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<document id="97567C1B0BB7E9C7988B092DBF3A2A05" ID-CLB-Dataset="7214" ID-DOI="10.11646/zootaxa.4963.1.2" ID-GBIF-Dataset="78e78beb-b1ed-43a4-84c7-8180e61463c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4711249" ID-ZooBank="E93AB178-3419-498B-9563-A704ABAE5EB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618560083953" checkinUser="plazi" docAuthor="Bálint, Zsolt, Boyer, Pierre, Cerdeña, José, Larico, Jackie Farfán, Brudecka, Jadwiga Lorenc-, Prieto, Carlos &amp; Pyrcz, Tomasz W." docDate="2021" docId="D5008F50FFFCB31BFF76FA3F9408FE8C" docLanguage="en" docName="zootaxa.4963.1.2.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Johnsonita subcunicula Balint, Cerdena &amp; Pyrcz 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="33" masterDocId="2939F728FFE9B30CFFE1FFA69629FFCD" masterDocTitle="Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)" masterLastPageNumber="57" masterPageNumber="11" pageNumber="32" updateTime="1698937134920" updateUser="ExternalLinkService">
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<mods:title id="AEA9B3C9127B105B8FABB0D967DAB0F7">Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)</mods:title>
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<mods:namePart id="228C4B4545212DC705FD27DC277176E6">Bálint, Zsolt</mods:namePart>
<mods:nameIdentifier id="7D19343B13679EC27694B38E79EED934" type="ORCID">0000-0001-8174-878X</mods:nameIdentifier>
<mods:affiliation id="C5357E04944C97CA22879EFEFAB7DC6B">Hungarian Natural History Museum, Department of Zoology, H- 1088 Budapest, Baross utca 13, Hungary. balint. zsolt @ nhmus. hu; https: // orcid. org / 0000 - 0001 - 8174 - 878 X</mods:affiliation>
<mods:nameIdentifier id="9FA15E66ECCEDB169D09119298541E4B" type="email">balint.zsolt@nhmus.hu</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="55D04AFF1132EA3E4D1C8192A8A03EFD">Boyer, Pierre</mods:namePart>
<mods:affiliation id="0A535A50D79F81F685CE0422F2285FD5">Lotissement l’Horizon, 13610 Le Puy Sainte Réparade, France.</mods:affiliation>
</mods:name>
<mods:name id="8A639B73338CBCB7D5D1E38EFBB6EC73" type="personal">
<mods:role id="0D047F91F1A2951CE9DBF5A80842257B">
<mods:roleTerm id="C8C111F6F03F8C4A984FD6643E4CAC20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="237737D1ACFD5AF872B54A1366A04F8D">Cerdeña, José</mods:namePart>
<mods:nameIdentifier id="39C8E0D249BF729AD08BEE640E0EA288" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="5E4448FA115E1DDFA9F3DC7739D4C771">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="6784720DAD651FBB044208C65B9EB51D" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="1F23E3F373D4F54AA644F4B5FA82465D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35B896F485139059AEC28305A703206B">Larico, Jackie Farfán</mods:namePart>
<mods:nameIdentifier id="3E7EE2923480C6E8CCBD9F5A92151D08" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="612D1A2F1877C36B57A7811BC6ABD580">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="369AF2FCB1D4E5A06AE3D9338EE1AAD2" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="00EEDD7EE8402F9DDEE0C280F69D9895">Brudecka, Jadwiga Lorenc-</mods:namePart>
<mods:affiliation id="E005C235FD3B2735A175203F2EE7125B">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:namePart id="670E85738CE655A98E98CF74D31A6783">Prieto, Carlos</mods:namePart>
