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<document id="37F23C47F7D42E4B8877B64B0DD9D5BF" ID-CLB-Dataset="7214" ID-DOI="10.11646/zootaxa.4963.1.2" ID-GBIF-Dataset="78e78beb-b1ed-43a4-84c7-8180e61463c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4711249" ID-ZooBank="E93AB178-3419-498B-9563-A704ABAE5EB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618560083953" checkinUser="plazi" docAuthor="Bálint, Zsolt, Boyer, Pierre, Cerdeña, José, Larico, Jackie Farfán, Brudecka, Jadwiga Lorenc-, Prieto, Carlos &amp; Pyrcz, Tomasz W." docDate="2021" docId="D5008F50FFE7B303FF76FAAA92E8FF34" docLanguage="en" docName="zootaxa.4963.1.2.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Johnsonita ianusca Balint, Lorenc-Brudecka, &amp; Pyrcz 2021, sp. n." docType="treatment" docVersion="6" lastPageNumber="26" masterDocId="2939F728FFE9B30CFFE1FFA69629FFCD" masterDocTitle="Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)" masterLastPageNumber="57" masterPageNumber="11" pageNumber="25" updateTime="1698937134920" updateUser="ExternalLinkService">
<mods:mods id="92C754CA2DC88D9A88CC5CFAC7F1BB4E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6DC39A21E7A8C5079FA8288D15A8167C">
<mods:title id="0BA7254870DDD3FEFF1381966DEFDD21">Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)</mods:title>
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<mods:roleTerm id="D6D307E9E9CBCCA06A21843EB483BE15">Author</mods:roleTerm>
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<mods:namePart id="87A88EA5AF02FF3D1740B49635082768">Bálint, Zsolt</mods:namePart>
<mods:nameIdentifier id="691FD40488ECD24148E1E589B2AF911F" type="ORCID">0000-0001-8174-878X</mods:nameIdentifier>
<mods:affiliation id="78E90B88E1FEF5161A890832B6DDE6B5">Hungarian Natural History Museum, Department of Zoology, H- 1088 Budapest, Baross utca 13, Hungary. balint. zsolt @ nhmus. hu; https: // orcid. org / 0000 - 0001 - 8174 - 878 X</mods:affiliation>
<mods:nameIdentifier id="F4595F7D28B7C9506EE9169009E70C9C" type="email">balint.zsolt@nhmus.hu</mods:nameIdentifier>
</mods:name>
<mods:name id="E7AFAED338A0D1023E14416D9A0FE575" type="personal">
<mods:role id="4DB97F05C31570C0274B715448621918">
<mods:roleTerm id="FD27B6AABF4C02ABCE5335407E5DDDE8">Author</mods:roleTerm>
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<mods:namePart id="400F0358BA1BD019DF7E2BDB14B0E118">Boyer, Pierre</mods:namePart>
<mods:affiliation id="3973A3E99765E79610CCEE76207E8EDD">Lotissement l’Horizon, 13610 Le Puy Sainte Réparade, France.</mods:affiliation>
</mods:name>
<mods:name id="2BD0AE60F842ED7319D8871EB1E367C1" type="personal">
<mods:role id="4AF8CEF40241F517676F85686BB7C47C">
<mods:roleTerm id="02D6EBF6B55D29AE4307D7DD21C96611">Author</mods:roleTerm>
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<mods:namePart id="83CF10AC57758D114170AA255802741E">Cerdeña, José</mods:namePart>
<mods:nameIdentifier id="F4D57F47BD755BEB0F7CAA559240178E" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="E27AD0896374B6721E7D41FCE1C7B423">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="618295DD5671054E45C281DD029DDF84" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
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<mods:roleTerm id="8BDD11FE53FAA7A02FC7EB5CFF123F1F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="96A2189895BBFC8C856929CC089D59D1">Larico, Jackie Farfán</mods:namePart>
<mods:nameIdentifier id="7095463EA5E8DBAAF99FBF0381508EC6" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="C5CBC1FFEFA978412D19EB47D9E333D4">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="96EACE1A70FB34E84970DCCF70FF7BF8" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
