">
<paragraph id="E808245CFFB3FFFD139524AA801ADEE0" blockId="23.[151,1437,1972,2027]" pageId="23" pageNumber="111">
<emphasis id="DAC3F84EFFB3FFFD139524AA827DDEC7" bold="true" box="[151,321,1972,1996]" pageId="23" pageNumber="111">Figures 16–17.</emphasis>
Habitats of
<taxonomicName id="2FB75FDFFFB3FFFD12CD24AA8112DEC7" authorityName="Zeller" authorityYear="1850" box="[463,558,1972,1996]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="DAC3F84EFFB3FFFD12CD24AA8112DEC7" box="[463,558,1972,1996]" italics="true" pageId="23" pageNumber="111">Triphysa</emphasis>
</taxonomicName>
: 16 –
<taxonomicName id="2FB75FDFFFB3FFFD117E24AB8058DEC7" authorityName="Staudinger" authorityYear="1901" box="[636,868,1972,1996]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="111" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="biocellata">
<emphasis id="DAC3F84EFFB3FFFD117E24AB8058DEC7" box="[636,868,1972,1996]" italics="true" pageId="23" pageNumber="111">T. nervosa biocellata</emphasis>
</taxonomicName>
, Central Mongolia, near Ulan-Baator (photo by A. Vaganov); 17 –
<taxonomicName id="2FB75FDFFFB3FFFD124624CA8119DEE0" authorityName="Staudinger" authorityYear="1901" box="[324,549,2003,2027]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="111" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="biocellata">
<emphasis id="DAC3F84EFFB3FFFD124624CA8119DEE0" box="[324,549,2003,2027]" italics="true" pageId="23" pageNumber="111">T. nervosa biocellata</emphasis>
</taxonomicName>
(photo by O. Kosterin).
</paragraph>
</caption>
<caption id="BCC874D4FFBCFFF2139524DF809ADEFD" ID-DOI="http://doi.org/10.5281/zenodo.13246770" ID-Zenodo-Dep="13246770" httpUri="https://zenodo.org/record/13246770/files/figure.png" pageId="24" pageNumber="112" startId="24.[151,237,1985,2009]" targetBox="[211,1373,190,1966]" targetPageId="24" targetType="figure">
<paragraph id="E808245CFFBCFFF2139524DF809ADEFD" blockId="24.[151,1437,1984,2038]" pageId="24" pageNumber="112">
<emphasis id="DAC3F84EFFBCFFF2139524DF827DDED2" bold="true" box="[151,321,1985,2009]" pageId="24" pageNumber="112">Figures 18–19.</emphasis>
Habitats of
<taxonomicName id="2FB75FDFFFBCFFF212D324DE810CDED3" authorityName="Zeller" authorityYear="1850" box="[465,560,1984,2008]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="DAC3F84EFFBCFFF212D324DE810CDED3" box="[465,560,1984,2008]" italics="true" pageId="24" pageNumber="112">Triphysa</emphasis>
</taxonomicName>
: 18 –
<taxonomicName id="2FB75FDFFFBCFFF2118224DF805ADED3" authorityName="Grum-Grshimailo" authorityYear="1889" box="[640,870,1984,2008]" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="112" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="tscherskii">
<emphasis id="DAC3F84EFFBCFFF2118224DF805ADED3" box="[640,870,1984,2008]" italics="true" pageId="24" pageNumber="112">T. nervosa tscherskii</emphasis>
</taxonomicName>
, near Yakutsk (photo by A. Burnasheva); 19 –
<taxonomicName id="2FB75FDFFFBCFFF2168524DF827ADEFD" authorityName="Motschoulsky" authorityYear="1866" class="Insecta" family="Nymphalidae" genus="Triphysa" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="112" phylum="Arthropoda" rank="subSpecies" species="nervosa" subSpecies="nervosa">
<emphasis id="DAC3F84EFFBCFFF2168524DF827ADEFD" italics="true" pageId="24" pageNumber="112">T. nervosa nervosa</emphasis>
</taxonomicName>
, E. Yakutiya, near Ust’-Nera vill. (photo by A. Samus’).