<mods:affiliation id="E6D45587B213682226609B6E400EA2CC">Corporación Universitaria Autónoma del Cauca, Popayán, Colombia.</mods:affiliation>
</mods:name>
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<mods:namePart id="35593930DA2C110150CE778D0C492856">Pyrcz, Tomasz W.</mods:namePart>
<mods:affiliation id="D5494935ED3EB6D16F80D75D27E57246">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:title id="2CEE9E655B6A3584CACC870F716B28E3">Zootaxa</mods:title>
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<mods:date id="0F74C351C994EA88F5323C51AE5C8EA8">2021</mods:date>
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<mods:number id="980B9FCAB3C86C6C30A04DB87B1F82C2">2021-04-16</mods:number>
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<mods:number id="BA092D2D29DA2A6FF0E41A767F6984D7">4963</mods:number>
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<mods:classification id="2FAA7F33E7C579F73A5AD4036D3E5BDD">journal article</mods:classification>
<mods:identifier id="3E34DEC0922111CDC32D8ABA9316B0F8" type="CLB-Dataset">7214</mods:identifier>
<mods:identifier id="B9BB6B34B46CB5821791B64BC4179326" type="DOI">10.11646/zootaxa.4963.1.2</mods:identifier>
<mods:identifier id="899F5A7170ED9849005C083571EB4A10" type="GBIF-Dataset">78e78beb-b1ed-43a4-84c7-8180e61463c5</mods:identifier>
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<treatment id="D5008F50FFFCB31BFF76FA3F9408FE8C" ID-DOI="http://doi.org/10.5281/zenodo.4696435" ID-GBIF-Taxon="181531936" ID-Zenodo-Dep="4696435" LSID="urn:lsid:plazi:treatment:D5008F50FFFCB31BFF76FA3F9408FE8C" httpUri="http://treatment.plazi.org/id/D5008F50FFFCB31BFF76FA3F9408FE8C" lastPageId="23" lastPageNumber="33" pageId="21" pageNumber="32">
<subSubSection id="15B36DCDFFFCB319FF76FA3F956EFA7E" box="[151,839,1432,1460]" pageId="21" pageNumber="32" type="nomenclature">
<paragraph id="5D163E46FFFCB319FF76FA3F956EFA7E" blockId="21.[151,839,1432,1494]" box="[151,839,1432,1460]" pageId="21" pageNumber="32">
<heading id="065E892AFFFCB319FF76FA3F956EFA7E" bold="true" box="[151,839,1432,1460]" fontSize="11" level="1" pageId="21" pageNumber="32" reason="1">
<emphasis id="6FDDE254FFFCB319FF76FA3F956EFA7E" bold="true" box="[151,839,1432,1460]" pageId="21" pageNumber="32">
<taxonomicName id="9AA945C5FFFCB319FF76FA3F94D1FA7E" authority="Balint, Cerdena &amp; Pyrcz" authorityName="Balint, Cerdena &amp; Pyrcz" authorityYear="2021" box="[151,760,1432,1460]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="32" phylum="Arthropoda" rank="species" species="subcunicula" status="sp. nov.">
<emphasis id="6FDDE254FFFCB319FF76FA3F9793FA79" bold="true" box="[151,442,1433,1460]" italics="true" pageId="21" pageNumber="32">Johnsonita subcunicula</emphasis>
Bálint, Cerdeña &amp; Pyrcz
</taxonomicName>
,
<taxonomicNameLabel id="74EE5F2FFFFCB319FCE4FA3F956EFA7E" box="[773,839,1433,1459]" pageId="21" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="15B36DCDFFFCB31BFF76FA1A9408FE8C" lastPageId="23" lastPageNumber="34" pageId="21" pageNumber="32" type="description">
<paragraph id="5D163E46FFFCB319FF76FA1A97F4FA1B" blockId="21.[151,839,1432,1494]" box="[151,477,1468,1494]" pageId="21" pageNumber="32">
(