</mods:name>
<mods:name id="D94AE6BFB4DDE82ADAA0E66945AFB9B2" type="personal">
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<mods:roleTerm id="16D7313F266B4F158DF5C2460C5793F9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A7913696CC521B7C5699896F6C13D227">Brudecka, Jadwiga Lorenc-</mods:namePart>
<mods:affiliation id="70DFB6F4409D85E69F5E1C6191AFAD27">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:namePart id="75DC15EB07B76F4B7257EE70D4E8FD6E">Prieto, Carlos</mods:namePart>
<mods:affiliation id="A85AF864C8C8EBB57086C89D31AA7893">Corporación Universitaria Autónoma del Cauca, Popayán, Colombia.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="BFA4216586B9E187F7646687178AC1F0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="886DC9B573E6C5687203D2F0FF5CB402">Pyrcz, Tomasz W.</mods:namePart>
<mods:affiliation id="2E8FF49FCD26EEB5E88C8854923BF254">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:title id="C4CC639189D2FDBECF7BAF3FE86239EF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7414489BC7CA57BCB9255029B8126A3F">
<mods:date id="07410B48BDEEED4D2E62FA581B33E4DE">2021</mods:date>
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<mods:number id="2374E8C67D4AA7288FCF914D14B3526E">2021-04-16</mods:number>
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<mods:number id="4063CB990505A7B9DB823E20DD6E1183">4963</mods:number>
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<mods:classification id="AEE22860A57B81EFD9DA8472D6F64895">journal article</mods:classification>
<mods:identifier id="96B027BF82DC11FDD7F0217DE07D5526" type="CLB-Dataset">7214</mods:identifier>
<mods:identifier id="D034DF7466D907EF0D6B0DDD6C13A0FB" type="DOI">10.11646/zootaxa.4963.1.2</mods:identifier>
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</mods:mods>
<treatment id="D5008F50FFE7B303FF76FAAA92E8FF34" ID-DOI="http://doi.org/10.5281/zenodo.4696420" ID-GBIF-Taxon="181531943" ID-Zenodo-Dep="4696420" LSID="urn:lsid:plazi:treatment:D5008F50FFE7B303FF76FAAA92E8FF34" httpUri="http://treatment.plazi.org/id/D5008F50FFE7B303FF76FAAA92E8FF34" lastPageId="15" lastPageNumber="26" pageId="14" pageNumber="25">
<subSubSection id="15B36DCDFFE7B302FF76FAAA95AEFAEA" box="[151,903,1292,1319]" pageId="14" pageNumber="25" type="nomenclature">
<paragraph id="5D163E46FFE7B302FF76FAAA95AEFAEA" blockId="14.[151,903,1292,1355]" box="[151,903,1292,1319]" pageId="14" pageNumber="25">
<heading id="065E892AFFE7B302FF76FAAA95AEFAEA" bold="true" box="[151,903,1292,1319]" fontSize="11" level="1" pageId="14" pageNumber="25" reason="1">
<emphasis id="6FDDE254FFE7B302FF76FAAA95AEFAEA" bold="true" box="[151,903,1292,1319]" pageId="14" pageNumber="25">
<taxonomicName id="9AA945C5FFE7B302FF76FAAA9469FAEA" authority="Balint, Lorenc" authorityName="Balint, Lorenc-Brudecka, &amp; Pyrcz" authorityYear="2021" box="[151,576,1292,1319]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="ianusca">
<emphasis id="6FDDE254FFE7B302FF76FAAA97ACFAEA" bold="true" box="[151,389,1292,1319]" italics="true" pageId="14" pageNumber="25">Johnsonita ianusca</emphasis>
Bálint, Lorenc
</taxonomicName>
–Brudecka &amp; Pyrcz,
<taxonomicNameLabel id="74EE5F2FFFE7B302FCA4FAAB95AEFAEA" box="[837,903,1293,1319]" pageId="14" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="15B36DCDFFE7B303FF76FA9792E8FF34" lastPageId="15" lastPageNumber="26" pageId="14" pageNumber="25" type="description">
<paragraph id="5D163E46FFE7B302FF76FA97979FFA86" blockId="14.[151,903,1292,1355]" box="[151,438,1329,1355]" pageId="14" pageNumber="25">