</paragraph>
</caption>
<subSubSection id="A0AD77D7FFBDFFF313F423DA82B1D8A2" pageId="25" pageNumber="113" type="materials_examined">
<paragraph id="E808245CFFBDFFF313F423DA82B1D8A2" blockId="25.[151,1437,195,458]" pageId="25" pageNumber="113">
<emphasis id="DAC3F84EFFBDFFF313F423DA82E6D9D5" bold="true" box="[246,474,196,222]" italics="true" pageId="25" pageNumber="113">Material examined</emphasis>
.
<materialsCitation id="58DF2E01FFBDFFF312E823DD86A4D9D5" box="[490,1432,195,222]" collectingDate="1990-05-24" collectionCode="ISEA" collectorName="Male, S. &amp; V. Dubatolov" location="Erzin" pageId="25" pageNumber="113" specimenCount="1" stateProvince="Tyva" typeStatus="holotype">
<typeStatus id="370C9AFEFFBDFFF312E823DD8167D9D5" box="[490,603,195,222]" pageId="25" pageNumber="113" type="holotype">Holotype</typeStatus>
.
<collectorName id="4542418AFFBDFFF3116723DD81F5D9D5" box="[613,713,195,222]" pageId="25" pageNumber="113">Male, S.</collectorName>
<collectingRegion id="2A73EABEFFBDFFF311CD23DA8032D9D5" box="[719,782,196,222]" country="Russia" name="Tyva" pageId="25" pageNumber="113">Tuva</collectingRegion>
,
<quantity id="2F4F89B9FFBDFFF3101923DD8058D9D5" box="[795,868,195,222]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="113" unit="km" value="15.0">15 km</quantity>
WSW
<location id="ED687287FFBDFFF310B923DD80C2D9D5" LSID="urn:lsid:plazi:treatment:601E954AFFB6FFF312A2269E8011D8C1:ED687287FFBDFFF310B923DD80C2D9D5" box="[955,1022,195,222]" name="Erzin" pageId="25" pageNumber="113" stateProvince="Tyva">Erzin</location>
,
<date id="9C09029CFFBDFFF3170823DD87B2D9D5" box="[1034,1166,195,222]" pageId="25" pageNumber="113" value="1990-05-24">
<collectingDate id="8C4DFB74FFBDFFF3170823DD87B2D9D5" box="[1034,1166,195,222]" pageId="25" pageNumber="113" value="1990-05-24">24-V-1990</collectingDate>
</date>
,
<collectorName id="4542418AFFBDFFF3179B23DA8605D9D5" box="[1177,1337,195,222]" pageId="25" pageNumber="113">V. Dubatolov</collectorName>
(
<collectionCode id="8EA6BC99FFBDFFF3164B23DD86ACD9D5" box="[1353,1424,195,222]" pageId="25" pageNumber="113">ISEA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="58DF2E01FFBDFFF3139523F887CCD80B" box="[151,1264,229,256]" collectingDate="1990-05-24" collectionCode="ISEA" collectorName="V. Dubatolov" location="15 km WSW Erzin" pageId="25" pageNumber="113" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Tyva" typeStatus="paratype">
<typeStatus id="370C9AFEFFBDFFF3139523F88231D80B" box="[151,269,230,256]" pageId="25" pageNumber="113" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="FEB1EFD5FFBDFFF3121B23FB8251D80B" box="[281,365,229,256]" count="1" pageId="25" pageNumber="113" type="male">1 male</specimenCount>
,
<specimenCount id="FEB1EFD5FFBDFFF3127B23FB82D3D80B" box="[377,495,229,256]" count="5" pageId="25" pageNumber="113" type="female">5 females</specimenCount>
, S.