<figureCitation id="C59222C3FFFCB319FF7EFA1A96DFFA1B" box="[159,246,1468,1494]" captionStart="FIGURE 13" captionStartId="22.[151,250,1414,1438]" captionTargetBox="[160,1391,226,1357]" captionTargetId="figure-23@22.[151,1436,181,1390]" captionTargetPageId="22" captionText="FIGURE 13. Johnsonita subcunicula Bálint, Cerdeña &amp; Pyrcz, sp. n. male imagines (left column: recto, right column: verso). A–B: holotype (MUSM), C–D: paratype (CEP–MZUJ) (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711277" httpUri="https://zenodo.org/record/4711277/files/figure.png" pageId="21" pageNumber="32">Figs 13</figureCitation>
,
<figureCitation id="C59222C3FFFCB319FEE3FA1A977CFA1B" box="[258,341,1468,1494]" captionStart="FIGURE 19" captionStartId="29.[151,250,1909,1933]" captionTargetBox="[313,1283,191,1883]" captionTargetId="figure-18@29.[200,1388,181,1885]" captionTargetPageId="29" captionText="FIGURE 19. Johnsonita genitalia armatures; entire male organs in lateral (left column) and ventral (right column) views.A–B: J. subcunicula, Peru, Huánuco, San Pedro de Carpish; C–D: J. turquisca, Ecuador, Napo, Papallacta, Chalpi Grande; E–F: J. oxalida, Peru, Pasco, Oxapampa; G–H: J. zubkova, Peru, Cuzco, Abra Acjanaco vers Pillcopata (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711295" httpUri="https://zenodo.org/record/4711295/files/figure.png" pageId="21" pageNumber="32">19A–B</figureCitation>
,
<figureCitation id="C59222C3FFFCB319FE80FA1A9785FA1B" box="[353,428,1468,1494]" captionStart="FIGURE 24" captionStartId="34.[151,250,1502,1526]" captionTargetBox="[400,1168,186,1466]" captionTargetId="figure-23@34.[396,1192,181,1478]" captionTargetPageId="34" captionText="FIGURE 24. Johnsonita mail traits in dorsal hindwing (left column) and ventral forewing subcubital patches (right column) indicated by arrow. A–B: J. oxalida, C–D: J. johnbanksi, E–F: J. subcunicula, G: J. auda." figureDoi="http://doi.org/10.5281/zenodo.4711307" httpUri="https://zenodo.org/record/4711307/files/figure.png" pageId="21" pageNumber="32">24E–F</figureCitation>
,
<figureCitation id="C59222C3FFFCB319FE56FA1A97FDFA1B" box="[439,468,1468,1494]" captionStart="FIGURE 31" captionStartId="40.[151,250,1927,1951]" captionTargetBox="[187,1436,207,1903]" captionTargetId="figure-23@40.[151,1436,181,1903]" captionTargetPageId="40" captionText="FIGURE 31. Johnsonita distributions based on museum material examinded. J. auda = red circle, J. iacinta = blue diamond, J. subcunicula = green square, J. turquisca = yellow trinagle, J. zubkova = brown trinagle." figureDoi="http://doi.org/10.5281/zenodo.4711327" httpUri="https://zenodo.org/record/4711327/files/figure.png" pageId="21" pageNumber="32">31</figureCitation>
)
</paragraph>
<paragraph id="5D163E46FFFCB319FF76F9A49796F96A" blockId="21.[151,1437,1537,1981]" pageId="21" pageNumber="32">
<emphasis id="6FDDE254FFFCB319FF76F9A49715F9D6" box="[151,316,1537,1563]" italics="true" pageId="21" pageNumber="32">Type material.</emphasis>
<materialsCitation id="EDC1341BFFFCB319FEA4F9A792CCF9AC" ID-GBIF-Occurrence="3083794336" collectingDate="2019-06-20" collectionCode="T, MUSM" collectorName="J. Cerdena Guiterrez &amp; T. Pyrcz " country="Peru" elevation="2425" location="San Pedro de Carpish" municipality="Carpish" pageId="21" pageNumber="32" specimenCount="1" stateProvince="Huanuco" typeStatus="holotype">
<typeStatus id="821280E4FFFCB319FEA4F9A79784F9D6" box="[325,429,1537,1563]" pageId="21" pageNumber="32" type="holotype">Holotype</typeStatus>
male in almost perfect condition (half of left antenna missing), set dorsally, with data:
<collectingCountry id="25BE7ED6FFFCB319FF76F98396C8F9F2" box="[151,225,1573,1599]" name="Peru" pageId="21" pageNumber="32">PERU</collectingCountry>
, Depto.