(
<figureCitation id="C59222C3FFE7B302FF7EFA9796CEFA86" box="[159,231,1329,1355]" captionStart="FIGURE 8" captionStartId="15.[151,250,993,1017]" captionTargetBox="[164,1422,280,959]" captionTargetId="figure-69@15.[151,1436,273,970]" captionTargetPageId="15" captionText="FIGURE 8. Johnsonita ianusca Bálint, Lorenc–Brudecka &amp; Pyrcz, sp. n. holotype male (Ecuador: Azuay), A: recto, B: verso (CEP–MZUJ specimen) (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711267" httpUri="https://zenodo.org/record/4711267/files/figure.png" pageId="14" pageNumber="25">Figs 8</figureCitation>
,
<figureCitation id="C59222C3FFE7B302FF15FA97976EFA86" box="[244,327,1329,1355]" captionStart="FIGURE 18" captionStartId="28.[151,249,1943,1967]" captionTargetBox="[227,1333,184,1900]" captionTargetId="figure-23@28.[175,1413,181,1920]" captionTargetPageId="28" captionText="FIGURE 18. Johnsonita genitalia armatures, entire male organs in lateral (left column) and ventral (right column) views. A–B: J. ianusca, Ecuador, Azuay, Cuenca; C–D: J. johnbanksi, Peru, Molinopampa; E–F: J. johnsoni, Ecuador, Riobamba-Macas via Atillo; G–H: J. pardoa, Venezuela, Táchira, P. N. Tama (CEP–MZUJ specimens) (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711293" httpUri="https://zenodo.org/record/4711293/files/figure.png" pageId="14" pageNumber="25">18A–B</figureCitation>
,
<figureCitation id="C59222C3FFE7B302FEB2FA9797AFFA86" box="[339,390,1329,1355]" captionStart="FIGURE 23" captionStartId="33.[151,250,1570,1594]" captionTargetBox="[390,1203,198,1539]" captionTargetId="figure-18@33.[378,1210,181,1545]" captionTargetPageId="33" captionText="FIGURE 23. Johnsonita mail trait ventral forewing subcubital scent patch indicated by arrow. A: J. carpia, B: J. catadupa, C: J. chlamydem, D: J. ianusca, E: J. johnsoni, F: J. pardoa, G: J. turquisca, H: J. zubkova." figureDoi="http://doi.org/10.5281/zenodo.4711305" httpUri="https://zenodo.org/record/4711305/files/figure.png" pageId="14" pageNumber="25">23D</figureCitation>
,
<figureCitation id="C59222C3FFE7B302FE70FA979787FA86" box="[401,430,1329,1355]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="14" pageNumber="25">30</figureCitation>
)
</paragraph>
<paragraph id="5D163E46FFE7B302FF76FADC94C2F9EE" blockId="14.[151,1437,1401,2039]" pageId="14" pageNumber="25">
<emphasis id="6FDDE254FFE7B302FF76FADC9713FA5E" box="[151,314,1401,1427]" italics="true" pageId="14" pageNumber="25">Type material.</emphasis>
<materialsCitation id="EDC1341BFFE7B302FEA0FADF9452F9EE" ID-GBIF-Occurrence="3083794347" collectingDate="2003-02-09" collectingDateMax="2016-05-17" collectingDateMin="2003-02-09" collectionCode="CEP, MZUJ" collectorName="J. Wojtusiak &amp; J. Lorenc" country="Ecuador" elevation="2600" location="Molleturo" municipality="Cuenca" pageId="14" pageNumber="25" specimenCount="1" stateProvince="Azuay" typeStatus="holotype">
<collectionCode id="3BB8A683FFE7B302FEA0FADF975DFA5E" box="[321,372,1401,1427]" pageId="14" pageNumber="25">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE7B302FE62FADF97E5FA5E" box="[387,460,1401,1427]" pageId="14" pageNumber="25">MZUJ</collectionCode>
<typeStatus id="821280E4FFE7B302FE35FADF941FFA5E" box="[468,566,1401,1427]" pageId="14" pageNumber="25" type="holotype">holotype</typeStatus>
male, in moderate condition (HWs broken, abdomen missing (genitalia with abdomen in microvial attached to the specimen), labelled as (1) “
<collectingCountry id="25BE7ED6FFE7B302FC89FA3B95DBFA7A" box="[872,1010,1437,1463]" name="Ecuador" pageId="14" pageNumber="25">ECUADOR</collectingCountry>
[//] Prov.