<collectingRegion id="2A73EABEFFBDFFF3111E23F88167D80B" box="[540,603,230,256]" country="Russia" name="Tyva" pageId="25" pageNumber="113">Tuva</collectingRegion>
,
<location id="ED687287FFBDFFF3116B23FB806CD80B" LSID="urn:lsid:plazi:treatment:601E954AFFB6FFF312A2269E8011D8C1:ED687287FFBDFFF3116B23FB806CD80B" box="[617,848,229,256]" name="15 km WSW Erzin" pageId="25" pageNumber="113" stateProvince="Tyva">
<quantity id="2F4F89B9FFBDFFF3116B23FB8188D80B" box="[617,692,229,256]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="113" unit="km" value="15.0">15 km</quantity>
WSW Erzin
</location>
,
<date id="9C09029CFFBDFFF3105F23FB80DDD80B" box="[861,993,229,256]" pageId="25" pageNumber="113" value="1990-05-24">
<collectingDate id="8C4DFB74FFBDFFF3105F23FB80DDD80B" box="[861,993,229,256]" pageId="25" pageNumber="113" value="1990-05-24">24-V-1990</collectingDate>
</date>
,
<collectorName id="4542418AFFBDFFF310EC23F887ACD80B" box="[1006,1168,229,256]" pageId="25" pageNumber="113">V. Dubatolov</collectorName>
(
<collectionCode id="8EA6BC99FFBDFFF317A223FB87DBD80B" box="[1184,1255,229,256]" pageId="25" pageNumber="113">ISEA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFBDFFF317FF23FB8178D829" collectingDate="2001-05-20" collectionCode="RYB" collectorName="V. Ivonin" location="Erzin" pageId="25" pageNumber="113" specimenCount="3" specimenCount-male="3" stateProvince="Tyva" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFBDFFF317FF23FB8660D80B" box="[1277,1372,229,256]" count="3" pageId="25" pageNumber="113" type="male">3 males</specimenCount>
, near
<location id="ED687287FFBDFFF31395221983E6D829" LSID="urn:lsid:plazi:treatment:601E954AFFB6FFF312A2269E8011D8C1:ED687287FFBDFFF31395221983E6D829" box="[151,218,263,290]" name="Erzin" pageId="25" pageNumber="113" stateProvince="Tyva">Erzin</location>
,
<date id="9C09029CFFBDFFF313E422198257D829" box="[230,363,263,290]" pageId="25" pageNumber="113" value="2001-05-20">
<collectingDate id="8C4DFB74FFBDFFF313E422198257D829" box="[230,363,263,290]" pageId="25" pageNumber="113" value="2001-05-20">20-V-2001</collectingDate>
</date>
.
<collectorName id="4542418AFFBDFFF31275221682D4D829" box="[375,488,263,290]" pageId="25" pageNumber="113">V. Ivonin</collectorName>
(
<collectionCode id="8EA6BC99FFBDFFF312FB22198107D829" box="[505,571,263,290]" pageId="25" pageNumber="113">RYB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFBDFFF31153221986A4D829" box="[593,1432,263,290]" collectingDate="2010-06-04" collectionCode="RYB" collectorName="R. &amp; A. Yakovlev" location="Erzin" pageId="25" pageNumber="113" specimenCount="22" specimenCount-female="9" specimenCount-male="13" stateProvince="Tyva" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFBDFFF3115322198182D829" box="[593,702,263,290]" count="13" pageId="25" pageNumber="113" type="male">13 males</specimenCount>
,
<specimenCount id="FEB1EFD5FFBDFFF311C92219807DD829" box="[715,833,263,290]" count="9" pageId="25" pageNumber="113" type="female">9 females</specimenCount>
, near
<location id="ED687287FFBDFFF3108A221980F6D829" LSID="urn:lsid:plazi:treatment:601E954AFFB6FFF312A2269E8011D8C1:ED687287FFBDFFF3108A221980F6D829" box="[904,970,263,290]" name="Erzin" pageId="25" pageNumber="113" stateProvince="Tyva">Erzin</location>
,
<date id="9C09029CFFBDFFF310D52219876AD829" box="[983,1110,263,290]" pageId="25" pageNumber="113" value="2010-06-04">
<collectingDate id="8C4DFB74FFBDFFF310D52219876AD829" box="[983,1110,263,290]" pageId="25" pageNumber="113" value="2010-06-04">4-VI-2010</collectingDate>
</date>
,
<collectorName id="4542418AFFBDFFF3176122168600D829" box="[1123,1340,263,290]" pageId="25" pageNumber="113">R. &amp; A. Yakovlev</collectorName>
(
<collectionCode id="8EA6BC99FFBDFFF3164F221986B3D829" box="[1357,1423,263,290]" pageId="25" pageNumber="113">RYB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFBDFFF31395223687EAD848" box="[151,1238,296,323]" collectingDate="1989-06-28" collectionCode="ISEAN" collectorName="V. Zinchenko" location="Mt. Onchalaan" municipality="Erzin distr." pageId="25" pageNumber="113" specimenCount="2" specimenCount-female="2" stateProvince="Tyva" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFBDFFF3139522368232D848" box="[151,270,296,323]" count="2" pageId="25" pageNumber="113" type="female">2 females</specimenCount>
, S.