<collectingRegion id="9F6DF0A4FFFCB319FEA4F9829784F9F3" box="[325,429,1572,1598]" country="Peru" name="Huanuco" pageId="21" pageNumber="32">Huánuco</collectingRegion>
,
<collectingMunicipality id="BD72A43CFFFCB319FE5DF982943FF9F3" box="[444,534,1572,1598]" pageId="21" pageNumber="32">Carpish</collectingMunicipality>
,
<location id="5876689DFFFCB319FDC5F982950BF9F3" LSID="urn:lsid:plazi:treatment:D5008F50FFFCB31BFF76FA3F9408FE8C:5876689DFFFCB319FDC5F982950BF9F3" box="[548,802,1572,1598]" country="Peru" municipality="Carpish" name="San Pedro de Carpish" pageId="21" pageNumber="32" stateProvince="Huanuco">San Pedro de Carpish</location>
,
<quantity id="9A5193A3FFFCB319FCD1F98295F9F9F2" box="[816,976,1572,1599]" metricMagnitude="3" metricUnit="m" metricValue="2.425" metricValueMax="2.45" metricValueMin="2.4" pageId="21" pageNumber="32" unit="m" value="2425.0" valueMax="2450.0" valueMin="2400.0">
<elevation id="D684D975FFFCB319FCD1F98295F9F9F2" box="[816,976,1572,1599]" metricMagnitude="3" metricUnit="m" metricValue="2.425" metricValueMax="2.45" metricValueMin="2.4" pageId="21" pageNumber="32" unit="m" value="2425.0" valueMax="2450.0" valueMin="2400.0">2400–2450 m</elevation>
</quantity>
,
<date id="29171886FFFCB319FC3EF9829248F9F3" box="[991,1121,1572,1598]" pageId="21" pageNumber="32" value="2019-06-20">
<collectingDate id="3953E16EFFFCB319FC3EF9829248F9F3" box="[991,1121,1572,1598]" pageId="21" pageNumber="32" value="2019-06-20">20.VI.2019</collectingDate>
</date>
,
<collectorName id="F05C5B90FFFCB319FB91F9839374F9F3" box="[1136,1373,1572,1599]" pageId="21" pageNumber="32">J. Cerdeña Guitérrez</collectorName>
&amp;
<collectionCode id="3BB8A683FFFCB319FA67F98393B1F9F2" box="[1414,1432,1573,1599]" name="Tavera, Department of Geology and Geophysics" pageId="21" pageNumber="32">T</collectionCode>
.
<collectorName id="F05C5B90FFFCB319FF76F9E196FEF9AC" box="[151,215,1607,1633]" pageId="21" pageNumber="32">Pyrcz</collectorName>
leg. (currently in the collection of
<collectorName id="F05C5B90FFFCB319FD86F9E194EDF9AC" box="[615,708,1607,1633]" pageId="21" pageNumber="32">Cerdeña</collectorName>
(
<collectingRegion id="9F6DF0A4FFFCB319FD35F9E19568F9AC" box="[724,833,1607,1633]" country="Peru" name="Arequipa" pageId="21" pageNumber="32">Arequipa</collectingRegion>
,
<collectorName id="F05C5B90FFFCB319FCAFF9E195AFF9AC" box="[846,902,1607,1633]" pageId="21" pageNumber="32">Peru</collectorName>
), will be deposited in
<collectionCode id="3BB8A683FFFCB319FB63F9E192F4F9AC" box="[1154,1245,1607,1633]" pageId="21" pageNumber="32">MUSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EDC1341BFFFCB319FB10F9E19792F96A" ID-GBIF-Occurrence="3083794354" collectionCode="CEP, MZUJ" collectorName="K. Florczyk" country="Peru" location="Arequipa" pageId="21" pageNumber="32" specimenCount="2" stateProvince="Arequipa" typeStatus="paratype">
<typeStatus id="821280E4FFFCB319FB10F9E1937AF9AC" box="[1265,1363,1607,1633]" pageId="21" pageNumber="32" type="paratype">Paratype</typeStatus>
males with
<typeStatus id="821280E4FFFCB319FF35F9CF971FF94E" box="[212,310,1641,1667]" pageId="21" pageNumber="32" type="holotype">holotype</typeStatus>
data (n=2) deposited in