<collectingRegion id="9F6DF0A4FFE7B302FB80FA3B9283FA7A" box="[1121,1194,1437,1463]" country="Ecuador" name="Azuay" pageId="14" pageNumber="25">Azuay</collectingRegion>
[//]
<collectingMunicipality id="BD72A43CFFE7B302FB38FA3B9319FA7A" box="[1241,1328,1437,1463]" pageId="14" pageNumber="25">Cuenca</collectingMunicipality>
,
<location id="5876689DFFE7B302FADBFA3B9687FA16" LSID="urn:lsid:plazi:treatment:D5008F50FFE7B303FF76FAAA92E8FF34:5876689DFFE7B302FADBFA3B9687FA16" country="Ecuador" municipality="Cuenca" name="Molleturo" pageId="14" pageNumber="25" stateProvince="Azuay">Molleturo</location>
[//]
<date id="29171886FFE7B302FF3EFA679749FA16" box="[223,352,1473,1499]" pageId="14" pageNumber="25" value="2003-02-09">
<collectingDate id="3953E16EFFE7B302FF3EFA679749FA16" box="[223,352,1473,1499]" pageId="14" pageNumber="25" value="2003-02-09">02.09.2003</collectingDate>
</date>
,
<quantity id="9A5193A3FFE7B302FE8DFA6797E8FA16" box="[364,449,1473,1499]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="25" unit="m" value="2600.0">
<elevation id="D684D975FFE7B302FE8DFA6797E8FA16" box="[364,449,1473,1499]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="25" unit="m" value="2600.0">2600 m</elevation>
</quantity>
[//] leg.
<collectorName id="F05C5B90FFE7B302FDC3FA679485FA16" box="[546,684,1473,1499]" pageId="14" pageNumber="25">J. Wojtusiak</collectorName>
” (white label, printed), (2) “
<taxonomicName id="9AA945C5FFE7B302FC09FA67924AFA16" authorityName="Salazar &amp; Constantino" authorityYear="1995" box="[1000,1123,1473,1499]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="25" phylum="Arthropoda" rank="genus">Johnsonita</taxonomicName>
M51 [//] Budapest 2010.IX [//] Bálint Zsolt” (white label, printed), (3) “prep. genit. 377 [//]
<date id="29171886FFE7B302FC8CFA4395C7F9CD" box="[877,1006,1509,1536]" pageId="14" pageNumber="25" value="2016-05-17">
<collectingDate id="3953E16EFFE7B302FC8CFA4395C7F9CD" box="[877,1006,1509,1536]" pageId="14" pageNumber="25" value="2016-05-17">17.05.2016</collectingDate>
</date>
/
<collectorName id="F05C5B90FFE7B302FC16FA439275FA32" box="[1015,1116,1509,1536]" pageId="14" pageNumber="25">J. Lorenc</collectorName>
[//]
<taxonomicName id="9AA945C5FFE7B302FB6CFA4392D2FA32" authorityName="Balint, Lorenc-Brudecka, &amp; Pyrcz" authorityYear="2021" box="[1165,1275,1509,1535]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="ianusca">J. ianusca</taxonomicName>
[//] Molleturo [//]
<date id="29171886FFE7B302FF21F9AF9766F9EE" box="[192,335,1545,1571]" pageId="14" pageNumber="25" value="2003-09-02">
<collectingDate id="3953E16EFFE7B302FF21F9AF9766F9EE" box="[192,335,1545,1571]" pageId="14" pageNumber="25" value="2003-09-02">02. 09. 2003</collectingDate>
</date>
” (white label, printed). No
</materialsCitation>
<typeStatus id="821280E4FFE7B302FD62F9AF94CEF9EE" box="[643,743,1545,1571]" pageId="14" pageNumber="25" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="5D163E46FFE7B302FF26F9889304F88E" blockId="14.[151,1437,1401,2039]" pageId="14" pageNumber="25">
<emphasis id="6FDDE254FFE7B302FF26F988977AF98A" box="[199,339,1582,1607]" italics="true" pageId="14" pageNumber="25">Description.</emphasis>
Wings. FW length:
<quantity id="9A5193A3FFE7B302FDDCF98B94A7F98A" box="[573,654,1581,1608]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="25" unit="mm" value="15.0">15 mm</quantity>
(
<typeStatus id="821280E4FFE7B302FD7FF98B952FF985" box="[670,774,1581,1608]" pageId="14" pageNumber="25" type="holotype">holotype</typeStatus>
), FW apical area of costa and outer margin is not straigth resulting in a somewhat rounded apex; male dorsal wing surface structural colouration bright violet; dorsal FW surface black border reaching the middle of vein M3; HW dorsal surface discal area structurally blue coloured; ventral wing surface submedian area with a conspicuous discal spot, medial pattern composed by light disrupted line supplemented distally with dark spots; ventral wing surface inner margin medial area with brown scaling. Genitalia. Male valva with pointed small costal margin flip and heavily sclerotized pointed valval terminus slightly downturning in lateral view (with valval length), gnathos tip extremely thin and pointed downturning at its very final point. Androconia present in ventral FW along Cubital 2 vein as a patch in submedian area. Female not known.