<collectingRegion id="2A73EABEFFBDFFF3123D22378242D848" box="[319,382,297,323]" country="Russia" name="Tyva" pageId="25" pageNumber="113">Tuva</collectingRegion>
,
<collectingMunicipality id="086CBE26FFBDFFF3128C2236812AD848" box="[398,534,296,323]" pageId="25" pageNumber="113">Erzin distr.</collectingMunicipality>
,
<location id="ED687287FFBDFFF31127223781E5D848" LSID="urn:lsid:plazi:treatment:601E954AFFB6FFF312A2269E8011D8C1:ED687287FFBDFFF31127223781E5D848" box="[549,729,296,323]" municipality="Erzin distr." name="Mt. Onchalaan" pageId="25" pageNumber="113" stateProvince="Tyva">Mt. Onchalaan</location>
,
<date id="9C09029CFFBDFFF311EA2236804AD848" box="[744,886,296,323]" pageId="25" pageNumber="113" value="1989-06-28">
<collectingDate id="8C4DFB74FFBDFFF311EA2236804AD848" box="[744,886,296,323]" pageId="25" pageNumber="113" value="1989-06-28">28-VI-1989</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFBDFFF310B822378762D848" box="[954,1118,296,323]" pageId="25" pageNumber="113">V. Zinchenko</collectorName>
(
<collectionCode id="8EA6BC99FFBDFFF31773223687F2D848" box="[1137,1230,296,323]" pageId="25" pageNumber="113">ISEAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFBDFFF317E7223682C7D86E" collectingDate="1990-05-24" collectionCode="ISEAN" collectorName="V. Zinchenko" location="Mt. Onchalaan" municipality="Erzin distr." pageId="25" pageNumber="113" specimenCount="1" specimenCount-female="1" stateProvince="Tyva" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFBDFFF317E72236866ED848" box="[1253,1362,296,323]" count="1" pageId="25" pageNumber="113" type="female">1 female</specimenCount>
, same locality,
<date id="9C09029CFFBDFFF31200225482BED86E" box="[258,386,330,357]" pageId="25" pageNumber="113" value="1990-05-24">
<collectingDate id="8C4DFB74FFBDFFF31200225482BED86E" box="[258,386,330,357]" pageId="25" pageNumber="113" value="1990-05-24">24-V-1990</collectingDate>
</date>
(
<collectionCode id="8EA6BC99FFBDFFF31294225482CFD86E" box="[406,499,330,357]" pageId="25" pageNumber="113">ISEAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFBDFFF3110922548238D88C" collectingDate="1989-06-01" collectionCode="ISEAN" collectorName="V. Zinchenko" location="15 km W Erzin" pageId="25" pageNumber="113" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tyva" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFBDFFF311092254815DD86E" box="[523,609,330,357]" count="1" pageId="25" pageNumber="113" type="male">1 male</specimenCount>
,
<specimenCount id="FEB1EFD5FFBDFFF31172225481E1D86E" box="[624,733,330,357]" count="1" pageId="25" pageNumber="113" type="female">1 female</specimenCount>
, S.