<collectionCode id="3BB8A683FFFCB319FDBBF9CF94A4F94E" box="[602,653,1641,1667]" pageId="21" pageNumber="32">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFFCB319FD7AF9CF94CDF94E" box="[667,740,1641,1667]" pageId="21" pageNumber="32">MZUJ</collectionCode>
(dissected: prep. gen. no. 1755/
<collectorName id="F05C5B90FFFCB319FBBEF9CF92C1F949" box="[1119,1256,1641,1668]" pageId="21" pageNumber="32">K. Florczyk</collectorName>
) and collection Cerdeña (
<collectingRegion id="9F6DF0A4FFFCB319FEE2F92A9759F96B" box="[259,368,1676,1702]" country="Peru" name="Arequipa" pageId="21" pageNumber="32">Arequipa</collectingRegion>
, Peru)
</materialsCitation>
.
</paragraph>
<paragraph id="5D163E46FFFCB319FF26F9099447F870" blockId="21.[151,1437,1537,1981]" pageId="21" pageNumber="32">
<emphasis id="6FDDE254FFFCB319FF26F909977AF905" box="[199,339,1711,1736]" italics="true" pageId="21" pageNumber="32">Description.</emphasis>
Male. Wings. FW costa length:
<quantity id="9A5193A3FFFCB319FD56F909952CF904" box="[695,773,1711,1737]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="21" pageNumber="32" unit="mm" value="14.0">14 mm</quantity>
(
<typeStatus id="821280E4FFFCB319FCF3F9099553F907" box="[786,890,1711,1738]" pageId="21" pageNumber="32" type="holotype">holotype</typeStatus>
); male dorsal wing surface bright blue with black border slightly widening towards apex; HW dorsal surface postbasal and medial area with a large androconial patch along costa; ventral wings surface medial pattern composed of irregular dashes, dark basally and gleaming marginally, HW costa unusually highly bent covering the FW scent patch. Genitalia. Male valva with high but rounded costal margin flip, valval terminus long and stout, uncus strong, falces relatively thin and from below supported by a membranous diaphragma, capsule thin and narrow with a relatively large saccus in dorso/ventral aspect, aedeagus apex upturned in dorsal view. Androconia present in ventral FW as a wide black spot below the cubital vein in postbasal and medial area. Female not known.
</paragraph>
<caption id="09D66ECEFFFFB31AFF76FA2095BAFA0E" ID-DOI="http://doi.org/10.5281/zenodo.4711277" ID-Zenodo-Dep="4711277" httpUri="https://zenodo.org/record/4711277/files/figure.png" pageId="22" pageNumber="33" startId="22.[151,250,1414,1438]" targetBox="[160,1391,226,1357]" targetPageId="22">
<paragraph id="5D163E46FFFFB31AFF76FA2095BAFA0E" blockId="22.[151,1437,1414,1475]" pageId="22" pageNumber="33">
<emphasis id="6FDDE254FFFFB31AFF76FA209709FA53" bold="true" box="[151,288,1414,1438]" pageId="22" pageNumber="33">FIGURE 13.</emphasis>
<taxonomicName id="9AA945C5FFFFB31AFEC6FA219533FA52" authority="Balint, Cerdena &amp; Pyrcz" authorityName="Balint, Cerdena &amp; Pyrcz" authorityYear="2021" box="[295,794,1415,1439]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="species" species="subcunicula" status="sp. nov.">
<emphasis id="6FDDE254FFFFB31AFEC6FA21943EFA52" box="[295,535,1415,1439]" italics="true" pageId="22" pageNumber="33">Johnsonita subcunicula</emphasis>
Bálint, Cerdeña &amp; Pyrcz
</taxonomicName>
,
<taxonomicNameLabel id="74EE5F2FFFFFB31AFCC4FA219575FA53" box="[805,860,1415,1438]" pageId="22" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
male imagines (left column: recto, right column: verso). A–B: holotype (MUSM), C–D: paratype (CEP–MZUJ) (scale bar: 10 mm).