</paragraph>
<paragraph id="5D163E46FFE7B302FF26F8E8927DF862" blockId="14.[151,1437,1401,2039]" pageId="14" pageNumber="25">
<emphasis id="6FDDE254FFE7B302FF26F8E89769F8AA" box="[199,320,1870,1895]" italics="true" pageId="14" pageNumber="25">Diagnosis.</emphasis>
The species is easily distinguished on the basis of the ventral HW discal pattern, where there is a discal spot (it is missing in all other congeners); somewhat resembles in wing shape and pattern to the sympatric
<taxonomicName id="9AA945C5FFE7B302FA68F8D496D6F862" authorityName="Balint" authorityYear="2003" baseAuthorityName="Hewitson" baseAuthorityYear="1869" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="catadupa">
<emphasis id="6FDDE254FFE7B302FA68F8D496D6F862" italics="true" pageId="14" pageNumber="25">J. catadupa</emphasis>
</taxonomicName>
, but the dorsal wing surfaces are deeper blue (
<taxonomicName id="9AA945C5FFE7B302FCEBF83095A5F862" authorityName="Balint" authorityYear="2003" baseAuthorityName="Hewitson" baseAuthorityYear="1869" box="[778,908,1941,1967]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="catadupa">
<emphasis id="6FDDE254FFE7B302FCEBF83095A5F862" box="[778,908,1941,1967]" italics="true" pageId="14" pageNumber="25">J. catadupa</emphasis>
</taxonomicName>
are bright azure).
</paragraph>
<paragraph id="5D163E46FFE7B302FF26F81F97ACF83A" blockId="14.[151,1437,1401,2039]" pageId="14" pageNumber="25">
<emphasis id="6FDDE254FFE7B302FF26F81F9771F81E" box="[199,344,1977,2003]" italics="true" pageId="14" pageNumber="25">Distribution.</emphasis>
Geographical:
<collectingCountry id="25BE7ED6FFE7B302FDE7F81F94A4F81E" box="[518,653,1977,2003]" name="Ecuador" pageId="14" pageNumber="25">ECUADOR</collectingCountry>
(
<collectingRegion id="9F6DF0A4FFE7B302FD7CF81F94C0F81E" box="[669,745,1977,2003]" country="Ecuador" name="Azuay" pageId="14" pageNumber="25">Azuay</collectingRegion>
) (
<figureCitation id="C59222C3FFE7B302FCE3F81F957EF81E" box="[770,855,1977,2003]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="14" pageNumber="25">Fig. 30</figureCitation>
); spatial: recorded from
<quantity id="9A5193A3FFE7B302FB8FF81F92ECF81E" box="[1134,1221,1977,2003]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="25" unit="m" value="2600.0">2600 m</quantity>
a.s.l; temporal: recorded in September.
</paragraph>
<paragraph id="5D163E46FFE6B303FF26FF3192E8FF34" blockId="15.[151,1436,151,249]" pageId="15" pageNumber="26">
<emphasis id="6FDDE254FFE6B303FF26FF31976CFF7C" box="[199,325,151,177]" italics="true" pageId="15" pageNumber="26">Etymology.</emphasis>
Dedicated to the late professor Janusz Wojtusiak (1942–2012), who was a great entomologist and our educator, mentor, Academic teacher, and showed us how to love tropical butterflies. The species name “ianusca” is freely formed by Latinising the word “Janusz” and treating it as a female noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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