<collectingRegion id="2A73EABEFFBDFFF3101222558073D86E" box="[784,847,331,357]" country="Russia" name="Tyva" pageId="25" pageNumber="113">Tuva</collectingRegion>
,
<location id="ED687287FFBDFFF3105D2254871ED86E" LSID="urn:lsid:plazi:treatment:601E954AFFB6FFF312A2269E8011D8C1:ED687287FFBDFFF3105D2254871ED86E" box="[863,1058,330,357]" name="15 km W Erzin" pageId="25" pageNumber="113" stateProvince="Tyva">
<quantity id="2F4F89B9FFBDFFF3105D22548090D86E" box="[863,940,330,357]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="113" unit="km" value="15.0">15 km</quantity>
W Erzin
</location>
,
<date id="9C09029CFFBDFFF317332254878DD86E" box="[1073,1201,330,357]" pageId="25" pageNumber="113" value="1989-06-01">
<collectingDate id="8C4DFB74FFBDFFF317332254878DD86E" box="[1073,1201,330,357]" pageId="25" pageNumber="113" value="1989-06-01">1-VI-1989</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFBDFFF317F5225586A0D86E" box="[1271,1436,330,357]" pageId="25" pageNumber="113">V. Zinchenko</collectorName>
(
<collectionCode id="8EA6BC99FFBDFFF3139D227283C0D88C" box="[159,252,364,391]" pageId="25" pageNumber="113">ISEAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58DF2E01FFBDFFF31210227282B5D8A2" collectingDate="1990-06-24" collectionCode="ISEAN" collectorName="V. Zinchenko" county="Erzin distr." location="Sook-Toj" municipality="Mts. Onchalaan" pageId="25" pageNumber="113" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Tyva" typeStatus="paratype">
<specimenCount id="FEB1EFD5FFBDFFF312102272824CD88C" box="[274,368,364,391]" count="2" pageId="25" pageNumber="113" type="male">2 males</specimenCount>
,
<specimenCount id="FEB1EFD5FFBDFFF3127C227282CFD88C" box="[382,499,364,391]" count="4" pageId="25" pageNumber="113" type="female">4 females</specimenCount>
, S.
<collectingRegion id="2A73EABEFFBDFFF3112222738163D88C" box="[544,607,365,391]" country="Russia" name="Tyva" pageId="25" pageNumber="113">Tuva</collectingRegion>
,
<collectingCounty id="01695CD0FFBDFFF3116E227281CFD88C" box="[620,755,364,391]" pageId="25" pageNumber="113">Erzin distr.</collectingCounty>
, near
<collectingMunicipality id="086CBE26FFBDFFF31039227380CAD88C" box="[827,1014,364,391]" pageId="25" pageNumber="113">Mts. Onchalaan</collectingMunicipality>
and
<location id="ED687287FFBDFFF3173022728799D88C" LSID="urn:lsid:plazi:treatment:601E954AFFB6FFF312A2269E8011D8C1:ED687287FFBDFFF3173022728799D88C" box="[1074,1189,364,391]" county="Erzin distr." municipality="Mts. Onchalaan" name="Sook-Toj" pageId="25" pageNumber="113" stateProvince="Tyva">Sook-Toj</location>
,
<date id="9C09029CFFBDFFF317B022728603D88C" box="[1202,1343,364,391]" pageId="25" pageNumber="113" value="1990-06-24">
<collectingDate id="8C4DFB74FFBDFFF317B022728603D88C" box="[1202,1343,364,391]" pageId="25" pageNumber="113" value="1990-06-24">24-VI-1990</collectingDate>
</date>
, leg.
<collectorName id="4542418AFFBDFFF3168222738228D8A2" pageId="25" pageNumber="113">V. Zinchenko</collectorName>
(
<collectionCode id="8EA6BC99FFBDFFF31226229082BDD8A2" box="[292,385,398,425]" pageId="25" pageNumber="113">ISEAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AD77D7FFBDFFF313F422AE8011D8C1" box="[246,813,431,458]" pageId="25" pageNumber="113" type="distribution">
<paragraph id="E808245CFFBDFFF313F422AE8011D8C1" blockId="25.[151,1437,195,458]" box="[246,813,431,458]" pageId="25" pageNumber="113">
<emphasis id="DAC3F84EFFBDFFF313F422AE82B5D8C1" bold="true" box="[246,393,432,458]" italics="true" pageId="25" pageNumber="113">Distribution</emphasis>
:
<collectingCountry id="90A064CCFFBDFFF3129A22B182D5D8C1" box="[408,489,431,458]" name="Russia" pageId="25" pageNumber="113">Russia</collectingCountry>
, S.
<collectingRegion id="2A73EABEFFBDFFF3111622AE816CD8C1" box="[532,592,432,458]" country="Russia" name="Tyva" pageId="25" pageNumber="113">Tuva</collectingRegion>
(Ubsunur Valley).
</paragraph>
</subSubSection>
</treatment>
</document>
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