</paragraph>
</caption>
<paragraph id="5D163E46FFFFB31AFF26FA549708F9BA" blockId="22.[151,1437,1521,2016]" pageId="22" pageNumber="33">
<emphasis id="6FDDE254FFFFB31AFF26FA549769F9C6" box="[199,320,1522,1547]" italics="true" pageId="22" pageNumber="33">Diagnosis.</emphasis>
The species is easily distinguished from other congeners on the basis of the distinctive male dorsal colouration and the androconia, which is associated with an unusually highly bent HW costa. The blue of
<taxonomicName id="9AA945C5FFFFB31AFACAF9B093B5F9E2" authorityName="Balint, Lorenc-Brudecka, &amp; Pyrcz" authorityYear="2021" box="[1323,1436,1558,1583]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="species" species="ianusca">
<emphasis id="6FDDE254FFFFB31AFACAF9B093B5F9E2" box="[1323,1436,1558,1583]" italics="true" pageId="22" pageNumber="33">J. ianusca</emphasis>
</taxonomicName>
is similar, but somewhat darker and the dorsal HW scent patch unique in the genus distinguishes immediately the two species.
</paragraph>
<paragraph id="5D163E46FFFFB31AFF26F927930CF956" blockId="22.[151,1437,1521,2016]" box="[199,1317,1665,1692]" pageId="22" pageNumber="33">
<emphasis id="6FDDE254FFFFB31AFF26F9279771F956" box="[199,344,1665,1691]" italics="true" pageId="22" pageNumber="33">Distribution.</emphasis>
Geographical:
<collectingCountry id="25BE7ED6FFFFB31AFDE5F9279465F956" box="[516,588,1665,1691]" name="Peru" pageId="22" pageNumber="33">PERU</collectingCountry>
(
<collectingRegion id="9F6DF0A4FFFFB31AFDBBF92794EBF956" box="[602,706,1665,1691]" country="Peru" name="Huanuco" pageId="22" pageNumber="33">Huánuco</collectingRegion>
) (
<figureCitation id="C59222C3FFFFB31AFD38F9279504F956" box="[729,813,1665,1691]" captionStart="FIGURE 31" captionStartId="40.[151,250,1927,1951]" captionTargetBox="[187,1436,207,1903]" captionTargetId="figure-23@40.[151,1436,181,1903]" captionTargetPageId="40" captionText="FIGURE 31. Johnsonita distributions based on museum material examinded. J. auda = red circle, J. iacinta = blue diamond, J. subcunicula = green square, J. turquisca = yellow trinagle, J. zubkova = brown trinagle." figureDoi="http://doi.org/10.5281/zenodo.4711327" httpUri="https://zenodo.org/record/4711327/files/figure.png" pageId="22" pageNumber="33">Fig. 31</figureCitation>
); spatial:
<quantity id="9A5193A3FFFFB31AFC7BF9279262F956" box="[922,1099,1665,1692]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="22" pageNumber="33" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">2400–2500 ma</quantity>
.s.l; temporal: June.
</paragraph>
<paragraph id="5D163E46FFFFB31AFF26F9039277F8CA" blockId="22.[151,1437,1521,2016]" pageId="22" pageNumber="33">
<emphasis id="6FDDE254FFFFB31AFF26F903976CF972" box="[199,325,1701,1727]" italics="true" pageId="22" pageNumber="33">Etymology.</emphasis>
The species is named after the collecting site of the
<typeStatus id="821280E4FFFFB31AFC6EF9039596F972" box="[911,959,1701,1727]" pageId="22" pageNumber="33">type</typeStatus>
specimens. The Latin word “subcunicula” means “under the tunnel”, since the specimens were collected only a couple of hundred metres away from the Carpish tunnel. The species name “subcunicula” is treated as a female noun in apposition.
</paragraph>
<paragraph id="5D163E46FFFFB31BFF26F8B49408FE8C" blockId="22.[151,1437,1521,2016]" lastBlockId="23.[151,1437,151,321]" lastPageId="23" lastPageNumber="34" pageId="22" pageNumber="33">
<emphasis id="6FDDE254FFFFB31AFF26F8B4972DF8E6" box="[199,260,1810,1835]" italics="true" pageId="22" pageNumber="33">Note.</emphasis>
The known specimens were collected while tree topping, there were interactions observed between the males of this species and other
<taxonomicName id="9AA945C5FFFFB31AFDE4F89394ACF882" authorityName="Leach" authorityYear="1815" box="[517,645,1845,1871]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="family">Lycaenidae</taxonomicName>
(
<taxonomicName id="9AA945C5FFFFB31AFD76F89395A4F882" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1819" box="[663,909,1845,1871]" class="Insecta" family="Lycaenidae" genus="Atlides" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFFB31AFD76F89394CCF882" box="[663,741,1845,1871]" italics="true" pageId="22" pageNumber="33">Atlides</emphasis>
Hübner, 1819
</taxonomicName>
;
<taxonomicName id="9AA945C5FFFFB31AFC7DF8909289F882" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1819" box="[924,1184,1845,1871]" class="Insecta" family="Lycaenidae" genus="Brangas" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFFB31AFC7DF89095D2F882" box="[924,1019,1846,1871]" italics="true" pageId="22" pageNumber="33">Brangas</emphasis>
Hübner, 1819
</taxonomicName>
and
<taxonomicName id="9AA945C5FFFFB31AFB3AF89396F9F8BE" authority="Hubner, 1818" authorityName="Hubner" authorityYear="1818" class="Insecta" family="Lycaenidae" genus="Theritas" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFFB31AFB3AF8939311F882" box="[1243,1336,1845,1871]" italics="true" pageId="22" pageNumber="33">Theritas</emphasis>
Hübner, 1818
</taxonomicName>
) found in the same spot. This is a species with unique characters in the genus suggesting a peculiar position in classification and phylogeny. The male HW shape with the highly bent costa is unique in the genus, but a widely distributed trait in the tribe. It is always associated with an extensive scent patch situated in the dorsal wing surface (for example in the genera
<emphasis id="6FDDE254FFFFB31AFE21F8639417F812" box="[448,574,1989,2015]" italics="true" pageId="22" pageNumber="33">Annamaria</emphasis>
d’Abrera &amp;
<bibRefCitation id="393843B7FFFFB31AFD2AF8639579F82D" author="Balint" box="[715,848,1989,2016]" firstAuthor="Balint" journalOrPublisher="Systematics and Biodiversity" pageId="22" pageNumber="33" pagination="255 - 373" part="2" refId="ref21503" refString="Balint, Zs. (2005) Frederick W. Goodson and his contribution to the taxonomy of neotropical hairstreak butterflies (Lepidoptera: Lycaenidae: Eumaeini). Systematics and Biodiversity, 2 (4), 255 - 373. https: // doi. org / 10.1017 / S 1477200004001550" title="Frederick W. Goodson and his contribution to the taxonomy of neotropical hairstreak butterflies (Lepidoptera: Lycaenidae: Eumaeini)" type="journal article" year="2005">Bálint 2005</bibRefCitation>
;
<taxonomicName id="9AA945C5FFFFB31AFCBAF863936AF812" authority="Johnson, Kruse &amp; Kroenlein, 1997" authorityName="Johnson, Kruse &amp; Kroenlein" authorityYear="1997" box="[859,1347,1989,2015]" class="Insecta" family="Lycaenidae" genus="Laothus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFFB31AFCBAF8639236F812" box="[859,1055,1989,2015]" italics="true" pageId="22" pageNumber="33">
Laothus
<bibRefCitation id="393843B7FFFFB31AFC5AF8639236F812" author="Johnson, Kruse &amp; Kroenlein" box="[955,1055,1989,2015]" firstAuthor="Johnson" journalOrPublisher="Revista de Theclinae Colombianos" pageId="22" pageNumber="33" pagination="1 - 37" part="2" refId="ref22931" refString="Johnson, K., Kruse, J. &amp; Kroenlein, K. R. (1997) The Macusiina, a New Infratribe of the Eumaeini with Description of Ten New Genera (Lycaenidae). Revista de Theclinae Colombianos, 2 (13), i - ii + 1 - 37." title="The Macusiina, a New Infratribe of the Eumaeini with Description of Ten New Genera (Lycaenidae)" type="journal article" year="1997">Johnson,</bibRefCitation>
</emphasis>
Kruse &amp; Kroenlein, 1997
</taxonomicName>
;,
<emphasis id="6FDDE254FFFFB31BFAB2F8639690FF7C" italics="true" lastPageId="23" lastPageNumber="34" pageId="22" pageNumber="33">Lathe- cla</emphasis>
<bibRefCitation id="393843B7FFFEB31BFF20FF31974CFF7C" author="Robbins" box="[193,357,151,177]" editor="Lamas, G." firstAuthor="Robbins" journalOrPublisher="Scientific Publishers, Gainesville, Florida" pageId="23" pageNumber="34" pagination="pp. xxiv - xxx" refId="ref23260" refString="Robbins, R. K. (2004 a) Introduction to the Checklist of Eumaeini (Lycaenidae). In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, Florida, pp. xxiv - xxx." title="Introduction to the Checklist of Eumaeini (Lycaenidae)" type="book chapter" volumeTitle="Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B" year="2004">Robbins, 2004</bibRefCitation>
,
<taxonomicName id="9AA945C5FFFEB31BFE92FF319454FF7C" authority="Balint, 2014" authorityName="Balint" authorityYear="2014" box="[371,637,151,177]" class="Insecta" family="Lycaenidae" genus="Meridaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFEB31BFE92FF3197CFFF7C" box="[371,486,151,177]" italics="true" pageId="23" pageNumber="34">Meridaria</emphasis>
Bálint, 2014
</taxonomicName>
;
<taxonomicName id="9AA945C5FFFEB31BFD69FF3192D2FF7C" authority="Johnson, Austin, Le Crom &amp; Salazar, 1997" authorityName="Johnson, Austin, Le Crom &amp; Salazar" authorityYear="1997" box="[648,1275,151,177]" class="Insecta" family="Lycaenidae" genus="Strephonota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFEB31BFD69FF319539FF7C" box="[648,784,151,177]" italics="true" pageId="23" pageNumber="34">Strephonota</emphasis>
Johnson, Austin, Le Crom &amp; Salazar, 1997
</taxonomicName>
and
<taxonomicName id="9AA945C5FFFEB31BFAD3FF319762FF18" authority="Hewitson, 1865" authorityName="Hewitson" authorityYear="1865" class="Insecta" family="Lycaenidae" genus="Trichonis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFEB31BFAD3FF3193B5FF7C" box="[1330,1436,151,177]" italics="true" pageId="23" pageNumber="34">Trichonis</emphasis>
Hewitson, 1865
</taxonomicName>
). The male genitalia with a long membranous apical valval process and upturned aedeagus terminus is also unique in the genus. Another interesting feature is the presence of the membraneous diaphragma. Enough to say that this species presents some unusual characters for
<taxonomicName id="9AA945C5FFFEB31BFCA1FEA59592FED0" authorityName="Salazar &amp; Constantino" authorityYear="1995" box="[832,955,259,285]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="6FDDE254FFFEB31BFCA1FEA59592FED0" box="[832,955,259,285]" italics="true" pageId="23" pageNumber="34">Johnsonita</emphasis>
</taxonomicName>
and we do not rule out that a phylogenetic study will alter its generic position.
</paragraph>
</subSubSection>
</treatment>
</document>
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